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<mods:title id="20C0858011E65D8996495ABFED38926F">Two new species of Brusqeulia Razowski &amp; Becker, 2000 from the Neotropics, with comments on the systematic position of the genus in relation to the Apolychrosis Amsel, 1962 group of genera (Lepidoptera, Tortricidae, Cochylini)</mods:title>
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<mods:namePart id="84BE2356F17C3D767549A79B55BE3E4C">Santa-Rita, Jose V. Perez</mods:namePart>
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<treatment id="C69BD6F99D5CAD12DE94A65BE3770A22" ID-GBIF-Taxon="156199977" LSID="urn:lsid:plazi:treatment:C69BD6F99D5CAD12DE94A65BE3770A22" httpUri="http://treatment.plazi.org/id/C69BD6F99D5CAD12DE94A65BE3770A22" lastPageId="2" lastPageNumber="194" pageId="1" pageNumber="194">
<subSubSection id="9411EA06B216DDE7D016D7F56CA07855" pageId="1" pageNumber="194" type="nomenclature">
<paragraph id="6DAE0698F03855D77866771E2C66328C" pageId="1" pageNumber="194">
<taxonomicName id="CAA27A9C727DE89955B46FE7C0719B4D" ID-CoL="B4B8D" authority="Razowski &amp; Becker, 2000" authorityName="Razowski &amp; Becker" authorityYear="2000" class="Insecta" family="Tortricidae" genus="Brusqeulia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brusqeulia" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="genus">Brusqeulia Razowski &amp; Becker, 2000</taxonomicName>
</paragraph>
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<subSubSection id="1F3F035883C07746CE8CA3358B81D60D" pageId="1" pageNumber="194" type="reference_group">
<paragraph id="61F69BB79F6D3D9566E473C7905EE804" pageId="1" pageNumber="194">
<taxonomicName id="4F4052C1314B3075429850D12040A940" class="Insecta" family="Tortricidae" genus="Brusqeulia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brusqeulia" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="genus">Brusqeulia</taxonomicName>
Razowski &amp; Becker, 2000, SHILAP Revista de
<normalizedToken id="F706754A5ABB572D524A346D90217685" originalValue="Lepidopterología">Lepidopterologia</normalizedToken>
28: 386; type-species:
<taxonomicName id="B4606248A6A500BB44F9A8E947B92748" class="Insecta" family="Tortricidae" genus="Brusqeulia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brusqeulia sebastiani" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="sebastiani">Brusqeulia sebastiani</taxonomicName>
Razowski &amp; Becker, 2000
</paragraph>
<paragraph id="3271DD3150D41613021BBA150EAA7A3B" pageId="1" pageNumber="194">
<taxonomicName id="E2F47AAD8E2CF785498D46EA16D0BFD7" class="Insecta" family="Tortricidae" genus="Pinhaisania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinhaisania" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="genus">Pinhaisania</taxonomicName>
Razowski &amp; Becker, 2000, SHILAP Revista de
<normalizedToken id="0CD8436368027B5DFA0E7E86FF3F7A72" originalValue="Lepidopterología">Lepidopterologia</normalizedToken>
28: 387; type-species:
<taxonomicName id="FE9A361C114206C0FA7D53D67CA17263" class="Insecta" family="Tortricidae" genus="Pinhaisania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinhaisania crispula" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="crispula">Pinhaisania crispula</taxonomicName>
Razowski &amp; Becker, 2000 - syn. n.
</paragraph>
</subSubSection>
<subSubSection id="B0C994F964904092711088657C3A2721" pageId="1" pageNumber="194" type="diagnosis">
<paragraph id="DE3417A0C3768701928397480D82C4D9" pageId="1" pageNumber="194">Diagnosis.</paragraph>
<paragraph id="4AE3E772248A542E81AB2A78F7DAB700" pageId="1" pageNumber="194">
Venation typically for
<taxonomicName id="B3318F209EF661A6172BD940632D85B6" lsidName="Cochylina" pageId="1" pageNumber="194" rank="subtribe" subtribe="Cochylina">Cochylina</taxonomicName>
(Fig. 1). Forewing (based on two slides) without costal fold; discal cell ca. 0.6 times length of wing, no
<normalizedToken id="4B23297CEC75AF8D1CB05F4ADD8AD74A" originalValue="Mstem">M-stem</normalizedToken>
, chorda obsolescent ca. 0.25 times length of wing, cross veins vestigial; all veins present except CuP; R4 to the costa near apex, R5 to the termen; distances between pairs of veins R5-M1, M1-M2 and M2-M3 on termen similar; distances between M3-CuA1, CuA1-CuA2 and CuA2-1A+2A similar; CuA2 opposite on discal cell ca. 0.3-0.5 the distance between R1 and R2 on the cell, approximately coincident with the point where the chorda meets Rs; anal loop ca. 0.3 times length of 1A+2A. Hindwing with Sc+R1 somewhat parallel to Rs basally, length ca. 0.8 times length of wing; M1 and Rs stalked in basal half; M2, M3 and CuA1 obsolescent basally; M3 and CuA1 connate; CuA2 well developed, CuP reduced, present only in distal portion; 1A+2A and 3A developed, anal loop ca. 0.4 times length of 1A+2A. Frenulum in males with one single bristle, three bristles in females. Male genitalia with transtilla broad and well developed; gnathos as two arms fused distally forming a short process, resulting in a plicate terminal plate; characteristic valva, elongate; cucullus with a more or less developed disc of hair like scales; sacculus with a free terminal process. Phallus with two distinctive sets of non-deciduous cornuti, one set as a ventral band of rather large aciculate cornuti basally attached; a second set in the inner part of vesica formed by microspinulate cornuti. Female genitalia with lobular lamella antevaginalis and postvaginalis; ventral spinous subpapillar sclerite on the 8-9 intersegmental membrane at the level of the ventral lobes of the anal papillae.
</paragraph>
<caption id="476BC67AECC5104B36368F613D44FB7B" pageId="1" pageNumber="194">
<paragraph id="70401FA208E2EC87F3C43C824F1A243E" pageId="1" pageNumber="194">
Figure 1. Venation in
<taxonomicName id="6397EDADA14334B3AA37AF7172406C5B" class="Insecta" family="Tortricidae" genus="Brusqeulia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brusqeulia yunkensis" order="Lepidoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="yunkensis">Brusqeulia yunkensis</taxonomicName>
. Based on slides JBA20815 and JBA20836. Dashed lines indicate obsolescent veins.
</paragraph>
</caption>
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<subSubSection id="724EC9D03720096E5766642B20BE830B" lastPageId="2" lastPageNumber="195" pageId="1" pageNumber="194" type="distribution">
<paragraph id="ED6BF130E4C8430051EF5820524A6B7B" pageId="1" pageNumber="194">Diversity and distribution.</paragraph>
<paragraph id="0811B6FE19E18BB698A5936B9A480449" lastPageId="2" lastPageNumber="195" pageId="1" pageNumber="194">
Fifteen species have been described from Brazil and one from Ecuador (
<bibRefCitation id="1A647A625C8BDBC2EE29A2FC02877F4C" author="Razowski, J" journalOrPublisher="SHILAP Revista de Lepidopterologia" pageId="11" pageNumber="204" pagination="385 - 393" title="Description of six Brazilian genera of Euliini and their species (Lepidoptera: Tortricidae)." volume="28" year="2000">Razowski and Becker 2000</bibRefCitation>
,
<bibRefCitation id="B28ED19E55A0E85DD16D4A0C36DC0597" author="Razowski, J" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="11" pageNumber="204" pagination="53 - 82" title="New species of Hynhamia Razowski and other genera close to Toreulia Razowski and Becker (Lepidoptera: Tortricidae)." url="https://doi.org/10.2478/v10200-011-0006-3" volume="80" year="2011">2011</bibRefCitation>
). To that we add one species
<pageBreakToken id="110818646CCA127441F559F9AED8BE51" pageId="2" pageNumber="195" start="start">from</pageBreakToken>
Bolivia and another from Venezuela. Given the broad geographical and elevational range (from near sea level to ca. 2000 m), we suspect that
<taxonomicName id="7A07787AD912A0F24BC47AFBE6677FA5" class="Insecta" family="Tortricidae" genus="Brusqeulia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brusqeulia" order="Lepidoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="genus">Brusqeulia</taxonomicName>
includes additional undiscovered species.
</paragraph>
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