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<mods:title id="77B0AFE3FA46E939B94B911B7E4F900C">A taxonomic revision of the genus Edosa Walker, 1886 from China (Lepidoptera, Tineidae, Perissomasticinae)</mods:title>
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<heading id="1E3B583F1268590378DAED4B0BABF849" bold="true" box="[151,671,1903,1930]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="82CC94D01268590378DAED4B0BABF849" ID-CoL="38T5S" authority="Walsingham, 1886" authorityName="Walsingham" authorityYear="1886" box="[151,671,1903,1930]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B833411268590378DAED4B0BABF849" bold="true" box="[151,671,1903,1930]" pageId="12" pageNumber="13">
<emphasis id="77B833411268590378DAED4B08ABF84A" bold="true" box="[151,415,1903,1929]" italics="true" pageId="12" pageNumber="13">Edosa subochraceella</emphasis>
(
<bibRefCitation id="215D92A21268590379E3ED540BA3F849" author="Walsingham" box="[430,663,1904,1930]" pageId="12" pageNumber="13" refString="Walsingham, T. de &amp; Lord, G. (1886) In: Swinhoe, C. C. (Ed.), 1. On the Lepidoptera of Mhow, in Central India. Proceedings of the Zoological Society of London, 1886, 421 - 465." type="book chapter" year="1886">Walsingham, 1886</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="4573EF531268590378DAEDB6084DF868" blockId="12.[151,671,1903,1963]" box="[151,377,1938,1963]" pageId="12" pageNumber="13">
(
<figureCitation id="DDF7F3D61268590378D2EDB609D1F868" box="[159,229,1938,1963]" captionStart="FIGURE 1" captionStartId="68.[151,250,1252,1274]" captionTargetBox="[158,1424,227,1227]" captionTargetId="figure@68.[158,1429,227,1231]" captionTargetPageId="68" captionText="FIGURE 1. Distribution of Chinese Edosa spp. hemichrysella - group: ■ E. bicolor sp. nov.; subochraceella - group: ● E. subochraceella, ▲ E. smithaella, ★ E. varians sp. nov., ♦ E. aurea sp. nov., ♥ E. torta sp. nov." httpUri="https://zenodo.org/record/249827/files/figure.png" pageId="12" pageNumber="13">Figs 1</figureCitation>
,
<figureCitation id="DDF7F3D61268590378BEEDB70824F868" box="[243,272,1939,1963]" captionStart="FIGURES 17" captionStartId="79.[151,264,1953,1975]" captionTargetBox="[219,1366,206,1907]" captionTargetId="figure@79.[204,1398,193,1927]" captionTargetPageId="79" captionText="FIGURES 17. Uncus and uncus lobes of Edosa spp. a, E. subochraceella; b, E. robinsoni sp. nov.; c, E. minuta sp. nov.; d, E. dentata sp. nov.; e, E. semicircularis sp. nov.; f, E. crayella; g, E. bicolor sp. nov.; h, E. uncusella sp. nov. (The number indicates character and its state as shown in Appendix 1; slide numbers same as male genitalia shown in figures 52 79)." httpUri="https://zenodo.org/record/249837/files/figure.png" pageId="12" pageNumber="13">17</figureCitation>
a, 21, 52)
</paragraph>
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<taxonomicName id="82CC94D01268590378DAEDFF0B5EF833" ID-CoL="572R5" authority="Walsingham, 1886: 464" authorityName="Walsingham" authorityPageNumber="464" authorityYear="1886" box="[151,618,2010,2032]" class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B833411268590378DAEDFF0844F833" box="[151,368,2010,2032]" italics="true" pageId="12" pageNumber="13">Tinea subochraceella</emphasis>
<treatmentCitation id="C46DC94212685903793BEDFE0B5EF833" author="Walsingham" box="[374,618,2010,2032]" page="464" pageId="12" pageNumber="13" year="1886">
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: 464
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;
<treatmentCitation id="C46DC942126859037A39EDFE0A06F833" author="Meyrick" box="[628,818,2010,2032]" page="42" pageId="12" pageNumber="13" year="1934">
<bibRefCitation id="215D92A2126859037A39EDFE0A39F833" author="Meyrick" box="[628,781,2010,2032]" pageId="12" pageNumber="13" refString="Meyrick, E. (1934) In: Caradja, A. &amp; Meyrick, E. (Eds.), Materialien zu einer Microlepidopteren - Fauna Kwangtungs. Pterophoridae, Tortricidae, Tineidae. Deutsche Entomologische Zeitschrift, Iris, 48, 28 - 43." type="journal article" year="1934">Meyrick, 1934</bibRefCitation>
: 42
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;
<treatmentCitation id="C46DC942126859037B70EDFE0AD5F833" author="Davis" box="[829,993,2010,2032]" page="65" pageId="12" pageNumber="13" year="1992">
<bibRefCitation id="215D92A2126859037B70EDFE0A8FF833" author="Davis" box="[829,955,2010,2032]" pageId="12" pageNumber="13" refString="Davis, D. R. (1992) Tineidae. In: Heppner, J. B. &amp; Inoue, H. (Eds.), Lepidoptera of Taiwan. Volume 1. Part 2. Checklist. Scientific Publishers Inc., Gainesville, Florida, 63 - 65." type="journal article" year="1992">Davis, 1992</bibRefCitation>
: 65
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.
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</paragraph>
<paragraph id="4573EF531269590278DAEABC0BACFF6D" blockId="13.[151,1224,152,204]" box="[151,664,152,174]" pageId="13" pageNumber="14">
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<taxonomicName id="82CC94D01269590278DAEABC0BA0FF6D" authority="Diakonoff, 1967: 286" authorityName="Diakonoff" authorityPageNumber="286" authorityYear="1967" box="[151,660,152,174]" class="Insecta" family="Tineidae" genus="Haplotinea" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B833411269590278DAEABC089EFF6D" box="[151,426,152,174]" italics="true" pageId="13" pageNumber="14">Haplotinea subochraceella</emphasis>
:
<treatmentCitation id="C46DC9421269590279FAEABC0BA0FF6D" author="Diakonoff" box="[439,660,152,174]" page="286" pageId="13" pageNumber="14" year="1967">
<bibRefCitation id="215D92A21269590279FAEABC0B55FF6D" author="Diakonoff" box="[439,609,152,174]" pageId="13" pageNumber="14" refString="Diakonoff, A. (1967) Microlepidoptera of the Philippine Islands. United States national museum bulletin, 257, 266 - 293. http: // dx. doi. org / 10.5479 / si. 03629236.257.1" type="journal article" year="1967">Diakonoff, 1967</bibRefCitation>
: 286
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.
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<taxonomicName id="82CC94D01269590278DAEA920BFDFF0F" ID-CoL="38T5S" authority="Robinson &amp; Nielsen, 1993: 198" authorityName="Robinson &amp; Nielsen" authorityPageNumber="198" authorityYear="1993" box="[151,713,182,204]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B833411269590278DAEA920843FF0F" box="[151,375,182,204]" italics="true" pageId="13" pageNumber="14">Edosa subochraceella</emphasis>
:
<treatmentCitation id="C46DC9421269590279C9EA920BFDFF0F" author="Robinson" box="[388,713,182,204]" page="198" pageId="13" pageNumber="14" year="1993">
<bibRefCitation id="215D92A21269590279C9EA920BA2FF0F" author="Robinson" box="[388,662,182,204]" pageId="13" pageNumber="14" refString="Robinson, G. S. &amp; Nielsen, E. S. (1993) Tineid genera of Australia (Lepidoptera). Monographs on Australian Lepidoptera, CSIRO Publications, Melbourne, 2, 190 - 199. http: // dx. doi. org / 10.1017 / s 0007485300039833" type="journal article" year="1993">Robinson &amp; Nielsen, 1993</bibRefCitation>
: 198
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;
<treatmentCitation id="C46DC942126959027A9EEA920ADBFF0F" author="Robinson" box="[723,1007,182,204]" page="11" pageId="13" pageNumber="14" year="1996">
<bibRefCitation id="215D92A2126959027A9EEA920AFEFF0F" author="Robinson" box="[723,970,182,204]" pageId="13" pageNumber="14" refString="Robinson, G. S. &amp; Tuck, K. R. (1996) A revisionary checklist of the Tineidae (Lepidoptera) of the Oriental Region. Occasional Papers on Systematic Entomology, 9, 1 - 29." type="journal article" year="1996">Robinson &amp; Tuck, 1996</bibRefCitation>
: 11
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;
<treatmentCitation id="C46DC942126959027BB7EA920DF0FF0F" author="Gaedike" box="[1018,1220,182,204]" httpUri="http://treatment.plazi.org/id/961887930E5FFFB6FF7AFD48FB7D57B7" page="174" pageId="13" pageNumber="14" year="2012">
<bibRefCitation id="215D92A2126959027BB7EA920DA5FF0F" author="Gaedike" box="[1018,1169,182,204]" pageId="13" pageNumber="14" refString="Gaedike, R. (2012) Beitrag zur Kenntnis der Tineiden von Nepal und benachbarter Gebiete. Beitrage zur Entomologie, 62, 165 - 206." type="journal article" year="2012">Gaedike, 2012</bibRefCitation>
: 174
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.
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</paragraph>
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<paragraph id="4573EF531269590278DAEADD0D10FEF5" blockId="13.[151,1436,249,814]" pageId="13" pageNumber="14">
<emphasis id="77B833411269590278DAEADD0BE1FED1" bold="true" box="[151,725,249,274]" pageId="13" pageNumber="14">
Material examined.
<collectingCountry id="3DDBAFC31269590279C3EADD08DBFED1" box="[398,495,249,274]" name="China" pageId="13" pageNumber="14">CHINA</collectingCountry>
: Hainan Province:
</emphasis>
<specimenCount id="53CA24DA126959027A92EADD0A38FED2" box="[735,780,249,273]" pageId="13" pageNumber="14" type="generic">1 ♂</specimenCount>
, Datian Conservation Zone (19°42΄N, 109°47΄E),
<quantity id="823442B6126959027D14EADD0CA3FED2" box="[1369,1431,249,274]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="m" value="25.0">25 m</quantity>
,
<date id="3172C9931269590278DAEB390860FEF5" box="[151,340,285,310]" pageId="13" pageNumber="14" value="2009-11-30" valueMax="2009-12-02" valueMin="2009-11-30">30.xi 2.xii.2009</date>
, leg. Zhaohui Du &amp; Linlin Yang, genitalia slide No. YLL10085.
</paragraph>
</subSubSection>
<subSubSection id="0DD6BCD812695902788AEB6508FFFDB9" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="4573EF5312695902788AEB6508FFFDB9" blockId="13.[151,1436,249,814]" pageId="13" pageNumber="14">
<emphasis id="77B8334112695902788AEB650875FE99" bold="true" box="[199,321,321,346]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Adult (
<figureCitation id="DDF7F3D61269590279D6EB6508C4FE9A" box="[411,496,321,346]" captionStart="FIGURES 20 27" captionStartId="82.[151,264,1944,1966]" captionTargetBox="[151,1436,224,1929]" captionTargetId="figure@82.[151,1436,193,1936]" captionTargetPageId="82" captionText="FIGURES 20 27. Adults of Edosa spp. 20, E. bicolor sp. nov., holotype, ♂; 21, E. subochraceella; 22, E. smithaella, ♂; 23, E. varians sp. nov., holotype, ♂; 24, E. aurea sp. nov., holotype, ♂; 25, E. torta sp. nov., holotype, ♂; 26, E. truncatula sp. nov., ♂; 27, E. hamata, sp. nov., holotype, ♂." httpUri="https://zenodo.org/record/249840/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
). Wingspan 12.0 mm.
<taxonomicName id="82CC94D0126959027AB9EB650ADBFE99" box="[756,1007,321,346]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B83341126959027AB9EB650ADBFE99" box="[756,1007,321,346]" italics="true" pageId="13" pageNumber="14">Edosa subochraceella</emphasis>
</taxonomicName>
is characterized by the twisted uncus lobes with distal 1/3 strongly contorted, forming a triangular section and a melanized nodule; the valva varying from subovate to rounded triangular, with finely coarse nodules below or on costal margin, the obvious ventral lobe, the absence of the basal flange; the trapezoidal juxta; the stout, double-walled aedeagus with inner wall tubular, outer wall forming shallow subapical carinae; the bulbus ejaculatorius 7.0× length of the aedeagus, subdistal section semicircularly hypertrophied, distal section hypertrophied, 0.5× width of the subdistal section, deep cup-shaped (
<figureCitation id="DDF7F3D612695902792EE83D088CFDF1" box="[355,440,537,562]" captionStart="FIGURES 52 53" captionStartId="86.[151,264,1969,1991]" captionTargetBox="[198,1395,215,1913]" captionTargetId="figure@86.[174,1410,193,1938]" captionTargetPageId="86" captionText="FIGURES 52 53. Male genitalia of Edosa spp. 52, E. subochraceella, slide No. YLL 10085 (52 a, aedeagus; 52 b, uncus lobes; 52 c, valva); 53, E. smithaella, slide No. YLL 10176 (53 a, aedeagus; 53 b, uncus lobes; 53 c, valva) (scale lines = 0.5 mm)." httpUri="https://zenodo.org/record/249844/files/figure.png" pageId="13" pageNumber="14">Fig. 52</figureCitation>
).
<taxonomicName id="82CC94D0126959027981E83D0BF0FDF1" box="[460,708,537,562]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B83341126959027981E83D0BF0FDF1" box="[460,708,537,562]" italics="true" pageId="13" pageNumber="14">Edosa subochraceella</emphasis>
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is similar to
<taxonomicName id="82CC94D0126959027B14E83E0AF9FDF2" box="[857,973,538,561]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="varians" status="sp. nov.">
<emphasis id="77B83341126959027B14E83E0AF9FDF2" box="[857,973,538,561]" italics="true" pageId="13" pageNumber="14">E. varians</emphasis>
</taxonomicName>
<emphasis id="77B83341126959027B99E83D0D1AFDF1" bold="true" box="[980,1070,537,562]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6C8B8E3A126959027B99E83D0D1AFDF1" box="[980,1070,537,562]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished by the valva without process on the costal margin, while the valva has a triangular to hillock-shaped process on the costal margin in the latter species.
</paragraph>
</subSubSection>
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<paragraph id="4573EF5312695902788AE8A10C33FD5D" blockId="13.[151,1436,249,814]" box="[199,1287,645,670]" pageId="13" pageNumber="14">
<emphasis id="77B8334112695902788AE8A10854FD5D" bold="true" box="[199,352,645,670]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingCountry id="3DDBAFC312695902792AE8A1089EFD5D" box="[359,426,645,670]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
(Guangdong, Hainan,
<collectingCountry id="3DDBAFC3126959027AE7E8A10A35FD5D" box="[682,769,645,670]" name="Taiwan" pageId="13" pageNumber="14">Taiwan</collectingCountry>
),
<collectingCountry id="3DDBAFC3126959027B5EE8A20A56FD5E" box="[787,866,646,669]" name="Myanmar" pageId="13" pageNumber="14">Burma</collectingCountry>
,
<collectingCountry id="3DDBAFC3126959027B20E8A10A9DFD5D" box="[877,937,645,670]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
,
<collectingCountry id="3DDBAFC3126959027BF8E8A10ACFFD5D" box="[949,1019,645,670]" name="Nepal" pageId="13" pageNumber="14">Nepal</collectingCountry>
,
<collectingCountry id="3DDBAFC3126959027C48E8A10DB2FD5D" box="[1029,1158,645,670]" name="Philippines" pageId="13" pageNumber="14">Philippines</collectingCountry>
,
<collectingCountry id="3DDBAFC3126959027CDFE8A10C37FD5D" box="[1170,1283,645,670]" name="Sri Lanka" pageId="13" pageNumber="14">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0DD6BCD812695902788AE88D0CA2FCED" pageId="13" pageNumber="14" type="discussion">
<paragraph id="4573EF5312695902788AE88D0AD7FD25" blockId="13.[151,1436,249,814]" pageId="13" pageNumber="14">
<emphasis id="77B8334112695902788AE88D080FFD01" bold="true" box="[199,315,681,706]" pageId="13" pageNumber="14">Remarks.</emphasis>
<bibRefCitation id="215D92A2126959027909E88D0B24FD01" author="Robinson" box="[324,528,681,706]" pageId="13" pageNumber="14" refString="Robinson, G. S. (2008 a) Hidden diversity in small brown moths - the systematics of Edosa (Lepidoptera: Tineidae) in Sundaland. Systematics and Biodiversity, 6 (3), 319 - 384. http: // dx. doi. org / 10.1017 / s 1477200008002685" type="journal article" year="2008" yearSuffix="a">Robinson (2008a)</bibRefCitation>
treated the male specimens of
<taxonomicName id="82CC94D0126959027B37E88D0CA8FD01" authority="Walsingham, 1886" authorityName="Walsingham" authorityYear="1886" box="[890,1436,681,706]" class="Insecta" family="Tineidae" genus="Haplotinea" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B83341126959027B37E88D0D98FD01" box="[890,1196,681,706]" italics="true" pageId="13" pageNumber="14">Haplotinea subochraceella</emphasis>
(
<bibRefCitation id="215D92A2126959027CF0E88D0CA0FD01" author="Walsingham" box="[1213,1428,681,706]" pageId="13" pageNumber="14" refString="Walsingham, T. de &amp; Lord, G. (1886) In: Swinhoe, C. C. (Ed.), 1. On the Lepidoptera of Mhow, in Central India. Proceedings of the Zoological Society of London, 1886, 421 - 465." type="book chapter" year="1886">Walsingham, 1886</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="3DDBAFC3126959027899E8E90866FD25" box="[212,338,717,742]" name="Philippines" pageId="13" pageNumber="14">Philippines</collectingCountry>
in
<bibRefCitation id="215D92A212695902793BE8E90B0EFD25" author="Diakonoff" box="[374,570,717,742]" pageId="13" pageNumber="14" refString="Diakonoff, A. (1967) Microlepidoptera of the Philippine Islands. United States national museum bulletin, 257, 266 - 293. http: // dx. doi. org / 10.5479 / si. 03629236.257.1" type="journal article" year="1967">Diakonoff (1967)</bibRefCitation>
as
<taxonomicName id="82CC94D0126959027A12E8EA0AD4FD25" authority="Robinson, 2008" authorityName="Robinson" authorityYear="2008" box="[607,992,717,742]" class="Insecta" family="Tineidae" genus="Edosa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="williamsonella">
<emphasis id="77B83341126959027A12E8EA0A10FD25" box="[607,804,717,742]" italics="true" pageId="13" pageNumber="14">E. williamsonella</emphasis>
Robinson, 2008
</taxonomicName>
.
</paragraph>
<paragraph id="4573EF5312695902788AE8D50CA2FCED" blockId="13.[151,1436,249,814]" pageId="13" pageNumber="14">
<bibRefCitation id="215D92A212695902788AE8D50843FCC9" author="Gaedike" box="[199,375,753,778]" pageId="13" pageNumber="14" refString="Gaedike, R. (2012) Beitrag zur Kenntnis der Tineiden von Nepal und benachbarter Gebiete. Beitrage zur Entomologie, 62, 165 - 206." type="journal article" year="2012">Gaedike (2012)</bibRefCitation>
figured the male genitalia of
<taxonomicName id="82CC94D0126959027A8BE8D60AA5FCC9" box="[710,913,753,778]" class="Insecta" family="Tineidae" genus="Haplotinea" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subochraceella">
<emphasis id="77B83341126959027A8BE8D60AA5FCC9" box="[710,913,753,778]" italics="true" pageId="13" pageNumber="14">E. subochraceella</emphasis>
</taxonomicName>
, which shows the variations of the uncus lobe and the valva (
<bibRefCitation id="215D92A2126959027974E9310B52FCED" author="Gaedike" box="[313,614,789,814]" pageId="13" pageNumber="14" refString="Gaedike, R. (2012) Beitrag zur Kenntnis der Tineiden von Nepal und benachbarter Gebiete. Beitrage zur Entomologie, 62, 165 - 206." type="journal article" year="2012">
Gaedike 2012,
<figureCitation id="DDF7F3D61269590279A9E9310B52FCED" box="[484,614,789,814]" captionStart-0="FIGURES 73 75" captionStart-1="FIGURES 76 79" captionStartId-0="95.[151,264,1942,1964]" captionStartId-1="96.[151,264,1942,1964]" captionTargetBox-0="[214,1367,200,1907]" captionTargetBox-1="[191,1396,205,1919]" captionTargetId-0="figure@95.[206,1381,193,1921]" captionTargetId-1="figure@96.[186,1401,193,1921]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionText-0="FIGURES 73 75. Male genitalia of Edosa spp. 73, E. robustispina sp. nov., paratype, slide No. YLL 10067; 74, E. uncusella sp. nov., paratype, slide No. YLL 10207 (74 a, left valva, slide No. XYL 04234); 75, E. semicircularis sp. nov., paratype, slide No. YLL 11094 (scale lines = 0.5 mm)." captionText-1="FIGURES 76 79. Male genitalia of Edosa spp. 76, E. dentata sp. nov., holotype, slide No. YLL 11180; 77, E. digitata sp. nov., holotype, slide No. YLL 11076; 78, E. longicornis sp. nov., holotype, slide No. YLL 10172; 79, E. minuta sp. nov., holotype, slide No. YLL 10219 (79 a, unrolling states, paratype, slide No. YLL 11016) (scale lines = 0.5 mm)." httpUri-0="https://zenodo.org/record/249853/files/figure.png" httpUri-1="https://zenodo.org/record/249854/files/figure.png" pageId="13" pageNumber="14">Figs 7379</figureCitation>
</bibRefCitation>
). The subovate valva in our specimen is within the range of the variation.
</paragraph>
</subSubSection>
</treatment>
</document>