168 lines
17 KiB
XML
168 lines
17 KiB
XML
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<mods:title id="00D9305A2847B01AC9458A0DACD64AB6">Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
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<mods:namePart id="7C8DD6755F162112E5A60509BE6FF9FE">Song, Zhi-Shun</mods:namePart>
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<mods:namePart id="5B1D3D6B3FB0B7E6CB67BC17F49A102A">Malenovsky, Igor</mods:namePart>
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<mods:namePart id="255629F5A7BA029D9137726ADFF01D2D">Chen, Jian-Qin</mods:namePart>
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<mods:namePart id="28E7F6D282F5DEFF404BDD8C891BFB11">Liang, Ai-Ping</mods:namePart>
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<mods:date id="17C879316AAEEB4D4F178D4E6F8D0005">2018</mods:date>
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<mods:number id="27256C853904E1109FF6FD6DE8F4F20B">94</mods:number>
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<treatment id="C6D9FE307B021118B52352433282E596" ID-GBIF-Taxon="147218784" LSID="urn:lsid:plazi:treatment:C6D9FE307B021118B52352433282E596" httpUri="http://treatment.plazi.org/id/C6D9FE307B021118B52352433282E596" lastPageNumber="377" pageId="8" pageNumber="377">
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<subSubSection id="238CC00FA20BD722C544CB2FAECC1352" pageId="8" pageNumber="377" type="nomenclature">
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<paragraph id="38C442F767107963E687F86469F5964E" pageId="8" pageNumber="377">
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<taxonomicName id="8BB727D9A3394862EF7A1485C47589C5" ID-CoL="6SWQ4" authority="Melichar, 1914" authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus Melichar, 1914</taxonomicName>
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Figs 2
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<normalizedToken id="E75B0D5242C35F988BBD8DBBA5FF0C72" originalValue="I–J">I-J</normalizedToken>
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, 3E, 9
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<normalizedToken id="0C48B1AD5EDD2CF01CA675ADD4ECD8A5" originalValue="A–J">A-J</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="C5AA6142EBB21371A91683D7E13D4489" pageId="8" pageNumber="377" type="reference_group">
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<paragraph id="AE4DD9BBD90D0B9FC7AE272F04DFDB29" pageId="8" pageNumber="377">
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<taxonomicName id="232DF2B864B6D7D32B4E0477E3579FA9" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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Melichar, 1914: 173, Pl. I, figs 1, 2. Lectotype (here designated) male, Los
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<normalizedToken id="B95B7DDC6527274A057225B60A2878F4" originalValue="Baños">Banos</normalizedToken>
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, Philippines (MMBC, examined).
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</paragraph>
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</subSubSection>
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<subSubSection id="99C9C3827D3B0E7FB834B531538D8E08" pageId="8" pageNumber="377" type="type material">
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<paragraph id="4E38CB8BFC8D2A041CA85C3A93D6959A" pageId="8" pageNumber="377">Type material examined.</paragraph>
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<paragraph id="DFF8792D5515AF3589C13F1146709CBB" pageId="8" pageNumber="377">
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Lectotype male (here designated), (1) Los Banos, P.I. Baker; (2) 1311; (3)
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<taxonomicName id="B2A138D7C973AC32AB602C0DA7A777B2" genus="Fulgoromorpha" lsidName="Fulgoromorpha philippinus" pageId="8" pageNumber="377" rank="species" species="philippinus">philippinus</taxonomicName>
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[
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<normalizedToken id="C4762BF31A19E708E2219CCCE1FF135B" originalValue="Melichar’s">Melichar's</normalizedToken>
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handwriting] det. Melichar; (4) Collectio Dr. L. Melichar,
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<normalizedToken id="6797D54078430736B4297A31BA96982A" originalValue="Moravské">Moravske</normalizedToken>
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museum Brno; (5)
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<taxonomicName id="5990ACE858DC7F673147385ECCBEE61C" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="genus">Orthopagus</taxonomicName>
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male
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<taxonomicName id="A1F68EC90138AB541E2EE8BF3CD22C57" genus="Fulgoromorpha" lsidName="Fulgoromorpha philippinus" pageId="8" pageNumber="377" rank="species" species="philippinus">philippinus</taxonomicName>
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sp.n., L. Melichar, 1914 [
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<normalizedToken id="CDDA065E0DA3D4E5243222911F135082" originalValue="Lauterer’s">Lauterer's</normalizedToken>
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handwriting], P. Lauterer det. 1991; (6) Syn-typus [red label]; (7) Invent.
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<normalizedToken id="2E22539C9CE6BEFEE1EDA9E48F2A39C9" originalValue="č">c</normalizedToken>
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. 4954/Ent., Mor. muzeum, Brno; (8) Lectotypus male,
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<taxonomicName id="45CC463AF560202BCB95BD960EAFABCA" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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Melichar, 1914, designated by Z. S. Song & I.
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<normalizedToken id="246B06A1C380AFA3F3183B13840D7732" originalValue="Malenovský">Malenovsky</normalizedToken>
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, 2018 [newly added red label] (MMBC).
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</paragraph>
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<paragraph id="39053F6790DDD8B6F52C8714A73F74C8" pageId="8" pageNumber="377">
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Paralectotypes, 4 females, same locality labels as holotype but Inv. nos 4949-4951, 4953;1 female, Mt. Makiling, Luzon, Baker (MMBC, Inv. no. 4952, all paralectotypes bearing the following label: Paralectotypus female,
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<taxonomicName id="0BC9DA3BDC44E49830CD25B173BA679D" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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Melichar, 1914, designated by Z. S. Song & I.
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<normalizedToken id="D2B519FAE17AB80200ED7FADD3E7032D" originalValue="Malenovský">Malenovsky</normalizedToken>
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, 2018).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C88047F3E88CFB9E3D603AFA31D372A4" pageId="8" pageNumber="377" type="materials_examined">
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<paragraph id="ECE8A937BA590347A13631D60BA6572B" pageId="8" pageNumber="377">Other material examined.</paragraph>
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<paragraph id="5ABFA0093FBEA298FF50D949C61F5A8C" pageId="8" pageNumber="377">
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PHILIPPINES: Luzon island: 1 male, Mt. Makiling, Baker leg. (USNM); 1 male, Los
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<normalizedToken id="E0D7C8CC4946E99CA645B53ADCB2FAC1" originalValue="Baños">Banos</normalizedToken>
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, 11.xii.1913, D. T. Fullaway leg. (BPBM); 1 male, Los
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<normalizedToken id="2A56968427BF7ADBA86548920D902592" originalValue="Baños">Banos</normalizedToken>
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, i.1913, P. Ledyard leg (LBOB); 2 males, Mt. Montalban, Rizal, Wa-wa Dam, 150-200 m, 6. and 17.iii.1965, H. M. Torrevillas leg. (BPBM); 1 male, 4 females, Manila, G. Boettcher leg. (MMBC); 1 male, 2 females, "B. M. 1925-491", E. M. Ledyard leg.; 1 female, "Acc. No. 6625, Lot, Bu. of Sci., P. I.,1908-228", C. S. Banks leg.; 1 male, "Acc. No. 5364, Lot, Govt. Lab. Coll., 1908-228", C. S. Banks leg. (all BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="BD9756D70F7E4984D44006577D6F89D3" pageId="8" pageNumber="377" type="redescription">
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<paragraph id="89DE559DB55118CDC92ADDEB402C2A2C" pageId="8" pageNumber="377">Redescription.</paragraph>
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<paragraph id="03519B4C335D1F66B065DCC278270AE4" pageId="8" pageNumber="377">Measurements (4 males, 9 females). Body length (from apex of head to tip of forewings): male 10.6-11.4 mm, female 12.9-13.9 mm; head length (from apex of cephalic process to base of eyes): male 1.25-1.33 mm, female 1.35-1.45 mm; head width (including eyes): male 1.30-1.43 mm, female 1.50-1.60 mm; forewing length: male 8.5-9.1 mm, female 10.3-11.4 mm.</paragraph>
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<paragraph id="041322762AA5F3543348EF1C72A49F74" pageId="8" pageNumber="377">
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Coloration. General coloration as in generic description (Figs 2
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<normalizedToken id="3E9BD72BFACEDC919910156E56EE0A6F" originalValue="I–J">I-J</normalizedToken>
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). Vertex predominantly dark brown, lateral margins with a pale ochraceous S-shaped streak in posterior three quarters (Fig. 9A). Frons light ochraceous with small dark brown spots along intermediate and lateral carinae and a large roundish dark brown spot at base (Fig. 9C). Forewing membrane pattern as in Fig. 3E. Hind wing membrane with a narrow dark brown infuscation along the apical portion of CuA1 vein, not extending along hind wing apical margin.
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</paragraph>
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<paragraph id="F5652A402B538F20C215112DA8F6EA87" pageId="8" pageNumber="377">
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Structure. Head with cephalic process moderately elongate (Figs 9
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<normalizedToken id="CA76DF58DFA54B74CF0D2AE9B92983C8" originalValue="A–B">A-B</normalizedToken>
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). Vertex (Fig. 9A) with ratio of length at midline to width between eyes 1.6-1.8. Transition of vertex to frons blunt, broadly rounded in lateral view, cephalic process thus appearing inflated and bulbous apically (Fig. 9B). Frons relatively narrow, with ratio of length at midline to maximum width 2.7-3.1.
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</paragraph>
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<paragraph id="AB06DC96EDE29CBB97043151AA83830F" pageId="8" pageNumber="377">
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Male genitalia. Pygofer, in lateral view, with dorso-posterior margin produced into a relatively large but narrow, tooth-like, apically obtuse process (Fig. 9E); in ventral view (Fig. 9F) much longer than in dorsal view (Fig. 9D) with ratio of ventral to dorsal width about 2.5. Gonostyles (Fig. 9G) large, broad medially, with dorsal margin weakly sinuate. Aedeagus (Fig. 9
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<normalizedToken id="DC7F9A7C64E5AA75E23155C624B38C07" originalValue="H–J">H-J</normalizedToken>
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) with endosomal processes covered with minute superficial spines, extended posteriad and strongly curved laterad and dorso-anteriad but not reaching the base of phallobase; phallobase with one pair of large, strongly inflated dorsolateral lobes, their apex convergent and tapering posteriad (Fig. 9I), and one pair of smaller, thumb-like ventral lobes, directed ventro-posteriad (Fig. 9
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<normalizedToken id="D5282ED2A91730CC87E8868279B43173" originalValue="I–J">I-J</normalizedToken>
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). Segment X, in lateral view, relatively narrow basally, widening to apex beyond middle, apex blunt (Fig. 9E); in dorsal view, relatively large and broad, widest in apical third, ratio of length to maximum width 1.0-1.1 (Fig. 9D).
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</paragraph>
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<paragraph id="12352AA4763696F7DD7F4D0EC9283C57" pageId="8" pageNumber="377">Female genitalia as in generic description.</paragraph>
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</subSubSection>
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<subSubSection id="E555F89061936A794E7201C9E9013A18" pageId="8" pageNumber="377" type="distribution">
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<paragraph id="D53BF17EE0757F33D2D00A5114977999" pageId="8" pageNumber="377">Distribution.</paragraph>
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<paragraph id="D9169CD21D515362FD296220B67018CA" pageId="8" pageNumber="377">So far only known from the Luzon island, Philippines.</paragraph>
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</subSubSection>
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<subSubSection id="96078DBD113711116B7607679904E46B" pageId="8" pageNumber="377" type="remarks">
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<paragraph id="0F56C601075DC71B61FE94DF0E1AF633" pageId="8" pageNumber="377">Remarks.</paragraph>
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<paragraph id="1048A50C50832EB630622C58DE806BF9" pageId="8" pageNumber="377">
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<taxonomicName id="00A0FC43E749268431CE6B12E32D80E6" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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can be easily recognized from other species of the genus by the shape of the head and the presence of a relatively large roundish dark spot at base of frons which is present in all specimens studied.
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</paragraph>
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<paragraph id="E363311872E6491644ECD34D91AFCF53" pageId="8" pageNumber="377">
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<bibRefCitation id="4DF0A11CCC4F1BE41368B6EBF5D8D138" author="Melichar, L" journalOrPublisher="Philippine Journal of Science" pageId="12" pageNumber="381" pagination="173 - 181" title="Neue Homopteren von den Philippinen." volume="9" year="1914">Melichar (1914)</bibRefCitation>
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indicated that he described this species based on five (male and female) specimens from "Luzon, Los
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<normalizedToken id="26F363120101A178B85D279EDB40AB68" originalValue="Baños">Banos</normalizedToken>
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, Mt. Maquiling (C. F. Baker)". Six specimens (1 male, 5 females) preserved in Melichar personal collection in MMBC and labelled as either from Los
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<normalizedToken id="36B68059A2367349AFB96B92F8E3DC89" originalValue="Baños">Banos</normalizedToken>
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or Mt. Makiling and collected by Baker are considered here as the original syntypes. The male specimen is designated here as the lectotype for
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<taxonomicName id="AE2C14331F8A2C215A4B30F93C75FE71" lsidName="O. philippinus" pageId="8" pageNumber="377" rank="species" species="philippinus">O. philippinus</taxonomicName>
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to stabilize the nomenclature according to Article 74 of
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<bibRefCitation id="6D454C102E03938A30ED02B745B26621" author="Germar, EF" journalOrPublisher="The International Trust for Zoological Nomenclature c / o the Natural History Museum, London" pageId="12" pageNumber="381" title="International Code of Zoological Nomenclature. Fourth Edition." url="https://doi.org/10.5962/bhl.title.50608" year="1999">ICZN (1999)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |