806 lines
70 KiB
XML
806 lines
70 KiB
XML
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<mods:title id="26B78BA4D1CD382061374789C2296919">Taxonomic resolution of the hillstream suck-loach Beaufortia pingi species group (Cypriniformes, Gastromyzontidae) and two new species from Southwest China – Beaufortia granulopinna and Beaufortia viridis</mods:title>
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<mods:namePart id="66457F72F87D19B1C40C17815A9DFC29">Chen, Jing-Chen</mods:namePart>
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<mods:nameIdentifier id="778626E6EE8BF497CD49BC8B1881E37A" type="ORCID">0000-0001-9164-6920</mods:nameIdentifier>
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<mods:affiliation id="B0172805E8B6F84B996ED29B2A8324D3">Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China</mods:affiliation>
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<mods:namePart id="E118A5C0D16C901224EB293CD4F329C3">Li, Jia-Jia</mods:namePart>
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<mods:affiliation id="37C94CDE9018242E9CAAFEDF2BD08D41">Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China</mods:affiliation>
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<mods:namePart id="CF3A0902B020A34ADDC8D96005DA6E02">Tang, Wen-Qiao</mods:namePart>
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<mods:affiliation id="34B9AEB1BD5A8272507D5683E975BC33">Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China</mods:affiliation>
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<mods:namePart id="55D970207BECCBC7EB699BD51B890673">Pu, Xin-Rui</mods:namePart>
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<mods:affiliation id="350FD57FBC6469829B90C615CA1F717B">Key Laboratory of Exploration and Utilization of Aquatic Genetic Resources, Ministry of Education, Shanghai Ocean University, Shanghai 201306, China</mods:affiliation>
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<mods:namePart id="3D9911242A1073DB41AA6BA7257AB28C">Lei, Hao-Tian</mods:namePart>
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<mods:affiliation id="525DD0026E50CBF3E66B6EC9E673CDCA">International College, Yunnan Agricultural University, Kunming 650500, China</mods:affiliation>
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<mods:title id="8B21BC1D560F26F8F75481F1050F566F">Zoosystematics and Evolution</mods:title>
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<mods:date id="28550AC58AF6B8E55E77D9D14001D4F4">2024</mods:date>
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<mods:number id="BF0CCA1310855C8CB4B9E6EDB6849996">2024-07-09</mods:number>
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<mods:number id="4A5036960A0EF4DE32E3767C37B421CE">100</mods:number>
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<mods:start id="34AC5A19FC3D68977FB8927FA534D688">941</mods:start>
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<mods:identifier id="89F8ED1BE5AFAB2ED97D200244A0E2ED" type="DOI">10.3897/zse.100.124370</mods:identifier>
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<treatment id="D024A262442F577EA888DBAF142AF987" ID-DOI="http://doi.org/10.5281/zenodo.12701249" ID-GBIF-Taxon="235025269" ID-Zenodo-Dep="12701249" ID-arpha="D024A262-442F-577E-A888-DBAF142AF987" ID-zoobank="https://zoobank.org/268BD22A-C14B-4662-BFFC-A8C3369DACC7" LSID="urn:lsid:zoobank.org:act:268BD22A-C14B-4662-BFFC-A8C3369DACC7" LSID-ZBK="urn:lsid:zoobank.org:act:268BD22A-C14B-4662-BFFC-A8C3369DACC7" httpUri="http://treatment.plazi.org/id/D024A262442F577EA888DBAF142AF987">
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<subSubSection id="B143E0FA37578ABC6139E567BF80ADCF" type="nomenclature">
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<paragraph id="E3F90496181D50ACE760939AFC5C98FB">
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<taxonomicName id="D070DAE0516396114A793BA07195684B" authority="Chen & Tang" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
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<emphasis id="53585C2DC589B19160488ECB3C4DB679" italics="true">Beaufortia viridis</emphasis>
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Chen & Tang
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</taxonomicName>
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<taxonomicNameLabel id="4E5CB174065FF9953CEF931F3C450756" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="756729BF19681ED5E4E4AA7D3815AB09" type="description">
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<paragraph id="9E579FFE0215B53B15D4CD9F70407505">
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<figureCitation id="C2465F01EEC2F0C2090A82A2E7ECA3BF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov., holotype, adult, SHOU 20240103101, 38.03 mm SL; from Wuming River, a stream tributary to the You River of the Pearl River basin, at Wuming District, Nanning City, Guangxi Province, China." figureDoi="10.3897/zse.100.124370.figure10" httpUri="https://binary.pensoft.net/fig/1088057">Figs 10</figureCitation>
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,
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<figureCitation id="D7FCBA139A82C8D7A61A5A8075515FF5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov. during early juvenile and sub-adult stages. А. Early juvenile stage, SHOU 20240103623, from type locality, 12.64 mm SL, pelvic fins completely separated; B. Sub-adult stage, SHOU 20240103102, paratype, 25.81 mm SL." figureDoi="10.3897/zse.100.124370.figure11" httpUri="https://binary.pensoft.net/fig/1088058">11</figureCitation>
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,
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<figureCitation id="3160F303050A861B7F31160724314EA8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lateral (top), dorsal (middle), and ventral (bottom) views of different regional phenotypes of Beaufortia viridis sp. nov. А. Regional phenotype from Lingyun County, Guangxi, SHOU 20240126104, paratype, 58.71 mm SL; B. Regional phenotype from Tuoniang River, Guangnan County, Yunnan, SHOU 20240109105, paratype, 40.15 mm SL (left); SHOU 20240109103, paratype, 39.36 mm SL (right)." figureDoi="10.3897/zse.100.124370.figure12" httpUri="https://binary.pensoft.net/fig/1088059">12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="511FA8295AF4B455B0D71158FFAF93AB" type="reference_group">
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<paragraph id="D9E6F790BB0588E8AD4B0708D1D78D59">
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||
<treatmentCitationGroup id="B19CEA477364A0A8FF134407092A7F49">
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||
<treatmentCitation id="2B1EAA6D97DF09E6D0E8850343D7F0F4" author="Yue ZH" page="170" year="1981">
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||
<taxonomicName id="E6C844C8DB5A2B61EDF21FE1B072F28E" ID-CoL="L5KW" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
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<emphasis id="1EAB85D9EDCC6055A94273E0EC6077DA" italics="true">Beaufortia pingi</emphasis>
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</taxonomicName>
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<comment id="357862304A664FF43C5E3DECD997146D">
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:
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<bibRefCitation id="4671C9DCC0A3A5111F316E70830F7717" author="Yue ZH" editor="Zheng BS" firstAuthor="Yue" journalOrPublisher="Guangxi People’s Publishing House, Nanning" pagination="166-175" refId="B26" refString="Yue ZH (1981) Gastromyzontidae. In: Zheng BS (Ed.) Freshwater fishes of Guangxi, China. Guangxi People’s Publishing House, Nanning, 166–175." title="Gastromyzontidae." volumeTitle="Freshwater fishes of Guangxi, China." year="1981">Yue (1981)</bibRefCitation>
|
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: 170–172, Lingyun County,
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<collectingRegion id="E5F61E328F5CC23075A113BC404A9673" country="China" name="Guangxi">Guangxi</collectingRegion>
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, admixed within
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<taxonomicName id="E9D5D3BD38925AB839051B9DB213F542" ID-CoL="L5KW" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
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<emphasis id="8D4F21B1BF9C0BAC2BF97ECF932BBEB1" italics="true">B. pingi</emphasis>
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</taxonomicName>
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||
.
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<treatmentCitation id="64714C3A9AD57B4311AB71AD43A034FC" author="Zheng CY" page="257" year="1989">
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||
<bibRefCitation id="1A2B97FD00D7345A5DBA66A23CDA7F87" author="Zheng CY" firstAuthor="Zheng" refId="B28" refString="Zheng CY (1989) The Fishes of Pearl River. Science Press, Beijing, 257–260." year="1989">Zheng (1989)</bibRefCitation>
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: 257
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</treatmentCitation>
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||
–258, Lingyun County,
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<collectingRegion id="F674FFAF31F12C84D163FD66477529D7" country="China" name="Guangxi">Guangxi</collectingRegion>
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, admixed within
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<taxonomicName id="92B2CA731EAA009CCAE04CBAD9E7FC05" ID-CoL="L5KW" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
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<emphasis id="EAEB66FC5C7C9E69BB92426F2934863D" italics="true">B. pingi</emphasis>
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</taxonomicName>
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||
.
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</comment>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="84023F17CED3CD4D36BE2983BFAD9BAB">
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||
<treatmentCitationGroup id="3F1D96F517091B1E43AD40928F51B421">
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||
<treatmentCitation id="AE2037370C50332B9D5DE251A7F1AA0C" author="Chen YR" page="85" year="1990">
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||
<taxonomicName id="B5D6ABCE443F20E38C2311A6CCFCF9E5" ID-CoL="L5KW" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
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<emphasis id="5A5D0E5AE6FF6D36246105977B6F6FBC" italics="true">Beaufortia pingi</emphasis>
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</taxonomicName>
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<comment id="EC549C3A6BEEE80BFE6616686853F4E7">
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:
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<bibRefCitation id="8213738236DF306E66E9285BBA9651DF" author="Chen YR" editor="Zhe LX & Chen YR" firstAuthor="Chen" journalOrPublisher="Science Press, Beijing" pagination="85-89" refId="B2" refString="Chen YR (1990) Beaufortia. In: Zhe LX, Chen YR (Eds) The Fishes of Yunnan, China Part II Cyprinidae. Science Press, Beijing, 85–89." title="Beaufortia." volumeTitle="The Fishes of Yunnan, China Part II Cyprinidae." year="1990">Chen (1990)</bibRefCitation>
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: 85–87, Xiyang River, Guangnan County,
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<collectingRegion id="4F2744C52F5736A0C25FAC705C588E7F" country="China" name="Yunnan">Yunnan</collectingRegion>
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.
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</comment>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="360CE3EBA6ED8F7FAEF2B36114EC1723">
|
||
<treatmentCitationGroup id="49F32D64BB074444C9A9619165B46161">
|
||
<treatmentCitation id="F4448046D909048BC04A59053D590A5E" author="Chen YY & Tang WQ" page="504" year="2000">
|
||
<taxonomicName id="D27819072F0C2C56581373C151352762" ID-CoL="L5LC" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="zebroidus">
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<emphasis id="3240626DA1015A740D83EBB28BF57FF7" italics="true">Beaufortia zebroidus</emphasis>
|
||
</taxonomicName>
|
||
<comment id="B80403A665FFEE4EAAA597FDD4FFB812">
|
||
:
|
||
<bibRefCitation id="806A892A8A1EAEA2697E89AFA0C472F1" author="Chen YY & Tang WQ" firstAuthor="Chen" refId="B4" refString="Chen YY, Tang WQ (2000) . Homalopteridae. In: Le PQ (Ed.) Fauna Sinica, Osteichthyes, Cypriniformes III. Science Press, Beijing 438–567." year="2000">Chen and Tang (2000)</bibRefCitation>
|
||
: 504–505, Tian′e County,
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<collectingRegion id="CDAE73E4B29535FD7324045689CCDB1B" country="China" name="Guangxi">Guangxi</collectingRegion>
|
||
.
|
||
<treatmentCitation id="69623BD0B6738F89142582DC2E181D62" author="Chen XY" page="292" year="2013">
|
||
<bibRefCitation id="D223BA45B983C4B0B0B5EE247DF1E6DD" author="Chen XY" firstAuthor="Chen" journalOrPublisher="Zoological Research" pagination="281-343" refId="B3" refString="Chen XY (2013) Checklist of Fishes of Yunnan. Zoological Research 34: 281–343." title="Checklist of Fishes of Yunnan." volume="34" year="2013">Chen (2013)</bibRefCitation>
|
||
: 292
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</treatmentCitation>
|
||
, Xiyang River,
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<collectingRegion id="F41760887306142CD55921324A0C94DF" country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
.
|
||
</comment>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2HBI" type="type material">
|
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<paragraph id="AB80A8631DAB797FF252FB6C76872196">
|
||
<heading id="5FD0185DB1CCE9AEE1DBAA8CB0D36D5F" reason="title">Type material.</heading>
|
||
</paragraph>
|
||
<paragraph id="D716D78329A240CF6ADBEB1217E85526">
|
||
<materialsCitation id="91C233D4819E2C3E14E123959B74E73B" ID-GBIF-Occurrence="4904609302" collectingDate="2023-12-27" collectionCode="SHOU" collectorName="Jing-Chen Chen & Qian-Yu Liang" country="China" county="Wuming District" elevation="160" latitude="24.339" location="Wuming River" longLatPrecision="9" longitude="106.3604" municipality="Nanning City" specimenCode="SHOU 20240103101" specimenCount="1" stateProvince="Guangxi Province" typeStatus="holotype">
|
||
<emphasis id="DF6A0630E47382E6B58EFEF0E11FD146" bold="true">
|
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<emphasis id="CF88121902D6D8B601A2E3D5C4C740B2" bold="true" italics="true">
|
||
<typeStatus id="E690002A987BA8DC7FF41CEFE82701B7" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="0B6A90ECBE71C598A7E4DDC64500DA8B" collectionCode="SHOU">SHOU 20240103101</specimenCode>
|
||
,
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||
<quantity id="76FC3A296FA40103BD1822CACAF921D7" metricMagnitude="-2" metricUnit="m" metricValue="4.842" unit="mm" value="48.42">48.42 mm</quantity>
|
||
total length (TL),
|
||
<quantity id="E72602AF59D3BEFAF08E549586B18DE8" metricMagnitude="-2" metricUnit="m" metricValue="3.803" unit="mm" value="38.03">38.03 mm</quantity>
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||
standard length (SL), adult (Fig.
|
||
<figureCitation id="D5BE9B5951647E1B29AB7132FD940C85" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov., holotype, adult, SHOU 20240103101, 38.03 mm SL; from Wuming River, a stream tributary to the You River of the Pearl River basin, at Wuming District, Nanning City, Guangxi Province, China." figureDoi="10.3897/zse.100.124370.figure10" httpUri="https://binary.pensoft.net/fig/1088057">10</figureCitation>
|
||
). Collected by
|
||
<collectorName id="DEBB962E7EAD0D42CF4095B18051BEF3">Jing-Chen Chen</collectorName>
|
||
and
|
||
<collectorName id="B8164D29C8387F3FF4CBA3426D65F6D9">Qian-Yu Liang</collectorName>
|
||
on
|
||
<date id="DE666701185DFC82B3B7A53EE04236DF" value="2023-12-27">
|
||
<collectingDate id="5345AAB42157FC8188DBA21B71A9E725" value="2023-12-27">27 December 2023</collectingDate>
|
||
</date>
|
||
, from
|
||
<location id="994CA1FF2F536A726B039AB94B3FA157" LSID="urn:lsid:plazi:treatment:D024A262442F577EA888DBAF142AF987:994CA1FF2F536A726B039AB94B3FA157" country="China" county="Wuming District" latitude="24.339" longLatPrecision="9" longitude="106.3604" municipality="Nanning City" name="Wuming River" stateProvince="Guangxi Province">Wuming River</location>
|
||
, a stream tributary to the You River of the Pearl River basin, at
|
||
<collectingCounty id="56668A952811C74CFBE9AAB8CC7104B2">Wuming District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="71B1755C3D21E6812D2EC4388A1C70B6">Nanning City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="34A13EDF48BEB1518FC14028E2986B81" country="China" name="Guangxi">Guangxi Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="B4D180777464E84150CE5E967BCEC7E1" name="China">China</collectingCountry>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENIBI" specific-use="{"type":"Point","coordinates":[106.360400,24.339000]}">
|
||
<geoCoordinate id="196718286BA9EAFABAE9CF5A56611A79" degrees="24" direction="north" minutes="20.34" orientation="latitude" precision="9" value="24.339">24 ° 20.34 ' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CA5D0E83D6A77C469B331B21C717782C" degrees="106" direction="east" minutes="21.624" orientation="longitude" precision="1" value="106.3604">106 ° 21.624 ' E</geoCoordinate>
|
||
</named-content>
|
||
; c.
|
||
<elevation id="A24A941F9778ABC4D9DBDD0018A1F64B" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">
|
||
<quantity id="2D2F19034454FFEFF126530E1446A3B3" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">160 m</quantity>
|
||
a. s. l.
|
||
</elevation>
|
||
) (Fig.
|
||
<figureCitation id="D17F4993DA6B51C5ED249E9D3C2B19CE" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Collection site of the holotype (SHOU 20240103101) of Beaufortia viridis sp. nov., from Wuming River, a stream tributary to the You River of the Pearl River basin, Wuming District, Nanning City, Guangxi Province, China. Photographed by Jing-Chen Chen on 27 December 2023." figureDoi="10.3897/zse.100.124370.figure13" httpUri="https://binary.pensoft.net/fig/1088060">13</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="78D98D0FF9FA6E7C747E6C6C4211BFFA" ID-DOI="10.3897/zse.100.124370.figure10" ID-arpha="8B08238C-D9C1-59BC-9F22-70B8ECB329D7" httpUri="https://binary.pensoft.net/fig/1088057" startId="F10">
|
||
<paragraph id="2C3190828363CFD04A799BDBFC1FBF86">
|
||
<label id="5423C99D475947BD82B46246168D107E">Figure 10.</label>
|
||
</paragraph>
|
||
<paragraph id="BD07589C01F349CAFA680710AA5D9033">
|
||
Lateral (top), dorsal (middle), and ventral (bottom) views of
|
||
<taxonomicName id="34CF017833A269D3ABC17DE0E27B68EE" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="EC0DE76EAE58CB8E4752ADE0AD81977F" italics="true">Beaufortia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E95458C1499783DA46FD9A6C6B962FE5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, adult, SHOU 20240103101, 38.03 mm SL; from Wuming River, a stream tributary to the You River of the Pearl River basin, at Wuming District, Nanning City, Guangxi Province, China.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5936D8E67F5C2FDF798B0D19E8E97AAD">
|
||
<emphasis id="F02087EBF203CFA9E0C38D920D76A90E" bold="true">
|
||
<emphasis id="9A3D93608A0E9FB804137EB1E319FC21" bold="true" italics="true">
|
||
<typeStatus id="86244F05E7BC3283F931C7A1233000AA" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="FAF9A57E7C7B03C13B8A02A9A315E617" count="15" type="generic">15 specimens</specimenCount>
|
||
from the same locality as
|
||
<typeStatus id="00BC3BB0EC63F3B3089062E6C29DFA34" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="066F7F9627BAF4E3B9EC37CA9E359ED7">SHOU</collectionCode>
|
||
20240103102-116,
|
||
<quantity id="6BA74F4E12BD10D004E5D04963CB12BB" metricMagnitude="-2" metricUnit="m" metricValue="3.273" metricValueMax="4.664" metricValueMin="1.882" unit="mm" value="32.73" valueMax="46.64" valueMin="18.82">18.82–46.64 mm</quantity>
|
||
SL, were collected by Qian-Yu Liang and Jing-Chen Chen on
|
||
<date id="6B026B3C1F4BA2FDE2EAD11CE0EBF465" value="2023-12-27">27 December 2023</date>
|
||
;
|
||
<specimenCount id="7A08ED77F6412ADF47DFFB0508C2BC9A" count="8" type="generic">eight specimens</specimenCount>
|
||
from Lingyun County, Baise City,
|
||
<collectingRegion id="A5FDD9D9E07889C42F512CE16A91568A" country="China" name="Guangxi">Guangxi Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="5994D80427398CFDF18F67171237C9B6" name="China">China</collectingCountry>
|
||
,
|
||
<collectionCode id="3BAA7243B3C29A60E0F72EF83AC6584D">SHOU</collectionCode>
|
||
20240126101-108,
|
||
<quantity id="E59BE8B6517058228A5D0B8F9CA49284" metricMagnitude="-2" metricUnit="m" metricValue="5.4925" metricValueMax="5.985" metricValueMin="5.0" unit="mm" value="54.925" valueMax="59.85" valueMin="50.0">50–59.85 mm</quantity>
|
||
SL, were collected by Qian-Yu Liang and Jing-Chen Chen on
|
||
<date id="98559E9204A29635413BFEA5A9E8F9A6" value="2024-01-24">24 January 2024</date>
|
||
;
|
||
<specimenCount id="22E5702417858F28232526763932C887" count="5" type="generic">five specimens</specimenCount>
|
||
from Guangnan County, Wenshan Zhuang, and Miao Autonomous Prefecture,
|
||
<collectingRegion id="27AEB40EA325702B9A3ABFAFFF11849C" country="China" name="Yunnan">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="595DCC43037910F5545D29E83F815DA2" name="China">China</collectingCountry>
|
||
,
|
||
<collectionCode id="4208F9998FB13C6F862238B1DE0C3373">SHOU</collectionCode>
|
||
20240109101-105,
|
||
<quantity id="715C3D3D3677EAF77712177C8B828CD9" metricMagnitude="-2" metricUnit="m" metricValue="4.5215" metricValueMax="5.42" metricValueMin="3.6229999999999998" unit="mm" value="45.215" valueMax="54.2" valueMin="36.23">36.23–54.20 mm</quantity>
|
||
SL, were collected by Lao Xing on
|
||
<date id="915592051F9FC62D84B0D86362451C7F" value="2023-12-28">28 December 2023</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EVJBI" type="material">
|
||
<paragraph id="1AAD455D978029CE463D53730B0A2DE7">
|
||
<heading id="6F3396D7245B4E7A7F8513371BA42C9D" reason="title">Additional materials.</heading>
|
||
</paragraph>
|
||
<paragraph id="C865039781E477FDF21BEA61194452FE">
|
||
<specimenCount id="FF2C60C03CFD47AACCE405A48ACD993B" count="7" type="generic">Seven specimens</specimenCount>
|
||
from Guangnan County, Wenshan Zhuang, and Miao Autonomous Prefecture,
|
||
<collectingRegion id="C14A57FCD5E97ED026E9559B077F46E9" country="China" name="Yunnan">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F1A70B15EC0FFA298166DA51EB1C5064" name="China">China</collectingCountry>
|
||
,
|
||
<collectionCode id="41E8971D982F7A1572CD2E3CC31176C6">SHOU</collectionCode>
|
||
20240112101-107,
|
||
<quantity id="0D53B1E14828364AFADCE37DB9652DD8" metricMagnitude="-2" metricUnit="m" metricValue="4.4305" metricValueMax="4.864" metricValueMin="3.997" unit="mm" value="44.305" valueMax="48.64" valueMin="39.97">39.97–48.64 mm</quantity>
|
||
SL, were collected by Lin Yang and Lao Xing from
|
||
<date id="F49FD222123B35C2C1FD2A2E51E8062D" value="2022-12">December 2022</date>
|
||
to
|
||
<date id="2FBB674B7F4770F0006BA1E0D72DB9E8" value="2023-02">February 2023</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1JBI" type="diagnosis">
|
||
<paragraph id="6CFD318E1A38739E4E9261F9D3BDD914">
|
||
<heading id="8E1D65F9F334B8E037107B585E7DC70A" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="58C581ECDF17217256CA494CA109DEDA">
|
||
<taxonomicName id="34B445951F9030C9AAD69E82BA4CB7EE" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="F9404F9CC8C47CFE8B004028794F1079" italics="true">B. viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="49765FC1EAD9C3EDD3D987DBC3586D83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shares the typical characteristics with members of the
|
||
<taxonomicName id="85B3A3CDDD8CF198E341A650377C39F3" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
|
||
<emphasis id="19998BA234B354555D9EB061FC44D832" italics="true">B. pingi</emphasis>
|
||
</taxonomicName>
|
||
species group, with distinct vertical stripes on lateral body and a pinnate-type lower lip (vs. lacking prominent vertical stripes and having a dicot-type lower lip in other congeneric species apart from this group) (see Figs
|
||
<figureCitation id="460DFCF0FE0CBD5781BE9F7161D9C404" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pinnate-type lower lip of the Beaufortia pingi species group, elongated, with the lower lip on both sides being multilobed, typically comprising more than three lobes, with deep divisions; the extremities of the lower lip extend to the corners of the mouth, with several wart-like protrusions present on the extended parts. The red line outlines the left side of the lower lip contour; the red arrow indicates the position of the ventral head sensory canal-pores located anterior to the origin of the pectoral fins. А. B. pingi, from the type locality; B. B. zebroida, from Napo County, Guangxi Province; C. B. viridis, from the type locality; D. B. granulopinna, from the type locality." figureDoi="10.3897/zse.100.124370.figure6" httpUri="https://binary.pensoft.net/fig/1088053">6 C</figureCitation>
|
||
,
|
||
<figureCitation id="6732D0021B18B602584AF4DF18793741" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dicot-type lower lip of species outside the Beaufortia pingi group, shorter, with the sides of the lower lip being smooth or featuring 1–2 shallow notches; the ends of the lower lip do not extend to the corners of the mouth, or the extension is poor-developed and lacks protrusions. The red line outlines the left side of the lower lip contour; the red arrow indicates the position of the ventral head sensory canal-pores, with the second pair of canal-pores located posterior to the origin of the pectoral fins in all species except B. liui. А. B. leveretti, from the type locality; B. B. kweichowensis, from Congjiang County, Guizhou Province, downstream near the type locality; C. B. cyclica, holotype; D. B. intermedia, paratype; E. B. polylepis, syntype; F. B. huangguoshuensis, syntype; G. B. szechuanensis, from Neijiang City, Sichuan Province; H. B. niulanensis, from the type locality; I. B. liui, syntype. (C-F and I were photographed by Yi-Yang Xu, Institute of Hydrobiology, Chinese Academy of Sciences.)" figureDoi="10.3897/zse.100.124370.figure7" httpUri="https://binary.pensoft.net/fig/1088054">7</figureCitation>
|
||
, Table
|
||
<tableCitation id="B4FAD5C1AEF885B1E1F9D6A1F4AE7047" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key diagnostic features and countable characteristics of species within the genus Beaufortia." httpUri="http://table.plazi.org/id/956DD01FFFB711969847EB8CE841D667" tableUuid="956DD01FFFB711969847EB8CE841D667">2</tableCitation>
|
||
).
|
||
<taxonomicName id="E21365A2A38EB12DB987C0984946E6F0" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="F57A43B2827182A344FBD733BAF101F8" italics="true">B. viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77FDFD0835784492364708F002DE893C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be clearly distinguished from
|
||
<taxonomicName id="ECA9196E1EF7B9BE979B1A26B23DEF94" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="granulopinna">
|
||
<emphasis id="1AA8227E4B0096907CE79B042860C8D2" italics="true">B. granulopinna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1B333ADD7945E65352EE5B68B89FF4A5" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
|
||
<emphasis id="64380A1395E900E6A9BDC417B1031198" italics="true">B. pingi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="0824CB2FF21E3B0186DC7F881FBD6DCD" authorityName="Kottelat" authorityYear="2012" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="zebroida">
|
||
<emphasis id="DE7B9314FB6288AE1CFE6CD4018AE51B" italics="true">B. zebroida</emphasis>
|
||
</taxonomicName>
|
||
by consistent vertical stripes with uniform length, width, and inter-spacing at all growth stages (stripes on mid-section lateral body occasionally shorten in individuals from the Xiayang and Tuoniang Rivers in
|
||
<collectingRegion id="EFCFE2FF42CC16639E6B3979BFD1F197" country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
) (vs. a certain proportion (54.76 %) of adults exhibit blurred or vanished vertical stripes on mid-section lateral body in
|
||
<taxonomicName id="2FE5B7FD9B555AC875314463826C9600" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="granulopinna">
|
||
<emphasis id="9CF6979C0DF401869D82182D40E7F84A" italics="true">B. granulopinna</emphasis>
|
||
</taxonomicName>
|
||
; stripe length and width uneven, with a high proportion (94.00 %) of adults showing blurred or vanished mid-section vertical stripes in
|
||
<taxonomicName id="FD29C20480CC702FDF01BB5EBBDF342C" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
|
||
<emphasis id="3CB25BAF53F8F3D6A4D8C2AB97E9C7BF" italics="true">B. pingi</emphasis>
|
||
</taxonomicName>
|
||
; stripe length and width uneven, typically narrower than inter-spacing, sometimes reduced to dots in
|
||
<taxonomicName id="DD7596564CE61448E00BB9F2544E03F2" authorityName="Kottelat" authorityYear="2012" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="zebroida">
|
||
<emphasis id="A505AFF2E923ED3B92767CFB7398903F" italics="true">B. zebroida</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="9103A5E69ED47FBB396A08B4FD6C9D69" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="84AECC8694B33ADBB47BFAD35944D9F2" italics="true">B. viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="635E69D4246FF0322F4B68493BA724F1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be further distinguished from
|
||
<taxonomicName id="900D1B1A0E8F24E700C243EA8279C0F6" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="granulopinna" status="sp. nov.">
|
||
<emphasis id="B23BA341A488F413EE4D2B50746C50C8" italics="true">B. granulopinna</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2534EA9F63CA0485CA472974160B8EDC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the lack of prominent tubercles on the branched rays of the pectoral fins (vs. well-developed tubercles on anterior 6–9 rays of the pectoral fins present). Moreover,
|
||
<taxonomicName id="7C57C947D4596841040734716195155E" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="A90C58AF2CCFDF1EBF50DB9E1C7EB7A6" italics="true">B. viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C96CFB077743268B948A5951474F2BD4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="42CB8ABC1CBCB8926891521259181D16" authorityName="Kottelat" authorityYear="2012" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="zebroida">
|
||
<emphasis id="7E7AF5821CFA59C7FD23E705B5B779A8" italics="true">B. zebroida</emphasis>
|
||
</taxonomicName>
|
||
by body dark cyan to green in dorsal profile (vs. brown yellow to golden).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENPBI" type="description">
|
||
<paragraph id="0E182C6693D3737A52DE83592BCD8800">
|
||
<heading id="85CBAB2F3CD1AF8DCE79F6F3D1224B31" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="C6467E64AC7DE498CE9F88B10EF41175">
|
||
Dorsal iii- 7, anal ii- 4, pectoral i- 19–23 (21.08 ± 1.02), pelvic i- 14.5–19 (17.11 ± 1.01). Lateral-line canal pores and scales: 65–83 (73.47 ± 4.21) (see Table
|
||
<tableCitation id="48A8EE8FE120EE20520E193A2D20474F" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key diagnostic features and countable characteristics of species within the genus Beaufortia." httpUri="http://table.plazi.org/id/956DD01FFFB711969847EB8CE841D667" tableUuid="956DD01FFFB711969847EB8CE841D667">2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="847568235F70A3F6F007B6FA76A5B00C">
|
||
Morphometric measurements for the specimens examined are given in Table
|
||
<tableCitation id="C179D6D1D62585F15306995EB7127E27" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric results for species within the Beaufortia pingi group." httpUri="http://table.plazi.org/id/0457113D80E4E69FE4FE3D0D3260E3AE" tableUuid="0457113D80E4E69FE4FE3D0D3260E3AE">3</tableCitation>
|
||
. See Fig.
|
||
<figureCitation id="185C71934D22381B6136633B656F46CF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov., holotype, adult, SHOU 20240103101, 38.03 mm SL; from Wuming River, a stream tributary to the You River of the Pearl River basin, at Wuming District, Nanning City, Guangxi Province, China." figureDoi="10.3897/zse.100.124370.figure10" httpUri="https://binary.pensoft.net/fig/1088057">10</figureCitation>
|
||
for lateral, dorsal, and ventral views of the body.
|
||
</paragraph>
|
||
<paragraph id="2F450A5C6B6B04C13CEB12C6FD0A7D26">Head-thorax cylindrical, dorsal slightly humped medially, flattened ventrally, body compressed from pelvic fins to caudal peduncle. Head broad, blunt, length slightly less than width, more than depth, nuptial tubercles well-developed on lower half of head in mature individuals, snout rounded, length about half of head length. Mouth inferior, narrow, width about one-fifth head width, horseshoe-shaped, angle about 94 ° from midpoint to ends. Upper lip smooth, without distinct papillae; lower lip pinnate-type, slightly concave medially with multilobed sides; jaw edges slightly protrude. Shallow groove between upper lip and snout, extending to mouth corners. Lateral grooves shallow or indistinct. Two pairs of rostral barbels present, with outer pair slightly longer, space between rostral barbels with leaf-like folds, edges of which rounded and poor developed. One pair of maxillary barbels, length about equal to outer rostral barbels. Nostrils with tubular nasal flaps, distance between nostrils equals one-third head width. Eyes supralateral, medium-sized, eye diameter about one-quarter head length, interorbital space flat, width about half head width. Gill opening small, about equal to eye diameter, originated about vertically above the second to third branched pectoral fin ray, limited to dorsal side of head. Scales small, diameter smaller than pupil, dorsal surface of head, base of paired fins, and ventral area before pelvic fin bases nude. Lateral line complete, at midlateral.</paragraph>
|
||
<paragraph id="FB13562233AB8DE9F02D0ADB91C60286">Dorsal fin base about equal to pre-pectoral length, starting around midpoint from snout to caudal fin base, adpressed extending to about midway between dorsal fin origin and caudal fin base. Anal fin base length about half of that of dorsal fin base, adpressed extending to slightly beyond caudal fin base. Paired fins extending outwards, forming disc-like structure with body. Pectoral fin base slightly longer than head length, starting at the posterior one-third point of head, pectoral fin length about twice to head length, tip of which reaching pelvic fin base midpoint, pectoral disc width about 1.5 times head width at pectoral origin, tubercles on rays absent. Pelvic fin shorter than pectoral, with well-developed fleshy flap at dorsal base, last 1–3 branched rays partially connected by fin membrane forming pelvic disc, connected part about two-thirds ray length, remaining parts separated, forming notch in middle rear edge, exposing anus. Pelvic disc width about equal to length. Anus at or near posterior edge of pelvic disc, distance to which less than to anal fin origin. Caudal fin length about equal to pelvic fin, slanted end, lower lobe slightly longer.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDQBI" type="Coloration in preservation">
|
||
<paragraph id="77C704312F3526EEAD0615D9DBD8FF0C">
|
||
<heading id="EFD94389B315BA5D6F669506FEDCCFB4" reason="title">Coloration in preservation.</heading>
|
||
</paragraph>
|
||
<paragraph id="90C5D8E23E5FE23622CD06556CCBC0A2">Preserved specimen from sub-adult to adult stage, body dark green to grey, white ventrally. Head with black spots or vermiculation dorsally, 2–5 larger black blotches along mid-dorsal body anterior to dorsal fin. Sides with 9–15 thick dark vertical stripes, with uniform length, width, and inter-spacing, easily recognizable at all ages. Paired fin with pale-white or hyaline margin, inner edges with continuous or dotted black arc. Dorsal fin hyaline with black stripes, one black spot present on the root before the second branched ray.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIQBI" type="Coloration in live">
|
||
<paragraph id="AC58114DD093FF1A8182AC4522DAC292">
|
||
<heading id="A7EF361F1AF783CB42D22F38BFDE9470" reason="title">Coloration in live.</heading>
|
||
</paragraph>
|
||
<paragraph id="9B5B09EE63A539723D3FB1709D59ED9D">In life, sides of body dark cyan to green. After maturity, black stripes and spots on body turn metallic green, inter-spacing and hyaline fin membranes pale yellow.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENQBI" type="Juvenile morphology">
|
||
<paragraph id="05CEC254B9A95B7DCD8B67DF355CEAD5">
|
||
<heading id="7BE750AA891868950AC6A47A3C141B7F" reason="title">Juvenile morphology.</heading>
|
||
</paragraph>
|
||
<paragraph id="B280B248B0A7B1AFDAF773DCAE4C03D0">
|
||
Pelvic fins completely separated, body dark green in dorsal profile, few wider vertical dark stripes on sides, 2–3 vertical black stripes on caudal fin, other fins hyaline or with inconspicuous black lines (see Fig.
|
||
<figureCitation id="01B61E6718290237BC19C96D247FA3E0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov. during early juvenile and sub-adult stages. А. Early juvenile stage, SHOU 20240103623, from type locality, 12.64 mm SL, pelvic fins completely separated; B. Sub-adult stage, SHOU 20240103102, paratype, 25.81 mm SL." figureDoi="10.3897/zse.100.124370.figure11" httpUri="https://binary.pensoft.net/fig/1088058">11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="76A3511436069FB65A1D60B79882F5CE" ID-DOI="10.3897/zse.100.124370.figure11" ID-arpha="DA07A11E-057B-5DFC-9DD5-6F971BDF715E" httpUri="https://binary.pensoft.net/fig/1088058" startId="F11">
|
||
<paragraph id="65FC1100EE9976BF9D8A84C68B8262D2">
|
||
<label id="C50E7E3551C028C254877CBA0AB298FB">Figure 11.</label>
|
||
</paragraph>
|
||
<paragraph id="28FAAF5EF93428EE2EC316B917C2D6EB">
|
||
Lateral (top), dorsal (middle), and ventral (bottom) views of
|
||
<taxonomicName id="4ABF3F697923CC442F31FC0F55419B1E" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="B8A264D1A5AD30B6D785975CD5B4B1D0" italics="true">Beaufortia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0F9E5122040C25F0CF6970677D9DFD59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
during early juvenile and sub-adult stages.
|
||
<emphasis id="0A530591A4CE3B2ED91CD4A50B924C77" bold="true">А.</emphasis>
|
||
Early juvenile stage, SHOU 20240103623, from type locality, 12.64 mm SL, pelvic fins completely separated;
|
||
<emphasis id="7A25DA582DB3A562D5B6994471F97019" bold="true">B.</emphasis>
|
||
Sub-adult stage, SHOU 20240103102, paratype, 25.81 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ETRBI" type="Sexual dimorphism">
|
||
<paragraph id="6DFBDDEE1BE1C49D1B00330F46A7432D">
|
||
<heading id="77E11C3E9C7FB9956FC154717D3466A9" reason="title">Sexual dimorphism.</heading>
|
||
</paragraph>
|
||
<paragraph id="86554190D1152D096B70565B879170C6">
|
||
In fully mature individuals, males generally slightly larger than females, with well-developed tubercles (see Fig.
|
||
<figureCitation id="F812B737342F3D6C171B3B5B119824C8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sexual dimorphism within the Beaufortia pingi species group, with males exhibiting well-developed and numerous nuptial tubercles on the head compared to females. A, B. B. viridis sp. nov.; C, D. B. pingi; and E, F. B. granulopinna sp. nov." figureDoi="10.3897/zse.100.124370.figure8" httpUri="https://binary.pensoft.net/fig/1088055">8 A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E4RBI" type="Geographic variation">
|
||
<paragraph id="00A76AFAD5D179CC319EC2B0D0B82863">
|
||
<heading id="A4D4B2BCEF1AD27B22007AAA768A5D20" reason="title">Geographic variation.</heading>
|
||
</paragraph>
|
||
<paragraph id="B5F0BCA35C4BDD8A14E9C92C597EC7A8">
|
||
From east to west, vertical stripes generally tend to become thinner. Specimens from type locality, inter-spacings narrower than dark stripes, sometimes appearing fissure-like (see Fig.
|
||
<figureCitation id="06B4A701B1F6708012E0A14ECE6227C5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lateral (top), dorsal (middle), and ventral (bottom) views of Beaufortia viridis sp. nov., holotype, adult, SHOU 20240103101, 38.03 mm SL; from Wuming River, a stream tributary to the You River of the Pearl River basin, at Wuming District, Nanning City, Guangxi Province, China." figureDoi="10.3897/zse.100.124370.figure10" httpUri="https://binary.pensoft.net/fig/1088057">10</figureCitation>
|
||
). In contrast, Lingyun County populations typically exhibit broader and taller bodies, shorter and thicker caudal peduncles, longer snouts, and wider inter-spacings (see Fig.
|
||
<figureCitation id="BFB6B5A4C49055F7970FBB74C1C26203" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lateral (top), dorsal (middle), and ventral (bottom) views of different regional phenotypes of Beaufortia viridis sp. nov. А. Regional phenotype from Lingyun County, Guangxi, SHOU 20240126104, paratype, 58.71 mm SL; B. Regional phenotype from Tuoniang River, Guangnan County, Yunnan, SHOU 20240109105, paratype, 40.15 mm SL (left); SHOU 20240109103, paratype, 39.36 mm SL (right)." figureDoi="10.3897/zse.100.124370.figure12" httpUri="https://binary.pensoft.net/fig/1088059">12 A</figureCitation>
|
||
, Table
|
||
<tableCitation id="61837874E7CB3B3310DC3EDC55448442" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological variations among populations of Beaufortia viridis sp. nov. and Beaufortia pingi from different distribution areas." httpUri="http://table.plazi.org/id/594A40DDB9649CE0146D4BB42738EE1B" tableUuid="594A40DDB9649CE0146D4BB42738EE1B">4</tableCitation>
|
||
). In populations from the Xiyang River and Tuoniang River, vertical stripes vary greatly, sometimes becoming finer, inter-spacings wider than stripes, few individuals with shorter stripes before caudal peduncle (see Fig.
|
||
<figureCitation id="762CAFDF3BF418B455EFAD37AAE586F0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lateral (top), dorsal (middle), and ventral (bottom) views of different regional phenotypes of Beaufortia viridis sp. nov. А. Regional phenotype from Lingyun County, Guangxi, SHOU 20240126104, paratype, 58.71 mm SL; B. Regional phenotype from Tuoniang River, Guangnan County, Yunnan, SHOU 20240109105, paratype, 40.15 mm SL (left); SHOU 20240109103, paratype, 39.36 mm SL (right)." figureDoi="10.3897/zse.100.124370.figure12" httpUri="https://binary.pensoft.net/fig/1088059">12 B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="4E4594310AA0D2FB2ECF54A4A6A16809" ID-DOI="10.3897/zse.100.124370.figure12" ID-arpha="44DD6FB0-675F-5E7B-AB42-1A84A3EFE953" httpUri="https://binary.pensoft.net/fig/1088059" startId="F12">
|
||
<paragraph id="E5BD2707AEEE25074FB9AD0138CE51CC">
|
||
<label id="5B052F1EFF633B607D259C535A87A440">Figure 12.</label>
|
||
</paragraph>
|
||
<paragraph id="BA77CAF55A158E794E042BE1F2DC36D1">
|
||
Lateral (top), dorsal (middle), and ventral (bottom) views of different regional phenotypes of
|
||
<taxonomicName id="D7261B7276D1E9085FC11035A1A4DC37" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="D90B3E79FEE166BB5D025FE87AE25C64" italics="true">Beaufortia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7A60FC1686890FEFB9331D27B6800C8E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="8AC558DB4A404EB87A99D01ECED4AF18" bold="true">А.</emphasis>
|
||
Regional phenotype from Lingyun County, Guangxi, SHOU 20240126104, paratype, 58.71 mm SL;
|
||
<emphasis id="2324229E9D4CD3796B144A02E874FC2F" bold="true">B.</emphasis>
|
||
Regional phenotype from Tuoniang River, Guangnan County, Yunnan, SHOU 20240109105, paratype, 40.15 mm SL (left); SHOU 20240109103, paratype, 39.36 mm SL (right).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6FE5EF6AE401BD35C44A3257F2592647" ID-DOI="10.3897/zse.100.124370.figure13" ID-arpha="52CF3365-9E46-5F3F-AFDF-77FA63E35F12" httpUri="https://binary.pensoft.net/fig/1088060" startId="F13">
|
||
<paragraph id="78206B5D9D87B16812E578697077CCB3">
|
||
<label id="1E9D47BD5BB4005E080F44B96C2703DA">Figure 13.</label>
|
||
</paragraph>
|
||
<paragraph id="37AAE7A4E09AA148590828CAB0F8591A">
|
||
Collection site of the holotype (SHOU 20240103101) of
|
||
<taxonomicName id="07B170C6C09F4EA65FCB91FD1EF408A6" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="42E74560F050B7299E427D8D8DF4CC7B" italics="true">Beaufortia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4C046B0DEDB011D7716BA096255BF460" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, from Wuming River, a stream tributary to the You River of the Pearl River basin, Wuming District, Nanning City, Guangxi Province, China. Photographed by Jing-Chen Chen on 27 December 2023.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="594A40DDB9649CE0146D4BB42738EE1B" ID-Table-UUID="594A40DDB9649CE0146D4BB42738EE1B" httpUri="http://table.plazi.org/id/594A40DDB9649CE0146D4BB42738EE1B" startId="T4">
|
||
<paragraph id="95A1ABD9664C47F3C9C60100681DFD5D">
|
||
<label id="44575772EE58C6EFC35D38F89E05FD5C">Table 4.</label>
|
||
</paragraph>
|
||
<paragraph id="27D74DEF3DEAAAC1E5C80BD61868134B">
|
||
Morphological variations among populations of
|
||
<taxonomicName id="6512C01BDE032F960BB69E0234EAFF49" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="BC717E20A2728BE199F452A0A9A47E08" italics="true">Beaufortia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="10F296A626C65E2A51D2DF9BDEE2F8DD" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="2F8A9BBBC1D14F568D76416CE2575ADE" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
|
||
<emphasis id="BEE38826C11E817903659A91B98E0B27" italics="true">Beaufortia pingi</emphasis>
|
||
</taxonomicName>
|
||
from different distribution areas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9EA50FFA2973EE9993A4EA3BFFD91068">
|
||
<table id="TID0EY6CI" rules="all">
|
||
<tbody id="0B07C2E7CC9040498D695C6104B697C5">
|
||
<tr id="C75A2D35C4130BC3E0E3F5A0A158C833">
|
||
<th id="0BB1B1D5557CA891280BF307B1955772" colspan="1" rowspan="1">
|
||
<taxonomicName id="398D1434A4A93C3B7A91A92A95EB2D8F" authorityName="Chen & Li & Tang & Pu & Lei" authorityYear="2024" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="CF08E22E59EE9FFEE5284837B3F48355" italics="true">B. viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12291390DA4691A43F9BD00F2CC857E2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="FF4835F99EFD6B062B4B814B9EFAEB44" colspan="2" rowspan="1">Wuming County (N = 16)</th>
|
||
<th id="1C580E3EE5AF79C30813469443B0DCE4" colspan="2" rowspan="1">Linyun County (N = 8)</th>
|
||
<th id="C833207931768E3A3AB7C591E2B0D43F" colspan="2" rowspan="1">Tuoniang River and Xiyang River (N = 12)</th>
|
||
</tr>
|
||
<tr id="7CB7D2E996DCDD3B014579643287D0F3">
|
||
<td id="BCDA18D11C58EBD0EBA33FA00C814858" colspan="1" rowspan="1">Characters</td>
|
||
<td id="934FA0D1B15EA82472639EE38C24E0E7" colspan="1" rowspan="1">Range</td>
|
||
<td id="C76DFBA7261AC8B4FD0092009BF8307F" colspan="1" rowspan="1">Mean ± SD</td>
|
||
<td id="A6017E7C84B36A733FAB041B3030AF33" colspan="1" rowspan="1">Range</td>
|
||
<td id="7C7F0A39B2CD96A38EB8053FCF9B2F39" colspan="1" rowspan="1">Mean ± SD</td>
|
||
<td id="CCEB791220698FECDC2D687F40B868A7" colspan="1" rowspan="1">Range</td>
|
||
<td id="7C419357F7C3B4CDE93ADD31E2B5601A" colspan="1" rowspan="1">Mean ± SD</td>
|
||
</tr>
|
||
<tr id="78B763D7922A0B3B0E34D435F988B702">
|
||
<td id="F2A22F712DA05293386C781C8AED29E5" colspan="1" rowspan="1">Pectoral branched rays</td>
|
||
<td id="385B2C1EC13F767893A9547FA98A9980" colspan="1" rowspan="1">19–21</td>
|
||
<td id="4C098C04087C862443820237D2C5FBE7" colspan="1" rowspan="1">20.25 ± 0.68</td>
|
||
<td id="4884A6154D04CDE046262BC6216E45FA" colspan="1" rowspan="1">21–22</td>
|
||
<td id="59177008A3ED622CEF12BABB51BAC199" colspan="1" rowspan="1">21.37 ± 0.44</td>
|
||
<td id="89952F49CFABFB34F884170BDEFD45B9" colspan="1" rowspan="1">21–23</td>
|
||
<td id="D35C14D762042D223D84F218E5044EEC" colspan="1" rowspan="1">22 ± 0.74</td>
|
||
</tr>
|
||
<tr id="19156EF79B814D6E3AE1D426C616A296">
|
||
<td id="82646F52632D6D2B4C92CE4D88FF721C" colspan="1" rowspan="1">Pelvic branched rays</td>
|
||
<td id="F41B92848863EBCC91D723C5D49E0752" colspan="1" rowspan="1">14.5–17</td>
|
||
<td id="1B7585DCB01B5AC09A584EBDA33D8B6E" colspan="1" rowspan="1">16.5 ± 0.71</td>
|
||
<td id="C26C2EF2750F7A1E620FDA97A56EC39F" colspan="1" rowspan="1">17–19</td>
|
||
<td id="81B26163253118DDDDDB2FBB46EE0833" colspan="1" rowspan="1">17.62 ± 0.74</td>
|
||
<td id="1057DFB93175EE69AAF63B6AE0347CFE" colspan="1" rowspan="1">16–19</td>
|
||
<td id="A006B0487BBB700EA441242FCA443771" colspan="1" rowspan="1">17.58 ± 1.10</td>
|
||
</tr>
|
||
<tr id="E2ED2796A89B0595E30503058494D2A7">
|
||
<td id="AB5E224A8006174061A2279E1D65747D" colspan="7" rowspan="1">Ratio of prepelvic length to</td>
|
||
</tr>
|
||
<tr id="860F86A50226B935CBB8B192075CE1C3">
|
||
<td id="D969CB26EF18E8EE71FC41D16CE448A4" colspan="1" rowspan="1">Head width</td>
|
||
<td id="1CCF5BF7E80201655BE6F0FEEF183EE9" colspan="1" rowspan="1">1.53–1.88</td>
|
||
<td id="7633CE5EE8BCF187CEEAF612AE155493" colspan="1" rowspan="1">1.65 ± 0.08</td>
|
||
<td id="4526A11791110BC6CAAB68B286431074" colspan="1" rowspan="1">1.40–1.65</td>
|
||
<td id="4D1E39C5D654F9E153CBA2808CB84FC8" colspan="1" rowspan="1">
|
||
<emphasis id="983B5B0CA97A634175D8D0741FDB5563" bold="true">1.53 ± 0.08</emphasis>
|
||
*
|
||
</td>
|
||
<td id="84E7B8B6561C4F05665F53166C651A8D" colspan="1" rowspan="1">1.54–1.85</td>
|
||
<td id="1557F7BA83B4C466971E7D1C4CFE5150" colspan="1" rowspan="1">1.67 ± 0.12</td>
|
||
</tr>
|
||
<tr id="25804899CC1E76F275DC01406B9F39B0">
|
||
<td id="E18CF9631B5D73DF6DE804EC72EA9D58" colspan="1" rowspan="1">Pectoral width</td>
|
||
<td id="33309F2CB81AF6B770CCFB40923B7871" colspan="1" rowspan="1">1.01–1.30</td>
|
||
<td id="D5F0E0BC7026556F77DA617CC2423D1F" colspan="1" rowspan="1">1.11 ± 0.07</td>
|
||
<td id="B5B6BF5910FC67F897D020A131F9626D" colspan="1" rowspan="1">0.94–1.03</td>
|
||
<td id="63D74271D7DB92EEDD69D74843B2698C" colspan="1" rowspan="1">
|
||
<emphasis id="39D866A785F389B90B55F9C41C338B40" bold="true">0.99 ± 0.03</emphasis>
|
||
*
|
||
</td>
|
||
<td id="9971856BB08C13ABF3A38061F53F9ECD" colspan="1" rowspan="1">0.96–1.19</td>
|
||
<td id="6E1A5362B352A632991088EF22164758" colspan="1" rowspan="1">1.08 ± 0.07</td>
|
||
</tr>
|
||
<tr id="45919C9D55CB4E94A720010BC897165F">
|
||
<td id="39D4732042BA3D849721661469F29EAB" colspan="1" rowspan="1">Body width</td>
|
||
<td id="77FFDA5BD3A9CA977CC0B0FF2210809C" colspan="1" rowspan="1">1.79–2.63</td>
|
||
<td id="411068CA9D755EF17D9D63FDA9F1FC80" colspan="1" rowspan="1">2.04 ± 0.22</td>
|
||
<td id="E12CAABDB3F502B4F138921F6C611F0B" colspan="1" rowspan="1">1.62–1.82</td>
|
||
<td id="AC3CE6AAA3EC3A7D4798DB1C96149862" colspan="1" rowspan="1">
|
||
<emphasis id="30C1291DD5A7290AED2B79951DE2C8AA" bold="true">1.73 ± 0.07</emphasis>
|
||
*
|
||
</td>
|
||
<td id="72D66BA75CE80177FEA9FADFDCE2070E" colspan="1" rowspan="1">1.79–2.29</td>
|
||
<td id="2B4A91B198725DE9C6F3A3960107F5C7" colspan="1" rowspan="1">1.97 ± 0.15</td>
|
||
</tr>
|
||
<tr id="8170F1977C6F65DA7379AFD286F36C17">
|
||
<td id="92898796029E7CC873497705F9747102" colspan="1" rowspan="1">Body depth</td>
|
||
<td id="A2251EEAA3AAB5B09E983547366A140C" colspan="1" rowspan="1">2.19–2.77</td>
|
||
<td id="5783C04CF727BA15F18418BE8ECB8509" colspan="1" rowspan="1">2.50 ± 0.17</td>
|
||
<td id="1FEE216DF29EED667FDC3242DD028E7E" colspan="1" rowspan="1">1.76–2.20</td>
|
||
<td id="C0694076F946A4436566504997C4028A" colspan="1" rowspan="1">
|
||
<emphasis id="ED99EE9E7C57711B3F81181E721110FF" bold="true">1.98 ± 0.15</emphasis>
|
||
*
|
||
</td>
|
||
<td id="2AAF3C30BE01D300CA4D8BE71D0B1BD8" colspan="1" rowspan="1">2.09–2.65</td>
|
||
<td id="93BEA15B82594370DD93D754EF1B2CA1" colspan="1" rowspan="1">2.34 ± 0.21</td>
|
||
</tr>
|
||
<tr id="D80DDB328949E28E536B6D3C6551F17A">
|
||
<td id="AE4CAAD2383B770AE93F5C5FFE03C59E" colspan="1" rowspan="1">Head depth</td>
|
||
<td id="CDA72B746FD3A2FA7090C7F952DCF520" colspan="1" rowspan="1">2.53–3.27</td>
|
||
<td id="D54E1A7D1302E3212A6F0A0315CBF6D6" colspan="1" rowspan="1">2.89 ± 0.22</td>
|
||
<td id="3E23361CBC5EDE5D7D4C09132D11B5A5" colspan="1" rowspan="1">2.30–2.67</td>
|
||
<td id="F778D9573FBF4C73DF1305641AA4E2DA" colspan="1" rowspan="1">
|
||
<emphasis id="AACDE879D80B002E553E51F48F821618" bold="true">2.48 ± 0.12</emphasis>
|
||
*
|
||
</td>
|
||
<td id="EAF751138DCF3A5DB7E1C63CEC29AA0F" colspan="1" rowspan="1">2.40–3.14</td>
|
||
<td id="818DC6A2D211D0A94D743ED92CF57719" colspan="1" rowspan="1">2.79 ± 0.25</td>
|
||
</tr>
|
||
<tr id="E6123667A5CDD70C510931A619E31E98">
|
||
<td id="1AEFC37FBDC271685174359B6C7071BB" colspan="1" rowspan="1">Caudal peduncle depth</td>
|
||
<td id="88890DD6B57DA66CC3B4300EDE4668AB" colspan="1" rowspan="1">4.07–5.16</td>
|
||
<td id="6F40317862D060ABC17233F53D74235E" colspan="1" rowspan="1">4.54 ± 0.29</td>
|
||
<td id="0E52ED11CE4868BC4E4DF5F7B004C7FB" colspan="1" rowspan="1">3.26–4.11</td>
|
||
<td id="AF914026723CC1BCB49B009410E7E0C2" colspan="1" rowspan="1">
|
||
<emphasis id="03207EA81911864FC686B3BC6308B3FA" bold="true">3.59 ± 0.28</emphasis>
|
||
*
|
||
</td>
|
||
<td id="254009BD28E1D8B2CC3695D66FC4CD2B" colspan="1" rowspan="1">3.90–5.05</td>
|
||
<td id="89458DCBFB80DF0D6E39896B357A738F" colspan="1" rowspan="1">4.55 ± 0.37</td>
|
||
</tr>
|
||
<tr id="9CC5200A0A4F3C5B043AA8340D5FF7E7">
|
||
<td id="E520531DE2915C699D0442C783A9A3E7" colspan="1" rowspan="1">Pelvic width</td>
|
||
<td id="2603770BFD95FC58F0ACFFBD6D904607" colspan="1" rowspan="1">1.31–2.02</td>
|
||
<td id="443F78A22DA70889A0FF88AA386E685B" colspan="1" rowspan="1">1.43 ± 0.17</td>
|
||
<td id="39BB28E5404C666EC57A4BF4F0D1D985" colspan="1" rowspan="1">1.15–1.30</td>
|
||
<td id="2F5DA02BC7CCCFCB918C6FF159BA3A60" colspan="1" rowspan="1">
|
||
<emphasis id="AAB41CB15BBAB7CF52D7A01AC339927F" bold="true">1.25 ± 0.05</emphasis>
|
||
*
|
||
</td>
|
||
<td id="8CCBDA905F38900C442D28A35328D024" colspan="1" rowspan="1">1.21–1.74</td>
|
||
<td id="06CF9C00684E119DB3C064266A12A3B6" colspan="1" rowspan="1">1.44 ± 0.17</td>
|
||
</tr>
|
||
<tr id="EE0EEC9FF32CBBEC8C16989A6B288FC2">
|
||
<td id="B0B465F17E74F1F107D985DA504CE376" colspan="7" rowspan="1">Ratio of head length to</td>
|
||
</tr>
|
||
<tr id="50260835294C0FFC1A7CF9AB4DB49C60">
|
||
<td id="3DCC322E4FD023091F3D31680441DE18" colspan="1" rowspan="1">Snout length</td>
|
||
<td id="865E24DE216D41FD59E8B0DF72660553" colspan="1" rowspan="1">2.23–2.59</td>
|
||
<td id="46DCEEB8A1B2F9FF1F9D5A08E86E9081" colspan="1" rowspan="1">
|
||
<emphasis id="C1EF2B04221FA7FF1BE73D6119083E4E" bold="true">2.40 ± 0.11</emphasis>
|
||
*
|
||
</td>
|
||
<td id="743D379561AABF72846FC4B2FE2C95A1" colspan="1" rowspan="1">1.89–2.12</td>
|
||
<td id="67B70FCD475BBBF9E18831F983F30D88" colspan="1" rowspan="1">
|
||
<emphasis id="1E61C2CCCB3CC8F9F0B5BE2974F0A541" bold="true">2.01 ± 0.08</emphasis>
|
||
*
|
||
</td>
|
||
<td id="2A0A863BA36C79494537612CEDA18563" colspan="1" rowspan="1">1.98–2.43</td>
|
||
<td id="56DA3B91E9419F1B5B05EB3925098BCA" colspan="1" rowspan="1">
|
||
<emphasis id="9D69488035398EF21E5A284024663A47" bold="true">2.19 ± 0.12</emphasis>
|
||
*
|
||
</td>
|
||
</tr>
|
||
<tr id="7CD3497DAFB02CBEEEE4D395D53CD4ED">
|
||
<td id="46BF8B6699729F29EECAA6DD328C5308" colspan="7" rowspan="1">Ratio of postanal length to</td>
|
||
</tr>
|
||
<tr id="DDBA2F1346B064A2B0FCBE4B487C3D9D">
|
||
<td id="90C6F8A759CB498EE6A672D617AA1211" colspan="1" rowspan="1">Caudal peduncle depth</td>
|
||
<td id="6A286F04DCC7AE24D4A804951BF20D46" colspan="1" rowspan="1">1.48–2.08</td>
|
||
<td id="08BF89FB859123DCCF3E0CA274CFA532" colspan="1" rowspan="1">1.10 ± 0.16</td>
|
||
<td id="0C6AF79A828D6BFF7F459BC3B90184CC" colspan="1" rowspan="1">1.17–1.39</td>
|
||
<td id="BF3A6B6D5A2B9E466865CA0EDA991F53" colspan="1" rowspan="1">
|
||
<emphasis id="00C7767C99C5BE78404D841F60B4EEFC" bold="true">1.28 ± 0.08</emphasis>
|
||
*
|
||
</td>
|
||
<td id="73AB42C0787026F2757348FEF02C8439" colspan="1" rowspan="1">1.37–1.94</td>
|
||
<td id="71CB1E973536C8D00629518AAE43B185" colspan="1" rowspan="1">1.70 ± 0.15</td>
|
||
</tr>
|
||
<tr id="0544BE56AA2DBF04C4F0E8D00582332C">
|
||
<td id="3660C7B2E78BFE93DFEF1DE8A4EF9328" colspan="7" rowspan="1">Ratio of prepelvic length to</td>
|
||
</tr>
|
||
<tr id="D45369DEFFDD66003E3D4B8BEDEE02B8">
|
||
<td id="9427F695B5A166ABD92528275C18F314" colspan="1" rowspan="1">Length of pelvic origin to anal origin</td>
|
||
<td id="D4556EA05324A20F38C3380BA80CAB10" colspan="1" rowspan="1">1.06–1.33</td>
|
||
<td id="D782E18E36B82DF4121531B1B36F2B1F" colspan="1" rowspan="1">
|
||
<emphasis id="21409D8F739A509764139CDC5F22BE7C" bold="true">1.19 ± 0.74</emphasis>
|
||
*
|
||
</td>
|
||
<td id="C71A22BA831A97307816C915F4A425C3" colspan="1" rowspan="1">0.95–1.17</td>
|
||
<td id="180E6F4AAD2D9E7BA98351EBBA41F6E2" colspan="1" rowspan="1">1.02 ± 0.71</td>
|
||
<td id="384D3BA6B20C815BCDDDD359A3F342EB" colspan="1" rowspan="1">0.96–1.17</td>
|
||
<td id="AE68A4BFBB8C6CDBF25D9E4D55390311" colspan="1" rowspan="1">1.08 ± 0.58</td>
|
||
</tr>
|
||
<tr id="357FBD6AB209E962377D19CACD7850D0">
|
||
<td id="C01C61FF99595E50184C100B08400F4E" colspan="1" rowspan="1">
|
||
<taxonomicName id="60AC175E10EC1DEED9E40A12D0ACCBA5" baseAuthorityName="Fang" baseAuthorityYear="1930" family="Balitoridae" genus="Beaufortia" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="pingi">
|
||
<emphasis id="01CEBE17647D75E1AE9E1E5E57CD5FFC" italics="true">B. pingi</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="FFDD96CACED58689F8656C6B64910059" colspan="2" rowspan="1">Linyun County & Youjiang County (N = 38)</td>
|
||
<td id="274AE6D31BBF4E266CCBCC8B7A7A63DB" colspan="2" rowspan="1">Debao County (N = 12)</td>
|
||
<td id="399B32E991AD30E1E930BF623EE2B6E7" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="EAE9841CB6F3A1CB54C4DF31774490EA" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
<tr id="7BF9534012A786FE244A4287055147C4">
|
||
<td id="C56397C664027A2AC746654B32BA7EDE" colspan="1" rowspan="1">Characters</td>
|
||
<td id="EAA48EA2238A02778205E689FB587339" colspan="1" rowspan="1">Range</td>
|
||
<td id="A1DB5F1EE6A25743A1EF7E4966900430" colspan="1" rowspan="1">Mean ± SD</td>
|
||
<td id="09B61BC945EF60470268B4A7994F91F5" colspan="1" rowspan="1">Range</td>
|
||
<td id="1EF23B9817B651EE927ACD8E9B8BC5B8" colspan="1" rowspan="1">Mean ± SD</td>
|
||
<td id="C38166C5B21968B59AEBFC24EBC1926E" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="C99A817D1B3016FA41BF7BB596E2E99B" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
<tr id="C6BFF77334C368893FE8D8AC279160AA">
|
||
<td id="1CC256E0C42584D4F02C583FEF5833AD" colspan="1" rowspan="1">Pelvic branched rays</td>
|
||
<td id="69873CAFB713DFD25376B99B83A02F1E" colspan="1" rowspan="1">17.5–21</td>
|
||
<td id="659A0036E05DCE8EF682458355E4219F" colspan="1" rowspan="1">18.95 ± 0.89</td>
|
||
<td id="3CF4AE83CB02F35142D0A027BB41B02A" colspan="1" rowspan="1">17–18.5</td>
|
||
<td id="2C564CFA12270D22B3AC361DD3938055" colspan="1" rowspan="1">17.67 ± 0.39</td>
|
||
<td id="4C7BD1CD48EA5409F9C8C0F95B91BBE6" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="7DE0EC4A862781453072AA52404AD18D" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
<tr id="30EC1D4DF42388DFF828EEE88240884F">
|
||
<td id="96E30C0C2963C94388B9B06EC099B631" colspan="7" rowspan="1">Ratio of pectoral width to</td>
|
||
</tr>
|
||
<tr id="BF1F398EBB1B16A694E80B94BA10E86B">
|
||
<td id="258BBD87A8A8985B8383BC8E81DC8434" colspan="1" rowspan="1">Body depth</td>
|
||
<td id="1BB2B7D76912492DFC2D26B00583A031" colspan="1" rowspan="1">1.67–2.50</td>
|
||
<td id="23B58A1EB865324E57B84295E33230F3" colspan="1" rowspan="1">2.02 ± 0.21</td>
|
||
<td id="42747CB463FEA97379896F1CDA9E5EEC" colspan="1" rowspan="1">1.92–2.49</td>
|
||
<td id="FF7E5F3950FFC61121733F0C22B8CB62" colspan="1" rowspan="1">
|
||
<emphasis id="372008445B2844D808A7EE101F2FA197" bold="true">2.32 ± 0.19</emphasis>
|
||
**
|
||
</td>
|
||
<td id="F79744A113F00645A6610439E930C18E" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="7E562E002E6114B2C001683E36CF8576" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
<tr id="84C26B1C63299051E322208A0CF3C8C2">
|
||
<td id="E4BE6751324220FAE6E798574FE1DFAD" colspan="7" rowspan="1">Ratio of postanal length to</td>
|
||
</tr>
|
||
<tr id="7228C15DE49FD158A00BF0F9B76A3AB9">
|
||
<td id="57CDDCA678DA381713B5FDF79CBA96EC" colspan="1" rowspan="1">Caudal peduncle depth</td>
|
||
<td id="6687115BC56FE39705E2CD16CF8999EB" colspan="1" rowspan="1">1.29–1.77</td>
|
||
<td id="9EA4C03135CD022A5879FD9E956B35A4" colspan="1" rowspan="1">1.50 ± 0.13</td>
|
||
<td id="488687A4D0223A91D1424911794E9A45" colspan="1" rowspan="1">1.58–1.86</td>
|
||
<td id="A4D9477A6217354F05ED172C77CAD0EE" colspan="1" rowspan="1">
|
||
<emphasis id="CCC88D279EB5E8ED1CC26E7A455AD857" bold="true">1.69 ± 0.09</emphasis>
|
||
**
|
||
</td>
|
||
<td id="9AD1459743EF2A05E2E884B89E2A6705" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="58842626F822C405D72ECAF968CF0D7F" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
<tr id="3E1D0349541FE908D324ADD8BFB945AF">
|
||
<td id="FBB428BF4CEB8DCC2A97FC010119252F" colspan="7" rowspan="1">Ratio of prepelvic length to</td>
|
||
</tr>
|
||
<tr id="E88F0DEB1FC1C24A3A8A7D8E5DF12A11">
|
||
<td id="20D007B31E85F05093A26D15D30C81B2" colspan="1" rowspan="1">Length of pelvic origin to anal origin</td>
|
||
<td id="AFBEC628AD306803CA970838BE90D0C3" colspan="1" rowspan="1">0.90–1.27</td>
|
||
<td id="A58EC1BC63D3DA7ADEAFBF8690B8B3CB" colspan="1" rowspan="1">
|
||
<emphasis id="5F996B8783242D18BEE86F092C72207E" bold="true">0.98 ± 0.66</emphasis>
|
||
**
|
||
</td>
|
||
<td id="C9526433451FCCD84C166E063585EB81" colspan="1" rowspan="1">0.97–1.14</td>
|
||
<td id="8BA06213CD5288CFC6FD05EED94EFA21" colspan="1" rowspan="1">1.07 ± 0.52</td>
|
||
<td id="A5424DB6A602CFD02E4A6144D97C4449" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
<td id="D51D68B0D88188658BBFFA51EA2381B6" colspan="1" isEmpty="true" rowspan="1">-</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A8E6ADCC71A2745C1A74A7A650472A47">
|
||
<tableNote id="3D684D02B80C293989F3D690A4B4EA10">“ * ” indicates that, in a one-way ANOVA test, post-hoc comparisons show significant differences in the trait in question from populations in other regions (p <0.05). “ ** ” indicates that, in an independent sample T-test, the trait in question significantly differs from populations in other regions (p <0.01).</tableNote>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EXFCI" type="Individual variation">
|
||
<paragraph id="E5F81082B06516AED9C4D513DBF858ED">
|
||
<heading id="06D9447BBC9B028222154D50ACCF8041" reason="title">Individual variation.</heading>
|
||
</paragraph>
|
||
<paragraph id="AF42E5EF230700AE6112D092CAAE6B7D">
|
||
<specimenCount id="57DA9A171CEA7CD17C14DE35263AF7F6" count="2" type="generic">Two specimens</specimenCount>
|
||
,
|
||
<collectionCode id="88CB6B3101AC300D7D4D8904EFFB525D">SHOU</collectionCode>
|
||
20240109101 and
|
||
<collectionCode id="4DD550C8C588CD866115454EBB89FAA6">SHOU</collectionCode>
|
||
20240112102, from Tuoniang River, Guangnan County,
|
||
<collectingRegion id="26E46B85BBD01D6268ECA7394B51927A" country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
, vertical stripes notably shortened.
|
||
<specimenCount id="EC72937E541307F79DDD950B5733D76A" count="1" type="generic">One specimen</specimenCount>
|
||
,
|
||
<collectionCode id="8AFC3756182347BD13C57F01B950C465">SHOU</collectionCode>
|
||
20240109701, from Chengbi River, Lingyun County,
|
||
<collectingRegion id="E6C2CCCC4B162F091AEC73DE4FADA014" country="China" name="Guangxi">Guangxi</collectingRegion>
|
||
, possesses completely separated pelvic fins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3FCI" type="Ethology">
|
||
<paragraph id="1F44E57F6955EC7A854EC582D8E89BEA">
|
||
<heading id="FA6FCC4A2F4CBF94DB6C26A95F4854D5" reason="title">Ethology.</heading>
|
||
</paragraph>
|
||
<paragraph id="1192B63A44F8B77485B8AE7E1E0880A1">Inhabits shallow streams with rapid currents and smooth pebble substrates that adhere to crevices between stones. Feeds on algae and small invertebrates and consumes mucus from fresh fish carcasses. Exhibits territorial behavior.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBGCI" type="distribution">
|
||
<paragraph id="07FB57E2E09EFB2C500B8F95951E14F0">
|
||
<heading id="3B489006A316FFFB4889EA527AFA0EF7" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="E0621E7064C8E5555D51BC9CDFFBF018">
|
||
Found exclusively in small tributaries from the Tuoniang River, Xiyang River, Leli River, and Wuming River to the You River basin, the upper reaches of the Pearl River system, central
|
||
<collectingRegion id="53325FE53BBE2DA535802607165BB5E9" country="China" name="Guangxi">Guangxi</collectingRegion>
|
||
, and eastern
|
||
<collectingRegion id="401FAAF0D235F6C6DDF1C59FF1B5ECD9" country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCountry id="135E440C2DBA97D5AC6C16E9F13D14CE" name="China">China</collectingCountry>
|
||
(see Fig.
|
||
<figureCitation id="D3768D6CCD859D53132D67444A4FF588" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution map of species within the Beaufortia pingi species group. Light red: B. granulopinna sp. nov.; light green: B. viridis sp. nov.; light blue: B. pingi; light yellow: B. zebroida. Rivers are labeled with their names in Chinese: Qingshui River (清水江), Nanpan River (南盘江), Beipan River (北盘江), Hongshui River (红水河), Tuoniang (驮娘江), Xiyang River (西洋江), Leli River (乐里河), Chengbi River (澄碧河), Bo’ai River (剥隘河), Jian River (鉴河), You River (右江), Wuming River (武鸣河), Yu River (郁江), Shuikou River (水口河, known as Sông B ằng in Vietnamese), and Red River (红河)." figureDoi="10.3897/zse.100.124370.figure9" httpUri="https://binary.pensoft.net/fig/1088056">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELGCI" type="etymology">
|
||
<paragraph id="9FFE8F6A59C8742C42BD324BED18C45F">
|
||
<heading id="F1DF6054CF226D17D86D8CFAC0505D0C" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="46FE1613F0F494EC5FEE31C26FB7D9DA">
|
||
Species epithet “
|
||
<emphasis id="F0388BDD98BF635CDE15A7D42441A20A" italics="true">viridis</emphasis>
|
||
, ” from Latin, meaning “ green, ” describes bright green coloration over dorsal body after sexual maturity, in nominative masculine. We propose the Chinese common specific name “ 绿斑爬岩鳅. ”
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |