treatments-xml/data/C6/7D/E7/C67DE7BC8EF7584384DB8FA4920F0285.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1156.98930" ID-Pensoft-Pub="1313-2970-1156-159" ID-Pensoft-UUID="A113628FABC65692A9B9D0C0FE204BA7" ID-ZooBank="2C1EAA9815154873872B426299C98FAD" ModsDocID="1313-2970-1156-159" checkinTime="1680231336932" checkinUser="pensoft" docAuthor="Murniati, Dewi Citra, Asakura, Akira &amp; Davie, Peter J. F." docDate="2023" docId="C67DE7BC8EF7584384DB8FA4920F0285" docLanguage="en" docName="ZooKeys 1156: 159-190" docOrigin="ZooKeys 1156" docPubDate="2023-03-30" docSource="http://dx.doi.org/10.3897/zookeys.1156.98930" docTitle="Tmethypocoelis celebensis Murniati &amp; Asakura &amp; Davie 2023, sp. nov." docType="treatment" docUuid="FA9B855D-5FE2-4007-BE15-EA27215DFD7E" docUuidSource="ZooBank" docVersion="2" id="A113628FABC65692A9B9D0C0FE204BA7" lastPageNumber="159" masterDocId="A113628FABC65692A9B9D0C0FE204BA7" masterDocTitle="Two new species of Tmethypocoelis Koelbel, 1897 (Decapoda, Brachyura, Dotillidae) from Sulawesi, Indonesia" masterLastPageNumber="190" masterPageNumber="159" pageNumber="159" updateTime="1680231638657" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of Tmethypocoelis Koelbel, 1897 (Decapoda, Brachyura, Dotillidae) from Sulawesi, Indonesia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Murniati, Dewi Citra</mods:namePart>
<mods:affiliation>Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta Bogor Km 46, Cibinong, Bogor, Indonesia &amp; Department of Zoology, Division of Biological Science, Graduate School of Science, Kyoto University, 606 - 8501, Yoshida-honmachi, Sakyo-ku, Kyoto-shi, Japan</mods:affiliation>
<mods:nameIdentifier type="email">dewicitramurniati@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asakura, Akira</mods:namePart>
<mods:affiliation>Seto Marine Biological Laboratory, Field Science Education and Research Center, Kyoto University, 459 Shirahama, Nishimuro, Wakayama 649 - 2211, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davie, Peter J. F.</mods:namePart>
<mods:affiliation>Queensland Museum, PO Box 3300, South Brisbane, Qld 4101, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1156</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>159</mods:start>
<mods:end>190</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1156.98930</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1156.98930</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1156-159</mods:identifier>
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<subSection pageId="0" pageNumber="159" type="taxonomy">
<treatment LSID="urn:lsid:zoobank.org:act:FA9B855D-5FE2-4007-BE15-EA27215DFD7E" httpUri="http://treatment.plazi.org/id/C67DE7BC8EF7584384DB8FA4920F0285" lastPageNumber="159" pageId="0" pageNumber="159">
<subSubSection pageId="0" pageNumber="159" type="nomenclature">
<paragraph pageId="0" pageNumber="159">
<taxonomicName LSID="https://zoobank.org/FA9B855D-5FE2-4007-BE15-EA27215DFD7E" authority="Murniati &amp; Asakura &amp; Davie, 2023" authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis" status="sp. nov.">Tmethypocoelis celebensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="159">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="description">
<paragraph pageId="0" pageNumber="159">
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus dorsal of Tmethypocoelis celebensis sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574) B paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180)." figureDoi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159">Figs 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A front area B left merus held against external orbital angle." figureDoi="10.3897/zookeys.1156.98930.figure11" httpUri="https://binary.pensoft.net/fig/831734" pageId="0" pageNumber="159">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Tmethypocoelis celebensis sp. nov. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang river (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left chela A inner surface B outer surface." figureDoi="10.3897/zookeys.1156.98930.figure14" httpUri="https://binary.pensoft.net/fig/831737" pageId="0" pageNumber="159">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">, 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">, 16</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Teeth of gastric mill (posterior portion on upper part). Tmethypocoelis simplex sp. nov., paratype, male (6.9 x 4.0 mm) (MZB. Cru. 5183) (A, C). Tmethypocoelis celebensis sp. nov., paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180) (B, D). A, B median tooth C, D lateral tooth." figureDoi="10.3897/zookeys.1156.98930.figure17" httpUri="https://binary.pensoft.net/fig/831740" pageId="0" pageNumber="159">, 17B, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="materials_examined">
<paragraph pageId="0" pageNumber="159">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="159">
<materialsCitation collectingDate="2020-09-12" collectorName="Murniati, D. Nurdiansyah, Cru." country="Indonesia" county="Moletang River" latitude="1.3665556" location="North Minahasa" longLatPrecision="1" longitude="125.07747" municipality="Kema Tiga" specimenCount="1" specimenCount-male="1" stateProvince="North Sulawesi" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="159">
<emphasis italics="true" pageId="0" pageNumber="159">
<typeStatus>Holotype</typeStatus>
</emphasis>
. 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
(7.4 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
); 
<collectingCounty>Moletang River</collectingCounty>
estuary, 
<collectingMunicipality>Kema Tiga</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:5BFAE0D1D48DAF175B3CCC2BAA2E8002" country="Indonesia" county="Moletang River" latitude="1.3665556" longLatPrecision="1" longitude="125.07747" municipality="Kema Tiga" name="North Minahasa" stateProvince="North Sulawesi">North Minahasa</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Utara">North Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="north" minutes="21" orientation="latitude" precision="1" seconds="59.6" value="1.3665556">1°21'59.6&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="125" direction="east" minutes="04" orientation="longitude" precision="1" seconds="38.9" value="125.07747">125°04'38.9&quot;E</geoCoordinate>
; 
<collectingDate value="2020-09-12">12 Sep. 2020</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
and 
<collectorName>D. Nurdiansyah</collectorName>
; MZB. 
<collectorName>Cru.</collectorName>
5574.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="159">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Indonesia" location="Indonesia" specimenCount="10" specimenCount-male="10" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="159">
<emphasis italics="true" pageId="0" pageNumber="159">
<typeStatus>Paratypes</typeStatus>
</emphasis>
. 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
</emphasis>
• 
<specimenCount type="male">10 ♂</specimenCount>
(2.8 
<normalizedToken originalValue="×">x</normalizedToken>
1.8 - 7.2 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation collectingDate="2020-09-12" collectorName="Murniati, D. Nurdiansyah, Cru." country="Indonesia" county="Moletang River" latitude="1.3665556" location="North Minahasa" longLatPrecision="1" longitude="125.07747" municipality="Kema Tiga" specimenCount="8" specimenCount-female="8" stateProvince="North Sulawesi" typeStatus="paratype">
<specimenCount type="female">8 ♀</specimenCount>
(4.8 
<normalizedToken originalValue="×">x</normalizedToken>
3.6 - 5.7 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.7" unit="mm" value="3.7">3.7 mm</quantity>
); 
<collectingCounty>Moletang River</collectingCounty>
estuary, 
<collectingMunicipality>Kema Tiga</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:40EB33D6F39FA55FC2FB08F2822AE077" country="Indonesia" county="Moletang River" latitude="1.3665556" longLatPrecision="1" longitude="125.07747" municipality="Kema Tiga" name="North Minahasa" stateProvince="North Sulawesi">North Minahasa</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Utara">North Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="north" minutes="21" orientation="latitude" precision="1" seconds="59.6" value="1.3665556">1°21'59.6&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="125" direction="east" minutes="04" orientation="longitude" precision="1" seconds="38.9" value="125.07747">125°04'38.9&quot;E</geoCoordinate>
; 
<collectingDate value="2020-09-12">12 Sep. 2020</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
and 
<collectorName>D. Nurdiansyah</collectorName>
; MZB. 
<collectorName>Cru.</collectorName>
5180
</materialsCitation>
• 
<materialsCitation collectingDate="2020-09-17" collectorName="D. Nurdiansyah, Cru." country="Indonesia" latitude="0.585" location="East Bolang Mongondow" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" specimenCount="10" specimenCount-male="10" stateProvince="North Sulawesi" typeStatus="paratype">
<specimenCount type="male">10 ♂</specimenCount>
(5.4 
<normalizedToken originalValue="×">x</normalizedToken>
3.2 - 6.5 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.7" unit="mm" value="3.7">3.7 mm</quantity>
); 
<collectingMunicipality>Iyok Beach</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:ABF2B43BE881D35DEAEDDE70AC05AF46" country="Indonesia" latitude="0.585" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" name="East Bolang Mongondow" stateProvince="North Sulawesi">East Bolang Mongondow</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Utara">North Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="0" direction="north" minutes="35" orientation="latitude" precision="1" seconds="06.0" value="0.585">0°35'06.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="124" direction="east" minutes="31" orientation="longitude" precision="1" seconds="58.6" value="124.53295">124°31'58.6&quot;E</geoCoordinate>
; 
<collectingDate value="2020-09-17">17 Sep. 2020</collectingDate>
; coll. 
<collectorName>D. Nurdiansyah</collectorName>
; MZB. 
<collectorName>Cru.</collectorName>
5181
</materialsCitation>
• 
<materialsCitation collectingDate="2021-06-10" collectorName="Murniati, Cru." country="Indonesia" latitude="0.4113889" location="Parigi Moutong" longLatPrecision="1" longitude="121.12886" municipality="Tuladenggi Sibatang" specimenCount="11" specimenCount-male="11" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">11 ♂</specimenCount>
(6.3 
<normalizedToken originalValue="×">x</normalizedToken>
3.8 - 7.9 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.8" unit="mm" value="4.8">4.8 mm</quantity>
); 
<collectingMunicipality>Tuladenggi Sibatang</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:0D5F04A96955BC346D46024B540D5929" country="Indonesia" latitude="0.4113889" longLatPrecision="1" longitude="121.12886" municipality="Tuladenggi Sibatang" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="0" direction="north" minutes="24" orientation="latitude" precision="1" seconds="41.0" value="0.4113889">0°24'41.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="121" direction="east" minutes="07" orientation="longitude" precision="1" seconds="43.9" value="121.12886">121°07'43.9&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-10">10 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
; MZB. 
<collectorName>Cru.</collectorName>
5575
</materialsCitation>
• 
<materialsCitation collectingDate="2021-06-25" collectorName="Murniati, Muslihun, M, Ikram, Cru." country="Indonesia" county="Maleyali" latitude="-1.0919445" location="Parigi Moutong" longLatPrecision="1" longitude="120.561005" municipality="Sausu" specimenCount="10" specimenCount-male="10" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">10 ♂</specimenCount>
(7.3 
<normalizedToken originalValue="×">x</normalizedToken>
3.8 - 7.4 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
); 
<collectingCounty>Maleyali</collectingCounty>
, 
<collectingMunicipality>Sausu</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:7E29F40484E862B230AEE0CE663FFEC5" country="Indonesia" county="Maleyali" latitude="-1.0919445" longLatPrecision="1" longitude="120.561005" municipality="Sausu" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="south" minutes="05" orientation="latitude" precision="1" seconds="31.0" value="-1.0919445">1°05'31.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="33" orientation="longitude" precision="1" seconds="39.6" value="120.561005">120°33'39.6&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-25">25 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
, 
<collectorName>Muslihun, M</collectorName>
. 
<collectorName>Ikram</collectorName>
; MZB. 
<collectorName>Cru.</collectorName>
5576
</materialsCitation>
• 
<materialsCitation collectingDate="2020-09-17" collectorName="D. Nurdiansyah" country="Indonesia" latitude="0.585" location="East Bolang Mongondow" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" specimenCount="5" specimenCount-male="5" stateProvince="North Sulawesi" typeStatus="paratype">
<specimenCount type="male">5 ♂</specimenCount>
(5.2 
<normalizedToken originalValue="×">x</normalizedToken>
3.0 - 6.0 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.4" unit="mm" value="3.4">3.4 mm</quantity>
); 
<collectingMunicipality>Iyok Beach</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:0DFB79913704AD6793D09B447916762D" country="Indonesia" latitude="0.585" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" name="East Bolang Mongondow" stateProvince="North Sulawesi">East Bolang Mongondow</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Utara">North Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="0" direction="north" minutes="35" orientation="latitude" precision="1" seconds="06.0" value="0.585">0°35'06.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="124" direction="east" minutes="31" orientation="longitude" precision="1" seconds="58.6" value="124.53295">124°31'58.6&quot;E</geoCoordinate>
; 
<collectingDate value="2020-09-17">17 Sep. 2020</collectingDate>
; coll. 
<collectorName>D. Nurdiansyah</collectorName>
; ZRC 2023.0056
</materialsCitation>
• 
<materialsCitation collectingDate="2021-06-25" collectorName="Murniati, Muslihun, M, Ikram" country="Indonesia" county="Maleyali" latitude="-1.0919445" location="Parigi Moutong" longLatPrecision="1" longitude="120.561005" municipality="Sausu" specimenCount="4" specimenCount-male="4" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">4 ♂</specimenCount>
(6.6 
<normalizedToken originalValue="×">x</normalizedToken>
3.8 - 7.2 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
); 
<collectingCounty>Maleyali</collectingCounty>
, 
<collectingMunicipality>Sausu</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:CF8E7CA231D34CAF1569951280BD6819" country="Indonesia" county="Maleyali" latitude="-1.0919445" longLatPrecision="1" longitude="120.561005" municipality="Sausu" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="south" minutes="05" orientation="latitude" precision="1" seconds="31.0" value="-1.0919445">1°05'31.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="33" orientation="longitude" precision="1" seconds="39.6" value="120.561005">120°33'39.6&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-25">25 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
, 
<collectorName>Muslihun, M</collectorName>
. 
<collectorName>Ikram</collectorName>
; ZRC. 2023.0057
</materialsCitation>
• 
<materialsCitation collectingDate="2020-09-17" collectorName="D. Nurdiansyah" country="Indonesia" latitude="0.585" location="East Bolang Mongondow" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" specimenCount="5" specimenCount-male="5" stateProvince="North Sulawesi" typeStatus="paratype">
<specimenCount type="male">5 ♂</specimenCount>
(4.7 
<normalizedToken originalValue="×">x</normalizedToken>
3.0 - 6.0 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.7" unit="mm" value="3.7">3.7 mm</quantity>
); 
<collectingMunicipality>Iyok Beach</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:7FF6DE7D072DBCD0B96F82CF13D1A64E" country="Indonesia" latitude="0.585" longLatPrecision="1" longitude="124.53295" municipality="Iyok Beach" name="East Bolang Mongondow" stateProvince="North Sulawesi">East Bolang Mongondow</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Utara">North Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="0" direction="north" minutes="35" orientation="latitude" precision="1" seconds="06.0" value="0.585">0°35'06.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="124" direction="east" minutes="31" orientation="longitude" precision="1" seconds="58.6" value="124.53295">124°31'58.6&quot;E</geoCoordinate>
; 
<collectingDate value="2020-09-17">17 Sep. 2020</collectingDate>
; coll. 
<collectorName>D. Nurdiansyah</collectorName>
; OMNH-Ar.12770-12774
</materialsCitation>
• 
<materialsCitation collectingDate="2021-06-25" collectorName="Murniati, Muslihun, M, Ikram" country="Indonesia" county="Maleyali" latitude="-1.0919445" location="Parigi Moutong" longLatPrecision="1" longitude="120.561005" municipality="Sausu" specimenCount="4" specimenCount-male="4" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">4 ♂</specimenCount>
(6.6 
<normalizedToken originalValue="×">x</normalizedToken>
3.8 - 7.8 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.4" unit="mm" value="4.4">4.4 mm</quantity>
); 
<collectingCounty>Maleyali</collectingCounty>
, 
<collectingMunicipality>Sausu</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:B66768E3E126EA7F33DC89775AA325D9" country="Indonesia" county="Maleyali" latitude="-1.0919445" longLatPrecision="1" longitude="120.561005" municipality="Sausu" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="south" minutes="05" orientation="latitude" precision="1" seconds="31.0" value="-1.0919445">1°05'31.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="33" orientation="longitude" precision="1" seconds="39.6" value="120.561005">120°33'39.6&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-25">25 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
, 
<collectorName>Muslihun, M</collectorName>
. 
<collectorName>Ikram</collectorName>
; OMNH-Ar. 12766-12769
</materialsCitation>
• 
<materialsCitation collectingDate="2021-06-25" collectorName="Murniati, Muslihun, M, Ikram" country="Indonesia" county="Maleyali" latitude="-1.0919445" location="Parigi Moutong" longLatPrecision="1" longitude="120.561005" municipality="Sausu" specimenCount="4" specimenCount-male="4" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">4 ♂</specimenCount>
(6.2 
<normalizedToken originalValue="×">x</normalizedToken>
3.7 - 7.4 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
); 
<collectingCounty>Maleyali</collectingCounty>
, 
<collectingMunicipality>Sausu</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:B77E2A7839B0C30F1CB26BDE59D830A3" country="Indonesia" county="Maleyali" latitude="-1.0919445" longLatPrecision="1" longitude="120.561005" municipality="Sausu" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="1" direction="south" minutes="05" orientation="latitude" precision="1" seconds="31.0" value="-1.0919445">1°05'31.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="33" orientation="longitude" precision="1" seconds="39.6" value="120.561005">120°33'39.6&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-25">25 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
, 
<collectorName>Muslihun, M</collectorName>
. 
<collectorName>Ikram</collectorName>
; RMNH.CRUS.D.58047
</materialsCitation>
• 
<materialsCitation accessionNumber="W29643" collectingDate="2021-06-10" collectorName="Murniati" country="Indonesia" latitude="0.4113889" location="Parigi Moutong" longLatPrecision="1" longitude="121.12886" municipality="Tuladenggi Sibatang" specimenCount="3" specimenCount-male="3" stateProvince="Central Sulawesi" typeStatus="paratype">
<specimenCount type="male">3 ♂</specimenCount>
(4.3 
<normalizedToken originalValue="×">x</normalizedToken>
3.8 - 4.9 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
); 
<collectingMunicipality>Tuladenggi Sibatang</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:267217DF67BA3299A89ECCD9E53728DE" country="Indonesia" latitude="0.4113889" longLatPrecision="1" longitude="121.12886" municipality="Tuladenggi Sibatang" name="Parigi Moutong" stateProvince="Central Sulawesi">Parigi Moutong</location>
, 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
; 
<geoCoordinate degrees="0" direction="north" minutes="24" orientation="latitude" precision="1" seconds="41.0" value="0.4113889">0°24'41.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="121" direction="east" minutes="07" orientation="longitude" precision="1" seconds="43.9" value="121.12886">121°07'43.9&quot;E</geoCoordinate>
; 
<collectingDate value="2021-06-10">10 Jun. 2021</collectingDate>
; coll. DC. 
<collectorName>Murniati</collectorName>
; QM 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/W29643">W29643</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="materials_examined">
<paragraph pageId="0" pageNumber="159">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="159">
<taxonomicName authorityName="Murniati, Asakura, Nugroho, Hernawan &amp; Dharmawan" authorityYear="2022" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis liki" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis liki</emphasis>
</taxonomicName>
<materialsCitation collectingDate="2018-11-21" collectorName="Murniati, Asakura, Nugroho, Hernawan, Dharmawan" country="Indonesia" county="Liki Village" latitude="-1.6236917" location="Sarmi Municipality" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" specimenCount="5" specimenCount-male="5" stateProvince="Papua Province" typeStatus="paratypes">
<collectorName>Murniati</collectorName>
, 
<collectorName>Asakura</collectorName>
, 
<collectorName>Nugroho</collectorName>
, 
<collectorName>Hernawan</collectorName>
&amp; 
<collectorName>Dharmawan</collectorName>
, 2022: 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
• 
<typeStatus>paratypes</typeStatus>
<specimenCount type="male">5 ♂</specimenCount>
(5.3 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.1" unit="mm" value="3.1">3.1 mm</quantity>
- 5.5 
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
); 
<collectingCounty>Liki Village</collectingCounty>
, 
<collectingMunicipality>Sarmi District</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:37420DC3C26E414F949836A16B63473A" country="Indonesia" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Sarmi Municipality" stateProvince="Papua Province">Sarmi Municipality</location>
, 
<location LSID="urn:lsid:plazi:treatment:C67DE7BC8EF7584384DB8FA4920F0285:4B89EE5BDA7F39385DBD33476D2040EF" country="Indonesia" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Liki Island" stateProvince="Papua Province">Liki Island</location>
, 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
; 
<geoCoordinate degrees="01" direction="south" minutes="37" orientation="latitude" precision="1" seconds="25.29" value="-1.6236917">01°37'25.29&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="138" direction="east" minutes="44" orientation="longitude" precision="1" seconds="26.54" value="138.7407">138°44'26.54&quot;E</geoCoordinate>
; 
<collectingDate value="2018-11-21">21 Nov. 2018</collectingDate>
; coll. DC. Murniati; MZB.Cru.5012
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="diagnosis">
<paragraph pageId="0" pageNumber="159">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="159">
Carapace pentagonal, ca. 1.6-1.7 
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus dorsal of Tmethypocoelis celebensis sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574) B paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180)." figureDoi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159">10A</figureCitation>
). Branchial region sloping, protobranchial, mesobranchial and metabranchial regions well-defined. Sub-branchial region bulging, bearing regular setae and tubercles. Posterior margin slightly concave, ca. 0.64 distance between exorbital angles. Exorbital angle triangular, acute, directed forward (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12A</figureCitation>
). Second anterolateral tooth less acute, slightly shorter. Male pleon ca. 2.0 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12E</figureCitation>
). Male chelipeds long. Palm bulky, ca. 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M, N</figureCitation>
). Fingers shorter than palm. Pollex short, triangular, cutting margin slightly oblique, without large differentiated tooth or lobe (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M</figureCitation>
); cutting margin of dactylus with large teeth over proximal half, small teeth on distal half, without median lobe, upper margin with row of fine tubercles; one triangular, upturned tooth subdistally (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M-O</figureCitation>
). G1 long, recurved, very slender; sub-proximal portion bulging (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15A, B</figureCitation>
); apical portion forming two lobes, with three short setae on outer margin, two or three long setae apically, four or five short setae on inner margin (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15C, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 10.</emphasis>
Habitus dorsal of 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
holotype, male (7.2 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5574) 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
paratype, female (5.5 
<normalizedToken originalValue="×">x</normalizedToken>
3.5 mm) (MZB.Cru.5180).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="description">
<paragraph pageId="0" pageNumber="159">Description.</paragraph>
<paragraph pageId="0" pageNumber="159">
Carapace (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus dorsal of Tmethypocoelis celebensis sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574) B paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180)." figureDoi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159">10A</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12A</figureCitation>
) pentagonal; weakly convex laterally and longitudinally; ca. 1.6-1.7 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Dorsal surface smooth, regions semi-defined; epigastric lobe poorly defined. Cervical grooves, well-marked; cardiac region slightly depressed. Branchial regions sloping, protobranchial, mesobranchial and metabranchial regions well-defined. Carapace widest between exorbital angles. Sub-branchial region bulging, bearing regular setae and tubercles. Intestinal and branchial regions well-defined. Posterior margin weakly concave, ca. 0.6 
<normalizedToken originalValue="×">x</normalizedToken>
distance between exorbital angles; fine ridge parallel with posterior margin forming broad rim. Lateral margin recurved with row of tubercles and short stout setae. Frontal margin rounded, moderately convergent, basal width ca. 0.19 
<normalizedToken originalValue="×">x</normalizedToken>
distance between exorbital angles, ca. 0.13 
<normalizedToken originalValue="×">x</normalizedToken>
at anterior margin; anterior margin with small central blunt prominence (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A front area B left merus held against external orbital angle." figureDoi="10.3897/zookeys.1156.98930.figure11" httpUri="https://binary.pensoft.net/fig/831734" pageId="0" pageNumber="159">11A</figureCitation>
). Exorbital angle triangular, acute, directed forwardly; anterior margin with microscopic tubercles, lateral margin slightly tubercular; posteriorly followed by broad U-shaped sinus. Epibranchial tooth less acute, slightly shorter. Posterolateral facet well-defined by a crest originating anteriorly from base of exorbital angle (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12A</figureCitation>
). Supra-orbital borders sinuous, sloping backward, microscopically tubercular. Infra-orbital border with medial notch; pterygostome with inwardly directed oblique channel. Inner part of infra-orbital border with two granular ridges separated by shallow channel; upper ridge with row of setae; granules on lower ridge larger than that of upper ridge, without setation. Outer part of infra-orbital border granular, concave, ending in broad notch below exorbital angle (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure11" httpUri="https://binary.pensoft.net/fig/831734" pageId="0" pageNumber="159" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 11.</emphasis>
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Holotype, male (7.2 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
front area 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
left merus held against external orbital angle.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 12.</emphasis>
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Holotype, male (7.2 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
exorbital angle 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
orbit area 
<emphasis bold="true" pageId="0" pageNumber="159">C</emphasis>
eyestalk 
<emphasis bold="true" pageId="0" pageNumber="159">D</emphasis>
third maxilliped 
<emphasis bold="true" pageId="0" pageNumber="159">E</emphasis>
pleon. Paratype, female (5.5 
<normalizedToken originalValue="×">x</normalizedToken>
3.5 mm) (MZB.Cru.5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">F</emphasis>
pleon.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
Eyestalks (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus dorsal of Tmethypocoelis celebensis sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574) B paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180)." figureDoi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159">10</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A front area B left merus held against external orbital angle." figureDoi="10.3897/zookeys.1156.98930.figure11" httpUri="https://binary.pensoft.net/fig/831734" pageId="0" pageNumber="159">11A</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12C</figureCitation>
) reaching exorbital angle, medial and distal diameters similar size; ocular style as long as cornea, tipped with setae; medial slope gives twisted appearance; cornea bulging.
</paragraph>
<paragraph pageId="0" pageNumber="159">
Third maxillipeds (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
) slightly vaulted, not completely covering buccal cavern. Ischium subquadrate, outer surface covered with spaced long setae, with one oblique row of long setae near upper margin, upper margin concave, upper-mesial angle with narrow, rounded lobe; lower-mesial angle curved; inner and lower margins with dense setae; lateral margin without setation (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
). Merus slightly larger than ischium, ca. 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
longer; outer surface with regularly scattered short setae; lateral margin convex, narrower distally, covered with short setae; mesial margin straight with long setae (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
). Carpus trihedral, subequal in length to propodus and dactylus together; mesial margin and distal portion with dense long setae (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
). Propodus shorter than dactylus; margins entire, with long dense setae (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
). Dactylus slender, with long dense setae laterally (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="159">
Male pleon (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12E</figureCitation>
) ca. 2.0 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Pl1 trapezoidal, ca. 8.0 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; ca. 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
wider than pl2. Pl2 very narrow, ca. 10 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Pl3 ca. 3 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Pl4 ca. 3.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, lateral margins convergent distally, distolateral angle pointed. Pl5 ca. 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long (at widest point), markedly constricted at base. Pl6 ca. 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; widest sub-distally; subequal in length to pl5. Male telson rounded, ca. 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="159">
Female pleon (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12F</figureCitation>
) conspicuously broad. Pl1 shortest; pl2 distinctly longer, as wide as pl1; pl3 trapezoidal, longer than pl2; pl4 rectangular, slightly longer than pl3, lateral margins convex; pl5 longer than pl4; pl6 distinctly longest. Female telson (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A exorbital angle B orbit area C eyestalk D third maxilliped E pleon. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi F pleon." figureDoi="10.3897/zookeys.1156.98930.figure12" httpUri="https://binary.pensoft.net/fig/831735" pageId="0" pageNumber="159">12F</figureCitation>
) triangular.
</paragraph>
<paragraph pageId="0" pageNumber="159">
Male chelipeds stout, long, equal. Merus triangular in cross-section; standing higher than exorbital angle (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A front area B left merus held against external orbital angle." figureDoi="10.3897/zookeys.1156.98930.figure11" httpUri="https://binary.pensoft.net/fig/831734" pageId="0" pageNumber="159">11B</figureCitation>
); lower margin with two rows of granules extending whole length of margin (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13A</figureCitation>
); upper margin narrowing proximally, with irregular rows of granules on distal half (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13K</figureCitation>
); outer margin with one row of granules extending whole length (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13L</figureCitation>
); upper surface flattened, ovate smooth tympanum, scattered long setae around tympanum, more setation distally, microscopically tuberculate (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13I</figureCitation>
); lower surface flattened, nearly smooth, with scattered granules, lacking tympanum (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13J</figureCitation>
); outer surface convex, tympanum smaller than that of upper surface, evenly distributed granules and setae (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13B</figureCitation>
). Carpus shorter than merus, elongate, ca. 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; upper and lower margins tubercular (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13C, D</figureCitation>
); outer surface rectangular, scattered microscopic granules near upper and lower margins, median portion without microscopic tubercles (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13E</figureCitation>
); lower surface smooth, with one longitudinal row of granules, one patch of tubercles on proximal part (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13F</figureCitation>
). Palm bulky, ca. 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; inner surface granular over upper half, with granules extending over upper margin and curved to sharply cut upper margin of outer surface, distally with one row of regular granules reaching pollex, smaller granules near lower margin, median portion smooth (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M</figureCitation>
); outer surface distinctly granular over upper half reaching to base of pollex, lower half smooth (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13N</figureCitation>
); upper margin with one row of granules, distinct groove extending below granular rows forming clear granular string (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13G</figureCitation>
); lower margin with granulation extending to lower part of inner surface (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13H</figureCitation>
). Fingers shorter than palm, lacking obvious basal gape, curved inwards, spooned-tip; cutting margins evenly serrated; inner margin at tip of both fingers with short row of stout setae. Pollex short, triangular, cutting margin slightly oblique, without large differentiated tooth or lobe, ca. 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as palm; inner surface nearly smooth, one row of granules over proximal half (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M</figureCitation>
); outer surface granular parallel to cutting margin, granules with similar size (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13N</figureCitation>
); lower margin granulated only along proximo-medially (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13H</figureCitation>
). Cutting margin of dactylus with teeth, larger teeth over proximal half, smaller teeth over distal half, without median lobe; inner surface with one row of granules parallel to upper margin, granulation extending from proximal to distal, one patch of granules proximally (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M</figureCitation>
); band of granules on outer surface near cutting margin and junction to palm, one tubercular ridge extending medially parallel to upper margin (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13N</figureCitation>
); upper margin with row of tubercles terminating with triangular upturned tooth subdistally (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13O</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 13.</emphasis>
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Holotype, male (7.2 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
lower margin 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
outer surface. Carpus 
<emphasis bold="true" pageId="0" pageNumber="159">C</emphasis>
upper margin 
<emphasis bold="true" pageId="0" pageNumber="159">D</emphasis>
lower margin 
<emphasis bold="true" pageId="0" pageNumber="159">E</emphasis>
outer surface 
<emphasis bold="true" pageId="0" pageNumber="159">F</emphasis>
inner surface. Chela 
<emphasis bold="true" pageId="0" pageNumber="159">G</emphasis>
upper margin 
<emphasis bold="true" pageId="0" pageNumber="159">H</emphasis>
lower margin. Merus 
<emphasis bold="true" pageId="0" pageNumber="159">I</emphasis>
upper surface 
<emphasis bold="true" pageId="0" pageNumber="159">J</emphasis>
lower surface 
<emphasis bold="true" pageId="0" pageNumber="159">K</emphasis>
upper margin 
<emphasis bold="true" pageId="0" pageNumber="159">L</emphasis>
outer margin. Chela 
<emphasis bold="true" pageId="0" pageNumber="159">M</emphasis>
inner surface 
<emphasis bold="true" pageId="0" pageNumber="159">N</emphasis>
outer surface 
<emphasis bold="true" pageId="0" pageNumber="159">O</emphasis>
dactylus upper margin.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
Female chelipeds small dotillid type (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus dorsal of Tmethypocoelis celebensis sp. nov. Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574) B paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180)." figureDoi="10.3897/zookeys.1156.98930.figure10" httpUri="https://binary.pensoft.net/fig/831733" pageId="0" pageNumber="159">10B</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Tmethypocoelis celebensis sp. nov. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang river (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left chela A inner surface B outer surface." figureDoi="10.3897/zookeys.1156.98930.figure14" httpUri="https://binary.pensoft.net/fig/831737" pageId="0" pageNumber="159">14</figureCitation>
). Not conspicuously different from 
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov. (see description for 
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure14" httpUri="https://binary.pensoft.net/fig/831737" pageId="0" pageNumber="159" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 14.</emphasis>
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Paratype, female (5.5 
<normalizedToken originalValue="×">x</normalizedToken>
3.5 mm) (MZB.Cru.5180), Moletang river (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left chela 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
inner surface 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
outer surface.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
Pereiopods (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15</figureCitation>
) slender, elongate, P2-P5 similar; smooth ovate tympanum on anterior and posterior surfaces of meri. Tympani on posterior surfaces becoming progressively smaller from P2-P5. Dactyli nearly straight, pointed, shorter than propodi.
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 15.</emphasis>
Left pereiopods of 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. Holotype, male (7.2 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">A, B</emphasis>
P2 
<emphasis bold="true" pageId="0" pageNumber="159">C, D</emphasis>
P3 
<emphasis bold="true" pageId="0" pageNumber="159">E, F</emphasis>
P4 
<emphasis bold="true" pageId="0" pageNumber="159">G, H</emphasis>
P5. Right side, anterior surfaces; left side, posterior surfaces.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
P2 (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15A, B</figureCitation>
) shorter than P3; merus ca. 2.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior surface bearing scattered granules outside tympanum, granules denser near lower margin, sparse setae near lower margin; posterior surface sparsely granulate, denser distal to tympanum; upper margin convex, sparse long setae, distally tubercular; lower margin smooth, sparse setae. Carpus shorter than propodus, surfaces smooth; margins without granules, sparse setae. Propodus with anterior and posterior surfaces with few small granules only; margins with sparse long setae.
</paragraph>
<paragraph pageId="0" pageNumber="159">
P3 (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15C, D</figureCitation>
) longest; merus ca. 2.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior surface scarcely granular; posterior surface sparsely granulate, denser near upper margin; upper and lower margins convex; upper margin tubercular distally, sparse long setae; lower margin smooth, sparse setae. Carpus shorter than propodus, surfaces nearly smooth, sparse setae distally; margins without tubercles, sparse setae. Propodus with anterior and posterior surfaces smooth; margins with sparse long setae.
</paragraph>
<paragraph pageId="0" pageNumber="159">
P4 (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15E, F</figureCitation>
) nearly as long as P2; merus ca. 2.6 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior surface scarcely granular; posterior surface with evenly distributed granules; upper and lower margins convex; upper margin tubercular distally, spaced long setae extending whole length; lower margin smooth, sparse setae. Carpus shorter than propodus, surfaces smooth; margins smooth, sparse setae. Propodus with anterior and posterior surfaces smooth; margins with sparse long setae.
</paragraph>
<paragraph pageId="0" pageNumber="159">
P5 (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left pereiopods of Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi A, B P 2 C, D P 3 E, F P 4 G, H P 5. Right side, anterior surfaces; left side, posterior surfaces." figureDoi="10.3897/zookeys.1156.98930.figure15" httpUri="https://binary.pensoft.net/fig/831738" pageId="0" pageNumber="159">15G, H</figureCitation>
) shortest; merus ca. 2.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior surface without granules; posterior surface granulate, granules denser near upper margin; upper and lower margins convex; upper margin sparsely tubercular, with sparse long setae, short setae proximally; lower margin smooth, with sparse setae. Carpus shorter than propodus, surfaces smooth; margins smooth, with sparse setae. Propodus with anterior and posterior surfaces smooth; margins with sparse long setae.
</paragraph>
<paragraph pageId="0" pageNumber="159">
Reproductive organs. G1 long, recurved, very slender; sub-proximal portion bulging (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">16A, B</figureCitation>
); apical portion forming two lobes, with three short setae on outer margin, two or three long setae apically, four or five short setae on mesial margin (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">16C, D</figureCitation>
). Vulva (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">16E</figureCitation>
) rounded, projecting.
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 16.</emphasis>
Reproductive organs of 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="159">b</emphasis>
paratype, male (7.3 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G1 
<emphasis bold="true" pageId="0" pageNumber="159">A</emphasis>
mesial view 
<emphasis bold="true" pageId="0" pageNumber="159">B</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="159">C, D</emphasis>
apical portion 
<emphasis bold="true" pageId="0" pageNumber="159">C</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="159">D</emphasis>
ventral view. Paratype, female (5.5 
<normalizedToken originalValue="×">x</normalizedToken>
3.5 mm) (MZB.Cru.5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi 
<emphasis bold="true" pageId="0" pageNumber="159">E</emphasis>
vulva.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
Gastric mill. Median tooth plate simple, without defined ridges. Urocardiac ossicle relatively narrower throughout length. Propyloric ossicle semi-circular, relatively narrow and protruding; posterior margin curved; anterior margin with one pointed lobe medially; lateral margins quadrate with anterior lobes discrete, prominent, and rounded (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Teeth of gastric mill (posterior portion on upper part). Tmethypocoelis simplex sp. nov., paratype, male (6.9 x 4.0 mm) (MZB. Cru. 5183) (A, C). Tmethypocoelis celebensis sp. nov., paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180) (B, D). A, B median tooth C, D lateral tooth." figureDoi="10.3897/zookeys.1156.98930.figure17" httpUri="https://binary.pensoft.net/fig/831740" pageId="0" pageNumber="159">17B</figureCitation>
). Lateral zygocardiac tooth plate with eight slender teeth, three anterior teeth large (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Teeth of gastric mill (posterior portion on upper part). Tmethypocoelis simplex sp. nov., paratype, male (6.9 x 4.0 mm) (MZB. Cru. 5183) (A, C). Tmethypocoelis celebensis sp. nov., paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180) (B, D). A, B median tooth C, D lateral tooth." figureDoi="10.3897/zookeys.1156.98930.figure17" httpUri="https://binary.pensoft.net/fig/831740" pageId="0" pageNumber="159">17D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure17" httpUri="https://binary.pensoft.net/fig/831740" pageId="0" pageNumber="159" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 17.</emphasis>
Teeth of gastric mill (posterior portion on upper part). 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis simplex" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis simplex</emphasis>
</taxonomicName>
sp. nov., paratype, male (6.9 
<normalizedToken originalValue="×">x</normalizedToken>
4.0 mm) (MZB.Cru. 5183) (
<emphasis bold="true" pageId="0" pageNumber="159">A, C</emphasis>
). 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov., paratype, male (7.3 
<normalizedToken originalValue="×">x</normalizedToken>
4.4 mm) (MZB.Cru.5180) (
<emphasis bold="true" pageId="0" pageNumber="159">B, D</emphasis>
). 
<emphasis bold="true" pageId="0" pageNumber="159">A, B</emphasis>
median tooth 
<emphasis bold="true" pageId="0" pageNumber="159">C, D</emphasis>
lateral tooth.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="habitat">
<paragraph pageId="0" pageNumber="159">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="159">
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. inhabits sandy substrata in estuarine areas (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitat of Tmethypocoelis celebensis sp. nov. at Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi." figureDoi="10.3897/zookeys.1156.98930.figure18" httpUri="https://binary.pensoft.net/fig/831741" pageId="0" pageNumber="159">18</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1156.98930.figure18" httpUri="https://binary.pensoft.net/fig/831741" pageId="0" pageNumber="159" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Figure 18.</emphasis>
Habitat of 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis celebensis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis celebensis</emphasis>
</taxonomicName>
sp. nov. at Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="etymology">
<paragraph pageId="0" pageNumber="159">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="159">The species name is derived from the type locality. Celebes is the former name of Sulawesi Island, one of the great islands in Indonesia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="159" type="remarks">
<paragraph pageId="0" pageNumber="159">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="159">
The two new species described here differ from each other and from the other known species by numerous characters compared below. In general, the species of 
<taxonomicName authorityName="Koelbel" authorityYear="1897" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis</emphasis>
</taxonomicName>
are all extremely similar in general morphology with only small differences in male cheliped shape and dentition (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of male chelipeds in the species of Tmethypocoelis." httpUri="http://table.plazi.org/id/73C8A668D175C5A2AA05A1FC2FC532D0" pageId="0" pageNumber="159" tableUuid="73C8A668D175C5A2AA05A1FC2FC532D0">1</tableCitation>
), differences in the apical setal ornamentation of the male first gonopod, and sometimes differences in the proportions of the somites of the male pleon.
</paragraph>
<caption ID-Table-UUID="73C8A668D175C5A2AA05A1FC2FC532D0" httpUri="http://table.plazi.org/id/73C8A668D175C5A2AA05A1FC2FC532D0" pageId="0" pageNumber="159" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Table 1.</emphasis>
Comparison of male chelipeds in the species of 
<taxonomicName authorityName="Koelbel" authorityYear="1897" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
<table pageId="0" pageNumber="159">
<tr pageId="0" pageNumber="159">
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. liki" pageId="0" pageNumber="159" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">T. liki</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. ceratophora" pageId="0" pageNumber="159" rank="species" species="ceratophora">
<emphasis italics="true" pageId="0" pageNumber="159">T. ceratophora</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. choreutes" pageId="0" pageNumber="159" rank="species" species="choreutes">
<emphasis italics="true" pageId="0" pageNumber="159">T. choreutes</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. odontodactylus" pageId="0" pageNumber="159" rank="species" species="odontodactylus">
<emphasis italics="true" pageId="0" pageNumber="159">T. odontodactylus</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Dactylus dorsal and outer armature</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Upper margin finely tuberculate; lacking differentiated subdistal tooth; outer surface with semi-defined granular row medially; irregular granulation near cutting margin (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tmethypocoelis simplex sp. nov. Holotype, male (7.7 x 4.4 mm) (MZB. Cru. 5573), Towale River, Central Banawa District, Donggala, Central Sulawesi. Left cheliped A merus lower margin B merus outer surface. Carpus C outer surface D inner surface E upper surface F lower surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure5" httpUri="https://binary.pensoft.net/fig/831728" pageId="0" pageNumber="159">5M-O</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Upper margin finely tuberculate; culminating in subdistal upward, upwardly projecting triangular tooth of variable size from high and distinct to low; size not correlated with crab size; outer surface with medial granular row over entire length (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tmethypocoelis celebensis sp. nov. Holotype, male (7.2 x 4.4 mm) (MZB. Cru. 5574), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi. Left cheliped. Merus A lower margin B outer surface. Carpus C upper margin D lower margin E outer surface F inner surface. Chela G upper margin H lower margin. Merus I upper surface J lower surface K upper margin L outer margin. Chela M inner surface N outer surface O dactylus upper margin." figureDoi="10.3897/zookeys.1156.98930.figure13" httpUri="https://binary.pensoft.net/fig/831736" pageId="0" pageNumber="159">13M-O</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Dorsal band of fine granules; superior border straight, terminating in overhanging triangular tooth at about 
<normalizedToken originalValue="¾">3/4</normalizedToken>
length; outer surface with 2 subregular lines of granules, superior one may extend 
<normalizedToken originalValue="¾">3/4</normalizedToken>
length to tip, lower one less than that of superior
</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Upper margin with median row of granules, culminating into 1 prominent tooth</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Finely tuberculate ridge on upper margin of dactylus terminates in outwardly directed flat triangular tooth (see 
<bibRefCitation author="Davie, PJF" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="159" pagination="207 - 215" refId="B7" refString="Davie, PJF, Kosuge, T, 1995. A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. The Raffles Bulletin of Zoology 43 (1): 207 - 215" title="A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan." volume="43" year="1995">Davie and Kosuge 1995</bibRefCitation>
: fig. 1 A, C)
</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Finely tubercular ridge on the upper margin continues evenly distally; outer surface has separate ridge ending in prominent triangular subdistal protrusion (see 
<bibRefCitation author="Davie, PJF" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="159" pagination="207 - 215" refId="B7" refString="Davie, PJF, Kosuge, T, 1995. A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. The Raffles Bulletin of Zoology 43 (1): 207 - 215" title="A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan." volume="43" year="1995">Davie and Kosuge 1995</bibRefCitation>
: fig. 1 B, D)
</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">
Full length medial granulate ridge; superior granulate crest terminating subdistally in strong upturned tooth (
<bibRefCitation author="Davie, PJF" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="159" pagination="463 - 473" refId="B5" refString="Davie, PJF, 1990. New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum 28: 463 - 473" title="New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea." volume="28" year="1990">Davie 1990</bibRefCitation>
: fig. 3C)
</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Dactylus cutting margin</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate; one wide enlarged convex tooth over proximal half</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate, larger over proximal half, then finger narrower over distal half</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate; smaller males with raised platform of teeth differentiated in proximal half, but less evident in mature chela</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Wide and blunt irregular serrations</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">One small low tooth proximally</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Low broad triangular convexity but lacking clearly differentiated tooth</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate; slightly deeper medially, but without obvious differentiated tooth</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Pollex cutting margin</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate; long flat enlarged dentate tooth over most of length</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate; straight, without differentiated tooth or lobe</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Evenly dentate</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Irregularly dentate</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Prominent enlarged convex tooth medially</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Lacking a defined tooth; slightly convex</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Weakly convex; evenly dentate</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Gape at base of fingers</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Large</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Poorly developed</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Moderate</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Poorly developed</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Not strongly developed</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Not strongly developed</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="159">
A comparison of male first gonopod setation patterns of described species suggests that the possession of two or three of markedly elongated apical setae (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Reproductive organs of Tmethypocoelis simplex sp. nov. Paratype, males (A-D 4.1 x 2.6 mm E 6.9 x 4.0 mm) (MZB. Cru. 5183), Tosale, Banawa District, Donggala, Central Sulawesi, left G 1 A dorsal view B apical (dorsal view) C, D apical (ventral view) E mesial view. Paratype female (5.8 x 3.6 mm) (MZB. Cru. 5182), Towale river, Central Banawa District, Donggala, Central Sulawesi F vulva." figureDoi="10.3897/zookeys.1156.98930.figure8" httpUri="https://binary.pensoft.net/fig/831731" pageId="0" pageNumber="159">8C-E</figureCitation>
, 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">16C, D</figureCitation>
; 
<bibRefCitation author="Davie, PJF" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="159" pagination="463 - 473" refId="B5" refString="Davie, PJF, 1990. New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum 28: 463 - 473" title="New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea." volume="28" year="1990">Davie 1990</bibRefCitation>
: fig. 2), common to both 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis simplex" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis simplex</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov., is so far shared with 
<taxonomicName lsidName="T. liki" pageId="0" pageNumber="159" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">T. liki</emphasis>
</taxonomicName>
from Papua and 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
from the Northern Territory, NW Australia. Therefore, these four species may be more closely related to each other than they are to 
<taxonomicName lsidName="T. ceratophora" pageId="0" pageNumber="159" rank="species" species="ceratophora">
<emphasis italics="true" pageId="0" pageNumber="159">T. ceratophora</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="T. choreutes" pageId="0" pageNumber="159" rank="species" species="choreutes">
<emphasis italics="true" pageId="0" pageNumber="159">T. choreutes</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="T. odontodactylus" pageId="0" pageNumber="159" rank="species" species="odontodactylus">
<emphasis italics="true" pageId="0" pageNumber="159">T. odontodactylus</emphasis>
</taxonomicName>
, which all share a coronet of shorter more evenly sized stout setae on the tip of the G1. A more thorough analysis of relationships within the genus will be undertaken as part of a larger revision of the genus, and with the help of DNA sequencing data.
</paragraph>
<paragraph pageId="0" pageNumber="159">
Both 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis simplex" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis simplex</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov. differ significantly from 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
in the shape of the male pleon, with that of 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
being relatively narrower, and in particular Pl5 being more constricted proximally (Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of pleonal somite proportions of Tmethypocoelis simplex sp. nov. and T. celebensis sp. nov. with the closely related T. koelbeli (proportions of latter taken from Davie 1990: fig. 1 A)." httpUri="http://table.plazi.org/id/8FC87AFC6FF2FFD917AA515AA3A8B65D" pageId="0" pageNumber="159" tableUuid="8FC87AFC6FF2FFD917AA515AA3A8B65D">2</tableCitation>
). The pleons of 
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov. are similar, however, both Pl6 and the telson are slightly proportionately wider in 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<caption ID-Table-UUID="8FC87AFC6FF2FFD917AA515AA3A8B65D" httpUri="http://table.plazi.org/id/8FC87AFC6FF2FFD917AA515AA3A8B65D" pageId="0" pageNumber="159" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="159">
<emphasis bold="true" pageId="0" pageNumber="159">Table 2.</emphasis>
Comparison of pleonal somite proportions of 
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis simplex" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis simplex</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov. with the closely related 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
(proportions of latter taken from 
<bibRefCitation author="Davie, PJF" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="159" pagination="463 - 473" refId="B5" refString="Davie, PJF, 1990. New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum 28: 463 - 473" title="New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea." volume="28" year="1990">Davie 1990</bibRefCitation>
: fig. 1A).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="159">
<table pageId="0" pageNumber="159">
<tr pageId="0" pageNumber="159">
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. liki" pageId="0" pageNumber="159" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">T. liki</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="159" rowspan="1">
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov.
</th>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Pleonite 5 width/length</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.1</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.5</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Pleonite 5 narrowest proximal width to distal width</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">0.7</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Pleonite 6 width/length</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.5</td>
</tr>
<tr pageId="0" pageNumber="159">
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">Telson width/length</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="159" rowspan="1">1.4</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="159">
<taxonomicName authorityName="Murniati &amp; Asakura &amp; Davie" authorityYear="2023" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tmethypocoelis simplex" order="Decapoda" pageId="0" pageNumber="159" phylum="Arthropoda" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">Tmethypocoelis simplex</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="T. liki" pageId="0" pageNumber="159" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">T. liki</emphasis>
</taxonomicName>
in the form and number of the apical setae of the G1. The G1 of 
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov. typically has two or three very long setae apically (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Reproductive organs of Tmethypocoelis simplex sp. nov. Paratype, males (A-D 4.1 x 2.6 mm E 6.9 x 4.0 mm) (MZB. Cru. 5183), Tosale, Banawa District, Donggala, Central Sulawesi, left G 1 A dorsal view B apical (dorsal view) C, D apical (ventral view) E mesial view. Paratype female (5.8 x 3.6 mm) (MZB. Cru. 5182), Towale river, Central Banawa District, Donggala, Central Sulawesi F vulva." figureDoi="10.3897/zookeys.1156.98930.figure8" httpUri="https://binary.pensoft.net/fig/831731" pageId="0" pageNumber="159">8C-E</figureCitation>
), and subapically there are three shorter stout setae on the outer margin increasing in length distally, and four short, downwardly reflexed setae on inner lobe. The G1 of 
<taxonomicName lsidName="T. celebensis" pageId="0" pageNumber="159" rank="species" species="celebensis">
<emphasis italics="true" pageId="0" pageNumber="159">T. celebensis</emphasis>
</taxonomicName>
sp. nov. has two or three very long recurved setae apically (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Reproductive organs of Tmethypocoelis celebensis b paratype, male (7.3 x 4.4 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi, left G 1 A mesial view B dorsal view C, D apical portion C dorsal view D ventral view. Paratype, female (5.5 x 3.5 mm) (MZB. Cru. 5180), Moletang River (estuary), Kema Tiga, North Minahasa, North Sulawesi E vulva." figureDoi="10.3897/zookeys.1156.98930.figure16" httpUri="https://binary.pensoft.net/fig/831739" pageId="0" pageNumber="159">16B-D</figureCitation>
), and subapically there are also three stout setae on the outer margin, though the proximal seta is much smaller and less prominent than on 
<taxonomicName lsidName="T. simplex" pageId="0" pageNumber="159" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="159">T. simplex</emphasis>
</taxonomicName>
sp. nov., and also four or five short, downwardly reflexed setae on the inner lobe. The G1 of 
<taxonomicName lsidName="T. koelbeli" pageId="0" pageNumber="159" rank="species" species="koelbeli">
<emphasis italics="true" pageId="0" pageNumber="159">T. koelbeli</emphasis>
</taxonomicName>
similarly has two long apical setae but lacks a row of outer subapical setae and has a row of five short distally pointed setae on the inner lobe (
<bibRefCitation author="Davie, PJF" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="159" pagination="463 - 473" refId="B5" refString="Davie, PJF, 1990. New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum 28: 463 - 473" title="New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea." volume="28" year="1990">Davie 1990</bibRefCitation>
: fig. 2). The G1 of 
<taxonomicName lsidName="T. liki" pageId="0" pageNumber="159" rank="species" species="liki">
<emphasis italics="true" pageId="0" pageNumber="159">T. liki</emphasis>
</taxonomicName>
has one long and five short apical setae (
<bibRefCitation DOI="https://doi.org/10.26107/RBZ-2022-0026" author="Murniati, DC" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="159" pagination="461 - 491" refId="B22" refString="Murniati, DC, Asakura, A, Nugroho, DA, Hernawan, UE, Dharmawan, IWE, 2022. On a collection of thoracotreme crabs (Crustacea: Brachyura: Ocypodidae, Macrophthalmidae, Dotillidae) from two offshore islands of Papua, eastern Indonesia, with descriptions of two new species. The Raffles Bulletin of Zoology 70: 461 - 491, DOI: https://doi.org/10.26107/RBZ-2022-0026" title="On a collection of thoracotreme crabs (Crustacea: Brachyura: Ocypodidae, Macrophthalmidae, Dotillidae) from two offshore islands of Papua, eastern Indonesia, with descriptions of two new species." url="https://doi.org/10.26107/RBZ-2022-0026" volume="70" year="2022">Murniati et al. 2022</bibRefCitation>
: fig. 20C).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>