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<document id="A9EFF9B99528D90EC554545DD2BD1F05" ID-CLB-Dataset="25363" ID-DOI="10.11646/zootaxa.4674.5.4" ID-GBIF-Dataset="18b8b529-9043-414e-b7f0-2a975e51c7ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="3465084" ID-ZooBank="2ECC423B-D0C0-467B-A637-68E8FCFA265E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569830583483" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Scholtz, Clarke H." docDate="2019" docId="CD0A87A2083AEA5C94DAF94CA2E81065" docLanguage="en" docName="zootaxa.4674.5.4.pdf" docOrigin="Zootaxa 4674 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phoberus lilianae Strümpher &amp; Scholtz 2019, new status" docType="treatment" docVersion="9" lastPageNumber="559" masterDocId="3133FFDA083DEA55944DFFB7A0401102" masterDocTitle="A new Phoberus MacLeay species (Coleoptera: Scarabaeoidea: Trogidae) from the Maloti-Drakensberg Mountains of southern Africa and the Phoberus caffer species group reviewed" masterLastPageNumber="563" masterPageNumber="551" pageNumber="558" updateTime="1698756544286" updateUser="ExternalLinkService">
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<mods:title id="44516C91C0438FA3D5EE6157286FCEA1">A new Phoberus MacLeay species (Coleoptera: Scarabaeoidea: Trogidae) from the Maloti-Drakensberg Mountains of southern Africa and the Phoberus caffer species group reviewed</mods:title>
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<paragraph id="451C36B4083AEA5294DAF94CA2FE1617" blockId="7.[151,702,1787,1849]" box="[151,702,1787,1814]" pageId="7" pageNumber="558">
<heading id="1E5481D8083AEA5294DAF94CA2FE1617" bold="true" box="[151,702,1787,1814]" fontSize="11" level="1" pageId="7" pageNumber="558" reason="1">
<emphasis id="77D7EAA6083AEA5294DAF94CA2FE1617" bold="true" box="[151,702,1787,1814]" pageId="7" pageNumber="558">
<taxonomicName id="82A34D37083AEA5294DAF94CA26E1617" ID-CoL="4GJWL" authority="(Scholtz, 1980)" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[151,558,1787,1814]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="558" phylum="Arthropoda" rank="species" species="lilianae" status="stat. nov.">
<emphasis id="77D7EAA6083AEA5294DAF94CA12D1614" bold="true" box="[151,365,1787,1814]" italics="true" pageId="7" pageNumber="558">Phoberus lilianae</emphasis>
(
<bibRefCitation id="21324B45083AEA529530F94CA2661617" author="Scholtz, C. H." box="[381,550,1787,1813]" pageId="7" pageNumber="558" pagination="1 - 104" refId="ref9058" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz, 1980</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="6CE457DD083AEA529671F94CA2FE1617" box="[572,702,1787,1813]" pageId="7" pageNumber="558">new status</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="451C36B4083AEA5294DAF8A8A118163B" blockId="7.[151,702,1787,1849]" box="[151,344,1822,1849]" pageId="7" pageNumber="558">
(
<figureCitation id="DD982A31083AEA5294D2F8A8A167163B" box="[159,295,1822,1849]" captionStart="FIGURES 1518" captionStartId="9.[151,264,1617,1642]" captionTargetBox="[357,1232,378,1587]" captionTargetId="figure@9.[348,1240,373,1597]" captionTargetPageId="9" captionText="FIGURES 1518. Phoberus lilianae (Scholtz, 1980), paratype (SAMC). 14, dorsal habitus; 15, lateral habitus; 16, dorsal and lateral views of aedeagus; 17, paratype labels." figureDoi="http://doi.org/10.5281/zenodo.3467964" httpUri="https://zenodo.org/record/3467964/files/figure.png" pageId="7" pageNumber="558">Figs. 1518</figureCitation>
,
<figureCitation id="DD982A31083AEA52957EF8A8A110163B" box="[307,336,1823,1849]" captionStart="FIGURE 19" captionStartId="10.[151,250,1131,1156]" captionTargetBox="[158,1433,177,1106]" captionTargetId="figure@10.[151,1439,174,1111]" captionTargetPageId="10" captionText="FIGURE 19. Distribution of species in the southern African Phoberus caffer species group." figureDoi="http://doi.org/10.5281/zenodo.3467966" httpUri="https://zenodo.org/record/3467966/files/figure.png" pageId="7" pageNumber="558">19</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="451C36B4083AEA5294DAF8D3A38F167E" blockId="7.[151,975,1891,1977]" box="[151,975,1891,1916]" pageId="7" pageNumber="558">
<treatmentCitationGroup id="65B3119A083AEA5294DAF8D3A38F167E" box="[151,975,1891,1916]" pageId="7" pageNumber="558">
<taxonomicName id="82A34D37083AEA5294DAF8D3A21A167E" ID-CoL="59DM4" authority="Scholtz, 1980: 96" authorityName="Scholtz" authorityPageNumber="96" authorityYear="1980" box="[151,602,1891,1916]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="558" phylum="Arthropoda" rank="subSpecies" species="caffer" subGenus="Trox" subSpecies="lilianae">
<emphasis id="77D7EAA6083AEA5294DAF8D3A0851679" box="[151,197,1892,1915]" italics="true" pageId="7" pageNumber="558">Trox</emphasis>
(
<emphasis id="77D7EAA6083AEA52949EF8D3A1411679" box="[211,257,1892,1915]" italics="true" pageId="7" pageNumber="558">Trox</emphasis>
)
<emphasis id="77D7EAA6083AEA52955DF8D4A1DE1679" box="[272,414,1891,1915]" italics="true" pageId="7" pageNumber="558">caffer lilianae</emphasis>
<treatmentCitation id="C40210A5083AEA5295E8F8D4A21A167E" author="Scholtz, C. H." box="[421,602,1891,1916]" page="96" pageId="7" pageNumber="558" year="1980">
<bibRefCitation id="21324B45083AEA5295E8F8D4A21A167E" author="Scholtz, C. H." box="[421,602,1891,1916]" pageId="7" pageNumber="558" pagination="1 - 104" refId="ref9058" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz, 1980: 96</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="C40210A5083AEA529629F8D4A3461679" author="Scholtz, C. H." box="[612,774,1891,1916]" page="3" pageId="7" pageNumber="558" year="1982">
<bibRefCitation id="21324B45083AEA529629F8D4A3461679" author="Scholtz, C. H." box="[612,774,1891,1916]" pageId="7" pageNumber="558" pagination="1 - 27" refId="ref9092" refString="Scholtz, C. H. (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agricul- ture and Fisheries, Republic of South Africa, 54, 1 - 27." type="journal article" year="1982">Scholtz 1982: 3</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C40210A5083AEA52975CF8D4A38F167E" author="Scholtz, C. H." box="[785,975,1891,1916]" page="206" pageId="7" pageNumber="558" year="1983">
<bibRefCitation id="21324B45083AEA52975CF8D4A38F167E" author="Scholtz, C. H." box="[785,975,1891,1916]" pageId="7" pageNumber="558" pagination="205 - 208" refId="ref9132" refString="Scholtz, C. H. (1983) A new species of Trox F. (Coleoptera: Trogidae) from Senegal with new distribution records for other African species. Journal of the Entomological Society of Southern Africa, 46 (2), 205 - 208." type="journal article" year="1983">Scholtz, 1983: 206</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="451C36B4083AEA5294DAF834A3D31698" blockId="7.[151,975,1891,1977]" box="[151,915,1921,1946]" pageId="7" pageNumber="558">
<treatmentCitationGroup id="65B3119A083AEA5294DAF834A3D31698" box="[151,915,1921,1946]" pageId="7" pageNumber="558">
<taxonomicName id="82A34D37083AEA5294DAF834A2681698" ID-CoL="59DM4" authority="Scholtz" authorityName="Scholtz" authorityYear="1980" box="[151,552,1921,1946]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="558" phylum="Arthropoda" rank="subSpecies" species="caffer" subGenus="Phoberus" subSpecies="lilianae">
<emphasis id="77D7EAA6083AEA5294DAF834A0851698" box="[151,197,1923,1946]" italics="true" pageId="7" pageNumber="558">Trox</emphasis>
(
<emphasis id="77D7EAA6083AEA52949EF835A1741698" box="[211,308,1922,1946]" italics="true" pageId="7" pageNumber="558">Phoberus</emphasis>
)
<emphasis id="77D7EAA6083AEA52950EF835A1921698" box="[323,466,1922,1946]" italics="true" pageId="7" pageNumber="558">caffer lilianae</emphasis>
Scholtz
</taxonomicName>
:
<treatmentCitation id="C40210A5083AEA52967FF836A2B41698" author="Scholtz, C. H." box="[562,756,1921,1946]" page="361" pageId="7" pageNumber="558" year="1986">
<bibRefCitation id="21324B45083AEA52967FF836A2B41698" author="Scholtz, C. H." box="[562,756,1921,1946]" pageId="7" pageNumber="558" pagination="355 - 363" refId="ref9182" refString="Scholtz, C. H. (1986) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology, 11 (3), 355 - 363." type="journal article" year="1986">Scholtz, 1986: 361</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C40210A5083AEA5296B3F836A3D31698" author="Zidek, J." box="[766,915,1921,1946]" page="9" pageId="7" pageNumber="558" year="2013">
<bibRefCitation id="21324B45083AEA5296B3F836A3D31698" author="Zidek, J." box="[766,915,1921,1946]" pageId="7" pageNumber="558" pagination="1 - 38" refId="ref9695" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek, 2013: 9</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="451C36B4083AEA5D94DAF816A4CD1417" blockId="7.[151,975,1891,1977]" lastBlockId="8.[151,1437,1167,2022]" lastPageId="8" lastPageNumber="559" pageId="7" pageNumber="558">
<taxonomicName id="82A34D37083AEA5294DAF816A1B416BB" ID-CoL="4GJWL" authority="(Scholtz)" authorityName="Scholtz" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[151,500,1952,1977]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="558" phylum="Arthropoda" rank="subSpecies" species="caffer" subSpecies="lilianae">
<emphasis id="77D7EAA6083AEA5294DAF816A1CD16BB" box="[151,397,1953,1977]" italics="true" pageId="7" pageNumber="558">Phoberus caffer lilianae</emphasis>
(Scholtz)
</taxonomicName>
:
<bibRefCitation id="21324B45083AEA5295B3F817A34416BB" author="Strumpher, W. P. &amp; Villet, M. H. &amp; Sole, C. L. &amp; Scholtz, C. H." box="[510,772,1952,1977]" pageId="7" pageNumber="558" pagination="53 - 82" refId="ref9533" refString="Strumpher, W. P., Villet, M. H., Sole, C. L. &amp; Scholtz, C. H. (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics &amp; Evolution, 47 (1), 53 - 82. https: // doi. org / 10.1163 / 1876312 X- 46052133" type="journal article" year="2016">
Strümpher
<emphasis id="77D7EAA6083AEA529622F816A2DC16BB" box="[623,668,1953,1977]" italics="true" pageId="7" pageNumber="558">et al</emphasis>
. 2016: 78
</bibRefCitation>
;
<bibRefCitation id="21324B45083AEA529742F817A3F016BB" author="Zidek, J." box="[783,944,1952,1977]" pageId="7" pageNumber="558" pagination="93 - 127" refId="ref9723" refString="Zidek, J. (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana, Series A, 25 (1), 93 - 127." type="journal article" year="2017">
Zidek, 2017:
<specimenCount id="53A5FD3D083AEA5297DAF816A3F016BB" box="[919,944,1953,1977]" pageId="7" pageNumber="558" type="generic">99</specimenCount>
</bibRefCitation>
<emphasis id="77D7EAA60835EA5D94DAFB27A09215A8" bold="true" box="[151,210,1168,1194]" pageId="8" pageNumber="559">Type</emphasis>
<materialsCitation id="F5CB3CE90835EA5D9496FB38A4C91417" ID-GBIF-Occurrence="2423311068" collectingDate="1962-07-05" collectionCode="SAMC, TMSA" collectorName="Struthers, J. K." country="Zimbabwe" location="Inyanga" municipality="Umtali" pageId="8" pageNumber="559" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Harare" typeStatus="paratype">
<emphasis id="77D7EAA60835EA5D9496FB38A29A15A8" bold="true" box="[219,730,1167,1194]" pageId="8" pageNumber="559">
material re-examined from
<bibRefCitation id="21324B450835EA5D9664FB38A29515A8" author="Scholtz, C. H." box="[553,725,1167,1194]" pageId="8" pageNumber="559" pagination="1 - 104" refId="ref9058" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz (1980)</bibRefCitation>
.
</emphasis>
Five
<typeStatus id="9A1888160835EA5D9753FB27A3CE15A8" box="[798,910,1168,1194]" pageId="8" pageNumber="559" type="paratype">paratypes</typeStatus>
:
<collectingMunicipality id="A578ACCE0835EA5D97D7FB38A3A815AB" box="[922,1000,1167,1193]" pageId="8" pageNumber="559">Umtali</collectingMunicipality>
[= Mutare], S. Rhodesia, 2.1951 (
<specimenCount id="53A5FD3D0835EA5D913EFB38A5D715AB" box="[1395,1431,1167,1193]" pageId="8" pageNumber="559" type="male">1♂</specimenCount>
,
<collectionCode id="23B2AE710835EA5D94DAFB04A0AB15CF" box="[151,235,1203,1229]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="8" pageNumber="559" type="Museum">SAMC</collectionCode>
);
<location id="407C606F0835EA5D94B2FB03A11915CC" LSID="urn:lsid:plazi:treatment:CD0A87A2083AEA5C94DAF94CA2E81065:407C606F0835EA5D94B2FB03A11915CC" box="[255,345,1204,1230]" country="Zimbabwe" municipality="Umtali" name="Inyanga" pageId="8" pageNumber="559" stateProvince="Harare">Inyanga</location>
[Nyanga Mountains], S.
<collectingCountry id="3DB476240835EA5D963AFB04A29F15CF" box="[631,735,1203,1229]" name="Zimbabwe" pageId="8" pageNumber="559">Rhodesia</collectingCountry>
[
<collectingCountry id="3DB476240835EA5D96A0FB04A32D15CC" box="[749,877,1203,1230]" name="Zimbabwe" pageId="8" pageNumber="559">Zimbabwe</collectingCountry>
],
<date id="311D10740835EA5D97CDFB04A3B215CC" box="[896,1010,1203,1230]" pageId="8" pageNumber="559" value="1962-07-05">
<collectingDate id="2159E99C0835EA5D97CDFB04A3B215CC" box="[896,1010,1203,1230]" pageId="8" pageNumber="559" value="1962-07-05">5.vii.1962</collectingDate>
</date>
(
<specimenCount id="53A5FD3D0835EA5D9048FB04A46615CF" box="[1029,1062,1203,1229]" pageId="8" pageNumber="559" type="female">1♀</specimenCount>
,
<collectionCode id="23B2AE710835EA5D9079FB04A4C715CF" box="[1076,1159,1203,1229]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="8" pageNumber="559" type="Museum">SAMC</collectionCode>
;
<specimenCount id="53A5FD3D0835EA5D90DEFB04A4F715CF" box="[1171,1207,1203,1229]" pageId="8" pageNumber="559" type="male">1♂</specimenCount>
,
<specimenCount id="53A5FD3D0835EA5D9088FB04A4A815CF" box="[1221,1256,1203,1229]" pageId="8" pageNumber="559" type="female">1♀</specimenCount>
,
<collectionCode id="23B2AE710835EA5D90BBFB04A50915CF" box="[1270,1353,1203,1229]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="559">TMSA</collectionCode>
); Salis- bury [=
<collectingRegion id="8767F8560835EA5D94BFFB6FA10215F0" box="[242,322,1240,1266]" country="Zimbabwe" name="Harare" pageId="8" pageNumber="559">Harare</collectingRegion>
; although the locality of this specimen is cited as Salisbury, it is more likely that this specimen was collected from Eastern Zimbabwean Highlands],
<date id="311D10740835EA5D96F2FB4CA2B11414" box="[703,753,1275,1302]" pageId="8" pageNumber="559">
<collectingDate id="2159E99C0835EA5D96F2FB4CA2B11414" box="[703,753,1275,1302]" pageId="8" pageNumber="559">3.iii.</collectingDate>
</date>
[19]66,
<collectorName id="E85653620835EA5D9706FB4CA3AC1414" box="[843,1004,1275,1302]" pageId="8" pageNumber="559">Struthers, J.K.</collectorName>
(
<specimenCount id="53A5FD3D0835EA5D97B0FB4CA4611417" box="[1021,1057,1275,1301]" pageId="8" pageNumber="559" type="male">1♂</specimenCount>
,
<collectionCode id="23B2AE710835EA5D9060FB4CA4C11417" box="[1069,1153,1275,1301]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="8" pageNumber="559" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="11DC663C0835EA5D94DAFB93A2C71562" ID-DOI="http://doi.org/10.5281/zenodo.3467962" ID-Zenodo-Dep="3467962" httpUri="https://zenodo.org/record/3467962/files/figure.png" pageId="8" pageNumber="559" startId="8.[151,264,1060,1085]" targetBox="[347,1239,178,1033]" targetPageId="8">
<paragraph id="451C36B40835EA5D94DAFB93A2C71562" blockId="8.[151,1437,1060,1121]" pageId="8" pageNumber="559">
<emphasis id="77D7EAA60835EA5D94DAFB93A117153F" bold="true" box="[151,343,1060,1085]" pageId="8" pageNumber="559">FIGURES 1314.</emphasis>
Invalid type of
<taxonomicName id="82A34D370835EA5D9649FB91A34A153F" ID-CoL="59DHR" authority="Harold, 1872" authorityName="Harold" authorityYear="1872" box="[516,778,1060,1085]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="caffer">
<emphasis id="77D7EAA60835EA5D9649FB91A236153F" box="[516,630,1061,1085]" italics="true" pageId="8" pageNumber="559">Trox caffer</emphasis>
<bibRefCitation id="21324B450835EA5D9632FB93A34A153F" author="Harold, E." box="[639,778,1060,1085]" pageId="8" pageNumber="559" pagination="1 - 192" refId="ref8678" refString="Harold, E. 1872. Monographie der Gattung Trox. Coleopterologische Hefte, 9, 1 - 192." type="journal article" year="1872">Harold, 1872</bibRefCitation>
</taxonomicName>
, designation unknown (MNHB). 12, dorsal habitus; 13, labels. Photographs by Bernd Jaeger, copyright MNHB.
</paragraph>
</caption>
<paragraph id="451C36B40835EA5D948AFA97A204145C" blockId="8.[151,1437,1167,2022]" pageId="8" pageNumber="559">
<typeStatus id="9A1888160835EA5D948AFA97A1741438" box="[199,308,1312,1338]" pageId="8" pageNumber="559" type="paratype">Paratypes</typeStatus>
examined bear new labels:
<taxonomicName id="82A34D370835EA5D9628FAA8A3D91438" ID-CoL="4GJWL" authority="(Scholtz)" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[613,921,1311,1338]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae" status="stat. nov.">
<emphasis id="77D7EAA60835EA5D9628FAA8A36D143B" box="[613,813,1311,1337]" italics="true" pageId="8" pageNumber="559">Phoberus lilianae</emphasis>
(Scholtz)
</taxonomicName>
<taxonomicNameLabel id="6CE457DD0835EA5D97D3FA97A4441438" box="[926,1028,1312,1338]" pageId="8" pageNumber="559">stat. nov.</taxonomicNameLabel>
, Strümpher &amp; Scholtz, 2019 (printed on white rectangle with black border).
</paragraph>
</subSubSection>
<subSubSection id="0DB9653F0835EA5D948AFAD0A4DC14C8" pageId="8" pageNumber="559" type="materials_examined">
<paragraph id="451C36B40835EA5D948AFAD0A4DC14C8" blockId="8.[151,1437,1167,2022]" pageId="8" pageNumber="559">
<emphasis id="77D7EAA60835EA5D948AFAD0A27D1480" bold="true" box="[199,573,1383,1410]" pageId="8" pageNumber="559">Additional material examined.</emphasis>
<materialsCitation id="F5CB3CE90835EA5D9605FAD0A34B14A7" ID-GBIF-Occurrence="2423311091" collectingDate="1958-07" collectionCode="TMSA" collectorName="A. Hoffmann" country="Zimbabwe" latitude="-18.283333" location="Mare Dam" longLatPrecision="1275" longitude="32.766666" municipality="Inyanga" pageId="8" pageNumber="559" specimenCount="1" specimenCount-female="1" stateProvince="Manicaland">
<emphasis id="77D7EAA60835EA5D9605FAD0A44B1480" bold="true" box="[584,1035,1383,1410]" pageId="8" pageNumber="559">
<collectingCountry id="3DB476240835EA5D9605FAD0A2A81480" box="[584,744,1383,1410]" name="Zimbabwe" pageId="8" pageNumber="559">ZIMBABWE</collectingCountry>
:
<collectingRegion id="8767F8560835EA5D96B0FAD0A4461480" box="[765,1030,1383,1410]" country="Zimbabwe" name="Manicaland" pageId="8" pageNumber="559">Manicaland Province</collectingRegion>
:
</emphasis>
S. Rhodesia,
<collectingMunicipality id="A578ACCE0835EA5D90FEFADFA5501480" box="[1203,1296,1384,1410]" pageId="8" pageNumber="559">Inyanga</collectingMunicipality>
,
<location id="407C606F0835EA5D9152FADFA5DC1480" LSID="urn:lsid:plazi:treatment:CD0A87A2083AEA5C94DAF94CA2E81065:407C606F0835EA5D9152FADFA5DC1480" box="[1311,1436,1384,1410]" country="Zimbabwe" latitude="-18.283333" longLatPrecision="1275" longitude="32.766666" municipality="Inyanga" name="Mare Dam" pageId="8" pageNumber="559" stateProvince="Manicaland">Mare Dam</location>
[
<geoCoordinate id="209750730835EA5D94D2FA3CA0BD14A7" box="[159,253,1419,1445]" degrees="18" direction="south" minutes="17" orientation="latitude" pageId="8" pageNumber="559" precision="925" value="-18.283333">18°17S</geoCoordinate>
<geoCoordinate id="209750730835EA5D954EFA3CA12514A4" box="[259,357,1419,1446]" degrees="32" direction="east" minutes="46" orientation="longitude" pageId="8" pageNumber="559" precision="925" value="32.766666">32°46E</geoCoordinate>
],
<date id="311D10740835EA5D953BFA3CA19514A4" box="[374,469,1419,1446]" pageId="8" pageNumber="559" value="1958-07">
<collectingDate id="2159E99C0835EA5D953BFA3CA19514A4" box="[374,469,1419,1446]" pageId="8" pageNumber="559" value="1958-07">vii.1958</collectingDate>
</date>
,
<collectorName id="E85653620835EA5D9593FA3BA23214A7" box="[478,626,1419,1446]" pageId="8" pageNumber="559">A. Hoffmann</collectorName>
(
<specimenCount id="53A5FD3D0835EA5D96C9FA3CA2E514A7" box="[644,677,1419,1445]" pageId="8" pageNumber="559" type="female">1♀</specimenCount>
,
<collectionCode id="23B2AE710835EA5D96FCFA3CA34414A7" box="[689,772,1419,1445]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="559">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F5CB3CE90835EA5D9758FA3CA4D814C8" ID-GBIF-Occurrence="2423311073" collectingDate="1951-01" collectionCode="SAMC" collectorName="J. C. Faure" country="South Africa" latitude="-24.533333" location="Mariepskop" longLatPrecision="1250" longitude="30.883333" pageId="8" pageNumber="559" specimenCount="1" specimenCount-female="1" stateProvince="Mpumalanga">
<emphasis id="77D7EAA60835EA5D9758FA3CA55114A4" bold="true" box="[789,1297,1419,1446]" pageId="8" pageNumber="559">
<collectingCountry id="3DB476240835EA5D9758FA3CA3AD14A4" box="[789,1005,1419,1446]" name="South Africa" pageId="8" pageNumber="559">SOUTH AFRICA</collectingCountry>
:
<collectingRegion id="8767F8560835EA5D97B5FA3BA54C14A4" box="[1016,1292,1419,1446]" country="South Africa" name="Mpumalanga" pageId="8" pageNumber="559">Mpumalanga Province</collectingRegion>
:
</emphasis>
<location id="407C606F0835EA5D915BFA3CA5DC14A7" LSID="urn:lsid:plazi:treatment:CD0A87A2083AEA5C94DAF94CA2E81065:407C606F0835EA5D915BFA3CA5DC14A7" box="[1302,1436,1419,1445]" country="South Africa" latitude="-24.533333" longLatPrecision="1250" longitude="30.883333" name="Mariepskop" pageId="8" pageNumber="559" stateProvince="Mpumalanga">Mariepskop</location>
[
<geoCoordinate id="209750730835EA5D94D3FA18A0BC14CB" box="[158,252,1455,1481]" degrees="24" direction="south" minutes="32" orientation="latitude" pageId="8" pageNumber="559" precision="925" value="-24.533333">24°32S</geoCoordinate>
<geoCoordinate id="209750730835EA5D9549FA18A12514C8" box="[260,357,1455,1482]" degrees="30" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="559" precision="925" value="30.883333">30°53E</geoCoordinate>
], Tvl. [Transvaal],
<date id="311D10740835EA5D9676FA18A2C414C8" box="[571,644,1455,1482]" pageId="8" pageNumber="559" value="1951-01">
<collectingDate id="2159E99C0835EA5D9676FA18A2C414C8" box="[571,644,1455,1482]" pageId="8" pageNumber="559" value="1951-01">i.1951</collectingDate>
</date>
,
<collectorName id="E85653620835EA5D96DDFA18A34214C8" box="[656,770,1455,1482]" pageId="8" pageNumber="559">J.C. Faure</collectorName>
(
<specimenCount id="53A5FD3D0835EA5D9758FA18A37614CB" box="[789,822,1455,1481]" pageId="8" pageNumber="559" type="female">1♀</specimenCount>
,
<collectionCode id="23B2AE710835EA5D970EFA18A3D614CB" box="[835,918,1455,1481]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="8" pageNumber="559" type="Museum">SAMC</collectionCode>
; see comments below)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DB9653F0835EA5D948AFA64A3BF17E8" pageId="8" pageNumber="559" type="diagnosis">
<paragraph id="451C36B40835EA5D948AFA64A3BF17E8" blockId="8.[151,1437,1167,2022]" pageId="8" pageNumber="559">
<emphasis id="77D7EAA60835EA5D948AFA64A10214EC" bold="true" box="[199,322,1491,1518]" pageId="8" pageNumber="559">Diagnosis.</emphasis>
<taxonomicName id="82A34D370835EA5D9506FA64A25614EF" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[331,534,1491,1517]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D9506FA64A25614EF" box="[331,534,1491,1517]" italics="true" pageId="8" pageNumber="559">Phoberus lilianae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="82A34D370835EA5D96BCFA63A3CA14EF" authority="Strümpher &amp; Scholtz, 2019" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" box="[753,906,1491,1517]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="ntlenyanae" status="sp. nov.">
<emphasis id="77D7EAA60835EA5D96BCFA63A3CA14EF" box="[753,906,1491,1517]" italics="true" pageId="8" pageNumber="559">P. ntlenyanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CE457DD0835EA5D97DEFA63A45C14EF" box="[915,1052,1491,1518]" pageId="8" pageNumber="559" rank="species">new species</taxonomicNameLabel>
in appearance, but can be distinguished from the latter by the smaller, evenly spaced, dull round tubercles on the even-numbered costae of the elytra.
<taxonomicName id="82A34D370835EA5D94DAF9ACA1C51737" authority="Strümpher &amp; Scholtz, 2019" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" box="[151,389,1563,1589]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="ntlenyanae" status="sp. nov.">
<emphasis id="77D7EAA60835EA5D94DAF9ACA1C51737" box="[151,389,1563,1589]" italics="true" pageId="8" pageNumber="559">Phoberus ntlenyanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CE457DD0835EA5D95C1F9ABA2521737" box="[396,530,1563,1590]" pageId="8" pageNumber="559" rank="species">new species</taxonomicNameLabel>
has larger, shiny tubercles, which are irregularly spaced. The overall appearance of these species also differs noticeably:
<taxonomicName id="82A34D370835EA5D9613F9F7A291175B" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[606,721,1599,1625]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D9613F9F7A291175B" box="[606,721,1599,1625]" italics="true" pageId="8" pageNumber="559">P. lilianae</emphasis>
</taxonomicName>
is dull, whereas
<taxonomicName id="82A34D370835EA5D97DEF9F7A46B175B" authority="Strümpher &amp; Scholtz, 2019" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" box="[915,1067,1599,1625]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="ntlenyanae" status="sp. nov.">
<emphasis id="77D7EAA60835EA5D97DEF9F7A46B175B" box="[915,1067,1599,1625]" italics="true" pageId="8" pageNumber="559">P. ntlenyanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CE457DD0835EA5D9079F9F7A4FC175B" box="[1076,1212,1599,1626]" pageId="8" pageNumber="559" rank="species">new species</taxonomicNameLabel>
has a characteristic shiny reflection.
<taxonomicName id="82A34D370835EA5D9515F9D4A263177F" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[344,547,1635,1661]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D9515F9D4A263177F" box="[344,547,1635,1661]" italics="true" pageId="8" pageNumber="559">Phoberus lilianae</emphasis>
</taxonomicName>
also bears some resemblance to the montane members of the
<taxonomicName id="82A34D370835EA5D90A0F9D3A5DC177F" baseAuthorityName="Kolbe" baseAuthorityYear="1904" box="[1261,1436,1635,1662]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="77D7EAA60835EA5D90A0F9D3A4BE177F" box="[1261,1278,1636,1661]" italics="true" pageId="8" pageNumber="559">P</emphasis>
.
<emphasis id="77D7EAA60835EA5D9143F9D4A5DC177F" box="[1294,1436,1635,1661]" italics="true" pageId="8" pageNumber="559">nigrociliatus</emphasis>
</taxonomicName>
species group which have similar elytral markings and a similar overall appearance, but can be separated on the basis of their different pronota and elytra, different male genitalia, and by their distribution ranges (see
<bibRefCitation id="21324B450835EA5D916BF91CA10717EB" author="Strumpher, W. P. &amp; Scholtz, C. H." pageId="8" pageNumber="559" pagination="292 - 298" refId="ref9394" refString="Strumpher, W. P. &amp; Scholtz, C. H. (2017) A new brachypterous species of Phoberus MacLeay (Coleoptera: Trogidae) and status changes of species in the East African P. nigrociliatus group. Zootaxa, 4216 (3), 292 - 298. https: // doi. org / 10.11646 / zootaxa. 4216.3.6" type="journal article" year="2017">Strümpher &amp; Scholtz 2017</bibRefCitation>
for a detailed treatment of the
<taxonomicName id="82A34D370835EA5D96ECF967A30E17EB" baseAuthorityName="Kolbe" baseAuthorityYear="1904" box="[673,846,1743,1770]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="77D7EAA60835EA5D96ECF967A2F217EB" box="[673,690,1744,1769]" italics="true" pageId="8" pageNumber="559">P</emphasis>
.
<emphasis id="77D7EAA60835EA5D968DF978A30E17EB" box="[704,846,1743,1769]" italics="true" pageId="8" pageNumber="559">nigrociliatus</emphasis>
</taxonomicName>
species group).
</paragraph>
</subSubSection>
<subSubSection id="0DB9653F0835EA5D948AF944A4E01633" pageId="8" pageNumber="559" type="distribution">
<paragraph id="451C36B40835EA5D948AF944A4E01633" blockId="8.[151,1437,1167,2022]" pageId="8" pageNumber="559">
<emphasis id="77D7EAA60835EA5D948AF944A1B8160C" bold="true" box="[199,504,1779,1806]" pageId="8" pageNumber="559">Distribution and habitat.</emphasis>
<taxonomicName id="82A34D370835EA5D964FF944A28E160F" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[514,718,1779,1805]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D964FF944A28E160F" box="[514,718,1779,1805]" italics="true" pageId="8" pageNumber="559">Phoberus lilianae</emphasis>
</taxonomicName>
shows a disjunct montane distribution, being restricted to the Eastern Highlands of
<collectingCountry id="3DB476240835EA5D95C6F8A0A2431633" box="[395,515,1815,1841]" name="Zimbabwe" pageId="8" pageNumber="559">Zimbabwe</collectingCountry>
and the north-eastern escarpment of
<collectingCountry id="3DB476240835EA5D97EFF8A0A4701633" box="[930,1072,1815,1841]" name="South Africa" pageId="8" pageNumber="559">South Africa</collectingCountry>
(
<figureCitation id="DD982A310835EA5D9072F8A0A4D21633" box="[1087,1170,1815,1841]" captionStart="FIGURE 19" captionStartId="10.[151,250,1131,1156]" captionTargetBox="[158,1433,177,1106]" captionTargetId="figure@10.[151,1439,174,1111]" captionTargetPageId="10" captionText="FIGURE 19. Distribution of species in the southern African Phoberus caffer species group." figureDoi="http://doi.org/10.5281/zenodo.3467966" httpUri="https://zenodo.org/record/3467966/files/figure.png" pageId="8" pageNumber="559">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DB9653F0835EA5C948AF88CA2E81065" lastPageId="9" lastPageNumber="560" pageId="8" pageNumber="559" type="discussion">
<paragraph id="451C36B40835EA5C948AF88CA2E81065" blockId="8.[151,1437,1167,2022]" lastBlockId="9.[151,1437,153,359]" lastPageId="9" lastPageNumber="560" pageId="8" pageNumber="559">
<emphasis id="77D7EAA60835EA5D948AF88CA10F1654" bold="true" box="[199,335,1851,1878]" pageId="8" pageNumber="559">Comments.</emphasis>
The
<typeStatus id="9A1888160835EA5D95CBF88CA1AB1657" box="[390,491,1851,1877]" pageId="8" pageNumber="559" type="holotype">holotype</typeStatus>
, which is deposited in the National Museum,
<collectingRegion id="8767F8560835EA5D97A9F88CA4181657" box="[996,1112,1851,1877]" country="Zimbabwe" name="Bulawayo" pageId="8" pageNumber="559">Bulawayo</collectingRegion>
,
<collectingCountry id="3DB476240835EA5D902FF88CA49A1657" box="[1122,1242,1851,1877]" name="Zimbabwe" pageId="8" pageNumber="559">Zimbabwe</collectingCountry>
(NMBZ;
<bibRefCitation id="21324B450835EA5D9105F88CA0BC1678" author="Scholtz, C. H." pageId="8" pageNumber="559" pagination="1 - 104" refId="ref9058" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980: 96</bibRefCitation>
), could not be studied, but
<taxonomicName id="82A34D370835EA5D9664F8D7A2DA167B" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[553,666,1887,1913]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D9664F8D7A2DA167B" box="[553,666,1887,1913]" italics="true" pageId="8" pageNumber="559">P. lilianae</emphasis>
</taxonomicName>
is a well-defined species and the five
<typeStatus id="9A1888160835EA5D900DF8D7A4EB1678" box="[1088,1195,1888,1914]" pageId="8" pageNumber="559" type="paratype">paratypes</typeStatus>
housed in the SAMC and TMSA collections were sufficient for accurate comparison.
<bibRefCitation id="21324B450835EA5D9722F834A456169F" author="Scholtz, C. H." box="[879,1046,1923,1949]" pageId="8" pageNumber="559" pagination="205 - 208" refId="ref9132" refString="Scholtz, C. H. (1983) A new species of Trox F. (Coleoptera: Trogidae) from Senegal with new distribution records for other African species. Journal of the Entomological Society of Southern Africa, 46 (2), 205 - 208." type="journal article" year="1983">Scholtz (1983)</bibRefCitation>
recorded new distribution records for
<taxonomicName id="82A34D370835EA5D948DF81FA17316C3" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[192,307,1959,1985]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60835EA5D948DF81FA17316C3" box="[192,307,1959,1985]" italics="true" pageId="8" pageNumber="559">P. lilianae</emphasis>
</taxonomicName>
(then regarded as a subspecies of
<taxonomicName id="82A34D370835EA5D96F4F81FA37416C0" authorityName="Harold" authorityYear="1872" box="[697,820,1960,1986]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="caffer">
<emphasis id="77D7EAA60835EA5D96F4F81FA37416C0" box="[697,820,1960,1986]" italics="true" pageId="8" pageNumber="559">Trox caffer</emphasis>
</taxonomicName>
) from Mariepskop in
<collectingCountry id="3DB476240835EA5D907DF810A48016C3" box="[1072,1216,1959,1985]" name="South Africa" pageId="8" pageNumber="559">South Africa</collectingCountry>
[
<geoCoordinate id="209750730835EA5D9082F810A56D16C3" box="[1231,1325,1959,1985]" degrees="24" direction="south" minutes="32" orientation="latitude" pageId="8" pageNumber="559" precision="925" value="-24.533333">24°32S</geoCoordinate>
<geoCoordinate id="209750730835EA5D9178F810A5D516C0" box="[1333,1429,1959,1986]" degrees="30" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="559" precision="925" value="30.883333">30°53E</geoCoordinate>
] and the Nyika Plateau in northern
<collectingCountry id="3DB476240835EA5D9642F87CA22516E7" box="[527,613,1995,2021]" name="Malawi" pageId="8" pageNumber="559">Malawi</collectingCountry>
[
<geoCoordinate id="209750730835EA5D963FF87CA29016E7" box="[626,720,1995,2021]" degrees="10" direction="south" minutes="48" orientation="latitude" pageId="8" pageNumber="559" precision="925" value="-10.8">10°48S</geoCoordinate>
<geoCoordinate id="209750730835EA5D9698F87CA37816E4" box="[725,824,1995,2022]" degrees="33" direction="east" minutes="48" orientation="longitude" pageId="8" pageNumber="559" precision="925" value="33.8">33°48E</geoCoordinate>
]. After re-examination of these specimens, it was found that only the one (female) specimen recorded from Mariepskop belongs to the
<taxonomicName id="82A34D370834EA5C9059FF2DA43511B1" authorityName="Harold" authorityYear="1872" box="[1044,1141,153,179]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="560" phylum="Arthropoda" rank="species" species="caffer">
<emphasis id="77D7EAA60834EA5C9059FF2DA46511B1" box="[1044,1061,154,179]" italics="true" pageId="9" pageNumber="560">P</emphasis>
.
<emphasis id="77D7EAA60834EA5C9079FF2EA43511B1" box="[1076,1141,153,179]" italics="true" pageId="9" pageNumber="560">caffer</emphasis>
</taxonomicName>
species group. The specimens recorded from the Nyika plateau clearly belong to the East African
<taxonomicName id="82A34D370834EA5C9787FF09A43311D5" baseAuthorityName="Kolbe" baseAuthorityYear="1904" box="[970,1139,189,215]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="560" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="77D7EAA60834EA5C9787FF09A43311D5" box="[970,1139,189,215]" italics="true" pageId="9" pageNumber="560">P. nigrociliatus</emphasis>
</taxonomicName>
species group (see
<bibRefCitation id="21324B450834EA5C9103FF0AA1C311F9" author="Strumpher, W. P. &amp; Scholtz, C. H." pageId="9" pageNumber="560" pagination="292 - 298" refId="ref9394" refString="Strumpher, W. P. &amp; Scholtz, C. H. (2017) A new brachypterous species of Phoberus MacLeay (Coleoptera: Trogidae) and status changes of species in the East African P. nigrociliatus group. Zootaxa, 4216 (3), 292 - 298. https: // doi. org / 10.11646 / zootaxa. 4216.3.6" type="journal article" year="2017">Strümpher &amp; Scholtz 2017</bibRefCitation>
). The
<taxonomicName id="82A34D370834EA5C958BFF55A27611F9" authorityName="Strümpher &amp; Scholtz" authorityYear="2019" baseAuthorityName="Scholtz" baseAuthorityYear="1980" box="[454,566,225,251]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="560" phylum="Arthropoda" rank="species" species="lilianae">
<emphasis id="77D7EAA60834EA5C958BFF55A27611F9" box="[454,566,225,251]" italics="true" pageId="9" pageNumber="560">P. lilianae</emphasis>
</taxonomicName>
series comprises specimens collected from two different mountain ranges, the Inyanga Highlands (eastern
<collectingCountry id="3DB476240834EA5C9582FEB2A2091022" box="[463,585,261,288]" name="Zimbabwe" pageId="9" pageNumber="560">Zimbabwe</collectingCountry>
) and Drakensberg Range (eastern escarpment of
<collectingCountry id="3DB476240834EA5C902AFEB2A4B71022" box="[1127,1271,261,288]" name="South Africa" pageId="9" pageNumber="560">South Africa</collectingCountry>
) (
<figureCitation id="DD982A310834EA5C9141FEB2A51E101D" box="[1292,1374,261,287]" captionStart="FIGURE 19" captionStartId="10.[151,250,1131,1156]" captionTargetBox="[158,1433,177,1106]" captionTargetId="figure@10.[151,1439,174,1111]" captionTargetPageId="10" captionText="FIGURE 19. Distribution of species in the southern African Phoberus caffer species group." figureDoi="http://doi.org/10.5281/zenodo.3467966" httpUri="https://zenodo.org/record/3467966/files/figure.png" pageId="9" pageNumber="560">Fig. 19</figureCitation>
). The different mountain populations may be isolated to such an extent that they may well represent genetically divergent lineages, but this requires further investigation.
</paragraph>
</subSubSection>
</treatment>
</document>