506 lines
70 KiB
XML
506 lines
70 KiB
XML
<document id="52602131BD9167444BBFEB836550B894" ID-DOI="10.26879/1294" ID-ISSN="1094-8074" ID-Zenodo-Dep="10962179" ID-ZooBank="62019DB5-8F62-4276-994C-8867124393A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712859367553" checkinUser="felipe" docAuthor="Varela-Hernández, Fernando, Riquelme, Francisco & Guerrero, Roberto J." docDate="2023" docId="CE1F87A5D711FFBAFF394991FF1DFF73" docLanguage="en" docName="PalaeontolElectron.26.3.a36.pdf" docOrigin="Palaeontologia Electronica (a 36) 26 (3)" docSource="http://dx.doi.org/10.26879/1294" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Forelius chenpauch Varela-Hernández & Riquelme & Guerrero 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="3226FFDDD713FFBCFFED4B6AFD0AFFAC" masterDocTitle="A Miocene ant species of the genus Forelius Emery, 1888 (Hymenoptera: Formicidae: Dolichoderinae) from Mexico" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1713374366597" updateUser="felipe" zenodo-license-document="CC-BY-NC-SA-4.0">
|
||
<mods:mods id="1026214D2F13E087EB2F569E1C0E4AD9" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3639B3030311F6A9FF5EE74140A478E7">
|
||
<mods:title id="18F8A554947CE54DBE0D9FDBECFA297F">A Miocene ant species of the genus Forelius Emery, 1888 (Hymenoptera: Formicidae: Dolichoderinae) from Mexico</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="80E78C4141A1BFBACC64338ED37FFAC9" type="personal">
|
||
<mods:role id="3B25BD3C844DCE435B2B0115C0E40277">
|
||
<mods:roleTerm id="E36351F533D5A4BD7C76D78DBBA5148D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8140F96E028D9C5254ADABE19158D907">Varela-Hernández, Fernando</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CD80BA739ED311F643A5BEF7F2C464F4" type="personal">
|
||
<mods:role id="8C6F63538F7A3D4DA9FF82D03DB97D0A">
|
||
<mods:roleTerm id="97E7BECF6230F86BD578064E41236E65">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="82A68E274B3FC03C0B2F497FAE2EE832">Riquelme, Francisco</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="653186AB54428433FD25DAEE4F9DB702" type="personal">
|
||
<mods:role id="812A44D3953F0C2173F47CCAECC41803">
|
||
<mods:roleTerm id="A027BEF4DBE08687243DE35A891DC886">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D75A7FB08C96416085E7A03D927202F2">Guerrero, Roberto J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D67E3BA433E28053734EE3508DD68C6C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7E183330338289DC39393D98A789970B" type="host">
|
||
<mods:titleInfo id="ED824AA200E521CFC9AD20BBDF92D0DA">
|
||
<mods:title id="61F08C3398E71AB3D0F256A4A55913F1">Palaeontologia Electronica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="FACF8F5DD1FE7DFBD0D01DE571994904">
|
||
<mods:date id="179F7D6DBC43F32A8FD651052E69E2C8">2023</mods:date>
|
||
<mods:detail id="C30DE0E0FB5610344571D2C0FBA099B6" type="series">
|
||
<mods:title id="CBF0F33A6FD32790944CBBE895B1D98A">a 36</mods:title>
|
||
</mods:detail>
|
||
<mods:detail id="4040C915CE79B725DC6D840E0D1F0817" type="volume">
|
||
<mods:number id="7D26F272C79C58C9A408F00743AEDBB7">26</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="9427BD1EC59C609CD7363CAC8DE4C1D6" type="issue">
|
||
<mods:number id="733946032762A176F49E280141F05E85">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A12DC7226D2BC78A4071A46365F5A99A" unit="page">
|
||
<mods:start id="517FCF1F6F1E8204AC176CDA97E52A72">1</mods:start>
|
||
<mods:end id="EDF49A7BA4449198C0A03EA53A3F9E5E">9</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="F3C2FD6ACC3D909CCD374223C75F8A80">
|
||
<mods:url id="3DE4AB07D8DD08D9A133CC57F5C97614">http://dx.doi.org/10.26879/1294</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="6E77B8AA4E1170A85FBC4F936BB408DF">journal article</mods:classification>
|
||
<mods:identifier id="63A663642E7AE942721BF28765FBC899" type="DOI">10.26879/1294</mods:identifier>
|
||
<mods:identifier id="D1D4FC5D7551CB7D8AEBCDB1F22ABD9C" type="ISSN">1094-8074</mods:identifier>
|
||
<mods:identifier id="4C4D8CFC2FCC2AD9388FD3C094D74BE9" type="Zenodo-Dep">10962179</mods:identifier>
|
||
<mods:identifier id="25719CDABB5078A73D89C00956E8BA5D" type="ZooBank">62019DB5-8F62-4276-994C-8867124393A6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CE1F87A5D711FFBAFF394991FF1DFF73" LSID="urn:lsid:plazi:treatment:CE1F87A5D711FFBAFF394991FF1DFF73" httpUri="http://treatment.plazi.org/id/CE1F87A5D711FFBAFF394991FF1DFF73" lastPageId="6" lastPageNumber="6" pageId="2" pageNumber="3">
|
||
<subSubSection id="0EAC6538D711FFBEFF394991FF7DFC98" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="460936B3D711FFBEFF394991FF7DFC98" blockId="2.[212,773,763,852]" pageId="2" pageNumber="3">
|
||
<heading id="1D4181DFD711FFBEFF394991FF7DFC98" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="7">
|
||
<taxonomicName id="81B64D30D711FFBEFF394991FCB6FCB8" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[212,444,763,788]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D711FFBEFF394991FCB6FCB8" box="[212,444,763,788]" italics="true" pageId="2" pageNumber="3">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD711FFBEFE294991FF2AFCB8" box="[452,544,763,788]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Varela-Hernández,
|
||
<collectingRegion id="8472F851D711FFBEFE8C4871FCC1FC99" box="[353,459,795,821]" country="Mexico" name="Guerrero" pageId="2" pageNumber="3">Guerrero</collectingRegion>
|
||
and Riquelme
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFE4D4851FFFCFCD6" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="460936B3D711FFBEFE4D4851FF32FCF8" blockId="2.[212,773,763,852]" box="[416,568,827,852]" pageId="2" pageNumber="3">
|
||
(
|
||
<figureCitation id="DE8D2A36D711FFBEFE4B4851FF25FCF8" box="[422,559,827,852]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[198,289,1790,1813]" captionStartId-1="4.[200,291,1752,1775]" captionTargetBox-0="[320,1315,164,1764]" captionTargetBox-1="[304,1334,167,1729]" captionTargetId-0="figure-25@3.[320,1315,164,1765]" captionTargetId-1="figure-13@4.[304,1334,166,1729]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." captionText-1="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi-0="http://doi.org/10.5281/zenodo.10962181" figureDoi-1="http://doi.org/10.5281/zenodo.10962183" httpUri-0="https://zenodo.org/record/10962181/files/figure.png" httpUri-1="https://zenodo.org/record/10962183/files/figure.png" pageId="2" pageNumber="3">Figures 1-2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFF2D480CFFFCFCD6" blockId="2.[192,758,870,890]" box="[192,758,870,890]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D480CFFFCFCD6" bold="true" box="[192,758,870,890]" pageId="2" pageNumber="3">
|
||
zoobank.org/
|
||
<uuid id="32100C66D711FFBEFEAB480CFFFCFCD6" box="[326,758,870,890]" pageId="2" pageNumber="3">795FBFFB-5934-4259-899E-E60A69DA1319</uuid>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFF2D48E1FCD4FBA8" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="460936B3D711FFBEFF2D48E1FCD4FBA8" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D48E1FC47FC08" bold="true" box="[192,333,907,932]" pageId="2" pageNumber="3">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="81B64D30D711FFBEFDD248E1FFCBFC08" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[575,705,907,932]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chenpauch">
|
||
<emphasis id="74C2EAA1D711FFBEFDD248E1FFCBFC08" box="[575,705,907,932]" italics="true" pageId="2" pageNumber="3">chenpauch</emphasis>
|
||
</taxonomicName>
|
||
means “from the amber cave” in the Tzotzil language, a name given by the local people of the Simojovel to the amber mines or pits.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFF2D4F61FF69FB68" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="460936B3D711FFBEFF2D4F61FC64FB28" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4F61FC7BFB88" bold="true" box="[192,369,1035,1060]" pageId="2" pageNumber="3">Type material.</emphasis>
|
||
<materialsCitation id="F6DE3CEED711FFBEFE944F61FC64FB28" collectionCode="CPAL, UAEM" collectorName="de Paleontologia" country="Mexico" location="Coleccion de Paleontologia" pageId="2" pageNumber="3" specimenCount="464" specimenCount-worker="464" stateProvince="Morelos" typeStatus="holotype">
|
||
<typeStatus id="990D8811D711FFBEFE944F61FCE8FB88" box="[377,482,1035,1060]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
<collectionCode id="20A7AE76D711FFBEFE074F61FF3BFB89" box="[490,561,1035,1061]" pageId="2" pageNumber="3">CPAL</collectionCode>
|
||
.
|
||
<specimenCount id="50B0FD3AD711FFBEFDD94F61FFC8FB88" box="[564,706,1035,1061]" count="464" pageId="2" pageNumber="3" type="worker">464 Worker</specimenCount>
|
||
, amber inclusion, complete specimen. It is deposited in the
|
||
<location id="43696068D711FFBEFF2D4F21FF02FBC9" LSID="urn:lsid:plazi:treatment:CE1F87A5D711FFBAFF394991FF1DFF73:43696068D711FFBEFF2D4F21FF02FBC9" box="[192,520,1099,1125]" country="Mexico" name="Coleccion de Paleontologia" pageId="2" pageNumber="3" stateProvince="Morelos">
|
||
Colección
|
||
<collectorName id="EB435365D711FFBEFED14F21FF02FBC9" box="[316,520,1099,1125]" pageId="2" pageNumber="3">de Paleontología</collectorName>
|
||
</location>
|
||
,
|
||
<collectionCode id="20A7AE76D711FFBEFDFE4F21FF6EFBC8" box="[531,612,1099,1124]" country="Mexico" httpUri="http://grbio.org/cool/r76y-vmgd" name="Univeridad Autonoma de Morelos" pageId="2" pageNumber="3">UAEM</collectionCode>
|
||
,
|
||
<collectingRegion id="8472F851D711FFBEFD824F21FFD8FBC8" box="[623,722,1099,1124]" country="Mexico" name="Morelos" pageId="2" pageNumber="3">Morelos</collectingRegion>
|
||
,
|
||
<collectingCountry id="3EA17623D711FFBEFD304F21FDE8FB28" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="DE8D2A36D711FFBEFF1D4F01FC6CFB28" box="[240,358,1131,1156]" captionStart="FIGURE 1" captionStartId="3.[198,289,1790,1813]" captionTargetBox="[320,1315,164,1764]" captionTargetId="figure-25@3.[320,1315,164,1765]" captionTargetPageId="3" captionText="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." figureDoi="http://doi.org/10.5281/zenodo.10962181" httpUri="https://zenodo.org/record/10962181/files/figure.png" pageId="2" pageNumber="3">Figure 1A</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFF2D4FE1FF69FB68" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4FE1FC20FB08" bold="true" box="[192,298,1163,1188]" pageId="2" pageNumber="3">Locality.</emphasis>
|
||
<materialsCitation id="F6DE3CEED711FFBEFEDE4FE1FF55FB68" country="Mexico" latitude="17.129442" location="Montecristo" longLatPrecision="14" longitude="-92.69528" municipality="Simojovel" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Chiapas">
|
||
<collectingCountry id="3EA17623D711FFBEFEDE4FE1FC86FB08" box="[307,396,1163,1188]" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="8472F851D711FFBEFE754FE1FCF4FB09" box="[408,510,1163,1189]" country="Mexico" name="Chiapas" pageId="2" pageNumber="3">Chiapas</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="A66DACC9D711FFBEFDE74FE1FF88FB09" box="[522,642,1163,1189]" pageId="2" pageNumber="3">Simojovel</collectingMunicipality>
|
||
,
|
||
<location id="43696068D711FFBEFD634FE1FE12FB08" LSID="urn:lsid:plazi:treatment:CE1F87A5D711FFBAFF394991FF1DFF73:43696068D711FFBEFD634FE1FE12FB08" box="[654,792,1163,1188]" country="Mexico" latitude="17.129442" longLatPrecision="14" longitude="-92.69528" municipality="Simojovel" name="Montecristo" pageId="2" pageNumber="3" stateProvince="Chiapas">Montecristo</location>
|
||
mine:
|
||
<geoCoordinate id="23825074D711FFBEFEE44FC1FCB7FB68" box="[265,445,1195,1220]" degrees="17" direction="north" minutes="7" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="45.99" value="17.129442">17° 7' 45.99" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="23825074D711FFBEFE244FC1FF55FB68" box="[457,607,1195,1220]" degrees="92" direction="west" minutes="41" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="43" value="-92.69528">92°41'43" W</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFF2D4FA1FCB1FAE8" pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph id="460936B3D711FFBEFF2D4FA1FFECFAA8" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4FA1FC57FB48" bold="true" box="[192,349,1227,1252]" pageId="2" pageNumber="3">Distribution.</emphasis>
|
||
It is only known from the Miocene strata of Simojovel,
|
||
<collectingRegion id="8472F851D711FFBEFE474F81FF1AFAA9" box="[426,528,1259,1285]" country="Mexico" name="Chiapas" pageId="2" pageNumber="3">Chiapas</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="3EA17623D711FFBEFD644F81FFE8FAA8" box="[649,738,1259,1284]" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFF2D4E61FCB1FAE8" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4E61FC20FA88" bold="true" box="[192,298,1291,1316]" pageId="2" pageNumber="3">Horizon.</emphasis>
|
||
The Mazantic and Balumtum strata, Early Miocene (ca. 23 Ma).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFF2D4E21FCB8F998" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="460936B3D711FFBEFF2D4E21FFFFF9A8" blockId="2.[192,792,907,1540]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4E21FC4DFAC8" bold="true" box="[192,327,1355,1380]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
|
||
Antennal scape reaching the posterior margin of the head; pronotum with six erect hairs; mesonotum with four erect hairs; propodeum with four erect thin hairs; front and dorsum of head, dorsum of pronotum, dorsum of mesonotum, and propodeum covered with fine longitudinal striae.
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFF2D4D71FCB8F998" blockId="2.[192,434,1563,1588]" box="[192,434,1563,1588]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4D71FCB8F998" bold="true" box="[192,434,1563,1588]" pageId="2" pageNumber="3">Worker description</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBEFF2D4D21F951FB93" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="460936B3D711FFBEFF2D4D21FE0BF908" blockId="2.[192,792,1611,1892]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4D21FCC4F9C8" bold="true" box="[192,462,1611,1636]" pageId="2" pageNumber="3">Measurements (mm).</emphasis>
|
||
TL: 2.0; HL: 0.6; HW: 0.6; EL: 0.1; EW: 0.1; IOD: 0.4; SL: 0.5; PSL: 0.05; PSW: 0.05; MsL: 0.8; CI: 100; OI: 100; SI: 83.33.
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFF2D4DC1F919FE33" blockId="2.[192,792,1611,1892]" lastBlockId="2.[840,1441,166,1887]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFF2D4DC1FC02F968" bold="true" box="[192,264,1707,1732]" pageId="2" pageNumber="3">Head.</emphasis>
|
||
As long as broad; vertex of head weakly concave; antennal scapes reaching the posterior margin of the head; mandible with a basal and masticator margin differentiated, with six observable teeth in the latter (
|
||
<figureCitation id="DE8D2A36D711FFBEFE264C41FF4FF8E8" box="[459,581,1835,1860]" captionStart="FIGURE 2" captionStartId="4.[200,291,1752,1775]" captionTargetBox="[304,1334,167,1729]" captionTargetId="figure-13@4.[304,1334,166,1729]" captionTargetPageId="4" captionText="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi="http://doi.org/10.5281/zenodo.10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="2" pageNumber="3">Figure 2B</figureCitation>
|
||
); clypeus with two slender, erect setae projecting onto the mandibles at the middle part, about the level of the antennal insertions; two other shorter setae on each side of the erect setae described previously: one at the lateral edge of the mandible and another between the latter and the one at the middle of the mandible; the dorsal face of mandibles covered with decumbent pilosity; dorsum of head finely striated (
|
||
<figureCitation id="DE8D2A36D711FFBEFA9D4A0CFEACFE33" captionStart="FIGURE 1" captionStartId="3.[198,289,1790,1813]" captionTargetBox="[320,1315,164,1764]" captionTargetId="figure-25@3.[320,1315,164,1765]" captionTargetPageId="3" captionText="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." figureDoi="http://doi.org/10.5281/zenodo.10962181" httpUri="https://zenodo.org/record/10962181/files/figure.png" pageId="2" pageNumber="3">Figures 1B</figureCitation>
|
||
,
|
||
<figureCitation id="DE8D2A36D711FFBEFC5E4AECF90DFE33" box="[947,1031,390,415]" captionStart="FIGURE 2" captionStartId="4.[200,291,1752,1775]" captionTargetBox="[304,1334,167,1729]" captionTargetId="figure-13@4.[304,1334,166,1729]" captionTargetPageId="4" captionText="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi="http://doi.org/10.5281/zenodo.10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="2" pageNumber="3">2A, 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFCA54ACCFEC6FCD3" blockId="2.[840,1441,166,1887]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFCA54ACCFED3FE13" bold="true" box="[840,985,422,447]" pageId="2" pageNumber="3">Mesosoma.</emphasis>
|
||
Dorsal profile discontinuous. In dorsal view, well-impressed promesonotal suture; weak mesonotal groove in lateral view; pronotum with six thin, erect hairs, four located anteriorly, longer than the other two located posteriorly, two at the anteriormost humeral and the other two at the middle; dorsal surface of pronotum and mesonotum with weak transversal striae; dorsal and posterior face of propodeum rounded, with no apparent angle that differentiates the two faces; one pair of thin, erect hairs located dorsolaterally on the propodeum dorsal face and another pair posterolaterally on the propodeum posterior face; propodeal spiracle small and rounded (
|
||
<figureCitation id="DE8D2A36D711FFBEFB8F482CF81AFCF3" box="[1122,1296,838,863]" captionStart="FIGURE 2" captionStartId="4.[200,291,1752,1775]" captionTargetBox="[304,1334,167,1729]" captionTargetId="figure-13@4.[304,1334,166,1729]" captionTargetPageId="4" captionText="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi="http://doi.org/10.5281/zenodo.10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="2" pageNumber="3">Figure 2A, 2B</figureCitation>
|
||
). Legs without pilosity.
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFCA548ECF9C4FC13" blockId="2.[840,1441,166,1887]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFCA548ECFEACFC33" bold="true" box="[840,934,902,927]" pageId="2" pageNumber="3">Petiole.</emphasis>
|
||
Scale present, inclined anteriorly, with a pair of erect setae, barely visible.
|
||
</paragraph>
|
||
<paragraph id="460936B3D711FFBEFCA548ACF951FB93" blockId="2.[840,1441,166,1887]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFCA548ACFEA8FC4C" bold="true" box="[840,930,966,992]" pageId="2" pageNumber="3">Gaster.</emphasis>
|
||
First and second tergite with a row of short, thin, erect setae running along the apical fringe of tergites, sternites lost; gaster concolorous yellow with mesosoma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EAC6538D711FFBAFCA54F2CFF1DFF73" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="discussion">
|
||
<paragraph id="460936B3D711FFBEFCA54F2CF89AF8F3" blockId="2.[840,1441,166,1887]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFCA54F2CFECAFBF3" bold="true" box="[840,960,1094,1119]" pageId="2" pageNumber="3">Remarks.</emphasis>
|
||
The worker (
|
||
<typeStatus id="990D8811D711FFBEFB894F2CF9DAFBF3" box="[1124,1232,1094,1119]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
<collectionCode id="20A7AE76D711FFBEFB374F2CF829FBCC" box="[1242,1315,1094,1120]" pageId="2" pageNumber="3">CPAL</collectionCode>
|
||
.464) studied here is classified in the genus
|
||
<taxonomicName id="81B64D30D711FFBEFB094F0CF849FBD3" authorityName="Emery" authorityYear="1888" box="[1252,1347,1126,1151]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D711FFBEFB094F0CF849FBD3" box="[1252,1347,1126,1151]" italics="true" pageId="2" pageNumber="3">Forelius</emphasis>
|
||
</taxonomicName>
|
||
since it presents the diagnostic characters of the genus, that is, the anterior clypeal margin with two long curved setae is observed in the specimen, which, although in the specimen the mandible is open, these clypeal hairs project reaching the external margin of the mandible. The configuration of the mandible and the dentition pattern on the masticatory margin agree with that of
|
||
<taxonomicName id="81B64D30D711FFBEFB444E0CF802FAD3" authorityName="Emery" authorityYear="1888" box="[1193,1288,1382,1407]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D711FFBEFB444E0CF802FAD3" box="[1193,1288,1382,1407]" italics="true" pageId="2" pageNumber="3">Forelius</emphasis>
|
||
</taxonomicName>
|
||
according to
|
||
<bibRefCitation id="22274B42D711FFBEFCA54EECF907FA33" author="Shattuck" box="[840,1037,1414,1440]" firstAuthor="Shattuck" pageId="2" pageNumber="3" refId="ref5427" refString="Shattuck, S. O. 1992. Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21 (1): 1 - 181." year="1992">Shattuck (1992)</bibRefCitation>
|
||
, that is, mandible with a smooth basal and masticatory margin differentiated, the latter armed with 6-7 teeth. The petiolar scale is reduced and strongly inclined anteriorly as well, however, in the specimen it appears relatively straight (
|
||
<figureCitation id="DE8D2A36D711FFBEFC5D4D4CF937F993" box="[944,1085,1574,1599]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[198,289,1790,1813]" captionStartId-1="4.[200,291,1752,1775]" captionTargetBox-0="[320,1315,164,1764]" captionTargetBox-1="[304,1334,167,1729]" captionTargetId-0="figure-25@3.[320,1315,164,1765]" captionTargetId-1="figure-13@4.[304,1334,166,1729]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." captionText-1="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi-0="http://doi.org/10.5281/zenodo.10962181" figureDoi-1="http://doi.org/10.5281/zenodo.10962183" httpUri-0="https://zenodo.org/record/10962181/files/figure.png" httpUri-1="https://zenodo.org/record/10962183/files/figure.png" pageId="2" pageNumber="3">Figures 1-2</figureCitation>
|
||
), but it is due to the position within the amber piece. The latter also makes it difficult to observe the first gastral segment projecting anteriorly and concealing petiole in dorsal view; however, an inclined view of the piece allowed us to observe a shallow depression on the anterior face of the first gastral tergite.
|
||
<taxonomicName id="81B64D30D711FFBEFB5A4D8CF8AAF953" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[1207,1440,1766,1791]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D711FFBEFB5A4D8CF8AAF953" box="[1207,1440,1766,1791]" italics="true" pageId="2" pageNumber="3">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD711FFBEFCA54C6CFEA2F8B3" box="[840,936,1798,1823]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a species with a rounded spiracle but differs from other
|
||
<taxonomicName id="81B64D30D711FFBEFBCF4C4CF9ECF893" box="[1058,1254,1830,1855]" pageId="2" pageNumber="3">
|
||
<emphasis id="74C2EAA1D711FFBEFBCF4C4CF98BF893" box="[1058,1153,1830,1855]" italics="true" pageId="2" pageNumber="3">Forelius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with this same condition by the characters listed below (
|
||
<tableCitation id="0B340308D711FFBEFACA4C2CF877F8F3" box="[1319,1405,1862,1887]" captionStart="TABLE 1" captionStartId="5.[188,268,164,187]" captionTargetPageId="5" captionText="TABLE 1. Morphological comparison between Forelius chenpauch sp. nov. and the other Forelius species with a rounded spiracle present in the New World." pageId="2" pageNumber="3">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="12C9663BD710FFBFFF2B4D94F995F8C2" ID-DOI="http://doi.org/10.5281/zenodo.10962181" ID-Zenodo-Dep="10962181" httpUri="https://zenodo.org/record/10962181/files/figure.png" pageId="3" pageNumber="4" startId="3.[198,289,1790,1813]" targetBox="[320,1315,164,1764]" targetPageId="3" targetType="figure">
|
||
<paragraph id="460936B3D710FFBFFF2B4D94F995F8C2" blockId="3.[198,1441,1790,1902]" pageId="3" pageNumber="4">
|
||
<emphasis id="74C2EAA1D710FFBFFF2B4D94FC53F8B9" bold="true" box="[198,345,1790,1813]" pageId="3" pageNumber="4">FIGURE 1. A</emphasis>
|
||
<taxonomicName id="81B64D30D710FFBFFE8E4D94FF3DF8B9" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[355,567,1790,1813]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D710FFBFFE8E4D94FF3DF8B9" box="[355,567,1790,1813]" italics="true" pageId="3" pageNumber="4">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD710FFBFFDAC4D94FF9DF8B9" box="[577,663,1790,1813]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc).
|
||
<emphasis id="74C2EAA1D710FFBFFE7C4C53FCABF8FC" bold="true" box="[401,417,1849,1872]" pageId="3" pageNumber="4">B</emphasis>
|
||
<taxonomicName id="81B64D30D710FFBFFE474C53FF77F8FC" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[426,637,1849,1872]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D710FFBFFE474C53FF77F8FC" box="[426,637,1849,1872]" italics="true" pageId="3" pageNumber="4">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD710FFBFFD694C53FFD3F8FC" box="[644,729,1849,1872]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="12C9663BD717FFB8FF254DB2FF19F8C9" ID-DOI="http://doi.org/10.5281/zenodo.10962183" ID-Zenodo-Dep="10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="4" pageNumber="5" startId="4.[200,291,1752,1775]" targetBox="[304,1334,167,1729]" targetPageId="4" targetType="figure">
|
||
<paragraph id="460936B3D717FFB8FF254DB2FF19F8C9" blockId="4.[200,1434,1752,1893]" pageId="4" pageNumber="5">
|
||
<emphasis id="74C2EAA1D717FFB8FF254DB2FC5FF95C" bold="true" box="[200,341,1752,1776]" pageId="4" pageNumber="5">FIGURE 2. A</emphasis>
|
||
<taxonomicName id="81B64D30D717FFB8FEB14DB3FF26F95C" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[348,556,1753,1776]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D717FFB8FEB14DB3FF26F95C" box="[348,556,1753,1776]" italics="true" pageId="4" pageNumber="5">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD717FFB8FDDE4DB3FF8CF95C" box="[563,646,1753,1776]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc).
|
||
<emphasis id="74C2EAA1D717FFB8FF104C79FC04F886" bold="true" box="[253,270,1811,1834]" pageId="4" pageNumber="5">B</emphasis>
|
||
<taxonomicName id="81B64D30D717FFB8FEF84C79FCECF886" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[277,486,1811,1834]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D717FFB8FEF84C79FCECF886" box="[277,486,1811,1834]" italics="true" pageId="4" pageNumber="5">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD717FFB8FE004C79FF4AF886" box="[493,576,1811,1834]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="12C9663BD716FFB9FF514BCEFF82FF75" pageId="5" pageNumber="6" startId="5.[188,268,164,187]" targetBox="[192,1435,239,1137]" targetIsTable="true" targetPageId="5" targetType="table">
|
||
<paragraph id="460936B3D716FFB9FF514BCEFF82FF75" blockId="5.[188,1444,164,217]" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FF514BCEFC21FF17" bold="true" box="[188,299,164,187]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
|
||
Morphological comparison between
|
||
<taxonomicName id="81B64D30D716FFB9FD2C4BCEFE9CFF17" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[705,918,164,187]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D716FFB9FD2C4BCEFE9CFF17" box="[705,918,164,187]" italics="true" pageId="5" pageNumber="6">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD716FFB9FC4C4BCEFEF3FF17" box="[929,1017,164,187]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and the other
|
||
<taxonomicName id="81B64D30D716FFB9FB494BCEF85CFF17" box="[1188,1366,164,187]" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FB494BCEF9F3FF17" box="[1188,1273,164,187]" italics="true" pageId="5" pageNumber="6">Forelius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with a rounded spiracle present in the New World.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="460936B3D716FFB9FF2D4B85F83DFBDD" pageId="5" pageNumber="6">
|
||
<table id="34B6C413D7160043FF2D4B85F891FBDD" box="[192,1435,239,1137]" gridcols="5" gridrows="8" pageId="5" pageNumber="6">
|
||
<tr id="F88634F1D7160043FF2D4B85F891FEAF" box="[192,1435,239,259]" gridrow="0" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4B85FC6DFEAF" box="[192,359,239,259]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FF2D4B85FC6DFEAF" bold="true" box="[192,359,239,259]" pageId="5" pageNumber="6">Taxon/Character</emphasis>
|
||
</th>
|
||
<th id="BB575D8DD7160043FE9C4B85FF7EFEAF" box="[369,628,239,259]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FE5D4B85FF3FFEAF" bold="true" box="[432,565,239,259]" pageId="5" pageNumber="6">Scape length</emphasis>
|
||
</th>
|
||
<th id="BB575D8DD7160043FD6C4B85FE58FEAF" box="[641,850,239,259]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FD6C4B85FE58FEAF" bold="true" box="[641,850,239,259]" pageId="5" pageNumber="6">Mandibular dentition</emphasis>
|
||
</th>
|
||
<th id="BB575D8DD7160043FCB24B85F97CFEAF" box="[863,1142,239,259]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FC9C4B85F96BFEAF" bold="true" box="[881,1121,239,259]" pageId="5" pageNumber="6">Hairs on the mesosoma</emphasis>
|
||
</th>
|
||
<th id="BB575D8DD7160043FB6F4B85F891FEAF" box="[1154,1435,239,259]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">
|
||
<emphasis id="74C2EAA1D716FFB9FB594B85F863FEAF" bold="true" box="[1204,1385,239,259]" pageId="5" pageNumber="6">Mesosoma profile</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D4A7DF891FED7" box="[192,1435,279,379]" gridrow="1" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4A7DFC6DFED7" box="[192,359,279,379]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B4A7DFC24FEEA" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[198,302,279,326]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B4A7DFC24FEEA" box="[198,302,279,326]" italics="true" pageId="5" pageNumber="6">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD716FFB9FED94A58FDE4FECD" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C4A7DFF7EFED7" box="[369,628,279,379]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">Reaching the posterior margin of the head (SI= 0.83)</td>
|
||
<td id="BB575D8DD7160043FD6C4A7DFE58FED7" box="[641,850,279,379]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">Six teeth and no denticles</td>
|
||
<td id="BB575D8DD7160043FCB24A7DF97CFED7" box="[863,1142,279,379]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">Six on the pronotum, mesonotum with four erect hairs, and propodeum with four erect ones.</td>
|
||
<td id="BB575D8DD7160043FB6F4A7DF891FED7" box="[1154,1435,279,379]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">Discontinuous, promesonotum convex, differentiated from the anterior face of the propodeum</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D4AE7F891FDA7" box="[192,1435,397,523]" gridrow="2" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4AE7FC6DFDA7" box="[192,359,397,523]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B4AE7FC4AFE7A" authority="(Forel, 1904) Exceeding" authorityName="Exceeding" baseAuthorityName="Forel" baseAuthorityYear="1904" box="[198,320,397,470]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="antarcticus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B4AE7FC27FE17" box="[198,301,397,443]" italics="true" pageId="5" pageNumber="6">Forelius antarcticus</emphasis>
|
||
(Forel, 1904)
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C4AE7FF7EFDA7" box="[369,628,397,523]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">Exceeding the posterior margin of the head capsule (SI = 0.98-1.04)</td>
|
||
<td id="BB575D8DD7160043FD6C4AE7FE58FDA7" box="[641,850,397,523]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">Five teeth and no denticles</td>
|
||
<td id="BB575D8DD7160043FCB24AE7F97CFDA7" box="[863,1142,397,523]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">Two on the pronotum, no erect hairs elsewhere on mesosoma</td>
|
||
<td id="BB575D8DD7160043FB6F4AE7F891FDA7" box="[1154,1435,397,523]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">Discontinuous, promesonotum almost continuous, slightly differentiated from the anterior face of the propodeum</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D4976F891FDCA" box="[192,1435,540,614]" gridrow="3" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4976FC6DFDCA" box="[192,359,540,614]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B4977FC68FD9D" box="[198,354,541,561]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="damiani">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B4977FC68FD9D" box="[198,354,541,561]" italics="true" pageId="5" pageNumber="6">Forelius damiani</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="22274B42D716FFB9FF2B495DFC6CFDCA" author="Guerrero and Fernandez" firstAuthor="Guerrero" pageId="5" pageNumber="6" pagination="51 - 60" refId="ref4681" refString="Guerrero, R. J. and Fernandez, F. 2008. A new species of the ant genus Forelius (Formicidae: Dolichoderinae) from the dry forest of Colombia. Zootaxa, 1958: 51 - 60. https: // doi. org / 10.11646 / zootaxa. 1958.1.5" year="2008">Guerrero and Fernández, 2008</bibRefCitation>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C4976FF7EFDCA" box="[369,628,540,614]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">Short, reaching posterior margin of head (SI= 0.95 – 1.32)</td>
|
||
<td id="BB575D8DD7160043FD6C4976FE58FDCA" box="[641,850,540,614]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">Four teeth and two denticles</td>
|
||
<td id="BB575D8DD7160043FCB24976F97CFDCA" box="[863,1142,540,614]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">Two on the pronotum, no erect hairs elsewhere on mesosoma</td>
|
||
<td id="BB575D8DD7160043FB6F4976F891FDCA" box="[1154,1435,540,614]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">Straight, pronotum in profile relatively low and flat</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D491DF891FD5A" box="[192,1435,631,758]" gridrow="4" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D491DFC6DFD5A" box="[192,359,631,758]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B491DFC4AFD0A" authority="(Forel, 1902) Short" authorityName="Short" baseAuthorityName="Forel" baseAuthorityYear="1902" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B491DFC5BFD27" box="[198,337,631,651]" italics="true" pageId="5" pageNumber="6">Forelius heyeri</emphasis>
|
||
(Forel, 1902)
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C491DFF7EFD5A" box="[369,628,631,758]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">Short, well below or barely reaching the posterior cephalic margin by a length less than the width of the pedicel (SI= 0.8 – 0.91).</td>
|
||
<td id="BB575D8DD7160043FD6C491DFE58FD5A" box="[641,850,631,758]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">Five teeth and no denticles</td>
|
||
<td id="BB575D8DD7160043FCB2491DF97CFD5A" box="[863,1142,631,758]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">Two on the pronotum, no erect hairs elsewhere on mesosoma</td>
|
||
<td id="BB575D8DD7160043FB6F491DF891FD5A" box="[1154,1435,631,758]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">Continuous, slightly flat; promesonotum continuous and slightly differentiated from the propodeum</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D486DF891FCC7" box="[192,1435,775,875]" gridrow="5" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D486DFC6DFCC7" box="[192,359,775,875]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B486DFC45FC9A" authority="Wheeler, 1934 Exceeding" authorityName="Wheeler" authorityYear="1934" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="keiferi">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B486DFC5AFCB7" box="[198,336,775,795]" italics="true" pageId="5" pageNumber="6">Forelius keiferi</emphasis>
|
||
Wheeler, 1934
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C486DFF7EFCC7" box="[369,628,775,875]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">Exceeding the posterior cephalic margin by 1/5 of its length (SI= 0.98 – 1.05)</td>
|
||
<td id="BB575D8DD7160043FD6C486DFE58FCC7" box="[641,850,775,875]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">Five teeth and one denticle</td>
|
||
<td id="BB575D8DD7160043FCB2486DF97CFCC7" box="[863,1142,775,875]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">Two to three on the pronotum</td>
|
||
<td id="BB575D8DD7160043FB6F486DF891FCC7" box="[1154,1435,775,875]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">Discontinuous, strongly convex promesonotum differentiated from anterior face of propodeum</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D4816F891FC57" box="[192,1435,892,1019]" gridrow="6" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4816FC6DFC57" box="[192,359,892,1019]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B4817FC57FC07" authority="(McCook, 1880) Exceeding" authorityName="Exceeding" baseAuthorityName="McCook" baseAuthorityYear="1880" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mccooki">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B4817FC69FC3D" box="[198,355,893,913]" italics="true" pageId="5" pageNumber="6">Forelius mccooki</emphasis>
|
||
(McCook, 1880)
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C4816FF7EFC57" box="[369,628,892,1019]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">Exceeding the posterior cephalic margin by 1/5 of its length (SI= 0.98 – 1.05)</td>
|
||
<td id="BB575D8DD7160043FD6C4816FE58FC57" box="[641,850,892,1019]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">Four teeth and three to four denticles</td>
|
||
<td id="BB575D8DD7160043FCB24816F97CFC57" box="[863,1142,892,1019]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">Pronotum usually with six erect setae, rest of mesosomal dorsum with bearing more than 10 erect setae</td>
|
||
<td id="BB575D8DD7160043FB6F4816F891FC57" box="[1154,1435,892,1019]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">Continuous, promesonotum continuous and slightly differentiated from anterior face of propodeum</td>
|
||
</tr>
|
||
<tr id="F88634F1D7160043FF2D4F66F891FBDD" box="[192,1435,1036,1137]" gridrow="7" pageId="5" pageNumber="6">
|
||
<th id="BB575D8DD7160043FF2D4F66FC6DFBDD" box="[192,359,1036,1137]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF2B4F67FC42FBFA" authority="(Roger, 1863) Exceeding" authorityName="Exceeding" baseAuthorityName="Roger" baseAuthorityYear="1863" box="[198,328,1037,1110]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2B4F67FC28FB97" box="[198,290,1037,1083]" italics="true" pageId="5" pageNumber="6">Forelius pruinosus</emphasis>
|
||
(Roger, 1863)
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BB575D8DD7160043FE9C4F66FF7EFBDD" box="[369,628,1036,1137]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">Exceeding the posterior cephalic margin by 1/5 of its length (SI= 1.07 – 1.14)</td>
|
||
<td id="BB575D8DD7160043FD6C4F66FE58FBDD" box="[641,850,1036,1137]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">Five teeth and one or two denticles</td>
|
||
<td id="BB575D8DD7160043FCB24F66F97CFBDD" box="[863,1142,1036,1137]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">Pronotum usually with four erect setae, rest of mesosomal dorsum with bearing 2 to 6 erect setae</td>
|
||
<td id="BB575D8DD7160043FB6F4F66F891FBDD" box="[1154,1435,1036,1137]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">Discontinuous, strongly convex promesonotum differentiated from strongly convex propodeum</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="460936B3D716FFB9FF174FCEFC0EF911" blockId="5.[192,793,1188,1917]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="81B64D30D716FFB9FF174FCEFCE3FB11" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[250,489,1188,1213]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D716FFB9FF174FCEFCE3FB11" box="[250,489,1188,1213]" italics="true" pageId="5" pageNumber="6">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD716FFB9FE154FCEFF51FB11" box="[504,603,1188,1213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
could be confused with extant dolichoderine ants of the genus
|
||
<taxonomicName id="81B64D30D716FFB9FF2D4F8EFCE8FB51" authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" box="[192,482,1252,1277]" class="Insecta" family="Formicidae" genus="Liometopum" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2D4F8EFC59FB51" box="[192,339,1252,1277]" italics="true" pageId="5" pageNumber="6">Liometopum</emphasis>
|
||
Mayr, 1861
|
||
</taxonomicName>
|
||
and the genus
|
||
<taxonomicName id="81B64D30D716FFB9FD494F8EFC7CFAB1" authority="Foerster, 1850" authorityName="Foerster" authorityYear="1850" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FD494F8EFE12FB51" box="[676,792,1252,1277]" italics="true" pageId="5" pageNumber="6">Tapinoma</emphasis>
|
||
Foerster, 1850
|
||
</taxonomicName>
|
||
, which can currently be found in
|
||
<collectingCountry id="3EA17623D716FFB9FF2D4E4EFC13FA91" box="[192,281,1316,1341]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
|
||
. However,
|
||
<taxonomicName id="81B64D30D716FFB9FE4F4E4EFF84FA91" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[418,654,1316,1341]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D716FFB9FE4F4E4EFF84FA91" box="[418,654,1316,1341]" italics="true" pageId="5" pageNumber="6">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD716FFB9FD774E4EFFF0FA91" box="[666,762,1316,1341]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is distinguished by having a mandible with a basal margin distinct from the masticatory margin (
|
||
<figureCitation id="DE8D2A36D716FFB9FD054E0EFC63FA31" captionStart="FIGURE 1" captionStartId="3.[198,289,1790,1813]" captionTargetBox="[320,1315,164,1764]" captionTargetId="figure-25@3.[320,1315,164,1765]" captionTargetPageId="3" captionText="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." figureDoi="http://doi.org/10.5281/zenodo.10962181" httpUri="https://zenodo.org/record/10962181/files/figure.png" pageId="5" pageNumber="6">Figures 1A, 1B</figureCitation>
|
||
,
|
||
<figureCitation id="DE8D2A36D716FFB9FE694EEEFCA3FA31" box="[388,425,1412,1437]" captionStart="FIGURE 2" captionStartId="4.[200,291,1752,1775]" captionTargetBox="[304,1334,167,1729]" captionTargetId="figure-13@4.[304,1334,166,1729]" captionTargetPageId="4" captionText="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi="http://doi.org/10.5281/zenodo.10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="5" pageNumber="6">2B</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="81B64D30D716FFB9FDA94EEEFFDDFA31" authorityName="Mayr" authorityYear="1861" box="[580,727,1412,1437]" class="Insecta" family="Formicidae" genus="Liometopum" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FDA94EEEFFDDFA31" box="[580,727,1412,1437]" italics="true" pageId="5" pageNumber="6">Liometopum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="81B64D30D716FFB9FF2D4ECEFC3EFA11" authorityName="Foerster" authorityYear="1850" box="[192,308,1444,1469]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2D4ECEFC3EFA11" box="[192,308,1444,1469]" italics="true" pageId="5" pageNumber="6">Tapinoma</emphasis>
|
||
</taxonomicName>
|
||
have an undifferentiated basal margin. In addition,
|
||
<taxonomicName id="81B64D30D716FFB9FEA74EAEFF38FA71" authority="Varela-Hernández & Riquelme & Guerrero, 2023" authorityName="Varela-Hernández & Riquelme & Guerrero" authorityYear="2023" box="[330,562,1476,1501]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chenpauch" status="sp. nov.">
|
||
<emphasis id="74C2EAA1D716FFB9FEA74EAEFF38FA71" box="[330,562,1476,1501]" italics="true" pageId="5" pageNumber="6">Forelius chenpauch</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6FF157DAD716FFB9FDD44EAEFF9FFA71" box="[569,661,1476,1501]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a petiole scale anteriorly inclined (
|
||
<figureCitation id="DE8D2A36D716FFB9FDE64E8EFF9BFA51" box="[523,657,1508,1533]" captionStart="FIGURE 1" captionStartId="3.[198,289,1790,1813]" captionTargetBox="[320,1315,164,1764]" captionTargetId="figure-25@3.[320,1315,164,1765]" captionTargetPageId="3" captionText="FIGURE 1. A Forelius chenpauch sp. nov. Holotype CPAL.464. Dorsolateral view of complete specimen showing: compound eye (ey); foreleg (fl); gaster (ga); mandible (mn), mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); scape (sc). B Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation showing: clypeus (clv); compound eye (ey); mandible (mn); mesonotum (mnt); pronotum (pn); scape (sc)." figureDoi="http://doi.org/10.5281/zenodo.10962181" httpUri="https://zenodo.org/record/10962181/files/figure.png" pageId="5" pageNumber="6">Figures 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DE8D2A36D716FFB9FD764E8EFFE5FA51" box="[667,751,1508,1533]" captionStart="FIGURE 2" captionStartId="4.[200,291,1752,1775]" captionTargetBox="[304,1334,167,1729]" captionTargetId="figure-13@4.[304,1334,166,1729]" captionTargetPageId="4" captionText="FIGURE 2. A Forelius chenpauch sp. nov. Holotype CPAL.464. Microphotograph augmentation of dorsolateral showing: head (h); foreleg (fl); gaster (ga); mesonotum (mnt); petiole (p); pronotum (pn); propodeal spiracle (ps); scape (sc). B Forelius chenpauch sp. nov. Drawing of Holotype CPAL.464 showing: clypeus (clv); compound eye (ey); foreleg (fl); gaster (ga); mandible (mn); mesonotum (mnt); petiole (p); pronotum (pn); propodeum (ppm); propodeal spiracle (ps); scape (sc); striae (str)." figureDoi="http://doi.org/10.5281/zenodo.10962183" httpUri="https://zenodo.org/record/10962183/files/figure.png" pageId="5" pageNumber="6">2A, 2B</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName id="81B64D30D716FFB9FEC24D6EFCC8F9B1" authorityName="Mayr" authorityYear="1861" box="[303,450,1540,1565]" class="Insecta" family="Formicidae" genus="Liometopum" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FEC24D6EFCC8F9B1" box="[303,450,1540,1565]" italics="true" pageId="5" pageNumber="6">Liometopum</emphasis>
|
||
</taxonomicName>
|
||
has a vertical petiolar scale, that is, never anteriorly inclined; and in the currently known Neotropical species of
|
||
<taxonomicName id="81B64D30D716FFB9FD984D2EFFE3F9F1" authorityName="Foerster" authorityYear="1850" box="[629,745,1604,1629]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FD984D2EFFE3F9F1" box="[629,745,1604,1629]" italics="true" pageId="5" pageNumber="6">Tapinoma</emphasis>
|
||
</taxonomicName>
|
||
the petiolar scale is strongly reduced, in most hidden by the first gastral segment (
|
||
<collectingRegion id="8472F851D716FFB9FDFE4DEEFF8AF932" box="[531,640,1668,1694]" country="Mexico" name="Guerrero" pageId="5" pageNumber="6">Guerrero</collectingRegion>
|
||
unpublished data).
|
||
</paragraph>
|
||
<paragraph id="460936B3D716FFBAFF174DAEFF1DFF73" blockId="5.[192,793,1188,1917]" lastBlockId="6.[192,792,166,223]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
On the other hand, the genus
|
||
<taxonomicName id="81B64D30D716FFB9FDB74DAEFFB3F971" authorityName="Emery" authorityYear="1888" box="[602,697,1732,1757]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FDB74DAEFFB3F971" box="[602,697,1732,1757]" italics="true" pageId="5" pageNumber="6">Forelius</emphasis>
|
||
</taxonomicName>
|
||
and the genus
|
||
<taxonomicName id="81B64D30D716FFB9FEFE4D8EFCF0F951" authority="Forel, 1878" authorityName="Forel" authorityYear="1878" box="[275,506,1764,1789]" class="Insecta" family="Formicidae" genus="Azteca" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FEFE4D8EFC6FF951" box="[275,357,1764,1789]" italics="true" pageId="5" pageNumber="6">Azteca</emphasis>
|
||
<bibRefCitation id="22274B42D716FFB9FE9D4D8EFCF0F951" author="Forel" box="[368,506,1764,1789]" firstAuthor="Forel" pageId="5" pageNumber="6" pagination="337 - 392" refId="ref4560" refString="Forel, A. 1878. Etudes myrmecologiques en 1878 (premiere partie) avec l'anatomie du gesier des fourmis. Bulletin de la Societe Vaudoise des Sciences Naturelles, 15: 337 - 392." year="1878">Forel, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
have been recorded in Mexican and Dominican amber (
|
||
<bibRefCitation id="22274B42D716FFB9FDAE4C6EFDF5F891" author="Duran-Ruiz" etAl="et al." firstAuthor="Duran-Ruiz" pageId="5" pageNumber="6" pagination="485 - 502" refId="ref4403" refString="Duran-Ruiz, C., Riquelme, F., Coutino-Jose, M., Carbot-Chanona, G., Castano-Meneses, G., and Ramos-Arias, M. 2013. Ants from Miocene Totolapa amber (Chiapas, Mexico) with the first record of the genus Forelius (Hymenoptera, Formicidae). Canadian Journal of Earth Sciences, 50: 485 - 502. https: // doi. org / 10.1139 / cjes- 2012 - 0166" year="2013">Duran-Ruiz et al., 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="22274B42D716FFB9FEE64C4EFCB8F891" author="Barden" box="[267,434,1828,1853]" firstAuthor="Barden" pageId="5" pageNumber="6" pagination="1 - 30" refId="ref4272" refString="Barden, P. 2017. Fossil ants (Hymenoptera: Formicidae): ancient diversity and the rise of modern lineages. Myrmecological News, 24: 1 - 30." year="2017">Barden, 2017</bibRefCitation>
|
||
). However, the fossil record of
|
||
<taxonomicName id="81B64D30D716FFB9FF2D4C2EFC15F8F1" authorityName="Emery" authorityYear="1888" box="[192,287,1860,1885]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2D4C2EFC15F8F1" box="[192,287,1860,1885]" italics="true" pageId="5" pageNumber="6">Forelius</emphasis>
|
||
</taxonomicName>
|
||
is uncommon in both deposits, whereas
|
||
<taxonomicName id="81B64D30D716FFB9FF2D4C0EFC1BF8D1" authorityName="Forel" authorityYear="1878" box="[192,273,1892,1917]" class="Insecta" family="Formicidae" genus="Azteca" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FF2D4C0EFC1BF8D1" box="[192,273,1892,1917]" italics="true" pageId="5" pageNumber="6">Azteca</emphasis>
|
||
</taxonomicName>
|
||
is more frequent.
|
||
<taxonomicName id="81B64D30D716FFB9FE184C0EFF5EF8D1" authorityName="Emery" authorityYear="1888" box="[501,596,1892,1917]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FE184C0EFF5EF8D1" box="[501,596,1892,1917]" italics="true" pageId="5" pageNumber="6">Forelius</emphasis>
|
||
</taxonomicName>
|
||
is composed of hyperactive thermophilic ground-nesting ants (
|
||
<bibRefCitation id="22274B42D716FFB9FCA24FAEFEEAFB71" author="Ward" box="[847,992,1220,1245]" firstAuthor="Ward" pageId="5" pageNumber="6" refId="ref5459" refString="Ward, P. S. 2005. A synoptic review of the ants of California (Hymenoptera: Formicidae). Zootaxa, 936: 1 - 68. https: // doi. org / 10.11646 / zootaxa. 936.1.1" year="2005">Ward, 2005</bibRefCitation>
|
||
) that are adapted to living in arid to semi-arid areas (
|
||
<bibRefCitation id="22274B42D716FFB9FBF64F8EF9D0FB51" author="Shattuck, S. O." box="[1051,1242,1252,1278]" pageId="5" pageNumber="6" refId="ref5427" refString="Shattuck, S. O. 1992. Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21 (1): 1 - 181." year="1992">Shattuck, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="22274B42D716FFB9FB014F8EF890FB51" author="Cuezzo, F." box="[1260,1434,1252,1278]" pageId="5" pageNumber="6" pagination="197 - 275" refId="ref4328" refString="Cuezzo, F. 2000. Revision del genero Forelius (Hymenoptera: Formicidae: Dolichoderinae). Sociobiology, 35 (2 A): 197 - 275" year="2000">Cuezzo, 2000</bibRefCitation>
|
||
) whilst
|
||
<taxonomicName id="81B64D30D716FFB9FC7B4E6EFEEDFAB1" authorityName="Forel" authorityYear="1878" box="[918,999,1284,1309]" class="Insecta" family="Formicidae" genus="Azteca" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FC7B4E6EFEEDFAB1" box="[918,999,1284,1309]" italics="true" pageId="5" pageNumber="6">Azteca</emphasis>
|
||
</taxonomicName>
|
||
encompasses ants that nest mainly in trees (
|
||
<bibRefCitation id="22274B42D716FFB9FC2E4E4EF98BFA91" author="Longino, J. & Rica" box="[963,1153,1316,1341]" pageId="5" pageNumber="6" pagination="1 - 63" refId="ref5067" refString="Longino, J. 2007. A taxonomic review of the genus Azteca (Hymenoptera: Formicidae) in Costa Rica and a global revision of the aurita group. Zootaxa, 1491: 1 - 63. https: // doi. org / 10.11646 / zootaxa. 1491.1.1" year="2007">Longino, 2007</bibRefCitation>
|
||
). Notably, the species
|
||
<taxonomicName id="81B64D30D716FFB9FCA54E2EF987FAF1" authority="Wilson, 1985" authorityName="Wilson" authorityYear="1985" box="[840,1165,1348,1373]" class="Insecta" family="Formicidae" genus="Azteca" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alpha">
|
||
<emphasis id="74C2EAA1D716FFB9FCA54E2EFEEFFAF1" box="[840,997,1348,1373]" italics="true" pageId="5" pageNumber="6">Azteca alpha</emphasis>
|
||
Wilson, 1985
|
||
</taxonomicName>
|
||
constitutes 30 to 50% of the total number of ants found in Dominican amber (
|
||
<bibRefCitation id="22274B42D716FFB9FC4A4EEEF95FFA31" author="Barden, P." box="[935,1109,1412,1437]" pageId="5" pageNumber="6" pagination="1 - 30" refId="ref4272" refString="Barden, P. 2017. Fossil ants (Hymenoptera: Formicidae): ancient diversity and the rise of modern lineages. Myrmecological News, 24: 1 - 30." year="2017">Barden, 2017</bibRefCitation>
|
||
). The highest frequency of
|
||
<taxonomicName id="81B64D30D716FFB9FCA54ECEFE90FA11" authorityName="Forel" authorityYear="1878" box="[840,922,1444,1469]" class="Insecta" family="Formicidae" genus="Azteca" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FCA54ECEFE90FA11" box="[840,922,1444,1469]" italics="true" pageId="5" pageNumber="6">Azteca</emphasis>
|
||
</taxonomicName>
|
||
records is consistent with an arboreal environment. The sedimentary data in Mexican amber deposits suggests a lowland fluvial environment close to the coastal plain (Allison, 1967;
|
||
<bibRefCitation id="22274B42D716FFB9FAC34D6EF925F991" author="Frost, S. H. & Langenheim, R. L." pageId="5" pageNumber="6" refId="ref4596" refString="Frost, S. H. and Langenheim, R. L. 1974. Cenozoic reef biofacies, tertiary larger Foraminifera and scleractinian corals from Chiapas, Mexico. Northern Illinois University Press, DeKalb." year="1974">Frost and Langenheim, 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="22274B42D716FFB9FBAD4D4EF9FFF991" author="Graham, A." box="[1088,1269,1572,1598]" pageId="5" pageNumber="6" pagination="17 - 31" refId="ref4633" refString="Graham, A. 1999. Studies in neotropical paleobotany. XIII. An Oligo-Miocene palynoflora from Simojovel (Chiapas, Mexico). American Journal of Botany, 86 (1): 17 - 31. https: // doi. org / 10.2307 / 2656951" year="1999">Graham, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="22274B42D716FFB9FAEB4D4EFE8DF9F1" author="Langenheim, J. H." pageId="5" pageNumber="6" refId="ref4935" refString="Langenheim, J. H. 2003. Plant resins: chemistry, evolution, ecology and ethnobotany. Timber Press, Portland, Cambridge, UK. https: // doi. org / 10.2307 / 4129644" year="2003">Langenheim, 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="22274B42D716FFB9FC784D2EF9A4F9F1" author="Riquelme, F. & Ruvalcaba-Sil, J. L. & Alvarado-Ortega, J. & Estrada-Ruiz, E. & Galicia-Chavez, M. & Porras-Muzquiz, H. & Stojanof, V. & Siddons, D. P. & Miller, L." box="[917,1198,1604,1629]" pageId="5" pageNumber="6" refId="ref5182" refString="Riquelme, F., Ruvalcaba-Sil, J. L., Alvarado-Ortega, J., Estrada-Ruiz, E., Galicia-Chavez, M., Porras-Muzquiz, H., Stojanof, V., Siddons, D. P., and Miller, L. 2014 a. Amber from Mexico: Coahuilite, Simojovelite and Bacalite. Cambridge / MRS Proceedings, 1618: 169 - 180. https: // doi. org / 10.1557 / opl. 2014.466" year="2014">Riquelme et al., 2014a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="22274B42D716FFB9FB564D2EF9F7F9F1" author="Riquelme, F. & Montejo-Cruz, M. & Luna-Castro, B. & Zuniga-Mijangos, L. & Abhandlungen" box="[1211,1277,1604,1629]" pageId="5" pageNumber="6" pagination="93 - 106" refId="ref5351" refString="Riquelme, F., Montejo-Cruz, M., Luna-Castro, B., and Zuniga-Mijangos, L. 2015. Fossil Jumping- bristletail from the Chiapas amber: Neomachilellus (Praeneomachilellus) ezetaelenensis sp. nov. (Microcoryphia: Meinertellidae). Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 275 (1): 93 - 106. https: // doi. org / 10.1127 / njgpa / 2015 / 0453" year="2015">2015</bibRefCitation>
|
||
). Accordingly, the leguminous trees of
|
||
<taxonomicName id="81B64D30D716FFB9FB9F4D0EF9F9F9D1" box="[1138,1267,1636,1661]" class="Magnoliopsida" family="Fabaceae" genus="Hymenaea" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="74C2EAA1D716FFB9FB9F4D0EF9F9F9D1" box="[1138,1267,1636,1661]" italics="true" pageId="5" pageNumber="6">Hymenaea</emphasis>
|
||
</taxonomicName>
|
||
(the botanical source of Mexican amber) were part of a tropical forest resembling modern mangroves (
|
||
<bibRefCitation id="22274B42D716FFB9FAD14DCEFEDBF971" author="Langenheim, J. H." pageId="5" pageNumber="6" refId="ref4935" refString="Langenheim, J. H. 2003. Plant resins: chemistry, evolution, ecology and ethnobotany. Timber Press, Portland, Cambridge, UK. https: // doi. org / 10.2307 / 4129644" year="2003">Langenheim, 2003</bibRefCitation>
|
||
) where a warmer (+1.99 °C) and drier (−
|
||
<quantity id="814E9B56D716FFB9FCB04D8EFED0F951" box="[861,986,1764,1789]" metricMagnitude="-1" metricUnit="m" metricValue="3.377" pageId="5" pageNumber="6" unit="mm" value="337.7">337.7 mm</quantity>
|
||
) tropical climate have been estimated than the current average records (
|
||
<bibRefCitation id="22274B42D716FFB9FAFF4C6EF962F891" author="Hernandez-Hernandez, M. J. & Cruz, J. A. & Castaneda-Posadas, C." pageId="5" pageNumber="6" pagination="102827" refId="ref4849" refString="Hernandez-Hernandez, M. J., Cruz, J. A., and Castaneda-Posadas, C. 2020. Paleoclimatic and vegetation reconstruction of the Miocene Southern Mexico using fossil flowers. Journal of South American Earth of Sciences, 104 (2020): 102827. https: // doi. org / 10.1016 / j. jsames. 2020.102827" year="2020">Hernández-Hernández et al., 2020</bibRefCitation>
|
||
). Although forested areas may have predominated, the Miocene tropical forest in southern
|
||
<collectingCountry id="3EA17623D716FFB9FBE04C0EF96BF8D1" box="[1037,1121,1892,1917]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
|
||
possibly presented minor arid to semi-arid zones that the
|
||
<taxonomicName id="81B64D30D715FFBAFD844BCCFFC2FF13" authorityName="Emery" authorityYear="1888" box="[617,712,166,191]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74C2EAA1D715FFBAFD844BCCFFC2FF13" box="[617,712,166,191]" italics="true" pageId="6" pageNumber="7">Forelius</emphasis>
|
||
</taxonomicName>
|
||
could exploit to locate the colonies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |