treatments-xml/data/C4/26/FC/C426FC3804F05675ADACBDAF58995DB5.xml

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<mods:title id="45B85E00BCE6FA8A8945204B1F7E0A8A">Taxonomical review of Prosymna angolensis Boulenger, 1915 (Elapoidea, Prosymnidae) with the description of two new species</mods:title>
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<mods:namePart id="13DB716FF4BD69A776904165DC32C581">Conradie, Werner</mods:namePart>
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<mods:affiliation id="0E899863C437D40A16FD5B58FFA6E281">Port Elizabeth Museum (Bayworld), P. O. Box 13147, Humewood 6013, Gqeberha, South Africa &amp; Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa</mods:affiliation>
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<mods:namePart id="382938B3983C26249918B8E723D15493">Keates, Chad</mods:namePart>
<mods:nameIdentifier id="9296FECEAABE593E85F4A907ABEB6DC3" type="ORCID">https://orcid.org/0000-0003-3125-9536</mods:nameIdentifier>
<mods:affiliation id="EB0818740947664315B3D3A96D93D2C3">Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa &amp; South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, South Africa &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa</mods:affiliation>
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<mods:namePart id="7A21E86CF27D3B6C2B47CF013352338C">Baptista, Ninda L.</mods:namePart>
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<mods:affiliation id="8D3100D5FA561FC3B4F1BE4E261DEDAF">National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa &amp; Instituto Superior de Ciencias da Educacao da Huila (ISCED-Huila), Rua Sarmento Rodrigues, Lubango, Angola &amp; CIBIO-InBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, Laboratorio Associado, Universidade do Porto, Campus Agrario de Vairao, Rua Padre Armando Quintas, 4485 - 661 Vairao, Portugal &amp; Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal</mods:affiliation>
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<mods:namePart id="FE7CDE9D12C46217F6B8B807887C3C24">Lobon-Rovira, Javier</mods:namePart>
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<mods:affiliation id="1043C904841C7578454F791FE2206BAF">CIBIO-InBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, Laboratorio Associado, Universidade do Porto, Campus Agrario de Vairao, Rua Padre Armando Quintas, 4485 - 661 Vairao, Portugal &amp; Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal</mods:affiliation>
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<treatment id="C426FC3804F05675ADACBDAF58995DB5" ID-GBIF-Taxon="200621706" LSID="urn:lsid:zoobank.org:act:70A338BA-ED44-440B-AE3B-863E84B3AFBE" httpUri="http://treatment.plazi.org/id/C426FC3804F05675ADACBDAF58995DB5" lastPageNumber="97" pageId="0" pageNumber="97" scope_family="Prosymnidae" scope_superFamily="Elapoidea">
<subSubSection id="3DC74D58D304BF3ED9E26A749466979C" pageId="0" pageNumber="97" type="nomenclature">
<paragraph id="73CF139995979BEBA16D04902943709E" pageId="0" pageNumber="97">
<taxonomicName id="D846DF58A024D45A51C8117C85C32BB3" LSID="https://zoobank.org/70A338BA-ED44-440B-AE3B-863E84B3AFBE" authority="Conradie &amp; Keates &amp; Baptista &amp; Lobón-Rovira, 2022" authorityName="Conradie &amp; Keates &amp; Baptista &amp; Lobón-Rovira" authorityYear="2022" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Prosymna lisima" order="Squamata" pageId="0" pageNumber="97" rank="species" species="lisima" status="sp. nov.">Prosymna lisima</taxonomicName>
<taxonomicNameLabel id="2EDC79AE8C9735EDFCC63FADE60D5956" pageId="0" pageNumber="97">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="2FDC1BDBF0EAF1B3E88F81E953D5A48A" pageId="0" pageNumber="97" type="description">
<paragraph id="87588FB2030ED3AC40BE5F79C742DDDA" pageId="0" pageNumber="97">
<figureCitation id="2969C537133D50FB21EB9299BE5995BB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of live P. lisima sp. nov. from eastern Angola A PEM R 23457 from Quembo River Source, Moxico Province, Angola B PEM R 23483 from Cuando River Source, Moxico Province, Angola C PEM R 23512 from Cuito Source Lake, Moxico Province, Angola D PEM R 27381 from Quembo River bridge camp, Moxico Province, Angola." figureDoi="10.3897/zookeys.1121.85693.figure5" httpUri="https://binary.pensoft.net/fig/744036" pageId="0" pageNumber="97">Figs 5</figureCitation>
<figureCitation id="3BA0F9EC6CC21A42D7E5D7BA19DEF93B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Lateral B ventral C medial D dorsal views of skulls of Prosymna angolensis subgroup. Red arrows depict variation characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure7" httpUri="https://binary.pensoft.net/fig/744038" pageId="0" pageNumber="97">, 7</figureCitation>
<figureCitation id="5D1B33D3501D49A10766168F1E7535BD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Frontal and B posterior views of skulls, and C lateral, dorsal, medial and ventral view of jaw of Prosymna angolensis subgroup. Red arrows depict variation in characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure8" httpUri="https://binary.pensoft.net/fig/744039" pageId="0" pageNumber="97">, 8</figureCitation>
<figureCitation id="99C31D0BF73486C0DAB039F693E6302D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Dorsal, frontal and ventral view of premaxillae B medial, posterior and dorsal views of palatine; and C lateral, dorsal, and ventral view of maxilla in P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis and P. janii (from left to right)." figureDoi="10.3897/zookeys.1121.85693.figure9" httpUri="https://binary.pensoft.net/fig/744040" pageId="0" pageNumber="97">, 9</figureCitation>
<figureCitation id="F1E62DFB0F8992ED9D0045839A27B40F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">, 11</figureCitation>
<figureCitation id="85D652E0004101FD30A7012D889DF3C0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Stomach and gut contents of Prosymna lisima sp. nov. (PEM R 23456) consisting of lizard egg shells. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1121.85693.figure12" httpUri="https://binary.pensoft.net/fig/744043" pageId="0" pageNumber="97">
, 12 Common names: Kalahari Shovel-snout snake (English); 
<normalizedToken id="F58C45247567AC19FE0A806798A0C3BD" originalValue="Cobra-de-focinho-de-pá-do-kalahari">Cobra-de-focinho-de-pa-do-kalahari</normalizedToken>
(Portuguese).
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4091E634B5167BE9A054E67E528C1319" pageId="0" pageNumber="97" type="chresonymy">
<paragraph id="4237DE8F11D8D84F165B3D7CFB79B640" pageId="0" pageNumber="97">Chresonymy.</paragraph>
<paragraph id="E15087E25241A0343F2224D28A573DB5" pageId="0" pageNumber="97">
<taxonomicName id="D9890070EAD3ED0B26F912E0000C4FA0" authorityName="Boulenger" authorityYear="1915" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna angolensis" order="Squamata" pageId="0" pageNumber="97" rank="species" species="angolensis">
<emphasis id="381F81138238E5CADE1E75E716D3AFDD" italics="true" pageId="0" pageNumber="97">Prosymna angolensis</emphasis>
</taxonomicName>
: 
<bibRefCitation id="C67808C38D272801193DF953EA2696D0" author="Broadley, DG" journalOrPublisher="The Puku" pageId="0" pageNumber="97" pagination="1 - 143" refId="B17" refString="Broadley, DG, 1971. The reptiles and amphibians of Zambia. The Puku 6: 1 - 143" title="The reptiles and amphibians of Zambia." volume="6" year="1971">Broadley 1971</bibRefCitation>
: 82, 
<bibRefCitation id="BC5A0D36BB9BE2D9AB0274791A097D48" author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">1980</bibRefCitation>
: 512 (in part); 
<bibRefCitation id="B51289FECD0E129C363CBA204D860C77" author="Broadley, DG" journalOrPublisher="Edition Chimaira, Frankfurt" pageId="0" pageNumber="97" refId="B23" refString="Broadley, DG, Doria, CT, Wigge, J, 2003. Snakes of Zambia. An Atlas and Field Guide. Edition Chimaira, Frankfurt" title="Snakes of Zambia. An Atlas and Field Guide." year="2003">Broadley et al. 2003</bibRefCitation>
: 187 (in part); 
<bibRefCitation id="375D4D5DD33B6E23FE21CEF71AF260B1" author="Pietersen, DW" journalOrPublisher="Struik Nature, Cape Town" pageId="0" pageNumber="97" refId="B86" refString="Pietersen, DW, Verburgt, L, Davies, J, 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature, Cape Town" title="Snakes and other reptiles of Zambia and Malawi." year="2021">Pietersen et al. 2021</bibRefCitation>
: 97, fig.; 
<bibRefCitation id="601FE5B6AC17122DBC8ABCE7C7084A33" author="Conradie, W" journalOrPublisher="African Journal of Herpetology" pageId="0" pageNumber="97" refId="B38" refString="Conradie, W, Baptista, NL, Verburgt, L, Keates, C, Harvey, J, Julio, T, Neef, G, 2021. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian &amp; Reptile Conservation 15(2)[General Section]: 244-278[e292]." title="Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian &amp; Reptile Conservation 15 (2) [General Section]: 244 - 278 [e 292]." year="2021">Conradie et al. 2021</bibRefCitation>
: 265.
</paragraph>
</subSubSection>
<subSubSection id="F028D6DC7E3FCFE0B541EDA341A4718D" pageId="0" pageNumber="97" type="materials_examined">
<paragraph id="DA58938FCC4AE0AD6168992EAE3D4B30" pageId="0" pageNumber="97">Material examined.</paragraph>
<paragraph id="A435F205D5C382EEE31C52CC8A0DEA32" pageId="0" pageNumber="97">
<materialsCitation id="BC67D397F6EB9951B8E8F18AED7CA293" accessionNumber="R23512" collectingDate="2016-11-26" country="Angola" county="James Harvey" elevation="1426" latitude="-12.68866" location="Cuito River" longLatPrecision="1" longitude="18.36025" municipality="Werner Conradie" specimenCount="1" stateProvince="Moxico Province" typeStatus="Holotype">
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<typeStatus id="AEB396E6635B459540F2EB1C4EBD2F6F">Holotype</typeStatus>
</emphasis>
(male). PEM 
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, collected from 
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source lake (
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, 
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, 
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<quantity id="90AF898DA36928553D0370A6DBAE8C2A" metricMagnitude="3" metricUnit="m" metricValue="1.426" unit="m" value="1426.0">1426 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="D4077057CDABCD50C998FDD18AED1FBE" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="8317C371D79EB93E5381C9068D3B2301" name="Angola">Angola</collectingCountry>
by 
<collectingMunicipality id="0FD6EC8BE94605BF8F7D27CAD5FA194C">Werner Conradie</collectingMunicipality>
and 
<collectingCounty id="F3739C1D82BA88D601C992C49EFBCB5D">James Harvey</collectingCounty>
on 
<collectingDate id="0DCAF4D948D00101D645C8FAAEB3F188" value="2016-11-26">26 November 2016</collectingDate>
</materialsCitation>
. 
<materialsCitation id="BE8B88110F12E8F00239C4A0FB275965" accessionNumber="R27381" collectingDate="2019-11-27" collectorName="Werner Conradie, Keates, Timoteo Julio" country="Angola" elevation="1248" latitude="-13.526579" location="Quembo River" longLatPrecision="1" longitude="19.278095" specimenCount="five" specimenCount-male="1" stateProvince="Moxico Province" typeStatus="Paratypes">
<emphasis id="33E88D280C6473A66326877BBE4BD8F2" bold="true" italics="true" pageId="0" pageNumber="97">
<typeStatus id="9AEFC53B949B99A48D2AA4FE7D6EFD4A">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="01272AA13A2569E273A5C3E02D6457FC" type="male">five males</specimenCount>
). PEM 
<accessionNumber id="FBC5B1308E0CB0CF0F840BD2405B5146" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R27381">R27381</accessionNumber>
, collected from 
<location id="7BCAEBB0F225FF09D26940367FC4A539" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:7BCAEBB0F225FF09D26940367FC4A539" country="Angola" latitude="-13.526579" longLatPrecision="1" longitude="19.278095" name="Quembo River" stateProvince="Moxico Province">Quembo River</location>
lower bridge (
<geoCoordinate id="D14D42F2BD29B7B1F2510C199444A898" degrees="13.526579" direction="south" orientation="latitude" precision="1" value="-13.526579">-13.526579</geoCoordinate>
, 
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, 
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<quantity id="06CAAF56BCC6BD388B8BEF0F9C45069B" metricMagnitude="3" metricUnit="m" metricValue="1.248" unit="m" value="1248.0">1248 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="3E8041FDDD1E685A3C48538D99D8BE65" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="C5400CA87B804610AED82077598EAA81" name="Angola">Angola</collectingCountry>
by 
<collectorName id="67AD68CFCE355BA205378127461ED818">Werner Conradie</collectorName>
, 
<collectingCountry id="070F8F9DD7DB89BA689A3F8CAD1EA5E3" name="Chad">Chad</collectingCountry>
<collectorName id="884F662BE4D92B562763077A629B18A8">Keates</collectorName>
and 
<collectorName id="E7A86903B6D615ECD008D0FB50CB07F7">
<normalizedToken id="B39EBE83AEF695AF8403A31D4F66017D" originalValue="Timóteo">Timoteo</normalizedToken>
<normalizedToken id="993242B6D56BCD508799EFE1B92A3C54" originalValue="Júlio">Julio</normalizedToken>
</collectorName>
on 
<collectingDate id="6B8916160ED2D468F1D72C21531D0917" value="2019-11-27">27 November 2019</collectingDate>
</materialsCitation>
; 
<materialsCitation id="40D392F66E5873C8FF29E59031AA9925" accessionNumber="R23457" collectingDate="2016-11-03" collectorName="Werner Conradie" country="Angola" elevation="1368" latitude="-13.13586" location="Quembo River" longLatPrecision="1" longitude="19.04709" specimenCount="1" stateProvince="Moxico Province" typeStatus="paratype">
PEM 
<accessionNumber id="F726B7A95C10AF2F7C4617B7589C94C8" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R23457">R23457</accessionNumber>
-8, collected from 
<location id="717696E5BD3F7AF939D995FD011B5A78" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:717696E5BD3F7AF939D995FD011B5A78" country="Angola" latitude="-13.13586" longLatPrecision="1" longitude="19.04709" name="Quembo River" stateProvince="Moxico Province">Quembo River</location>
source (
<geoCoordinate id="FEE34196BD54CF76FD2ACF05D982DAC8" degrees="13.13586" direction="south" orientation="latitude" precision="1" value="-13.13586">-13.13586</geoCoordinate>
, 
<geoCoordinate id="AAF595C2433D6283E193E68C6E8E670D" degrees="19.04709" direction="east" orientation="longitude" precision="1" value="19.04709">19.04709</geoCoordinate>
, 
<elevation id="81309D639BB17954DA95BC1C5FD9EC9D" metricMagnitude="3" metricUnit="m" metricValue="1.368" unit="m" value="1368.0">
<quantity id="E45AB6D4A281CCEAA946B7A6D4A7F2FD" metricMagnitude="3" metricUnit="m" metricValue="1.368" unit="m" value="1368.0">1368 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="53792261AD8DD18622F79DF3BCF2FAA8" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="8A28921F46E5B4B58101D15E933077C6" name="Angola">Angola</collectingCountry>
by 
<collectorName id="13AA436949EE41122BC26F8344185F50">Werner Conradie</collectorName>
on 
<collectingDate id="B70ADE6831AEB89521088A4DB5DD51D7" value="2016-11-03">3 November 2016</collectingDate>
</materialsCitation>
; 
<materialsCitation id="F22C2B04F793B24B7D4DAB029D3435A2" accessionNumber="R23483" collectingDate="2016-11-17" collectorName="Werner Conradie, James Harvey" country="Angola" elevation="1372" latitude="-13.00164" location="Cuando River" longLatPrecision="5" longitude="19.1296" specimenCount="1" stateProvince="Moxico Province" typeStatus="paratype">
PEM 
<accessionNumber id="C71A69337E9ADDAC2FD043E5568CEF58" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R23483">R23483</accessionNumber>
, 
<location id="0113516850DFA828BA3EF1A33A27DF42" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:0113516850DFA828BA3EF1A33A27DF42" country="Angola" latitude="-13.00164" longLatPrecision="5" longitude="19.1296" name="Cuando River" stateProvince="Moxico Province">Cuando River</location>
source (
<geoCoordinate id="706F9A0B6766827B7842EE05114F2673" degrees="13.00164" direction="south" orientation="latitude" precision="1" value="-13.00164">-13.00164</geoCoordinate>
, 
<geoCoordinate id="368A52936F046D8250359A57184D03C6" degrees="19.1296" direction="east" orientation="longitude" precision="5" value="19.1296">19.1296</geoCoordinate>
, 
<elevation id="BE0ACFB33E91A4BBCA07783D19A30012" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" unit="m" value="1372.0">
<quantity id="FB0E4FF40104084623829BFCE7110A23" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" unit="m" value="1372.0">1372 m</quantity>
a.s.l.
</elevation>
) 
<collectingRegion id="80D46D531599F9B2CD479BF045535C7A" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="0663C8E76BDDF048A6DEC5D617892FAA" name="Angola">Angola</collectingCountry>
by 
<collectorName id="A9080B5BFFFE8DE88EFDA3BB93D535B4">Werner Conradie</collectorName>
and 
<collectorName id="11B833011C065C6F24F1780924CE125E">James Harvey</collectorName>
on 
<collectingDate id="91F0CE56B2F54DBB6F2A1E839E4C8C65" value="2016-11-17">17 November 2016</collectingDate>
</materialsCitation>
; 
<materialsCitation id="23851F6F763CBB610EF5E00DFFA6FAA1" accessionNumber="R23510" collectingDate="2016-11-26" collectorName="Werner Conradie, James Harvey" country="Angola" elevation="1426" latitude="-12.68866" location="Cuito River" longLatPrecision="1" longitude="18.36025" specimenCount="1" stateProvince="Moxico Province" typeStatus="paratype">
PEM 
<accessionNumber id="0E0625CAAD8647E00C3A8DE32B335891" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R23510">R23510</accessionNumber>
, collected from 
<location id="B4E9506D6C3C013CE5A150B68DD641CE" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:B4E9506D6C3C013CE5A150B68DD641CE" country="Angola" latitude="-12.68866" longLatPrecision="1" longitude="18.36025" name="Cuito River" stateProvince="Moxico Province">Cuito River</location>
source lake (
<geoCoordinate id="33955CDC1D0F206485CDF45DAEECD7A3" degrees="12.68866" direction="south" orientation="latitude" precision="1" value="-12.68866">-12.68866</geoCoordinate>
, 
<geoCoordinate id="4DB36A48DF40C735DF33C464A9DACB2E" degrees="18.36025" direction="east" orientation="longitude" precision="1" value="18.36025">18.36025</geoCoordinate>
, 
<elevation id="D3C9FAF36D2E5431E12911B472030836" metricMagnitude="3" metricUnit="m" metricValue="1.426" unit="m" value="1426.0">
<quantity id="231958610C5B8946D514051CDE1C0C08" metricMagnitude="3" metricUnit="m" metricValue="1.426" unit="m" value="1426.0">1426 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="23348BA0533BC71C9177999C4451AC39" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="6464CE54C59AA583B2D35A465966747B" name="Angola">Angola</collectingCountry>
by 
<collectorName id="F2327B3FF0885DE837F45E22DFB8D02B">Werner Conradie</collectorName>
and 
<collectorName id="B3C3F066783EBC0617DE7B1FFF1979DE">James Harvey</collectorName>
on 
<collectingDate id="597CD1127915BC7FC23ED6D6E3D9CBA9" value="2016-11-26">26 November 2016</collectingDate>
</materialsCitation>
. 
<materialsCitation id="0910CB7D4D82087264C20E786426D710" accessionNumber="R23456" collectingDate="2016-11-03" country="Angola" elevation="1368" latitude="-13.13586" location="Quembo River" longLatPrecision="1" longitude="19.04709" municipality="Werner Conradie" specimenCount="two" specimenCount-female="1" stateProvince="Moxico Province" typeStatus="Paratypes">
<emphasis id="B4A88FE83E6F3D02A14F827D4AD37970" bold="true" italics="true" pageId="0" pageNumber="97">
<typeStatus id="9B3A73A878E14CBBC04506CD77972B9D">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="83323B156B5110545D966EEBED2FE468" type="female">two females</specimenCount>
). PEM 
<accessionNumber id="94A0CA86B7E370B0B78A46505E15EB05" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R23456">R23456</accessionNumber>
, collected from 
<location id="29DCC537B4EE39E6FCDD02C37976A93F" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:29DCC537B4EE39E6FCDD02C37976A93F" country="Angola" latitude="-13.13586" longLatPrecision="1" longitude="19.04709" municipality="Werner Conradie" name="Quembo River" stateProvince="Moxico Province">Quembo River</location>
source (
<geoCoordinate id="85927151061CC534CA175B4D3413B7DF" degrees="13.13586" direction="south" orientation="latitude" precision="1" value="-13.13586">-13.13586</geoCoordinate>
, 
<geoCoordinate id="96674CA75A12718698278818759992D2" degrees="19.04709" direction="east" orientation="longitude" precision="1" value="19.04709">19.04709</geoCoordinate>
, 
<elevation id="4FFDD3FE99FB788375D8984CE1BB0248" metricMagnitude="3" metricUnit="m" metricValue="1.368" unit="m" value="1368.0">
<quantity id="D9506323B58A1928F59E338686FF7404" metricMagnitude="3" metricUnit="m" metricValue="1.368" unit="m" value="1368.0">1368 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="7B7A6356C88B27A180677DBF60A846DE" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="00D5608DBA3AFED906ED2D3CF0792772" name="Angola">Angola</collectingCountry>
by 
<collectingMunicipality id="14165B1200E9B1A1C53D7A8A27C949CB">Werner Conradie</collectingMunicipality>
on 
<collectingDate id="0A735D5FDEE4E91A065BB49C8FE727D8" value="2016-11-03">3 November 2016</collectingDate>
</materialsCitation>
; 
<materialsCitation id="7895970446F9A0A5CD765288D054EC14" accessionNumber="R23511" collectingDate="2016-11-26" country="Angola" county="James Harvey" elevation="1426" latitude="-12.68866" location="Cuito River" longLatPrecision="1" longitude="18.36025" municipality="Werner Conradie" specimenCount="1" stateProvince="Moxico Province" typeStatus="paratype">
PEM 
<accessionNumber id="7FB25D3B41FB3372B86CFE98567D96FF" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R23511">R23511</accessionNumber>
, 
<location id="6871A5E322DEBBEB9EC23FB1B4C0157E" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:6871A5E322DEBBEB9EC23FB1B4C0157E" country="Angola" county="James Harvey" latitude="-12.68866" longLatPrecision="1" longitude="18.36025" municipality="Werner Conradie" name="Cuito River" stateProvince="Moxico Province">Cuito River</location>
source lake (
<geoCoordinate id="5CAB8881D010F1CFE1E1E5C58962B10A" degrees="12.68866" direction="south" orientation="latitude" precision="1" value="-12.68866">-12.68866</geoCoordinate>
, 
<geoCoordinate id="83590FBF58100EB989A772CE941D197F" degrees="18.36025" direction="east" orientation="longitude" precision="1" value="18.36025">18.36025</geoCoordinate>
, 
<elevation id="8AA2D955078755094A3A72A60834D74B" metricMagnitude="3" metricUnit="m" metricValue="1.426" unit="m" value="1426.0">
<quantity id="4EEC3CADEA076E2F862D0F6A7ED17253" metricMagnitude="3" metricUnit="m" metricValue="1.426" unit="m" value="1426.0">1426 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="8D2BD78F40AFF447E04C5252ABDC41E4" country="Angola" name="Moxico">Moxico Province</collectingRegion>
, 
<collectingCountry id="924645F8DAC88FBB4ABC0FFD3DD4B77A" name="Angola">Angola</collectingCountry>
by 
<collectingMunicipality id="FA910715D6325428E75988F807D4BE2D">Werner Conradie</collectingMunicipality>
and 
<collectingCounty id="F7008981CE5EFBC73F916B203818C292">James Harvey</collectingCounty>
on 
<collectingDate id="49652EB41E14FFDFD53954890F6274A7" value="2016-11-26">26 November 2016</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D82EC69BE6465187828B4CF8858B263E" pageId="0" pageNumber="97" type="materials_examined">
<paragraph id="75F79331FAE31F9E2276F07BDBB7DCF4" pageId="0" pageNumber="97">Additional material assigned to the new species.</paragraph>
<paragraph id="F99052792E44E7EBC0CF0E3A977139C7" pageId="0" pageNumber="97">
<materialsCitation id="CB4646BAC1D3201F851CD5EF75FA546C" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Zambia" latitude="-14.46667" location="Katima Mulilo" longLatPrecision="5" longitude="16.3" specimenCount="1">
NMZB-UM 10096, Kalabo (approx. 
<geoCoordinate id="6DBD43BDC07B79E7732990B191D228FA" degrees="14.99391" direction="south" orientation="latitude" precision="1" value="-14.99391">-14.99391</geoCoordinate>
, 
<geoCoordinate id="51FDCAE17169BFF2A59D1F447C7F8311" degrees="22.67795" direction="east" orientation="longitude" precision="1" value="22.67795">22.67795</geoCoordinate>
), 
<collectingCountry id="59FFF8DFBD92EA47B3652A1AA2639E77" name="Zambia">Zambia</collectingCountry>
; NMZB-UM 21272, 15 km WSW of 
<location id="EA278B3E248E8F17BA0BF8AB509895EC" LSID="urn:lsid:plazi:treatment:C426FC3804F05675ADACBDAF58995DB5:EA278B3E248E8F17BA0BF8AB509895EC" country="Zambia" latitude="-14.46667" longLatPrecision="5" longitude="16.3" name="Katima Mulilo">Katima Mulilo</location>
(approx. 
<geoCoordinate id="FEFB0B54531C0C5F395164D0C22EAABF" degrees="17.61448" direction="south" orientation="latitude" precision="1" value="-17.61448">-17.61448</geoCoordinate>
, 
<geoCoordinate id="D637D8688B40F8B52385D9259D36A477" degrees="24.20593" direction="east" orientation="longitude" precision="1" value="24.20593">24.20593</geoCoordinate>
), 
<collectingCountry id="38C7508A91B2A36DBEA658A79EAEC248" name="Namibia">Namibia</collectingCountry>
. A specimen from Kuvangu [= Vila-da-Ponte], 
<geoCoordinate id="18C9529EDCC1DB22ADAFB729C58EB2A1" degrees="14.46667" direction="south" orientation="latitude" precision="1" value="-14.46667">-14.46667</geoCoordinate>
, 
<geoCoordinate id="C4A55B515E536C5327876D9C1E7694E7" degrees="16.3000" direction="east" orientation="longitude" precision="5" value="16.3">16.3000</geoCoordinate>
(
<bibRefCitation id="F2C06FA4177CA961C05DD48EA166729B" author="Monard, A" journalOrPublisher="Arquivos do Museu Bocage" pageId="0" pageNumber="97" pagination="19 - 154" refId="B79" refString="Monard, A, 1937. Contribution a l'Herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154" title="Contribution a l'Herpetologie d'Angola." volume="8" year="1937">Monard 1937</bibRefCitation>
: 123) has two postoculars and the characteristic confluent blotched dorsal pattern and might belong to this new species, but this needs verification and is thus tentatively referred to the new species
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2672AE0373BD69BD1698D75E6A1738ED" pageId="0" pageNumber="97" type="diagnosis">
<paragraph id="606C144E47989A5C60AE9D79541226E6" pageId="0" pageNumber="97">Diagnosis.</paragraph>
<paragraph id="51B685C3C5335E7DB6CB7785B8016704" pageId="0" pageNumber="97">
The new species differs from other 
<taxonomicName id="B4E8A8A535108082EDD5F1238342A5E6" authorityName="Gray" authorityYear="1849" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna" order="Squamata" pageId="0" pageNumber="97" rank="genus">
<emphasis id="97613ECA27715D1B050FB4900654ACB9" italics="true" pageId="0" pageNumber="97">Prosymna</emphasis>
</taxonomicName>
in the following characters: rostral sharply depressed and angular (vs. rounded in 
<taxonomicName id="5F66B10B63F37B9505E9A2E14BD7548D" lsidName="P. visseri" pageId="0" pageNumber="97" rank="species" species="visseri">
<emphasis id="18E8CE09688D93FDFC2B4BF7F5CEB195" italics="true" pageId="0" pageNumber="97">P. visseri</emphasis>
</taxonomicName>
); presences of a single band-like internasals (vs. paired internasals in 
<taxonomicName id="AE5E2E2B171019C947F727847FDEA11F" lsidName="P. somalica" pageId="0" pageNumber="97" rank="species" species="somalica">
<emphasis id="59BA02A4950FB2F9DE0B9AE38AE43A97" italics="true" pageId="0" pageNumber="97">P. somalica</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7B850AD422B13D644AE81998AE27F5D" lsidName="P. bivittata" pageId="0" pageNumber="97" rank="species" species="bivittata">
<emphasis id="AB70C00FE6FF3423DF809D3323086168" italics="true" pageId="0" pageNumber="97">P. bivittata</emphasis>
</taxonomicName>
, 
<taxonomicName id="2C05B6EF020DD4AF294F30178315EA66" lsidName="P. sudevalli" pageId="0" pageNumber="97" rank="species" species="sudevalli">
<emphasis id="E945ED610F6FB35A64384D0013D3204E" italics="true" pageId="0" pageNumber="97">P. sudevalli</emphasis>
</taxonomicName>
, 
<taxonomicName id="EF38694163F53AC1EF485F86809630D3" lsidName="P. lineata" pageId="0" pageNumber="97" rank="species" species="lineata">
<emphasis id="2A6F2124A107D117E69EFE0DE4A65A80" italics="true" pageId="0" pageNumber="97">P. lineata</emphasis>
</taxonomicName>
); dorsal scales smooth (keeled in 
<taxonomicName id="17E754FF2BAD199E4D95BCB5729FA701" lsidName="P. janii" pageId="0" pageNumber="97" rank="species" species="janii">
<emphasis id="E3217B32ADDF10D5E6FBBB42B799F72C" italics="true" pageId="0" pageNumber="97">P. janii</emphasis>
</taxonomicName>
); midbody scale rows 15-17 (vs. 19-21 in 
<taxonomicName id="F7925CAD0A9AD8A803242C7B75BC465D" lsidName="P. pitmani" pageId="0" pageNumber="97" rank="species" species="pitmani">
<emphasis id="AE8A738CB64E347B139AE92313714986" italics="true" pageId="0" pageNumber="97">P. pitmani</emphasis>
</taxonomicName>
); 6 supralabials, with 3rd and 4th entering orbit (vs. 5 supralabials, with 2nd and 3rd entering orbit in 
<taxonomicName id="EBE07D11C7CFAEB9FA35E86C2AFAB9AB" lsidName="P. meleagris" pageId="0" pageNumber="97" rank="species" species="meleagris">
<emphasis id="41DD1E3668BB94C929E7AD4E8E9145CF" italics="true" pageId="0" pageNumber="97">P. meleagris</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8E3874BD91DFAD0C4C08093901B5652" lsidName="P. greigerti" pageId="0" pageNumber="97" rank="species" species="greigerti">
<emphasis id="084C6412B270F6BC10959AF0DAE90362" italics="true" pageId="0" pageNumber="97">P. greigerti</emphasis>
</taxonomicName>
); single apical pits on dorsal scales (vs. paired apical pits in 
<taxonomicName id="2C5AEFF0ED2839B7F1D0422FD8C482A6" lsidName="P. ruspolii" pageId="0" pageNumber="97" rank="species" species="ruspolii">
<emphasis id="E80FB0B7A16C5DFFAABC2445B2ED0BBA" italics="true" pageId="0" pageNumber="97">P. ruspolii</emphasis>
</taxonomicName>
); lower number of ventral scales in both sexes (116-129 vs. 153-199 in 
<taxonomicName id="89F43324E67718B1BBED836AC8B50931" lsidName="P. frontalis" pageId="0" pageNumber="97" rank="species" species="frontalis">
<emphasis id="B065954E97375C51E3FC510C2782F67D" italics="true" pageId="0" pageNumber="97">P. frontalis</emphasis>
</taxonomicName>
); dorsum with dark black spots (vs. scarlet head and dark body in 
<taxonomicName id="40904DB5C7862520907854170EF6D8FC" lsidName="P. ornatissima" pageId="0" pageNumber="97" rank="species" species="ornatissima">
<emphasis id="B093E0684CB05DEF75A13A06CEF4D289" italics="true" pageId="0" pageNumber="97">P. ornatissima</emphasis>
</taxonomicName>
; uniform dark brown to grey in 
<taxonomicName id="687FCB330FDA9935F00F1366220AB2DE" lsidName="P. ambigua" pageId="0" pageNumber="97" rank="species" species="ambigua">
<emphasis id="F7ECAADE26E172FEE4C4C96D54E4DBFB" italics="true" pageId="0" pageNumber="97">P. ambigua</emphasis>
</taxonomicName>
and 
<taxonomicName id="8AC9E90C0A10C59327F16AA31D2443BE" lsidName="P. stuhlmanni" pageId="0" pageNumber="97" rank="species" species="stuhlmanni">
<emphasis id="BECCD9A55CD115F8396A406EEB3EFF3B" italics="true" pageId="0" pageNumber="97">P. stuhlmanni</emphasis>
</taxonomicName>
). It further differs from its closest congener, 
<taxonomicName id="316A4EA48AF197FC645C2CEE1E542BE5" lsidName="P. angolensis" pageId="0" pageNumber="97" rank="species" species="angolensis">
<emphasis id="789F94AB6E586F11EEAC01A51C2AA904" italics="true" pageId="0" pageNumber="97">P. angolensis</emphasis>
</taxonomicName>
, in having two post oculars (vs. one), dorsal large black blotches mostly fused (vs. mostly small paired dorsal grey to black spots), postorbital bone present (vs. absent) and by the presence of four to five well-developed palatine teeth (vs. three reduced teeth).
</paragraph>
</subSubSection>
<subSubSection id="0EE132B26BEC94A69BD420967CFA636F" pageId="0" pageNumber="97" type="etymology">
<paragraph id="C09ADA72A183BEFE9BF39BF7A5BF7342" pageId="0" pageNumber="97">Etymology.</paragraph>
<paragraph id="C3418FBFF84F82F8A106F97C526DDA05" pageId="0" pageNumber="97">
The name 
<emphasis id="B7E3B68696901ABC4009A2C5FCDD197E" italics="true" pageId="0" pageNumber="97">lisima</emphasis>
is derived from the locally spoken Luchaze language in the region of the type locality meaning 
<normalizedToken id="76D0420B37BA94C91F9015FA2E5BE7CF" originalValue="‘source’">'source'</normalizedToken>
. The full phrase used, ' 
<emphasis id="C5221A288F7948A4D6803625C224BFFC" italics="true" pageId="0" pageNumber="97">Lisima Lwa Mwondo</emphasis>
', is translated as &quot;source of life&quot;. This is a reference to central Angola, a high rainfall area where some of the most important rivers in Angola arise. This water makes it its way to the Okavango Delta, sustaining wildlife and local communities in Angola, Namibia and Botswana.
</paragraph>
</subSubSection>
<subSubSection id="190A0415B81EEA97D41C91B275CBEABA" pageId="0" pageNumber="97" type="description">
<paragraph id="86CBB261B11CD16EF3C6FED4605B34D6" pageId="0" pageNumber="97">Description of holotype</paragraph>
<paragraph id="05D112AC3B520947BF9A0E5292C2E470" pageId="0" pageNumber="97">
<emphasis id="460316FFA62DB2F2CDC5FA14B42B0EA0" bold="true" pageId="0" pageNumber="97">
(Fig. 
<figureCitation id="83285375A677A9E53E660F1A7455FA69" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11</figureCitation>
).
</emphasis>
See Table 
<tableCitation id="3FB5D4C52C9A755524C8B752588241E3" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological features and measurements for the type series of Prosymna lisima sp. nov. (SVL = snout-vent length, t = truncated)." httpUri="http://table.plazi.org/id/2ED70644E9B2B616C3BE33FAE3A24FD4" pageId="0" pageNumber="97" tableUuid="2ED70644E9B2B616C3BE33FAE3A24FD4">4</tableCitation>
for further details and meristic data for the holotype. The body is cylindrical and elongated, tapering gradually to a very short tail, 13% total length, tail tip with a prominent spike. Dorsal scales smooth with single apical pits (some suprasubcaudal scales have two apical pits) in 17-15-15 scale rows, scale row reduction from 17 to 16 take place at ventral number 17 with the fusion of 3rd and 4th dorsal scale rows on left side and from 16 to 15 at ventral 23 with the fusion of 3rd and 4th dorsal scale rows on right side; 122 ventral scales; cloaca entire; 13 paired subcaudal scales. Head in dorsal view (Fig. 
<figureCitation id="95864912E1BE6E72B0A638606D851026" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11B</figureCitation>
): head narrow and rounded, barely wider than 
<normalizedToken id="81796ACBC8D807589872857E5A00E8E6" originalValue="‘neck’">'neck'</normalizedToken>
; rostral clearly visible from above, much broader than long (3.39 
<normalizedToken id="9EAEA80306F91FBAA28B4D0412F4136D" originalValue="×">x</normalizedToken>
1.06 mm); a single narrow internasal, which is much longer than wide (2.73 
<normalizedToken id="1E780DFBC6C10845CFF36AFF3DDE365C" originalValue="×">x</normalizedToken>
0.67 mm) and in broad contact with the rostral anteriorly, posteriorly in broad contact with prefrontal and laterally with nasals; single band-like prefrontal which is longer than wide (3.70 
<normalizedToken id="4D7D62AE51A45973B3DDBEB042B98728" originalValue="×">x</normalizedToken>
1.30 mm), in contact laterally with loreal, and posteriorly with the frontal and supraocular scales; frontal pentangular, almost as long as wide (3.09 
<normalizedToken id="43AB837609A066BF1B4D83C44D78B142" originalValue="×">x</normalizedToken>
3.00 mm), nearly equal distance to snout (3.30 mm), shorter than prefrontals (3.0 vs. 3.70 mm), but nearly equal in length to the parietal scales (3.00 vs. 3.04 mm), in contact laterally with narrow supraoculars, and posteriorly with two very large parietals; paired parietals as wide as long (3.04 
<normalizedToken id="658BDB76F38F958284458658DFCA1893" originalValue="×">x</normalizedToken>
3.04 mm), in contact posteriorly with each other and laterally with temporals. Head in ventral view (Fig. 
<figureCitation id="593EA58ECB5098DD1E53217A859861C8" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11C</figureCitation>
): rostral clearly visible from below, protruding well past jawline; mental small, triangular; infralabials seven, first three in contact with single paired chin shields, 1st infralabials in contact with each other; additional three or four rows of smaller gular scales present before start of ventral scales. Head in lateral view (Fig. 
<figureCitation id="63626CD534BC65CBAEE9AFE40CE25CF7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11D</figureCitation>
): snout sharply pointed, longer than the horizontal diameter of eye (ED/SL = 0.49); rostral large with acutely horizontal angular edge, excavated below; nostril is oval shaped, piercing a fully-divided nasal, and directed backwards; nasal scale longer than wide, with anterior part in full contact with rostral, posterior lower corner in contact with 1st supralabial and above with internasal scale and prefrontal; nasal suture present and intersecting 1st supralabial in uppermost corner; single small loreal as long as wide (0.7 
<normalizedToken id="7F5278B4AD69646583DB74FBFD30B1DA" originalValue="×">x</normalizedToken>
0.7 mm), in contact below with 1st and 2nd supralabial, above with prefrontal, anteriorly with nasal and posteriorly with single preocular; a single preocular on the right side and two on the left side in contact anteriorly with loreal and prefrontal and above with supraocular, posteriorly protruding of loreal overlap with preoculars to create a small flap; eye large 19.36% headlight, vertical diameter (1.47 mm) two thirds as deep as distance between eye and lip (0.99 mm); pupil round; two postoculars, the lower one largest and in contact with 4th and 5th supralabials, first temporal scale and parietal, the upper smaller in contact with both supraocular and parietal; temporals 1+2 on both sides; narrow elongated supraocular in contact anteriorly with preocular, posteriorly with upper postocular and parietal and above with frontal; six supralabials, 3rd and 4th contacting eye, 5th and 6th supralabial the largest.
</paragraph>
<caption id="2ED70644E9B2B616C3BE33FAE3A24FD4" ID-Table-UUID="2ED70644E9B2B616C3BE33FAE3A24FD4" httpUri="http://table.plazi.org/id/2ED70644E9B2B616C3BE33FAE3A24FD4" pageId="0" pageNumber="97" start="Table 4" startId="T4">
<paragraph id="F0569B0F677A7D842BE226B72BF4AD6C" pageId="0" pageNumber="97">
<emphasis id="C2A5B263F410917E5F90023D09E75F6B" bold="true" pageId="0" pageNumber="97">Table 4.</emphasis>
Morphological features and measurements for the type series of 
<taxonomicName id="1DFAAD70549F6BB36F98D52F6496349D" authorityName="Conradie &amp; Keates &amp; Baptista &amp; Lobón-Rovira" authorityYear="2022" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna lisima" order="Squamata" pageId="0" pageNumber="97" rank="species" species="lisima">
<emphasis id="376531F48F9CF89FEBA6833C93D047D9" italics="true" pageId="0" pageNumber="97">Prosymna lisima</emphasis>
</taxonomicName>
sp. nov. (SVL = snout-vent length, t = truncated).
</paragraph>
</caption>
<paragraph id="5C6378232D5F62D054EFC97E2A8980A0" pageId="0" pageNumber="97">
<table id="5D6D5778926F146C57BACA4D9D895A16" pageId="0" pageNumber="97">
<tr id="ECC3DC5F64A355D85531A3620E9A54CA" pageId="0" pageNumber="97">
<th id="1E52D360A3F71C9566C116EFA4B5DF3F" colspan="1" pageId="0" pageNumber="97" rowspan="1">Catalogue number</th>
<th id="4877DC713128BBEDD9078D051CA32777" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23512</th>
<th id="25335D4FB9B3E473820AC2471268B7FC" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23456</th>
<th id="36AA287430C31689323CB8EA97430969" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23457</th>
<th id="902EF671F0118492932D5B7CEE6CBD5C" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23458</th>
<th id="5C5D00BC0A2EF01879D1A8A50CDE7BE8" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23483</th>
<th id="DBD06044CA889CD25DEC95D50BA1131C" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23510</th>
<th id="C61F035ED7C227B87A8CDC1178411F59" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R23511</th>
<th id="AD3D27FBAD29EB85F5B64F6D7367D291" colspan="1" pageId="0" pageNumber="97" rowspan="1">PEM R27381</th>
</tr>
<tr id="4167482C28619F594F9E36D3D9C867A5" pageId="0" pageNumber="97">
<td id="1A385050B8900763441C572647DD6E84" colspan="1" pageId="0" pageNumber="97" rowspan="1">Type status</td>
<td id="F78CC79754AAE2C03B2E09F2A50964C3" colspan="1" pageId="0" pageNumber="97" rowspan="1">Holotype</td>
<td id="43504CA1BE98BCEF5FB4C5230733FDBD" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="FB636959C16AEC9275C9EE802BFB1486" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="112B4A6C08CCD4579453B2DF78F68141" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="BA8057F2E85A6E44F7B4DE5670B1F291" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="26030618E9AA44BF3DB75FD3CE647B4A" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="83207601401440077D2437E70688BB8B" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
<td id="5436A81451981A05D58F0FB2BEC46BEE" colspan="1" pageId="0" pageNumber="97" rowspan="1">Paratype</td>
</tr>
<tr id="8FAF126E5C5ADDA046589A29EBE4987B" pageId="0" pageNumber="97">
<td id="2B056747E0AE91FAD5E72F3BA8C4087A" colspan="1" pageId="0" pageNumber="97" rowspan="1">Sex</td>
<td id="48F73861DF971542549BC587856D2764" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
<td id="6A74EDDD2E39360C50FB8C9B01D24283" colspan="1" pageId="0" pageNumber="97" rowspan="1">Female</td>
<td id="AFF5381816AA4D4B2521EEF923D1160C" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
<td id="6F51BD5C3F09F6665F69B0639C8773AE" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
<td id="204A458A57EE06042AD6321BBD0843FB" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
<td id="E8550F1563798B84C5CDA4198F0A431F" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
<td id="DC5A49E17E937C4788020F6DC21FA0B1" colspan="1" pageId="0" pageNumber="97" rowspan="1">Female</td>
<td id="6578EDE8F647A33772BDB352C27E19B2" colspan="1" pageId="0" pageNumber="97" rowspan="1">Male</td>
</tr>
<tr id="140644C21B299229C8DB23953A853B03" pageId="0" pageNumber="97">
<td id="925B8659B52C9E24B629F469746F39CF" colspan="1" pageId="0" pageNumber="97" rowspan="1">SVL+TL = total length mm</td>
<td id="5712DA77327E62889FFA84C18DDE36C0" colspan="1" pageId="0" pageNumber="97" rowspan="1">193+28.9 = 221.9</td>
<td id="2CDFE9FBAC2FE2B25322A5C73347D007" colspan="1" pageId="0" pageNumber="97" rowspan="1">200+19.3 = 219.3</td>
<td id="A426EBD9A3BB2498E5AF7317A5549D79" colspan="1" pageId="0" pageNumber="97" rowspan="1">194+25.8 = 219.8</td>
<td id="08DDC330ECCC0A6DCD03ACBF1EC79158" colspan="1" pageId="0" pageNumber="97" rowspan="1">198+24.6 = 222.6</td>
<td id="857E55228EC1445E5B3C493D4D783F85" colspan="1" pageId="0" pageNumber="97" rowspan="1">181+23.8 = 204.8</td>
<td id="B202B63B89D16EDE90FBC2FA757FBC2B" colspan="1" pageId="0" pageNumber="97" rowspan="1">186+26.3 = 212.3</td>
<td id="EC33308553D2D526F2F8D9EF01981A36" colspan="1" pageId="0" pageNumber="97" rowspan="1">168+20.6 = 188.6</td>
<td id="7CCD6CA69107DAD6DD021F7FB8529105" colspan="1" pageId="0" pageNumber="97" rowspan="1">138+17t = 155</td>
</tr>
<tr id="BB31399D0575C2E19E4DA7B189A7F846" pageId="0" pageNumber="97">
<td id="17B587FEFB9C652C852CA84CECF797E7" colspan="1" pageId="0" pageNumber="97" rowspan="1">TL/total length ratio (%)</td>
<td id="32C92EBA65A66786E2A1A38F139C4175" colspan="1" pageId="0" pageNumber="97" rowspan="1">13.0</td>
<td id="ED73F8E2CE52B9F2F861F97CC87E476D" colspan="1" pageId="0" pageNumber="97" rowspan="1">8.8</td>
<td id="ADEF4CEFC7446959B6255FF1B1612E56" colspan="1" pageId="0" pageNumber="97" rowspan="1">11.7</td>
<td id="E9697766E26F571D257DCF0BCAE1E294" colspan="1" pageId="0" pageNumber="97" rowspan="1">11.1</td>
<td id="621644076CC80730657BE4886440D55D" colspan="1" pageId="0" pageNumber="97" rowspan="1">11.6</td>
<td id="6F3619C81079819D21D09E2A9490A23F" colspan="1" pageId="0" pageNumber="97" rowspan="1">12.4</td>
<td id="12B33F443D3D37A88353904B1ED0C615" colspan="1" pageId="0" pageNumber="97" rowspan="1">10.9</td>
<td id="966A467BB5AAF7F1B2006B25FF067F69" colspan="1" pageId="0" pageNumber="97" rowspan="1">11.0</td>
</tr>
<tr id="E960DFF0E87B404C22DCB6856671278F" pageId="0" pageNumber="97">
<td id="FB5A7E89CD34A726165828DFC160330C" colspan="1" pageId="0" pageNumber="97" rowspan="1">Midbody scale rows</td>
<td id="E3AB46CBD3A8A9B6DE21E4027D418F03" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-17-15</td>
<td id="B623BD431640E751BB6F2EE4A9F840C1" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-17-15</td>
<td id="8780894466FFB59C6946F9423D9903A9" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-15-15</td>
<td id="48C36A47FFB60477B03D930AB605210E" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-17-15</td>
<td id="CF3F2F8C95D5BBD8B3B086CB1014E4C3" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-15-15</td>
<td id="95611ADD52C918F3028E73AA9041E06B" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-17-15</td>
<td id="7E6D987BB9E81364D8071889A50EA6D7" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-15-15</td>
<td id="2DB59FA5D5427E4162DD7F6C3F8AC7F8" colspan="1" pageId="0" pageNumber="97" rowspan="1">17-15-15</td>
</tr>
<tr id="501260451AEE6C3F5C225D445AB29D29" pageId="0" pageNumber="97">
<td id="8C931B9B5C97594CB08F47A70E97E4ED" colspan="1" pageId="0" pageNumber="97" rowspan="1">Ventral scales</td>
<td id="AFB6270DBB6CBF5881D8572A78864459" colspan="1" pageId="0" pageNumber="97" rowspan="1">122</td>
<td id="0A9B216B0ABC07254F962A54F59E83B9" colspan="1" pageId="0" pageNumber="97" rowspan="1">121</td>
<td id="EE03544143CB395BA9B2026DA97DC344" colspan="1" pageId="0" pageNumber="97" rowspan="1">117</td>
<td id="9F0E141DADC36F356873E0970F0EF3D5" colspan="1" pageId="0" pageNumber="97" rowspan="1">124</td>
<td id="2209B3D124B9C58208841EF65D8C970D" colspan="1" pageId="0" pageNumber="97" rowspan="1">121</td>
<td id="BDC448177B377FDE73A5FEDAE045294D" colspan="1" pageId="0" pageNumber="97" rowspan="1">116</td>
<td id="282E0A3549C41D4B30E20A28FBC39F65" colspan="1" pageId="0" pageNumber="97" rowspan="1">117</td>
<td id="E34BA16AA0425A95B16EF37B7F4B10FD" colspan="1" pageId="0" pageNumber="97" rowspan="1">122</td>
</tr>
<tr id="6FB4FC4FF544822F3187D5BA0DEDFFFC" pageId="0" pageNumber="97">
<td id="A027AC7E5F8FF46B6F0A5115DE90AC80" colspan="1" pageId="0" pageNumber="97" rowspan="1">Subcaudal scales</td>
<td id="87F1E8189E7F8643499108AACEAE8C2D" colspan="1" pageId="0" pageNumber="97" rowspan="1">26</td>
<td id="34306E09B5F9300E3A437151C7B34AD7" colspan="1" pageId="0" pageNumber="97" rowspan="1">18</td>
<td id="9E248B23EB1B88D673B0B737C32DB1DF" colspan="1" pageId="0" pageNumber="97" rowspan="1">22</td>
<td id="0B246761BEF4B63A6469BD584E4772BE" colspan="1" pageId="0" pageNumber="97" rowspan="1">23</td>
<td id="5CED93FC1CC9CBFA4927DEFD49381894" colspan="1" pageId="0" pageNumber="97" rowspan="1">22</td>
<td id="AB4B8C1D51C811A475AE28CFC3A053BA" colspan="1" pageId="0" pageNumber="97" rowspan="1">23</td>
<td id="DE6C7B4F8C3B4806FDC5341D42F869AF" colspan="1" pageId="0" pageNumber="97" rowspan="1">24</td>
<td id="E74A260A907FD35171EBAC6C74A0D2D9" colspan="1" pageId="0" pageNumber="97" rowspan="1">21t</td>
</tr>
<tr id="A20541EC0A60A20F75B0CC77491DF6F3" pageId="0" pageNumber="97">
<td id="6418E47D61A4C4680FC1CD1EBA908827" colspan="1" pageId="0" pageNumber="97" rowspan="1">Cloacal Scale</td>
<td id="B6C3F09B40DCEA4CB62256030C95342A" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="C41493E60DC36A7743237CC927F041D5" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="8B64F6C36ACC07783200C531D0C93910" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="08413AD079A56324BC46E194DA5BE217" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="1507C4EBFEC8F41FB35EA8662756C060" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="F8AB991DB6605C603BEDD0CF3A5DACB1" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="3D7D4006232D32E8E2DA8D8756515927" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
<td id="A4E246A6E018AE02DE230A6CB2615942" colspan="1" pageId="0" pageNumber="97" rowspan="1">Entire</td>
</tr>
<tr id="CA41EA9B05A91ECAADEB269D120DBF3C" pageId="0" pageNumber="97">
<td id="4DACEE2C8495D9D8BC518205258F7F33" colspan="1" pageId="0" pageNumber="97" rowspan="1">Preoculars</td>
<td id="2D0EE0BE6F0DFC3C0D081A8C73608FF4" colspan="1" pageId="0" pageNumber="97" rowspan="1">1/2</td>
<td id="48CA61930A91773687BC42822FBF306C" colspan="1" pageId="0" pageNumber="97" rowspan="1">1</td>
<td id="EF2403DA2C5FF055EAB8AED9537AD00C" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="AFE98482D2D14EE8FD65B94A7763D7AE" colspan="1" pageId="0" pageNumber="97" rowspan="1">1</td>
<td id="A821F355263542E32C86B78A762BF7BD" colspan="1" pageId="0" pageNumber="97" rowspan="1">1</td>
<td id="EBBB4A460BFEACAB5452760C9A7FAFAC" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="B190AF013F5090711ED67A182BC17A0F" colspan="1" pageId="0" pageNumber="97" rowspan="1">1</td>
<td id="6C28441772462B060731AC1A7F9E67D0" colspan="1" pageId="0" pageNumber="97" rowspan="1">1</td>
</tr>
<tr id="080D006A537CD149C4A17076535FFC5A" pageId="0" pageNumber="97">
<td id="1EB07CA773B97EAA9A6A025613CCCBB9" colspan="1" pageId="0" pageNumber="97" rowspan="1">Postoculars</td>
<td id="0B1607EFEFB85A60CC151926E6DAB2FD" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="73E79E087279063A3F8D7EBDBE9B826B" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="0C7C8AB29C9A3D402916416E791B9845" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="051B9EED1D992388772A9A7815213A32" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="E079DE61D3FFABE096A9769250037290" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="2B1E743019D7A34B1206CB8A9502D63D" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="DF698FCCFAFA456D803A8B3102D9DE9C" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
<td id="172C83BC7305B84832BC1113707D72E7" colspan="1" pageId="0" pageNumber="97" rowspan="1">2</td>
</tr>
<tr id="0A2B2152F908711313CE5DABC633D30F" pageId="0" pageNumber="97">
<td id="3F01D5C08F454EEA30BE4F1DB47855BC" colspan="1" pageId="0" pageNumber="97" rowspan="1">Temporals</td>
<td id="42227E20458B2BAC7E154C9DCE685231" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="2D0C413D9D04BE507DB6870A51961E27" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="C5D9FF4B8485ED6D9164067771893ED9" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="05C310DEB4B41BD6EE52398FD14FCBDF" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="634E1FB40B968C80588BA121F1D78991" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="5325DD362753B60907717F3FC0097004" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="10E33BC7E846419A569826B119C32AEC" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2</td>
<td id="E755993BB2AE7C0F30B6A4B5153B9DEB" colspan="1" pageId="0" pageNumber="97" rowspan="1">1+2+3</td>
</tr>
<tr id="DDEBFA0B4089337C7479043F126DA010" pageId="0" pageNumber="97">
<td id="E7620E299472568EED765747228DF0A4" colspan="1" pageId="0" pageNumber="97" rowspan="1">Supralabials (contacting eye)</td>
<td id="6BE53986268F2E6CDD8107C01DACDE74" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="4F2C43ACB4F0B6F87D1BAF55D3023A36" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="6152A2FA38D16CA84BD8DA021C96F3D2" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="7F71916862BAE25783D9A52843243186" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="873639EBA76894C085C421B20B8FC74E" colspan="1" pageId="0" pageNumber="97" rowspan="1">5 (2,3)/6 (3,4)</td>
<td id="A6FE56CA57F3BC859CD2BD64FB31B3FA" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="82388A8777A552F20CE1BC7ADCD8B0D2" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
<td id="18049636026D18E0F69DFA14544FEFCD" colspan="1" pageId="0" pageNumber="97" rowspan="1">6 (3,4)</td>
</tr>
<tr id="586951541EEE326D4035BB345AA9DB99" pageId="0" pageNumber="97">
<td id="DA8BF0844786D5655B2B5D3FD1C80C17" colspan="1" pageId="0" pageNumber="97" rowspan="1">Infralabials (in contact with 1st chin shield)</td>
<td id="34AFB9BFCDDD9B93722D0E32F9C6A8F4" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="0D14BDD83CC9455344178A1F773F238A" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="18D2907404A61C29B37A0E91C59E883D" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="E45640C18E5AB02BDC5651E164F3A6E2" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="6F2C069349BF5E7B53D751AFCEEEF905" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="5B2330B98C1AEA36104D38FE43047C09" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="AD05E2AF1BCF747859AE73A7C5E5AC72" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
<td id="E37018F316994B760D4A422444B76EEE" colspan="1" pageId="0" pageNumber="97" rowspan="1">7 (3)</td>
</tr>
<tr id="76853CAE77D95BBC71A1C5E8A8CEFACE" pageId="0" pageNumber="97">
<td id="A97F487DE4CA8A10D15F8BBCCCC5C915" colspan="1" pageId="0" pageNumber="97" rowspan="1">Loreal</td>
<td id="F55B21718CB6DF9416FE0EE2B03E2251" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="3A3350B6DB743D21ACA48CF6FBA724D4" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="52F5BB5674CA3DF1F610C7BFE1C50418" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="9D814F1B6F750D15D4DEFF77B0EDD149" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="8770DC8B0F033454C9548539F2245456" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="36097F664BB7D6666C86C293C8935C00" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="EE1C083C501859AA3295787C600672B6" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
<td id="F6D3D2FDB3FB43D29E38451B9D5F1553" colspan="1" pageId="0" pageNumber="97" rowspan="1">Yes</td>
</tr>
<tr id="50AB578BA510EA1C50EF3524F03E5FF1" pageId="0" pageNumber="97">
<td id="C02C8ABD5D59A50CAD8142C2193EA477" colspan="1" pageId="0" pageNumber="97" rowspan="1">Dorsal colouration</td>
<td id="5A04D761400569D614AB2947F4888596" colspan="1" pageId="0" pageNumber="97" rowspan="1">26 fused blotches</td>
<td id="F4AD1013B59106D51E16E217DCA80044" colspan="1" pageId="0" pageNumber="97" rowspan="1">30 fused blotches</td>
<td id="A1B93C156838C3C1920B749805D1450B" colspan="1" pageId="0" pageNumber="97" rowspan="1">21 fused blotches</td>
<td id="97658A6BD212D192526E800D460B61A7" colspan="1" pageId="0" pageNumber="97" rowspan="1">31 fused blotches</td>
<td id="C12107E6265CC92DD3F70BA11AFC82C0" colspan="1" pageId="0" pageNumber="97" rowspan="1">22 fused blotches</td>
<td id="841EEDF27FD4735542BDB61D623204D6" colspan="1" pageId="0" pageNumber="97" rowspan="1">26 fused blotches</td>
<td id="23148142E62C3777EF8454032533A294" colspan="1" pageId="0" pageNumber="97" rowspan="1">27 fused blotches</td>
<td id="92D480AF2CE9DA552D5767FB0EB6CE2A" colspan="1" pageId="0" pageNumber="97" rowspan="1">36 paired blotches</td>
</tr>
</table>
</paragraph>
<caption id="B74512EBA4DBC4E06172E5D9D77227B8" doi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97" start="Figure 11" startId="F11">
<paragraph id="7F5215EDE644A89A596FA6DF9AB9AA6C" pageId="0" pageNumber="97">
<emphasis id="328307F48F4013BABE716641993C8008" bold="true" pageId="0" pageNumber="97">Figure 11.</emphasis>
Holotype of 
<taxonomicName id="E3E681720B16C2A3DF7A92A788B37701" authorityName="Conradie &amp; Keates &amp; Baptista &amp; Lobón-Rovira" authorityYear="2022" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna lisima" order="Squamata" pageId="0" pageNumber="97" rank="species" species="lisima">
<emphasis id="6DAD67F3D8C72BBD8553704A57FDAC96" italics="true" pageId="0" pageNumber="97">Prosymna lisima</emphasis>
</taxonomicName>
sp. nov. (PEM R23512) from Cuito River source, Moxico Province, Angola 
<emphasis id="9BFD1564000C1566356EC92AAA592543" bold="true" pageId="0" pageNumber="97">A</emphasis>
dorsal and ventral full body 
<emphasis id="1C9DAA1F40280BD64B11F6612A6BAA26" bold="true" pageId="0" pageNumber="97">B</emphasis>
dorsal head 
<emphasis id="D5358ABE735701F44C5D87601B61EDAD" bold="true" pageId="0" pageNumber="97">C</emphasis>
ventral head 
<emphasis id="0D488B869E4981B7451DCDBCD4D8F7B2" bold="true" pageId="0" pageNumber="97">D</emphasis>
lateral head.
</paragraph>
</caption>
<paragraph id="A86B708E951C9C4BE4883477F1AF5118" pageId="0" pageNumber="97">
<emphasis id="C9A7A03019A8E032BDE04DBD39E0A6CA" bold="true" pageId="0" pageNumber="97">
<emphasis id="CCFAB5D67DFA393D588A61EF5004467D" italics="true" pageId="0" pageNumber="97">Colouration</emphasis>
.
</emphasis>
In life (Figs 
<figureCitation id="2642404F21098C0810FF22B187A87787" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of live P. lisima sp. nov. from eastern Angola A PEM R 23457 from Quembo River Source, Moxico Province, Angola B PEM R 23483 from Cuando River Source, Moxico Province, Angola C PEM R 23512 from Cuito Source Lake, Moxico Province, Angola D PEM R 27381 from Quembo River bridge camp, Moxico Province, Angola." figureDoi="10.3897/zookeys.1121.85693.figure5" httpUri="https://binary.pensoft.net/fig/744036" pageId="0" pageNumber="97">5C</figureCitation>
, 
<figureCitation id="9AF97FF50BF3A9EED63501F0FDFEF150" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11B-D</figureCitation>
). Dorsum bright yellow-brown with 27 irregular fused black blotches that extend along the back from the nape onto the tail. Each dorsal scale has a darker edge giving it a faint reticulated pattern. Dorsolaterally, between the black vertebral blotches, there is a cluster of 2-4 scales with black edges. The large black nape blotch originates at the posterior margin of the frontal and runs through the parietal onto the dorsal scales, and is approximately nine scale rows deep and eleven scale rows wide. Each vertebral black blotch varies from 4-8 scale rows deep and 3-7 scale row wide. Some of the blotches are fused to form a continuous zig-zag pattern. Frontal and prefrontal sutures have a dark edge forming a pale grey crossbar. Eyes black. Ventrum cream-white, with the two outermost dorsal scale rows same colour as ventrum. In preservative (Fig. 
<figureCitation id="BAA96A43AE642D91E16DF2210A86982E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Prosymna lisima sp. nov. (PEM R 23512) from Cuito River source, Moxico Province, Angola A dorsal and ventral full body B dorsal head C ventral head D lateral head." figureDoi="10.3897/zookeys.1121.85693.figure11" httpUri="https://binary.pensoft.net/fig/744042" pageId="0" pageNumber="97">11A</figureCitation>
). Same as in life, but yellow-brown colouration faded and the dark edges became more noticeable. Ventrum beige.
</paragraph>
<paragraph id="6FF691B838E2EB64A077023161A938AE" pageId="0" pageNumber="97">
<emphasis id="0299D2245C24E4E92CE15CDFCF7C309D" bold="true" pageId="0" pageNumber="97">
<emphasis id="E8C8CAB044BB1885D4A3B0764516D44B" italics="true" pageId="0" pageNumber="97">Paratype and additional material variation</emphasis>
.
</emphasis>
See Table 
<tableCitation id="54C1403982A51B0F6B33E92D0F2FE5EE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of morphological features and measurements for Prosymna angolensis group. For abbreviations see Materials and methods. Notes: * including data on the Ebanga (Monard 1937) and Capelongo (Bogert 1940) specimens." httpUri="http://table.plazi.org/id/CE661A2298586FEF7FBADB3D50769868" pageId="0" pageNumber="97" tableUuid="CE661A2298586FEF7FBADB3D50769868">2</tableCitation>
and 
<tableCitation id="81907BEC2C9123965420759C70F9A40C" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological features and measurements for the type series of Prosymna lisima sp. nov. (SVL = snout-vent length, t = truncated)." httpUri="http://table.plazi.org/id/2ED70644E9B2B616C3BE33FAE3A24FD4" pageId="0" pageNumber="97" tableUuid="2ED70644E9B2B616C3BE33FAE3A24FD4">4</tableCitation>
for full meristic data. Dorsal scales smooth and in 17-17-15 rows at midbody; 116-124 (116-124 males, 117-129 females) smooth ventral scales; 18-26 (22-26 males, 18-24 females) paired subcaudal scales; one (rarely two) preoculars; two postoculars; temporals mostly 1+2; mostly six supralabials, with 3rd and 4th entering the orbit; seven infralabials, with first three in contact with the chin shield, cloacal scale entire; 21-36 fused dark dorsal spots. Largest female: 275+28 mm (NMZ UM 21272: 15 km WSW of Katima Mulilo); largest male: 198+25 mm (PEM R23458: Quembo River source). The colouration of the type material is in general in agreement with the holotype, except that the dorsal fused blotches vary in size, number and arrangement (Fig. 
<figureCitation id="60379B27F6FB14F27A9901FC8A65CB51" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of live P. lisima sp. nov. from eastern Angola A PEM R 23457 from Quembo River Source, Moxico Province, Angola B PEM R 23483 from Cuando River Source, Moxico Province, Angola C PEM R 23512 from Cuito Source Lake, Moxico Province, Angola D PEM R 27381 from Quembo River bridge camp, Moxico Province, Angola." figureDoi="10.3897/zookeys.1121.85693.figure5" httpUri="https://binary.pensoft.net/fig/744036" pageId="0" pageNumber="97">5A-C</figureCitation>
). The nape black blotch always originates at the anterior part of the frontal extending through the parietals to 7-9 dorsal scale rows deep, 11-15 scales wide and start from the 3rd-5th lateral dorsal scale row. The dorsum consists of 21-36 confluent black blotches that are 7-11 scales wide and three to four scales deep. One specimen (PEM R23456) exhibits a distinct dark interorbital band and internasal band. The only juvenile collected (PEM R27381, Fig. 
<figureCitation id="6CD78EFF27581C20B379B44165476754" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of live P. lisima sp. nov. from eastern Angola A PEM R 23457 from Quembo River Source, Moxico Province, Angola B PEM R 23483 from Cuando River Source, Moxico Province, Angola C PEM R 23512 from Cuito Source Lake, Moxico Province, Angola D PEM R 27381 from Quembo River bridge camp, Moxico Province, Angola." figureDoi="10.3897/zookeys.1121.85693.figure5" httpUri="https://binary.pensoft.net/fig/744036" pageId="0" pageNumber="97">5C</figureCitation>
) has small paired black blotches (two scales deep and four scales wide) on a lighter yellow ground colour, large head blotch starts at posterior frontal through parietals, seven scales wide.
</paragraph>
<paragraph id="21833B0D43FAAD7A59CB78304107750E" pageId="0" pageNumber="97">
<emphasis id="B2D1BC56217106524FD1D7FAB8D92911" bold="true" pageId="0" pageNumber="97">
<emphasis id="805C62627682DB7FF109330C2CA422CD" italics="true" pageId="0" pageNumber="97">Skull osteology and teeth</emphasis>
(Figs 
<figureCitation id="E9C84DEB6FFC88D6EBBCE21EE914F56A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Lateral B ventral C medial D dorsal views of skulls of Prosymna angolensis subgroup. Red arrows depict variation characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure7" httpUri="https://binary.pensoft.net/fig/744038" pageId="0" pageNumber="97">7</figureCitation>
- 
<figureCitation id="06FDF9B0D60FE8B76B08FCA2E77E3600" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Dorsal, frontal and ventral view of premaxillae B medial, posterior and dorsal views of palatine; and C lateral, dorsal, and ventral view of maxilla in P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis and P. janii (from left to right)." figureDoi="10.3897/zookeys.1121.85693.figure9" httpUri="https://binary.pensoft.net/fig/744040" pageId="0" pageNumber="97">9</figureCitation>
).
</emphasis>
This species presents a compact and rigid skull, common among 
<taxonomicName id="6426EB2022907E98DEBF975C6917698F" authorityName="Gray" authorityYear="1849" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna" order="Squamata" pageId="0" pageNumber="97" rank="genus">
<emphasis id="18A52B4934E4F613747E0127A296AE34" italics="true" pageId="0" pageNumber="97">Prosymna</emphasis>
</taxonomicName>
species with unfused braincase and nasal bones. Parietals are fused. Postorbital bone is present and contributes to the posterior edge of the orbital rim. Premaxilla has a short but robust ascending nasal process that lies between the ventral laminae of the nasals with low profile of the anterior portion which gradually slope ending in a moderate narrow tip and two elongated maxillary processes. Maxilla and premaxilla are in contact. Nasal bones are medium large bones in contact with frontal and premaxilla. Septomaxilla is a well-developed bone, in broad contact with the premaxilla, frontal, vomer, prefrontal and frontal bones. Vomer is well developed with a perforated dorsolateral portion. Maxillary is reduced anteriorly with an elongated pick-shaped palatine process with five or six laterally reduced curved tooth loci, followed by four to five enlarged lancet-shaped tooth loci, on same disposition. Palatine with four to five well developed teeth and an enlarged dorsal curved vomerine process. Pterygoid is a thin elongated bone. Supratemporal is in broad contact with the quadrate and participates in the lateral movement of the lower jaw. The lower jaw presents a compound bone, splenial, coronoid and dentary. Coronoid and splenial are reduced, almost vestigial. Dentary with eight tooth loci.
</paragraph>
<paragraph id="A8BBBADE307DEE2E4B1C899E413F7FB0" pageId="0" pageNumber="97">
<emphasis id="EB1CECA7DF5254F8D92E97CBDB1D5230" bold="true" pageId="0" pageNumber="97">
<taxonomicName id="2890A6356C1ABE0CDDACF87C3FB4B704" genus="Hemipenis" lsidName="Hemipenis" pageId="0" pageNumber="97" rank="genus">
<emphasis id="720E3949FC3357E0259B4C76FAB05AF6" italics="true" pageId="0" pageNumber="97">Hemipenis</emphasis>
</taxonomicName>
.
</emphasis>
Short simple structure, only reaching the 6-9th ventral scale. Single non-bifurcated sulcus. Ornamentation is flounced. Proximal third is smooth. Distal portion with four to five flounces that starts at the sulcal fold and encircle the whole organ, the most proximal often branched, forming a pocket of which the edges is smooth, tapering into a distal point. Retractor muscle is straight.
</paragraph>
</subSubSection>
<subSubSection id="0DAFBFF96B43636F5558090EE92B4B4E" pageId="0" pageNumber="97" type="natural history">
<paragraph id="046E0ABA45011E79EF6C5B210EE5F9C6" pageId="0" pageNumber="97">Natural history notes.</paragraph>
<paragraph id="03A61DEC9CA5DC549D21388133CE802C" pageId="0" pageNumber="97">
All specimens were caught in late November during the rainy season. At this time, many adult lacertids, 
<taxonomicName id="E07D658B5D6ABBE0E446E55AC47D33B0" baseAuthorityName="Smith" baseAuthorityYear="1838" class="Reptilia" family="Lacertidae" genus="Ichnotropis" kingdom="Animalia" lsidName="Ichnotropis capensis" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="capensis">
<emphasis id="C7A82CFD6A4F1274FA3EA9B7FA14C69D" italics="true" pageId="0" pageNumber="97">Ichnotropis capensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E66FC91ED6E8EF5A8FBC3E89E91CDB6E" lsidName="I. grandiceps" pageId="0" pageNumber="97" rank="species" species="grandiceps">I. cf. grandiceps</taxonomicName>
, were also observed mating in the same habitat. 
<taxonomicName id="6D271DF2B177A7281DFFC045891FF2B9" authorityName="Gray" authorityYear="1849" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna" order="Squamata" pageId="0" pageNumber="97" rank="genus">
<emphasis id="E5AAA4DD98E43A7DEE367D45243EF93B" italics="true" pageId="0" pageNumber="97">Prosymna</emphasis>
</taxonomicName>
are well known to prey on soft-shell lizard eggs, and 
<taxonomicName id="6670A0C89DCD080A77B82E60676ED3E9" lsidName="P. lisima" pageId="0" pageNumber="97" rank="species" species="lisima">
<emphasis id="D01F3F30F9F1C3D4FFCAD118AA716E18" italics="true" pageId="0" pageNumber="97">P. lisima</emphasis>
</taxonomicName>
sp. nov. may actively seek out these 
<normalizedToken id="9EB34DB71254D5CA4E6EA20B3E699FF0" originalValue="lacertids’">lacertids'</normalizedToken>
eggs. Only two of the females had stomach contents, while all the males had empty stomachs. The largest female (PEM R23456) had three empty lizard egg shells in the hind gut, three empty egg shells at the rear end of the stomach, and four undigested lizard eggs in the main stomach (Fig. 
<figureCitation id="153FC89D89E229522608D7B9647784BF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Stomach and gut contents of Prosymna lisima sp. nov. (PEM R 23456) consisting of lizard egg shells. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1121.85693.figure12" httpUri="https://binary.pensoft.net/fig/744043" pageId="0" pageNumber="97">12</figureCitation>
). Another female (PEM R23511) had four undigested lizard eggs in the main stomach. All eggs measured ~ 11.0 mm in length and each had a lateral cut. The eggs in the main stomach also all had a lateral cut but still maintained their shape and were surrounded by calcified leaked yolk (due to the preservation process). The eggs at the rear and the hindgut were all undecomposed and compressed. One of the paratype females (PEM R23456) was gravid, and had three eggs in early developmental stages (16.8 
<normalizedToken id="AEAB4EBB768E12DB17557B3D0550DC3D" originalValue="×">x</normalizedToken>
4.2 mm). Interestingly, on two different occasions, three specimens (two males and one female) were caught on the same night in the same trap array. This may indicate that males were following females to breed.
</paragraph>
<caption id="561A54204E4FEFC254B40B4B6B82533E" doi="10.3897/zookeys.1121.85693.figure12" httpUri="https://binary.pensoft.net/fig/744043" pageId="0" pageNumber="97" start="Figure 12" startId="F12">
<paragraph id="6A6371563F010129C475A9CFD08B7912" pageId="0" pageNumber="97">
<emphasis id="F2ACD5C65D89ED9C1A391643188A96C5" bold="true" pageId="0" pageNumber="97">Figure 12.</emphasis>
Stomach and gut contents of 
<taxonomicName id="D3AAEBC5D68C6D6FE0EFD780AF464610" authorityName="Conradie &amp; Keates &amp; Baptista &amp; Lobón-Rovira" authorityYear="2022" class="Reptilia" family="Prosymnidae" genus="Prosymna" kingdom="Animalia" lsidName="Prosymna lisima" order="Squamata" pageId="0" pageNumber="97" rank="species" species="lisima">
<emphasis id="6806BAF14AECDF34FAB638B892C38CA6" italics="true" pageId="0" pageNumber="97">Prosymna lisima</emphasis>
</taxonomicName>
sp. nov. (PEM R23456) consisting of lizard egg shells. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4BE2C33F7AB0A02BCACDE041D8EAA6AE" pageId="0" pageNumber="97" type="distribution">
<paragraph id="0DFDC7EF24F739404515F4D8361DAB08" pageId="0" pageNumber="97">Distribution and habitat.</paragraph>
<paragraph id="636660E8762000B4D471539432C3F136" pageId="0" pageNumber="97">
Currently only known from east-central Angola, western Zambia and the Zambezi Region of north-eastern Namibia. In the region of Katima Mulilo (eastern Zambezi Region, Namibia) it occurs in sympatry with its sister species, 
<taxonomicName id="AC748861AFCEFF94FE79BE5662A87055" lsidName="P. angolensis" pageId="0" pageNumber="97" rank="species" species="angolensis">
<emphasis id="5F7844FF71F6DDA53C028C48F67B8F3E" italics="true" pageId="0" pageNumber="97">P. angolensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="A67E74DEB49AA41DE8E9294381264C55" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a Maximum likelihood (IQTREE) concatenated phylogeny with BI BEAST and BI MrBayes support overlain. Each of the species groups are demarcated by coloured vertical bars at the right margin b maximum likelihood (IQTREE) phylogenies (16 S and cyt-b) with gene-specific single-locus species delimitation analyses results overlain. The bars to the right of the phylogeny represent the putative taxa assignments for each analysis and the values beneath the bars denote the total number of putative taxa for each analysis." figureDoi="10.3897/zookeys.1121.85693.figure2" httpUri="https://binary.pensoft.net/fig/744033" pageId="0" pageNumber="97">2</figureCitation>
). The tentative assigned material from Kuvango (
<bibRefCitation id="4259DD96C17E2E9DC7F7490E7292B179" author="Monard, A" journalOrPublisher="Bulletin de la Societe Neuchateloise des sciences Naturelles" pageId="0" pageNumber="97" pagination="89 - 111" refId="B78" refString="Monard, A, 1931. Mission scientifique Suisse dans l'Angola. Resultats scientifiques. Reptiles. Bulletin de la Societe Neuchateloise des sciences Naturelles 33: 89 - 111" title="Mission scientifique Suisse dans l'Angola. Resultats scientifiques. Reptiles." volume="33" year="1931">Monard 1931</bibRefCitation>
) needs verification, but it agrees in colouration and morphology to the new species. This species is expected to be much more widely distributed in the Kalahari basin, as it seems to be associated with the deep Kalahari sands. The Angolan material occurs in Angolan moist miombo woodland, while the Zambian and Namibian material occurs in dry miombo woodland. The elevation ranges between 950 and 1450 m a.s.l. All newly collected specimens were captured in trap arrays set in sandy areas next to river source lakes or main rivers in eastern Angola (
<bibRefCitation id="696BB01302ECEAB5B71CA323C0AB88A2" author="Conradie, W" journalOrPublisher="African Journal of Herpetology" pageId="0" pageNumber="97" refId="B38" refString="Conradie, W, Baptista, NL, Verburgt, L, Keates, C, Harvey, J, Julio, T, Neef, G, 2021. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian &amp; Reptile Conservation 15(2)[General Section]: 244-278[e292]." title="Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian &amp; Reptile Conservation 15 (2) [General Section]: 244 - 278 [e 292]." year="2021">Conradie et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>