262 lines
29 KiB
XML
262 lines
29 KiB
XML
<document id="A14DA372D72660D96E0CDB642378D822" ID-CLB-Dataset="3655" ID-DOI="10.11646/zootaxa.5067.1.10" ID-GBIF-Dataset="6a67bcc8-1bb3-41b2-8419-641126a9e64e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5656369" ID-ZooBank="27C8082E-56C5-4556-A47B-D6C866F3B41D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636444132939" checkinUser="plazi" docAuthor="Lukashevich, Elena D." docDate="2021" docId="C92E87B8FF95A935FF46FEB0D0B9FA62" docLanguage="en" docName="zootaxa.5067.1.10.pdf" docOrigin="Zootaxa 5067 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bashkonia Lukashevich 2021, gen. nov." docType="treatment" docVersion="7" lastPageNumber="137" masterDocId="3517FFC0FF97A937FFD1FF97D322FF87" masterDocTitle="The oldest Diptera (Insecta) from the Upper Buntsandstein (early Middle Triassic of Europe" masterLastPageNumber="143" masterPageNumber="135" pageNumber="137" updateTime="1699468025339" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F6E5D079E84AE68C54BEBD2101F1EA8E">
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<mods:title id="6B689A9D2D7E8285231D09AB3DF56A1B">The oldest Diptera (Insecta) from the Upper Buntsandstein (early Middle Triassic of Europe</mods:title>
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<mods:name id="A6C9962DE755AF10A4F63F274547A931" type="personal">
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<mods:roleTerm id="33125BDB1BD287BC505ED52A806255D3">Author</mods:roleTerm>
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<mods:namePart id="07FB7C80C0122D7C406961B000547001">Lukashevich, Elena D.</mods:namePart>
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<mods:title id="4C862E23FFEDFB94D2FA313E1627E80B">Zootaxa</mods:title>
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<mods:part id="222BCA4C748670F24238277F7BCA6BF8">
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<mods:date id="8ED8A998830CD6A8E391185963B4AEC2">2021</mods:date>
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<mods:number id="92E70680477B3030BE2E2AE87EFEF98E">2021-11-09</mods:number>
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<mods:number id="8A1731E802CBFD29E4FC9AEB16053730">5067</mods:number>
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<mods:classification id="223C21722776E01D9E29371D50E7BD09">journal article</mods:classification>
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<mods:identifier id="0E153815785CB25942166335B072BCF5" type="DOI">10.11646/zootaxa.5067.1.10</mods:identifier>
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<treatment id="C92E87B8FF95A935FF46FEB0D0B9FA62" ID-DOI="http://doi.org/10.5281/zenodo.5656385" ID-GBIF-Taxon="190244033" ID-Zenodo-Dep="5656385" LSID="urn:lsid:plazi:treatment:C92E87B8FF95A935FF46FEB0D0B9FA62" httpUri="http://treatment.plazi.org/id/C92E87B8FF95A935FF46FEB0D0B9FA62" lastPageNumber="137" pageId="2" pageNumber="137">
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<subSubSection id="099D6525FF95A935FF46FEB0D2ADFEC6" box="[151,399,295,322]" pageId="2" pageNumber="137" type="nomenclature">
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<paragraph id="413836AEFF95A935FF46FEB0D2ADFEC6" blockId="2.[151,399,295,357]" box="[151,399,295,322]" pageId="2" pageNumber="137">
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<heading id="1A7081C2FF95A935FF46FEB0D2ADFEC6" bold="true" box="[151,399,295,322]" fontSize="11" level="1" pageId="2" pageNumber="137" reason="1">
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<emphasis id="73F3EABCFF95A935FF46FEB0D2ADFEC6" bold="true" box="[151,399,295,322]" pageId="2" pageNumber="137">
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<taxonomicName id="86874D2DFF95A935FF46FEB0D23BFEC5" ID-CoL="9HLNV" authority="Lukashevich, 2021" authorityName="Lukashevich" authorityYear="2021" box="[151,281,295,322]" class="Insecta" family="Nadipteridae" genus="Bashkonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="73F3EABCFF95A935FF46FEB0D23BFEC5" bold="true" box="[151,281,295,322]" italics="true" pageId="2" pageNumber="137">Bashkonia</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="68C057C7FF95A935FEF0FEB0D2ADFEC6" box="[289,399,295,321]" pageId="2" pageNumber="137" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="099D6525FF95A935FF46FEDCD22AFEE2" box="[151,264,331,357]" pageId="2" pageNumber="137" type="description">
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<paragraph id="413836AEFF95A935FF46FEDCD22AFEE2" blockId="2.[151,399,295,357]" box="[151,264,331,357]" pageId="2" pageNumber="137">
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(
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<figureCitation id="D9BC2A2BFF95A935FF4EFEDCD3DFFEE2" box="[159,253,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[173,1414,181,1912]" captionTargetId="figure-16@3.[173,1414,181,1920]" captionTargetPageId="3" captionText="FIGURE 1. Bashkonia franconica gen. et sp. nov., holotype SMTE 6123/1-61 (Lengfurt, Germany, Tr2): A, B. Positive and negative impression of isolated wing. C. Line drawing of wing. Scale bars equal 1mm." figureDoi="http://doi.org/10.5281/zenodo.5656371" httpUri="https://zenodo.org/record/5656371/files/figure.png" pageId="2" pageNumber="137">Figure 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="099D6525FF95A935FF46FE04D1ABFE2A" box="[151,649,403,429]" pageId="2" pageNumber="137" type="type_taxon">
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<paragraph id="413836AEFF95A935FF46FE04D1ABFE2A" blockId="2.[151,1437,403,1510]" box="[151,649,403,429]" pageId="2" pageNumber="137">
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||
<emphasis id="73F3EABCFF95A935FF46FE04D209FE2A" bold="true" box="[151,299,403,429]" pageId="2" pageNumber="137">
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<typeStatus id="9E3C880CFF95A935FF46FE04D3F0FE2A" box="[151,210,403,429]" pageId="2" pageNumber="137">Type</typeStatus>
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species
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</emphasis>
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<taxonomicName id="86874D2DFF95A935FEE3FE04D10AFE2A" ID-CoL="9HLRX" authority="Lukashevich, 2021" authorityName="Lukashevich" authorityYear="2021" box="[306,552,403,429]" class="Insecta" family="Nadipteridae" genus="Bashkonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="franconica" status="sp. nov.">
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<emphasis id="73F3EABCFF95A935FEE3FE04D10AFE2A" box="[306,552,403,429]" italics="true" pageId="2" pageNumber="137">Bashkonia franconica</emphasis>
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</taxonomicName>
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<emphasis id="73F3EABCFF95A935FDE1FE04D1ABFE2A" bold="true" box="[560,649,403,429]" pageId="2" pageNumber="137">
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<taxonomicNameLabel id="68C057C7FF95A935FDE1FE04D1ABFE2A" box="[560,649,403,429]" pageId="2" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="099D6525FF95A935FF16FE20D2F4FE72" pageId="2" pageNumber="137" type="etymology">
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<paragraph id="413836AEFF95A935FF16FE20D2F4FE72" blockId="2.[151,1437,403,1510]" pageId="2" pageNumber="137">
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<emphasis id="73F3EABCFF95A935FF16FE20D26CFE56" bold="true" box="[199,334,439,465]" pageId="2" pageNumber="137">Etymology.</emphasis>
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The generic epithet refers to Alexey Bashkuev, a recognized collector and expert on Triassic insects. The name is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="099D6525FF95A935FF16FE68D26FFBDE" pageId="2" pageNumber="137" type="diagnosis">
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<paragraph id="413836AEFF95A935FF16FE68D021FCDA" blockId="2.[151,1437,403,1510]" pageId="2" pageNumber="137">
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<emphasis id="73F3EABCFF95A935FF16FE68D260FD9E" bold="true" box="[199,322,511,537]" pageId="2" pageNumber="137">Diagnosis.</emphasis>
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Wings elongate, pterostigma large. Crossvein
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<emphasis id="73F3EABCFF95A935FC8FFD97D0A8FD9E" box="[862,906,512,537]" italics="true" pageId="2" pageNumber="137">sc-r</emphasis>
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far from Rs origin. R kink long and shallow, opposed to long M+CuA fusion; arculus inclined; R
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<subScript id="DD0334EBFF95A935FD31FDA1D1DBFDC3" attach="left" box="[736,761,566,580]" fontSize="6" pageId="2" pageNumber="137">2+3</subScript>
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fork short and wide; Rs relatively long, more than 2/3 wing length; R
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<subScript id="DD0334EBFF95A935FF2FFDCED23AFDE0" attach="left" box="[254,280,601,615]" fontSize="6" pageId="2" pageNumber="137">4+5</subScript>
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fork long, 10 times longer than R
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<subScript id="DD0334EBFF95A935FD58FDCED181FDE0" attach="left" box="[649,675,601,615]" fontSize="6" pageId="2" pageNumber="137">4+5</subScript>
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stem; R
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<subScript id="DD0334EBFF95A935FD2AFDCED021FDE0" attach="left" box="[763,771,601,615]" fontSize="6" pageId="2" pageNumber="137">5</subScript>
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convexity continued along Rs up to MA, forming false vein
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<emphasis id="73F3EABCFF95A935FF46FDFCD389FD02" box="[151,171,619,645]" italics="true" pageId="2" pageNumber="137">fv</emphasis>
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between R and M sectors;
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<emphasis id="73F3EABCFF95A935FE0DFDFBD126FD02" box="[476,516,620,645]" italics="true" pageId="2" pageNumber="137">r-m</emphasis>
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jointing R
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<subScript id="DD0334EBFF95A935FDAAFDEAD1A1FD0C" attach="left" box="[635,643,637,651]" fontSize="6" pageId="2" pageNumber="137">5</subScript>
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base and basal 1/3 of M
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<subScript id="DD0334EBFF95A935FC45FDE9D08FFD0B" attach="left" box="[916,941,638,652]" fontSize="6" pageId="2" pageNumber="137">1+2</subScript>
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stem. M forking before Rs bifurcation, near wing midlength; M
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<subScript id="DD0334EBFF95A935FEA2FD35D2AEFD37" attach="left" box="[371,396,674,688]" fontSize="6" pageId="2" pageNumber="137">1+2</subScript>
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and M
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<subScript id="DD0334EBFF95A935FE0DFD35D2D4FD37" attach="left" box="[476,502,674,688]" fontSize="6" pageId="2" pageNumber="137">3+4</subScript>
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forks rather long and subequal;
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<emphasis id="73F3EABCFF95A935FCBBFD07D0A4FD2E" box="[874,902,656,681]" italics="true" pageId="2" pageNumber="137">im</emphasis>
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connecting M
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<subScript id="DD0334EBFF95A935FBFAFD35D767FD37" attach="left" box="[1067,1093,674,688]" fontSize="6" pageId="2" pageNumber="137">1+2</subScript>
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stem and M
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<subScript id="DD0334EBFF95A935FB03FD35D7F8FD37" attach="left" box="[1234,1242,674,688]" fontSize="6" pageId="2" pageNumber="137">3</subScript>
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; M
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<subScript id="DD0334EBFF95A935FAD2FD35D63FFD37" attach="left" box="[1283,1309,674,688]" fontSize="6" pageId="2" pageNumber="137">1+2</subScript>
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forking far beyond
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<emphasis id="73F3EABCFF95A935FF20FD23D235FD4A" box="[241,279,692,717]" italics="true" pageId="2" pageNumber="137">im;</emphasis>
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M
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<subScript id="DD0334EBFF95A935FEE9FD51D273FD53" attach="left" box="[312,337,710,724]" fontSize="6" pageId="2" pageNumber="137">3+4</subScript>
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forking clearly distal to R
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<subScript id="DD0334EBFF95A935FDA8FD52D1B1FD54" attach="left" box="[633,659,709,723]" fontSize="6" pageId="2" pageNumber="137">4+5</subScript>
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furcation; d cell subequal to M
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<subScript id="DD0334EBFF95A935FC22FD51D72FFD53" attach="left" box="[1011,1037,710,724]" fontSize="6" pageId="2" pageNumber="137">3+4</subScript>
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fork;
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<emphasis id="73F3EABCFF95A935FB81FD23D7AAFD4A" box="[1104,1160,692,717]" italics="true" pageId="2" pageNumber="137">m-cu</emphasis>
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in distal wing half, near midlength of d cell, ending on straight M
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<subScript id="DD0334EBFF95A935FDBDFD7DD1A7FD7F" attach="left" box="[620,645,746,760]" fontSize="6" pageId="2" pageNumber="137">3+4</subScript>
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. CuA bent at
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<emphasis id="73F3EABCFF95A935FCF5FD4FD07EFD76" box="[804,860,728,753]" italics="true" pageId="2" pageNumber="137">m-cu</emphasis>
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; CuA apex closer to R
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||
<subScript id="DD0334EBFF95A935FBB1FD7DD74AFD7F" attach="left" box="[1120,1128,746,760]" fontSize="6" pageId="2" pageNumber="137">1</subScript>
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apex than to Sc apex; CuP entering margin near wing midlength, strongly diverging from CuA in its distal 1/4, CuP being weak after its divergence with cubital fold
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<emphasis id="73F3EABCFF95A935FE71FCB7D2E0FCBE" box="[416,450,800,825]" italics="true" pageId="2" pageNumber="137">icu</emphasis>
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. Two cubital folds well developed: curved one behind entire length of CuA and straight one
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<emphasis id="73F3EABCFF95A935FF16FCD3D3CBFCDA" box="[199,233,836,861]" italics="true" pageId="2" pageNumber="137">icu</emphasis>
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before CuP to CuA apex. A
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||
<subScript id="DD0334EBFF95A935FDF0FCC1D10BFCE3" attach="left" box="[545,553,854,868]" fontSize="6" pageId="2" pageNumber="137">1</subScript>
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||
distal to
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<emphasis id="73F3EABCFF95A935FD40FCD3D1E1FCDA" box="[657,707,836,861]" italics="true" pageId="2" pageNumber="137">cu-a</emphasis>
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long.
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</paragraph>
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<paragraph id="413836AEFF95A935FF16FCF0D13CFC06" blockId="2.[151,1437,403,1510]" box="[199,542,871,897]" pageId="2" pageNumber="137">
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<emphasis id="73F3EABCFF95A935FF16FCF0D2B1FC06" bold="true" box="[199,403,871,897]" pageId="2" pageNumber="137">Species included.</emphasis>
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Monobasic.
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</paragraph>
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<paragraph id="413836AEFF95A935FF16FC1CD26FFBDE" blockId="2.[151,1437,403,1510]" pageId="2" pageNumber="137">
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<emphasis id="73F3EABCFF95A935FF16FC1CD240FC22" bold="true" box="[199,354,907,933]" pageId="2" pageNumber="137">Comparison.</emphasis>
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The new genus differs from both other genera of
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<taxonomicName id="86874D2DFF95A935FC54FC1CD73BFC22" authorityName="Lukashevich" authorityYear="1995" box="[901,1049,907,933]" class="Insecta" family="Nadipteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="family">Nadipteridae</taxonomicName>
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||
,
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||
<taxonomicName id="86874D2DFF95A935FBF3FC1CD64CFC21" authority="Lukashevich, 1995" authorityName="Lukashevich" authorityYear="1995" box="[1058,1390,907,934]" class="Insecta" family="Nadipteridae" genus="Nadiptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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||
<emphasis id="73F3EABCFF95A935FBF3FC1CD7B6FC22" box="[1058,1172,907,933]" italics="true" pageId="2" pageNumber="137">Nadiptera</emphasis>
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Lukashevich, 1995
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||
</taxonomicName>
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||
and
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||
<taxonomicName id="86874D2DFF95A935FF46FC27D16DFC4E" authority="Krzeminski & Krzeminska, 2003" authorityName="Krzeminski & Krzeminska" authorityYear="2003" box="[151,591,943,969]" class="Insecta" family="Nadipteridae" genus="Tanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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<emphasis id="73F3EABCFF95A935FF46FC27D3FBFC4E" box="[151,217,944,969]" italics="true" pageId="2" pageNumber="137">Tanus</emphasis>
|
||
<bibRefCitation id="25164B5FFF95A935FF0EFC38D16DFC4E" author="Krzeminski, W. & Krzeminska, E." box="[223,591,943,969]" pageId="2" pageNumber="137" pagination="153 - 184" refId="ref4497" refString="Krzeminski, W. & Krzeminska, E. (2003) Triassic Diptera: descriptions, revisions and phylogenetic relations. Acta Zoologica Cracoviensia, 46, 153 - 184." type="journal article" year="2003">
|
||
Krzemiński & Krzemińska,
|
||
<quantity id="867F9B4BFF95A935FDC6FC38D16DFC4E" box="[535,591,943,969]" metricMagnitude="1" metricUnit="m" metricValue="5.087619999999999" pageId="2" pageNumber="137" unit="in" value="2003.0">2003</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in
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||
<emphasis id="73F3EABCFF95A935FDA1FC27D1BEFC4E" box="[624,668,944,969]" italics="true" pageId="2" pageNumber="137">sc-r</emphasis>
|
||
position (closer to Rs origin than to Sc apex in
|
||
<taxonomicName id="86874D2DFF95A935FB75FC38D634FC4E" authorityName="Lukashevich" authorityYear="1995" box="[1188,1302,943,969]" class="Insecta" family="Nadipteridae" genus="Nadiptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FB75FC38D634FC4E" box="[1188,1302,943,969]" italics="true" pageId="2" pageNumber="137">Nadiptera</emphasis>
|
||
</taxonomicName>
|
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and
|
||
<taxonomicName id="86874D2DFF95A935FA9BFC27D6AEFC4E" authorityName="Krzeminski & Krzeminska" authorityYear="2003" box="[1354,1420,944,969]" class="Insecta" family="Nadipteridae" genus="Tanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FA9BFC27D6AEFC4E" box="[1354,1420,944,969]" italics="true" pageId="2" pageNumber="137">Tanus</emphasis>
|
||
</taxonomicName>
|
||
), shorter Rs and in CuA, which is longer (instead of apex, closer to level of Sc apex than to R
|
||
<subScript id="DD0334EBFF95A935FB77FC71D78CFC73" attach="left" box="[1190,1198,998,1012]" fontSize="6" pageId="2" pageNumber="137">1</subScript>
|
||
apex) and bent at
|
||
<emphasis id="73F3EABCFF95A935FAAEFC43D393FB96" italics="true" pageId="2" pageNumber="137">mcu</emphasis>
|
||
. The new genus shares with
|
||
<taxonomicName id="86874D2DFF95A935FE2BFC60D14EFB96" authorityName="Lukashevich" authorityYear="1995" box="[506,620,1015,1041]" class="Insecta" family="Nadipteridae" genus="Nadiptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FE2BFC60D14EFB96" box="[506,620,1015,1041]" italics="true" pageId="2" pageNumber="137">Nadiptera</emphasis>
|
||
</taxonomicName>
|
||
false vein
|
||
<emphasis id="73F3EABCFF95A935FD36FC60D1D9FB96" box="[743,763,1015,1041]" italics="true" pageId="2" pageNumber="137">fv</emphasis>
|
||
between R and M sectors and CuP strongly diverging from CuA (
|
||
<figureCitation id="D9BC2A2BFF95A935FF0BFB8CD210FBB2" box="[218,306,1051,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1220,1244]" captionTargetBox="[151,1435,204,1172]" captionTargetId="figure-24@4.[151,1435,204,1172]" captionTargetPageId="4" captionText="FIGURE 2. SEM images (uncoated, BSE) of Diptera wings:A. Nadiptera anachrona Lukashevich, 1995 (Nadipteridae), holo- type PIN No. 371/926,?male, left wing (Sogyuty, Kyrgyzstan, J 1). B. Vladiptera kovalevi Shcherbakov, 1995 (Vladipteridae), holotype PIN No. 2497/305, isolated wing (Kendyrlik, Kazakhstan, Tr 3). Scale bars equal 1mm." figureDoi="http://doi.org/10.5281/zenodo.5656373" httpUri="https://zenodo.org/record/5656373/files/figure.png" pageId="2" pageNumber="137">Fig. 2A</figureCitation>
|
||
), whereas it shares with
|
||
<taxonomicName id="86874D2DFF95A935FD96FB8BD1ABFBB2" authorityName="Krzeminski & Krzeminska" authorityYear="2003" box="[583,649,1052,1077]" class="Insecta" family="Nadipteridae" genus="Tanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FD96FB8BD1ABFBB2" box="[583,649,1052,1077]" italics="true" pageId="2" pageNumber="137">Tanus</emphasis>
|
||
</taxonomicName>
|
||
M furcation and CuP apex in midlength of the wing and
|
||
<emphasis id="73F3EABCFF95A935FAD9FB8BD612FBB2" box="[1288,1328,1052,1077]" italics="true" pageId="2" pageNumber="137">r-m</emphasis>
|
||
proximal to
|
||
<emphasis id="73F3EABCFF95A935FF65FBD7D3CEFBDE" box="[180,236,1088,1113]" italics="true" pageId="2" pageNumber="137">m-cu</emphasis>
|
||
(
|
||
<figureCitation id="D9BC2A2BFF95A935FF2AFBA8D262FBDE" box="[251,320,1087,1113]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-16@5.[178,1409,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Photos of Tanus triassicus Krzemiński & Krzemińska, 2003 (Nadipteridae), holotype No. 5336, male: A. Total view. B, C. Left and right wings (Vosges, France, Tr2). Arrows mark crossvein m-cu. Scale bars equal 1mm." figureDoi="http://doi.org/10.5281/zenodo.5656375" httpUri="https://zenodo.org/record/5656375/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="099D6525FF95A935FF16FBF4D0B9FA62" pageId="2" pageNumber="137" type="discussion">
|
||
<paragraph id="413836AEFF95A935FF16FBF4D7C2FA8A" blockId="2.[151,1437,403,1510]" pageId="2" pageNumber="137">
|
||
<emphasis id="73F3EABCFF95A935FF16FBF4D219FBFA" bold="true" box="[199,315,1123,1149]" pageId="2" pageNumber="137">Remarks.</emphasis>
|
||
Characters typical of the family are observed in the wing venation of the new genus: Sc apex beyond wing midlength, distal to Rs bifurcation, the crossvein
|
||
<emphasis id="73F3EABCFF95A935FD2DFB1FD00AFB26" box="[764,808,1160,1185]" italics="true" pageId="2" pageNumber="137">sc-r</emphasis>
|
||
not terminal; R distinctly kinked in horizontal plane, Rs forking symmetrically before Sc apex, free R
|
||
<subScript id="DD0334EBFF95A935FD45FB29D1BEFB4B" attach="left" box="[660,668,1214,1228]" fontSize="6" pageId="2" pageNumber="137">2</subScript>
|
||
reaching wing margin, R
|
||
<subScript id="DD0334EBFF95A935FC6AFB2AD0F7FB4C" attach="left" box="[955,981,1213,1227]" fontSize="6" pageId="2" pageNumber="137">4+5</subScript>
|
||
fork several times longer than short R
|
||
<subScript id="DD0334EBFF95A935FA52FB2AD6BFFB4C" attach="left" box="[1411,1437,1213,1227]" fontSize="6" pageId="2" pageNumber="137">4+5</subScript>
|
||
stem; long convex M base aligned with CuA, M stem forking symmetrically, d cell narrow and long with
|
||
<emphasis id="73F3EABCFF95A935FAFDFB47D646FB6E" box="[1324,1380,1232,1257]" italics="true" pageId="2" pageNumber="137">m-cu</emphasis>
|
||
ending on straight М
|
||
<subScript id="DD0334EBFF95A935FE88FA91D251FA93" attach="left" box="[345,371,1286,1300]" fontSize="6" pageId="2" pageNumber="137">3+4</subScript>
|
||
; A
|
||
<subScript id="DD0334EBFF95A935FE45FA91D2BEFA93" attach="left" box="[404,412,1286,1300]" fontSize="6" pageId="2" pageNumber="137">1</subScript>
|
||
not reaching margin, A
|
||
<subScript id="DD0334EBFF95A935FD74FA91D18FFA93" attach="left" box="[677,685,1286,1300]" fontSize="6" pageId="2" pageNumber="137">2</subScript>
|
||
forming anal loop, anal lobe somewhat produced.
|
||
</paragraph>
|
||
<paragraph id="413836AEFF95A935FF16FA80D0B9FA62" blockId="2.[151,1437,403,1510]" pageId="2" pageNumber="137">
|
||
The cubital fold
|
||
<emphasis id="73F3EABCFF95A935FE55FA8FD284FAB6" box="[388,422,1304,1329]" italics="true" pageId="2" pageNumber="137">icu</emphasis>
|
||
reaching CuA apex, found in the new genus, is also present in other Triassic
|
||
<taxonomicName id="86874D2DFF95A935FAC6FA80D64CFAB6" baseAuthorityName="Cumming & Wood" baseAuthorityYear="2017" box="[1303,1390,1303,1329]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
|
||
, the tipulomorph
|
||
<taxonomicName id="86874D2DFF95A935FEFBFAACD1A3FAD1" authority="Shcherbakov, 1995" authorityName="Shcherbakov" authorityYear="1995" box="[298,641,1339,1366]" class="Insecta" family="Vladipteridae" genus="Vladiptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FEFBFAACD280FAD2" box="[298,418,1339,1365]" italics="true" pageId="2" pageNumber="137">Vladiptera</emphasis>
|
||
<bibRefCitation id="25164B5FFF95A935FE78FAACD1A3FAD1" author="Shcherbakov, D. E. & Lukashevich, E. D. & Blagoderov, V. A." box="[425,641,1339,1366]" pageId="2" pageNumber="137" pagination="75 - 115" refId="ref5181" refString="Shcherbakov, D. E., Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6 (2), 75 - 115." type="journal article" year="1995">Shcherbakov, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="86874D2DFF95A935FD41FAACD00CFAD2" authorityName="Shcherbakov" authorityYear="1995" box="[656,814,1339,1365]" class="Insecta" family="Vladipteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="family">Vladipteridae</taxonomicName>
|
||
;
|
||
<figureCitation id="D9BC2A2BFF95A935FCEBFAACD0B2FAD2" box="[826,912,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[151,250,1220,1244]" captionTargetBox="[151,1435,204,1172]" captionTargetId="figure-24@4.[151,1435,204,1172]" captionTargetPageId="4" captionText="FIGURE 2. SEM images (uncoated, BSE) of Diptera wings:A. Nadiptera anachrona Lukashevich, 1995 (Nadipteridae), holo- type PIN No. 371/926,?male, left wing (Sogyuty, Kyrgyzstan, J 1). B. Vladiptera kovalevi Shcherbakov, 1995 (Vladipteridae), holotype PIN No. 2497/305, isolated wing (Kendyrlik, Kazakhstan, Tr 3). Scale bars equal 1mm." figureDoi="http://doi.org/10.5281/zenodo.5656373" httpUri="https://zenodo.org/record/5656373/files/figure.png" pageId="2" pageNumber="137">Fig. 2B</figureCitation>
|
||
) and the psychodomorph
|
||
<taxonomicName id="86874D2DFF95A935FB66FAACD199FAFD" authority="Lukashevich & Shcherbakov, 1999" authorityName="Lukashevich & Shcherbakov" authorityYear="1999" class="Insecta" family="Tillyardipteridae" genus="Tillyardiptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="73F3EABCFF95A935FB66FAACD670FAD2" box="[1207,1362,1339,1365]" italics="true" pageId="2" pageNumber="137">Tillyardiptera</emphasis>
|
||
<bibRefCitation id="25164B5FFF95A935FA88FAACD2C7FAFE" author="Lukashevich, E. D. & Shcherbakov, D. E." pageId="2" pageNumber="137" pagination="81 - 89" refId="ref4805" refString="Lukashevich, E. D. & Shcherbakov, D. E. (1999) A new Triassic family of Diptera from Australia. In: Proceedings of the 1 st International Palaeoentomological Conference, Moscow 1998. AMBA project, Bratislava, pp. 81 - 89." type="book chapter" year="1999">Lukashevich & Shcherbakov, 1999</bibRefCitation>
|
||
(Tillyardipteridae)
|
||
</taxonomicName>
|
||
, but they were missed in the original descriptions (
|
||
<bibRefCitation id="25164B5FFF95A935FB3FFAC8D3D5FA19" author="Shcherbakov, D. E. & Lukashevich, E. D. & Blagoderov, V. A." pageId="2" pageNumber="137" pagination="75 - 115" refId="ref5181" refString="Shcherbakov, D. E., Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6 (2), 75 - 115." type="journal article" year="1995">
|
||
Shcherbakov
|
||
<emphasis id="73F3EABCFF95A935FA59FAF7D38FFA1A" italics="true" pageId="2" pageNumber="137">et al</emphasis>
|
||
. 1995
|
||
</bibRefCitation>
|
||
: fig. 24;
|
||
<bibRefCitation id="25164B5FFF95A935FE88FA14D1C3FA1A" author="Lukashevich, E. D. & Shcherbakov, D. E." box="[345,737,1411,1437]" pageId="2" pageNumber="137" pagination="81 - 89" refId="ref4805" refString="Lukashevich, E. D. & Shcherbakov, D. E. (1999) A new Triassic family of Diptera from Australia. In: Proceedings of the 1 st International Palaeoentomological Conference, Moscow 1998. AMBA project, Bratislava, pp. 81 - 89." type="book chapter" year="1999">Lukashevich & Shcherbakov 1999</bibRefCitation>
|
||
: fig.1). Such cubital fold is not a common character of
|
||
<taxonomicName id="86874D2DFF95A935FA99FA14D6BEFA1A" baseAuthorityName="Cumming & Wood" baseAuthorityYear="2017" box="[1352,1436,1411,1437]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
|
||
but it is well known in extant and extinct
|
||
<taxonomicName id="86874D2DFF95A935FDA0FA30D03BFA46" baseAuthorityName="Lukashevich" baseAuthorityYear="2012" box="[625,793,1447,1473]" class="Insecta" family="Ptychopteridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="family">Ptychopteridae</taxonomicName>
|
||
as well as false vein
|
||
<emphasis id="73F3EABCFF95A935FBDCFA30D703FA46" box="[1037,1057,1447,1473]" italics="true" pageId="2" pageNumber="137">fv</emphasis>
|
||
between R and M sectors and A
|
||
<subScript id="DD0334EBFF95A935FA45FA2DD6BEFA4F" attach="left" box="[1428,1436,1466,1480]" fontSize="6" pageId="2" pageNumber="137">1</subScript>
|
||
retaining distal to
|
||
<emphasis id="73F3EABCFF95A935FEB3FA5BD2A4FA62" box="[354,390,1484,1509]" italics="true" pageId="2" pageNumber="137">cu-</emphasis>
|
||
a as a rather long spur (e.g.
|
||
<bibRefCitation id="25164B5FFF95A935FD6DFA5CD0ADFA62" author="Lukashevich, E. D." box="[700,911,1483,1509]" pageId="2" pageNumber="137" pagination="476 - 484" refId="ref4584" refString="Lukashevich, E. D. (2012) Phylogeny of Ptychopteroidea (Insecta: Diptera). Paleontologicheskii Zhurnal, 5, 32 - 40. [translated into English in Paleontological Journal, 46 (5), 476 - 484] https: // doi. org / 10.1134 / S 003103011205005 X" type="journal article" year="2012">Lukashevich 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |