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<document id="CA97945389668AD9500DC8E414B8CEEF" ID-CLB-Dataset="51458" ID-DOI="10.5281/zenodo.155781" ID-GBIF-Dataset="9f9ad059-7f30-42dd-91d3-324ef0ba9c24" ID-ISSN="11755326" ID-Zenodo-Dep="155781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459432365550" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="C419FA02FFBA6270CB19A0FEB130FE52" docLanguage="en" docName="zt00034.pdf" docOrigin="Zootaxa 34" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Microlia Casey 1910" docType="treatment" docVersion="15" lastPageNumber="7" masterDocId="3820827AFFBB6277CA11A34FB41BFFFC" masterDocTitle="A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="2" updateTime="1698192266583" updateUser="ExternalLinkService">
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<mods:title id="91C3021E6935D1E3C1FA3DBE2F02EFDE">A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini)</mods:title>
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<taxonomicName id="8BB03097FFBA6276CB19A0FEB610FC37" ID-CoL="8VZDB" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[264,523,945,971]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A0FEB56BFC37" bold="true" box="[264,368,945,971]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
<bibRefCitation id="282136E5FFBA6276CB69A0FEB610FC37" author="Casey" box="[376,523,945,971]" pageId="1" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
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(
<figureCitation id="D48B5791FFBA6276C80BA0FEB68EFC37" box="[538,661,945,971]" captionStart-0="FIGURES 1  5" captionStart-1="FIGURES 6  12" captionStart-10="FIGURES 62  67" captionStart-11="FIGURES 68  76" captionStart-12="FIGURES 77  79" captionStart-13="FIGURE 80" captionStart-2="FIGURES 13  20" captionStart-3="FIGURES 21  25" captionStart-4="FIGURES 26  33" captionStart-5="FIGURES 34  37" captionStart-6="FIGURES 38  44" captionStart-7="FIGURES 45  48" captionStart-8="FIGURES 49  52" captionStart-9="FIGURES 53  61" captionStartId-0="2.[264,383,1788,1812]" captionStartId-1="3.[264,383,1352,1376]" captionStartId-10="19.[264,383,1428,1452]" captionStartId-11="20.[264,383,1277,1301]" captionStartId-12="21.[264,382,1006,1030]" captionStartId-13="22.[264,368,1184,1208]" captionStartId-2="5.[264,383,1693,1717]" captionStartId-3="9.[264,383,1766,1790]" captionStartId-4="10.[264,383,1674,1698]" captionStartId-5="12.[264,383,1107,1131]" captionStartId-6="13.[264,383,1768,1792]" captionStartId-7="15.[264,383,1232,1256]" captionStartId-8="16.[264,382,1284,1308]" captionStartId-9="17.[264,383,1492,1516]" captionTargetBox-0="[332,1234,835,1756]" captionTargetBox-10="[360,1260,320,1405]" captionTargetBox-11="[351,1226,513,1257]" captionTargetBox-12="[400,1162,317,972]" captionTargetBox-13="[360,1265,358,1173]" captionTargetBox-3="[324,1235,612,1747]" captionTargetBox-4="[363,1225,458,1639]" captionTargetBox-5="[339,1231,452,1085]" captionTargetBox-6="[362,1223,685,1753]" captionTargetBox-7="[335,1229,335,1210]" captionTargetBox-8="[302,1286,479,1262]" captionTargetBox-9="[350,1231,590,1462]" captionTargetId-0="figure@2.[329,1237,830,1756]" captionTargetId-10="figure@19.[354,1262,320,1409]" captionTargetId-11="figure@20.[348,1227,513,1257]" captionTargetId-12="figure@21.[400,1163,315,974]" captionTargetId-13="figure@22.[358,1266,353,1174]" captionTargetId-3="figure@9.[320,1236,612,1749]" captionTargetId-4="figure@10.[359,1226,452,1639]" captionTargetId-5="figure@12.[337,1231,452,1086]" captionTargetId-6="figure@13.[358,1230,683,1753]" captionTargetId-7="figure@15.[329,1237,329,1216]" captionTargetId-8="figure@16.[296,1290,474,1264]" captionTargetId-9="figure@17.[350,1231,585,1464]" captionTargetPageId-0="2" captionTargetPageId-10="19" captionTargetPageId-11="20" captionTargetPageId-12="21" captionTargetPageId-13="22" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="15" captionTargetPageId-8="16" captionTargetPageId-9="17" captionText-0="FIGURES 1  5. Mouthparts of Microlia meticola (Casey) (Arizona). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." captionText-10="FIGURES 62  67. Details of Microlia tetramera Gusarov, sp. nov. (62  63 – holotype; 64  67 – possible female of M. tetramera from Costa Rica, 64  65 – type A, 66  67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." captionText-11="FIGURES 68  76. Details of Microlia tetramera Gusarov, sp. nov. (68  72, 75  76 – holotype; 73  74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73  74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73  74), 0.1 mm (69, 71  72, 75), 0.55 mm (76)." captionText-12="FIGURES 77  79. Details of Microlia panamensis Gusarov, sp. nov. (holotype). 77 – female sternum 8; 78 – female tergum 8; 79 – spermatheca. Scale bar 0.2 mm." captionText-13="FIGURE 80. Geographical distribution of Microlia Casey. 1 – M. silacea (Erichson); 2 – M. meticola (Casey); 3 – M. azteca (Casey); 4 – M. pentamera Gusarov, sp. nov.; 5 – M. tetramera Gusarov, sp. nov.; 6 – M. panamensis Gusarov, sp. nov." captionText-2="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." captionText-3="FIGURES 21  25. Abdominal segment 8 and female accessory sclerites of Microlia silacea (Erichson) (21 – paralectotype of Homalota silacea; 22, 25 – North Carolina; 23  24 – lectotype of Homalota silacea). 21 – male sternum 8; 22 – male tergum 8; 23 – female sternum 8; 24 – female tergum 8; 25 – female accessory sclerites. Scale bar 0.25 mm (21  24), 0.1 mm (25)." captionText-4="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." captionText-5="FIGURES 34  37. Details of Microlia meticola (Casey) (Arizona). 34 – male sternum 8; 35 – male tergum 8; 36 – female sternum 8; 37 – female tergum 8. Scale bar 0.4 mm." captionText-6="FIGURES 38  44. Details of Microlia meticola (Casey) (Arizona). 38 – median lobe of aedeagus, lateral view; 39 – apex of median lobe of aedeagus, lateral view; 40 – median lobe of aedeagus, ventral view; 41 – apex of median lobe of aedeagus, ventral view; 42  43 – spermatheca; 44 – apical lobe of right paramere, internal view. Scale bar 0.2 mm (38, 40, 42  43), 0.1 mm (39, 41, 44)." captionText-7="FIGURES 45  48. Details of Microlia azteca (Casey) (paralectotype). 45 – female sternum 8; 46 – female tergum 8; 47 – spermatheca; 48 – right antenna. Scale bar 0.2 mm." captionText-8="FIGURES 49  52. Details of Microlia pentamera Gusarov, sp. nov. (49  50 – holotype; 51  52 – paratype). 49 – male sternum 8; 50 – male tergum 8; 51 – female sternum 8; 52 – female tergum 8. Scale bar 0.2 mm." captionText-9="FIGURES 53  61. Details of Microlia pentamera Gusarov, sp. nov. (53  58, 61 – holotype; 59  60 – paratypes). 53 – median lobe of aedeagus, lateral view; 54 – apex of median lobe of aedeagus, lateral view; 55 – median lobe of aedeagus, ventral view; 56 – apex of median lobe of aedeagus, ventral view; 57 – copulatory piece of internal sac; 58 – apical lobe of right paramere, internal view; 59  60 – spermatheca; 61 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (53, 55, 59  60), 0.1 mm (54, 56  58), 0.55 mm (61)." httpUri-0="https://zenodo.org/record/155782/files/figure.png" httpUri-10="https://zenodo.org/record/155790/files/figure.png" httpUri-11="https://zenodo.org/record/155791/files/figure.png" httpUri-12="https://zenodo.org/record/155792/files/figure.png" httpUri-13="https://zenodo.org/record/155793/files/figure.png" httpUri-3="https://zenodo.org/record/155783/files/figure.png" httpUri-4="https://zenodo.org/record/155784/files/figure.png" httpUri-5="https://zenodo.org/record/155785/files/figure.png" httpUri-6="https://zenodo.org/record/155786/files/figure.png" httpUri-7="https://zenodo.org/record/155787/files/figure.png" httpUri-8="https://zenodo.org/record/155788/files/figure.png" httpUri-9="https://zenodo.org/record/155789/files/figure.png" pageId="1" pageNumber="2">Figs. 180</figureCitation>
)
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<taxonomicName id="8BB03097FFBA6276CB19A0B6B571FBEC" ID-CoL="8XKZB" box="[264,362,1017,1040]" class="Insecta" family="Staphylinidae" genus="Dolosota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A0B6B571FBEC" box="[264,362,1017,1040]" italics="true" pageId="1" pageNumber="2">Dolosota</emphasis>
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(
<taxonomicName id="8BB03097FFBA6276CB68A0B6B67FFBEC" ID-CoL="8VZDB" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[377,612,1017,1040]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB68A0B6B5CEFBEC" box="[377,469,1017,1040]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
<bibRefCitation id="282136E5FFBA6276CBCCA0B6B67FFBEC" author="Casey" box="[477,612,1017,1040]" pageId="1" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
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): 144 (subtribe
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Athetina
<bibRefCitation id="282136E5FFBA6276C962A0B6B7E2FBEC" author="Casey" box="[883,1017,1017,1040]" pageId="1" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
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).
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<emphasis id="7EC49706FFBA6276CB19A750B54DFBCA" box="[264,342,1055,1078]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
<treatmentCitation id="CD116D05FFBA6276CB4FA750B600FBCA" author="Casey" box="[350,539,1055,1078]" page="145" pageId="1" pageNumber="2" year="1911">
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: 145
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(tribe
<taxonomicName id="8BB03097FFBA6276C873A750B7B5FBCA" authority="Thomson, 1867" authorityName="Thomson" authorityYear="1867" box="[610,942,1055,1078]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Myrmedoniini">Myrmedoniini Thomson, 1867</taxonomicName>
),
<emphasis id="7EC49706FFBA6276C9D0A751B03FFBCA" bold="true" box="[961,1060,1054,1078]" pageId="1" pageNumber="2">syn. nov.</emphasis>
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<taxonomicName id="8BB03097FFBA6276CB19A70BB639FBA7" ID-CoL="8XN8C" authority="Leng, 1920: 122" authorityName="Leng" authorityPageNumber="122" authorityYear="1920" box="[264,546,1092,1115]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A70BB54DFBA7" box="[264,342,1092,1115]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA70BB639FBA7" author="Leng" box="[364,546,1092,1115]" page="122" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="282136E5FFBA6276CB7DA70BB5F2FBA7" author="Leng" box="[364,489,1092,1115]" pageId="1" pageNumber="24" refString="Leng, C. (1920) Catalogue of the Coleoptera of America, North of Mexico. Mt. Vernon, New York: J. D. Sherman, Jr., x + 470 pp." type="book" year="1920">Leng, 1920</bibRefCitation>
: 122
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</taxonomicName>
(as valid genus; tribe
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).
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<taxonomicName id="8BB03097FFBA6276CB19A726B5C3FB7C" ID-CoL="8XNJR" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,472,1129,1152]" class="Insecta" family="Staphylinidae" genus="Pancota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A726B57AFB7C" box="[264,353,1129,1152]" italics="true" pageId="1" pageNumber="2">Pancota</emphasis>
(
<emphasis id="7EC49706FFBA6276CB60A726B5D6FB7C" box="[369,461,1129,1152]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276CBFAA726B6BAFB7C" author="Leng" box="[491,673,1129,1152]" page="122" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="282136E5FFBA6276CBFAA726B673FB7C" author="Leng" box="[491,616,1129,1152]" pageId="1" pageNumber="24" refString="Leng, C. (1920) Catalogue of the Coleoptera of America, North of Mexico. Mt. Vernon, New York: J. D. Sherman, Jr., x + 470 pp." type="book" year="1920">Leng, 1920</bibRefCitation>
: 122
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(as valid subgenus; tribe
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).
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</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A7C0B036FB5A" box="[264,1069,1167,1190]" pageId="1" pageNumber="2" type="description">
<paragraph id="4C0F4B14FFBA6276CB19A7C0B036FB5A" blockId="1.[264,1323,1017,1712]" box="[264,1069,1167,1190]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A7C0B036FB5A" box="[264,1069,1167,1190]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A7C0B5DEFB5A" ID-CoL="8VRBL" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,453,1167,1190]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A7C0B556FB5A" box="[264,333,1167,1190]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="7EC49706FFBA6276CB4CA7C0B5A2FB5A" box="[349,441,1167,1190]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276CBC9A7C0B6B9FB5A" author="Fenyes" box="[472,674,1167,1190]" page="202" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="282136E5FFBA6276CBC9A7C0B672FB5A" author="Fenyes" box="[472,617,1167,1190]" pageId="1" pageNumber="24" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. Bruxelles: L. Desmet-Verteneuil, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 202
</treatmentCitation>
(as a synonym of
<taxonomicName id="8BB03097FFBA6276C977A7C0B7B0FB5A" box="[870,939,1167,1190]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276C977A7C0B7B0FB5A" box="[870,939,1167,1190]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
</taxonomicName>
(
<taxonomicName id="8BB03097FFBA6276C9AAA7C0B00FFB5A" box="[955,1044,1167,1190]" class="Insecta" family="Staphylinidae" genus="Pancota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276C9AAA7C0B00FFB5A" box="[955,1044,1167,1190]" italics="true" pageId="1" pageNumber="2">Pancota</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A7FBB564FA57" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="4C0F4B14FFBA6276CB19A7FBB05FFB37" blockId="1.[264,1323,1017,1712]" box="[264,1092,1204,1227]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A7FBB05FFB37" box="[264,1092,1204,1227]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A7FBB62DFB37" ID-CoL="8XN8C" authority="Fenyes, 1920: 308" authorityName="Fenyes" authorityPageNumber="308" authorityYear="1920" box="[264,566,1204,1227]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A7FBB54DFB37" box="[264,342,1204,1227]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA7FBB62DFB37" author="Fenyes" box="[364,566,1204,1227]" page="308" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="282136E5FFBA6276CB7DA7FBB5E6FB37" author="Fenyes" box="[364,509,1204,1227]" pageId="1" pageNumber="24" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. Bruxelles: L. Desmet-Verteneuil, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 308
</treatmentCitation>
</taxonomicName>
(as valid genus; tribe
<taxonomicName id="8BB03097FFBA6276C932A7FBB02CFB37" ID-CoL="8VR8M" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[803,1079,1204,1227]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">
Hoplandriini
<bibRefCitation id="282136E5FFBA6276C9A3A7FBB02CFB37" author="Casey" box="[946,1079,1204,1227]" pageId="1" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A796B0F4FB0C" blockId="1.[264,1323,1017,1712]" box="[264,1263,1241,1264]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A796B0F4FB0C" box="[264,1263,1241,1264]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A796B5DEFB0C" ID-CoL="8VRBL" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,453,1241,1264]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A796B556FB0C" box="[264,333,1241,1264]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="7EC49706FFBA6276CB4CA796B5A2FB0C" box="[349,441,1241,1264]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276CBC9A796B77FFB0C" author="Bernhauer" box="[472,868,1241,1264]" page="660" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="282136E5FFBA6276CBC9A796B730FB0C" author="Bernhauer" box="[472,811,1241,1264]" pageId="1" pageNumber="24" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 660
</treatmentCitation>
(as a synonym of
<taxonomicName id="8BB03097FFBA6276CE39A796B075FB0C" ID-CoL="8VRBL" box="[1064,1134,1241,1264]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CE39A796B075FB0C" box="[1064,1134,1241,1264]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
</taxonomicName>
(
<taxonomicName id="8BB03097FFBA6276CE6CA796B0CDFB0C" box="[1149,1238,1241,1264]" class="Insecta" family="Staphylinidae" genus="Pancota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CE6CA796B0CDFB0C" box="[1149,1238,1241,1264]" italics="true" pageId="1" pageNumber="2">Pancota</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A7B0B0BDFAEA" blockId="1.[264,1323,1017,1712]" box="[264,1190,1279,1302]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A7B0B0BDFAEA" box="[264,1190,1279,1302]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A7B0B6E3FAEA" ID-CoL="8XN8C" authority="Bernhauer &amp; Scheerpeltz, 1926: 717" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="717" authorityYear="1926" box="[264,760,1279,1302]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A7B0B54DFAEA" box="[264,342,1279,1302]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA7B0B6E3FAEA" author="Bernhauer" box="[364,760,1279,1302]" page="717" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="282136E5FFBA6276CB7DA7B0B6A4FAEA" author="Bernhauer" box="[364,703,1279,1302]" pageId="1" pageNumber="24" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 717
</treatmentCitation>
</taxonomicName>
(as valid genus; subtribe
<taxonomicName id="8BB03097FFBA6276CE18A7B0B080FAEA" ID-CoL="8VRCT" box="[1033,1179,1279,1302]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subTribe" subTribe="Hoplandriina">Hoplandriina</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A66BB755FAC7" blockId="1.[264,1323,1017,1712]" box="[264,846,1316,1339]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A66BB755FAC7" box="[264,846,1316,1339]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A66BB68CFAC7" ID-CoL="8XN8C" authority="Leng &amp; Mutchler, 1927: 23" authorityName="Leng &amp; Mutchler" authorityPageNumber="23" authorityYear="1927" box="[264,663,1316,1339]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A66BB54DFAC7" box="[264,342,1316,1339]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA66BB68CFAC7" author="Leng" box="[364,663,1316,1339]" page="23" pageId="1" pageNumber="2" year="1927">
<bibRefCitation id="282136E5FFBA6276CB7DA66BB677FAC7" author="Leng" box="[364,620,1316,1339]" pageId="1" pageNumber="24" refString="Leng, C. &amp; Mutchler, A. J. (1927) Supplement 1919 to 1924 (inclusive) to catalogue of the Coleoptera of America, North of Mexico. Mt. Vernon, New York: J. D. Sherman, Jr., 52 pp." type="book" year="1927">Leng &amp; Mutchler, 1927</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A606B733FA9C" blockId="1.[264,1323,1017,1712]" box="[264,808,1353,1376]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A606B733FA9C" box="[264,808,1353,1376]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A606B66AFA9C" ID-CoL="8XN8C" authority="Blackwelder, 1952: 262" authorityName="Blackwelder" authorityPageNumber="262" authorityYear="1952" box="[264,625,1353,1376]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A606B54DFA9C" box="[264,342,1353,1376]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA606B66AFA9C" author="Blackwelder" box="[364,625,1353,1376]" page="262" pageId="1" pageNumber="2" year="1952">
<bibRefCitation id="282136E5FFBA6276CB7DA606B623FA9C" author="Blackwelder" box="[364,568,1353,1376]" pageId="1" pageNumber="24" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
: 262
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A620B564FA57" blockId="1.[264,1323,1017,1712]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A620B564FA57" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A620B5E5FA7A" ID-CoL="8W3JB" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,510,1391,1414]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A620B59FFA7A" box="[264,388,1391,1414]" italics="true" pageId="1" pageNumber="2">Ischnopoda</emphasis>
(
<emphasis id="7EC49706FFBA6276CB86A620B5E8FA7A" box="[407,499,1391,1414]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276C806A620B733FA7A" author="Blackwelder" box="[535,808,1391,1414]" page="202" pageId="1" pageNumber="2" year="1952">
<bibRefCitation id="282136E5FFBA6276C806A620B6FDFA7A" author="Blackwelder" box="[535,742,1391,1414]" pageId="1" pageNumber="24" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
: 202
</treatmentCitation>
, 246, 288 (as a synonym of
<taxonomicName id="8BB03097FFBA6276CE79A620B0FFFA7A" box="[1128,1252,1391,1414]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CE79A620B0FFFA7A" box="[1128,1252,1391,1414]" italics="true" pageId="1" pageNumber="2">Ischnopoda</emphasis>
</taxonomicName>
(
<taxonomicName id="8BB03097FFBA6276CEE6A620B57DFA57" class="Insecta" family="Staphylinidae" genus="Pancota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CEE6A620B57DFA57" italics="true" pageId="1" pageNumber="2">Pancota</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A6F6B08EFA2C" box="[264,1173,1465,1488]" pageId="1" pageNumber="2" type="description">
<paragraph id="4C0F4B14FFBA6276CB19A6F6B08EFA2C" blockId="1.[264,1323,1017,1712]" box="[264,1173,1465,1488]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A6F6B08EFA2C" box="[264,1173,1465,1488]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A6F6B5DEFA2C" ID-CoL="8VRBL" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,453,1465,1488]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A6F6B556FA2C" box="[264,333,1465,1488]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="7EC49706FFBA6276CB4CA6F6B5A2FA2C" box="[349,441,1465,1488]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276CBC9A6F6B710FA2C" author="Moore" box="[472,779,1465,1488]" page="351" pageId="1" pageNumber="2" year="1975">
<bibRefCitation id="282136E5FFBA6276CBC9A6F6B6C9FA2C" author="Moore" box="[472,722,1465,1488]" pageId="1" pageNumber="24" refString="Moore, I. &amp; Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp." type="book" year="1975">Moore &amp; Legner, 1975</bibRefCitation>
: 351
</treatmentCitation>
(as a synonym of
<taxonomicName id="8BB03097FFBA6276C9DFA6F6B008FA2C" box="[974,1043,1465,1488]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276C9DFA6F6B008FA2C" box="[974,1043,1465,1488]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
</taxonomicName>
(
<taxonomicName id="8BB03097FFBA6276CE32A6F6B067FA2C" box="[1059,1148,1465,1488]" class="Insecta" family="Staphylinidae" genus="Pancota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CE32A6F6B067FA2C" box="[1059,1148,1465,1488]" italics="true" pageId="1" pageNumber="2">Pancota</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A690B042F99A" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="4C0F4B14FFBA6276CB19A690B74EFA0A" blockId="1.[264,1323,1017,1712]" box="[264,853,1503,1526]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A690B74EFA0A" box="[264,853,1503,1526]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A690B684FA0A" ID-CoL="8XN8C" authority="Moore &amp; Legner, 1975: 456" authorityName="Moore &amp; Legner" authorityPageNumber="456" authorityYear="1975" box="[264,671,1503,1526]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A690B54DFA0A" box="[264,342,1503,1526]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA690B684FA0A" author="Moore" box="[364,671,1503,1526]" page="456" pageId="1" pageNumber="2" year="1975">
<bibRefCitation id="282136E5FFBA6276CB7DA690B67EFA0A" author="Moore" box="[364,613,1503,1526]" pageId="1" pageNumber="24" refString="Moore, I. &amp; Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp." type="book" year="1975">Moore &amp; Legner, 1975</bibRefCitation>
: 456
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A54BB024F9E7" blockId="1.[264,1323,1017,1712]" box="[264,1087,1540,1563]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A54BB024F9E7" box="[264,1087,1540,1563]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A54BB5FAF9E7" ID-CoL="8VRFB" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,481,1540,1563]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A54BB571F9E7" box="[264,362,1540,1563]" italics="true" pageId="1" pageNumber="2">Acrotona</emphasis>
(
<emphasis id="7EC49706FFBA6276CB6BA54BB5CDF9E7" box="[378,470,1540,1563]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="CD116D05FFBA6276CBE5A54BB6DDF9E7" author="Seevers" box="[500,710,1540,1563]" page="100" pageId="1" pageNumber="2" year="1978">
<bibRefCitation id="282136E5FFBA6276CBE5A54BB695F9E7" author="Seevers" box="[500,654,1540,1563]" pageId="1" pageNumber="24" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
: 100
</treatmentCitation>
(as valid subgenus; tribe
<taxonomicName id="8BB03097FFBA6276C9C9A54BB02FF9E7" ID-CoL="8VR6Z" box="[984,1076,1540,1563]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A566B7D9F9BC" blockId="1.[264,1323,1017,1712]" box="[264,962,1577,1600]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A566B7D9F9BC" box="[264,962,1577,1600]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A566B625F9BC" ID-CoL="8XN8C" authority="Seevers, 1978: 142" authorityName="Seevers" authorityPageNumber="142" authorityYear="1978" box="[264,574,1577,1600]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A566B54DF9BC" box="[264,342,1577,1600]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA566B625F9BC" author="Seevers" box="[364,574,1577,1600]" page="142" pageId="1" pageNumber="2" year="1978">
<bibRefCitation id="282136E5FFBA6276CB7DA566B61EF9BC" author="Seevers" box="[364,517,1577,1600]" pageId="1" pageNumber="24" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
: 142
</treatmentCitation>
</taxonomicName>
(as valid genus; tribe
<taxonomicName id="8BB03097FFBA6276C93BA566B7ACF9BC" ID-CoL="8VR8M" box="[810,951,1577,1600]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">Hoplandriini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4C0F4B14FFBA6276CB19A500B042F99A" blockId="1.[264,1323,1017,1712]" box="[264,1113,1615,1638]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A500B042F99A" box="[264,1113,1615,1638]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A500B5B8F99A" ID-CoL="8XN8C" authority="Ashe" authorityName="Ashe" box="[264,419,1615,1638]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A500B54DF99A" box="[264,342,1615,1638]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
: Ashe
</taxonomicName>
in
<treatmentCitation id="CD116D05FFBA6276CBD4A500B6CFF99A" author="Newton" box="[453,724,1615,1638]" page="360" pageId="1" pageNumber="2" year="2000">
<bibRefCitation id="282136E5FFBA6276CBD4A500B680F99A" author="Newton" box="[453,667,1615,1638]" pageId="1" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter" year="2000">
Newton
<emphasis id="7EC49706FFBA6276C831A500B64EF99A" box="[544,597,1615,1638]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
, 2000
</bibRefCitation>
: 360
</treatmentCitation>
(as valid genus; tribe
<taxonomicName id="8BB03097FFBA6276C9D1A500B055F99A" ID-CoL="8VR8M" box="[960,1102,1615,1638]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">Hoplandriini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A53BB0CEF977" box="[264,1237,1652,1675]" pageId="1" pageNumber="2" type="description">
<paragraph id="4C0F4B14FFBA6276CB19A53BB0CEF977" blockId="1.[264,1323,1017,1712]" box="[264,1237,1652,1675]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A53BB0CEF977" box="[264,1237,1652,1675]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A53BB5FAF977" ID-CoL="8VRFB" authority="(Microlia)" baseAuthorityName="Microlia" box="[264,481,1652,1675]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A53BB571F977" box="[264,362,1652,1675]" italics="true" pageId="1" pageNumber="2">Acrotona</emphasis>
(
<emphasis id="7EC49706FFBA6276CB6BA53BB5CDF977" box="[378,470,1652,1675]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
)
</taxonomicName>
: Ashe in
<treatmentCitation id="CD116D05FFBA6276C85CA53BB746F977" author="Newton" box="[589,861,1652,1675]" page="368" pageId="1" pageNumber="2" year="2000">
<bibRefCitation id="282136E5FFBA6276C85CA53BB73FF977" author="Newton" box="[589,804,1652,1675]" pageId="1" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter" year="2000">
Newton
<emphasis id="7EC49706FFBA6276C8B9A53BB6C5F977" box="[680,734,1652,1675]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
, 2000
</bibRefCitation>
: 368
</treatmentCitation>
(as valid subgenus; tribe
<taxonomicName id="8BB03097FFBA6276CE7FA53BB0D0F977" box="[1134,1227,1652,1675]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6276CB19A5D6B7A0F94C" box="[264,955,1689,1712]" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="4C0F4B14FFBA6276CB19A5D6B7A0F94C" blockId="1.[264,1323,1017,1712]" box="[264,955,1689,1712]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="6CA06C3AFFBA6276CB19A5D6B7A0F94C" box="[264,955,1689,1712]" pageId="1" pageNumber="2">
<taxonomicName id="8BB03097FFBA6276CB19A5D6B62DF94C" ID-CoL="8XN8C" authority="Hanley, 2001: 221" authorityName="Hanley" authorityPageNumber="221" authorityYear="2001" box="[264,566,1689,1712]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB19A5D6B54DF94C" box="[264,342,1689,1712]" italics="true" pageId="1" pageNumber="2">Nosora</emphasis>
:
<treatmentCitation id="CD116D05FFBA6276CB7DA5D6B62DF94C" author="Hanley" box="[364,566,1689,1712]" page="221" pageId="1" pageNumber="2" year="2001">
<bibRefCitation id="282136E5FFBA6276CB7DA5D6B5E6F94C" author="Hanley" box="[364,509,1689,1712]" pageId="1" pageNumber="24" refString="Hanley, R. S. (2001) Generic revision and phylogeny of the Aleocharine beetle tribe Hoplandriini Casey 1910 (Coleoptera: Staphylinidae). Ph. D. thesis. Lawrence: University of Kansas, xxiii + 537 pp." type="book" year="2001">Hanley, 2001</bibRefCitation>
: 221
</treatmentCitation>
</taxonomicName>
(as valid genus; tribe
<taxonomicName id="8BB03097FFBA6276C933A5D6B7B4F94C" ID-CoL="8VR8M" box="[802,943,1689,1712]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">Hoplandriini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBA6275CB19A5ACB087FD8A" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="4C0F4B14FFBA6275CB19A5ACB087FD8A" blockId="1.[264,1324,1763,1870]" lastBlockId="2.[264,1324,284,750]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="7EC49706FFBA6276CB19A5ACB593F901" bold="true" box="[264,392,1763,1789]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="8BB03097FFBA6276CB88A5ABB5E4F901" box="[409,511,1764,1789]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBA6276CB88A5ABB5E4F901" box="[409,511,1764,1789]" italics="true" pageId="1" pageNumber="2">Microlia</emphasis>
</taxonomicName>
can be distinguished from other aleocharine genera by the combination of the following characters: body broadest at elytra; antennal articles 510 strongly transverse (
<figureCitation id="D48B5791FFBA6276CB80A47BB5EEF8B2" box="[401,501,1844,1870]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="1" pageNumber="2">Figs. 12</figureCitation>
,
<figureCitation id="D48B5791FFBA6276C812A47BB638F8B2" box="[515,547,1844,1870]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="1" pageNumber="2">30</figureCitation>
,
<figureCitation id="D48B5791FFBA6276C820A47BB649F8B2" box="[561,594,1844,1870]" captionStart="FIGURES 45  48" captionStartId="15.[264,383,1232,1256]" captionTargetBox="[335,1229,335,1210]" captionTargetId="figure@15.[329,1237,329,1216]" captionTargetPageId="15" captionText="FIGURES 45  48. Details of Microlia azteca (Casey) (paralectotype). 45 – female sternum 8; 46 – female tergum 8; 47 – spermatheca; 48 – right antenna. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155787/files/figure.png" pageId="1" pageNumber="2">48</figureCitation>
); labial and maxillary palpi with pseudosegment (
<figureCitation id="D48B5791FFBA6276CEB6A47BB107F8B2" box="[1191,1308,1844,1870]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="1" pageNumber="2">Figs. 6, 9</figureCitation>
); apical half of ligula divided into two lobes (
<figureCitation id="D48B5791FFB96275C90DA253B76AFECA" box="[796,881,284,310]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="2" pageNumber="3">Figs. 9</figureCitation>
); pronotum with microsetae directed posteriorly along the midline of the disc (
<typeStatus id="930BF5B6FFB96275C8FFA20BB731FEA2" box="[750,810,324,350]" pageId="2" pageNumber="3">Type</typeStatus>
V,
<bibRefCitation id="282136E5FFB96275C943A20BB04DFEA2" author="Benick" box="[850,1110,324,350]" pageId="2" pageNumber="24" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="D48B5791FFB96275CE7EA20BB0DDFEA2" box="[1135,1222,324,350]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="2" pageNumber="3">Fig. 13</figureCitation>
); pronotal macrosetae short; pronotal hypomera invisible in lateral view; medial microseta of mesotibia inconspicuous, shorter than tibial width; tarsal formula 445 or 455; metatarsal segment 1 not longer than segment 2; one empodial setae; posterior margin of male abdominal sternum 8 extended posteriorly as large triangular lobe (
<figureCitation id="D48B5791FFB96275CE41A2ABB0A3FE02" box="[1104,1208,484,510]" captionStart="FIGURES 21  25" captionStartId="9.[264,383,1766,1790]" captionTargetBox="[324,1235,612,1747]" captionTargetId="figure@9.[320,1236,612,1749]" captionTargetPageId="9" captionText="FIGURES 21  25. Abdominal segment 8 and female accessory sclerites of Microlia silacea (Erichson) (21 – paralectotype of Homalota silacea; 22, 25 – North Carolina; 23  24 – lectotype of Homalota silacea). 21 – male sternum 8; 22 – male tergum 8; 23 – female sternum 8; 24 – female tergum 8; 25 – female accessory sclerites. Scale bar 0.25 mm (21  24), 0.1 mm (25)." httpUri="https://zenodo.org/record/155783/files/figure.png" pageId="2" pageNumber="3">Figs. 21</figureCitation>
,
<figureCitation id="D48B5791FFB96275CEDBA2ABB0F1FE02" box="[1226,1258,484,510]" captionStart="FIGURES 34  37" captionStartId="12.[264,383,1107,1131]" captionTargetBox="[339,1231,452,1085]" captionTargetId="figure@12.[337,1231,452,1086]" captionTargetPageId="12" captionText="FIGURES 34  37. Details of Microlia meticola (Casey) (Arizona). 34 – male sternum 8; 35 – male tergum 8; 36 – female sternum 8; 37 – female tergum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155785/files/figure.png" pageId="2" pageNumber="3">34</figureCitation>
,
<figureCitation id="D48B5791FFB96275CEEDA2ABB106FE02" box="[1276,1309,484,510]" captionStart="FIGURES 62  67" captionStartId="19.[264,383,1428,1452]" captionTargetBox="[360,1260,320,1405]" captionTargetId="figure@19.[354,1262,320,1409]" captionTargetPageId="19" captionText="FIGURES 62  67. Details of Microlia tetramera Gusarov, sp. nov. (62  63 – holotype; 64  67 – possible female of M. tetramera from Costa Rica, 64  65 – type A, 66  67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155790/files/figure.png" pageId="2" pageNumber="3">62</figureCitation>
); median lobe of aedeagus with long and narrow apical process (
<figureCitation id="D48B5791FFB96275CE20A143B0D9FDDA" box="[1073,1218,524,550]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="2" pageNumber="3">Figs. 2629</figureCitation>
,
<figureCitation id="D48B5791FFB96275CEC4A143B105FDDA" box="[1237,1310,524,550]" captionStart="FIGURES 68  76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68  76. Details of Microlia tetramera Gusarov, sp. nov. (68  72, 75  76 – holotype; 73  74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73  74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73  74), 0.1 mm (69, 71  72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="2" pageNumber="3">6872</figureCitation>
); parameres with two long and two short macrosetae (
<figureCitation id="D48B5791FFB96275C992A17BB7FCFDB2" box="[899,999,564,590]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="2" pageNumber="3">Figs. 31</figureCitation>
,
<figureCitation id="D48B5791FFB96275C9E6A17BB003FDB2" box="[1015,1048,564,590]" captionStart="FIGURES 38  44" captionStartId="13.[264,383,1768,1792]" captionTargetBox="[362,1223,685,1753]" captionTargetId="figure@13.[358,1230,683,1753]" captionTargetPageId="13" captionText="FIGURES 38  44. Details of Microlia meticola (Casey) (Arizona). 38 – median lobe of aedeagus, lateral view; 39 – apex of median lobe of aedeagus, lateral view; 40 – median lobe of aedeagus, ventral view; 41 – apex of median lobe of aedeagus, ventral view; 42  43 – spermatheca; 44 – apical lobe of right paramere, internal view. Scale bar 0.2 mm (38, 40, 42  43), 0.1 mm (39, 41, 44)." httpUri="https://zenodo.org/record/155786/files/figure.png" pageId="2" pageNumber="3">44</figureCitation>
); spermatheca forming two to many coils (
<figureCitation id="D48B5791FFB96275CBF8A113B668FD8A" box="[489,627,604,630]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="2" pageNumber="3">Figs. 3233</figureCitation>
,
<figureCitation id="D48B5791FFB96275C86EA113B685FD8A" box="[639,670,604,630]" captionStart="FIGURES 45  48" captionStartId="15.[264,383,1232,1256]" captionTargetBox="[335,1229,335,1210]" captionTargetId="figure@15.[329,1237,329,1216]" captionTargetPageId="15" captionText="FIGURES 45  48. Details of Microlia azteca (Casey) (paralectotype). 45 – female sternum 8; 46 – female tergum 8; 47 – spermatheca; 48 – right antenna. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155787/files/figure.png" pageId="2" pageNumber="3">47</figureCitation>
) or numerous irregular loops (
<figureCitation id="D48B5791FFB96275CE15A113B094FD8A" box="[1028,1167,604,630]" captionStart="FIGURES 38  44" captionStartId="13.[264,383,1768,1792]" captionTargetBox="[362,1223,685,1753]" captionTargetId="figure@13.[358,1230,683,1753]" captionTargetPageId="13" captionText="FIGURES 38  44. Details of Microlia meticola (Casey) (Arizona). 38 – median lobe of aedeagus, lateral view; 39 – apex of median lobe of aedeagus, lateral view; 40 – median lobe of aedeagus, ventral view; 41 – apex of median lobe of aedeagus, ventral view; 42  43 – spermatheca; 44 – apical lobe of right paramere, internal view. Scale bar 0.2 mm (38, 40, 42  43), 0.1 mm (39, 41, 44)." httpUri="https://zenodo.org/record/155786/files/figure.png" pageId="2" pageNumber="3">Figs. 4243</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFB96273CB29A1CCB647FA42" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="4C0F4B14FFB96275CB29A1CCB74FFD12" blockId="2.[264,1324,284,750]" pageId="2" pageNumber="3">
<emphasis id="7EC49706FFB96275CB29A1CCB5CBFD61" bold="true" box="[312,464,643,669]" pageId="2" pageNumber="3">Description.</emphasis>
Length
<quantity id="8B48E6F1FFB96275C82AA1CBB6D2FD62" box="[571,713,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="2" pageNumber="3" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.82.3 mm</quantity>
. Body from brownish yellow to brown with darker apex of antennae and abdominal segments 46, in some species with darker elytra or head; elytra much broader than pronotum and abdomen.
</paragraph>
<caption id="18CF1B9CFFB96275CB19A5B3B592F8A5" httpUri="https://zenodo.org/record/155782/files/figure.png" pageId="2" pageNumber="3" targetBox="[332,1234,835,1756]" targetPageId="2">
<paragraph id="4C0F4B14FFB96275CB19A5B3B592F8A5" blockId="2.[264,1323,1788,1881]" pageId="2" pageNumber="3">
<emphasis id="7EC49706FFB96275CB19A5B3B5B3F8E8" bold="true" box="[264,424,1788,1812]" pageId="2" pageNumber="3">FIGURES 15</emphasis>
. Mouthparts of
<taxonomicName id="8BB03097FFB96275C848A5B2B7F9F8E8" authority="(Casey) (Arizona)" authorityName="Casey" baseAuthorityName="Casey" box="[601,994,1789,1812]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFB96275C848A5B2B70CF8E8" box="[601,791,1789,1812]" italics="true" pageId="2" pageNumber="3">Microlia meticola</emphasis>
(Casey) (Arizona)
</taxonomicName>
. 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption id="18CF1B9CFFB86274CB19A607B520FA34" pageId="3" pageNumber="4">
<paragraph id="4C0F4B14FFB86274CB19A607B520FA34" blockId="3.[264,1323,1352,1480]" pageId="3" pageNumber="4">
<emphasis id="7EC49706FFB86274CB19A607B5A3FA9C" bold="true" box="[264,440,1352,1376]" pageId="3" pageNumber="4">FIGURES 612</emphasis>
. Mouthparts and antenna of
<taxonomicName id="8BB03097FFB86274C8EBA606B093FA9C" authority="(Casey) (Arizona)" authorityName="Casey" baseAuthorityName="Casey" box="[762,1160,1353,1376]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFB86274C8EBA606B7A1FA9C" box="[762,954,1353,1376]" italics="true" pageId="3" pageNumber="4">Microlia meticola</emphasis>
(Casey) (Arizona)
</taxonomicName>
. 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (711), 0.28 mm (12).
</paragraph>
</caption>
<paragraph id="4C0F4B14FFB86273CB29A6B3B7A7FE02" blockId="3.[264,1323,1532,1878]" lastBlockId="4.[264,1324,284,1870]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Head transverse; eyes large, 23 times as long as temples; infraorbital carina complete. Antennal article 2 longer than article 3, 4 subquadrate or transverse, 510 transverse or strongly transverse (ratio 2.02.4), apical article with two coeloconic sensilla (
<figureCitation id="D48B5791FFB86274CE89A503B0E1F99A" box="[1176,1274,1612,1638]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="3" pageNumber="4">Figs. 12</figureCitation>
,
<figureCitation id="D48B5791FFB86274CF17A503B13DF99A" box="[1286,1318,1612,1638]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="3" pageNumber="4">30</figureCitation>
,
<figureCitation id="D48B5791FFB86274CB19A53BB532F972" box="[264,297,1652,1678]" captionStart="FIGURES 45  48" captionStartId="15.[264,383,1232,1256]" captionTargetBox="[335,1229,335,1210]" captionTargetId="figure@15.[329,1237,329,1216]" captionTargetPageId="15" captionText="FIGURES 45  48. Details of Microlia azteca (Casey) (paralectotype). 45 – female sternum 8; 46 – female tergum 8; 47 – spermatheca; 48 – right antenna. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155787/files/figure.png" pageId="3" pageNumber="4">48</figureCitation>
). Labrum (
<figureCitation id="D48B5791FFB86274CBA0A53BB5EEF972" box="[433,501,1652,1678]" captionStart="FIGURES 1  5" captionStartId="2.[264,383,1788,1812]" captionTargetBox="[332,1234,835,1756]" captionTargetId="figure@2.[329,1237,830,1756]" captionTargetPageId="2" captionText="FIGURES 1  5. Mouthparts of Microlia meticola (Casey) (Arizona). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155782/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
) transverse, anterior margin with four shallow emarginations. Adoral surface of labrum (epipharynx) without transverse row of pores (
<figureCitation id="D48B5791FFB86274C9EEA5D3B05CF94A" box="[1023,1095,1692,1718]" captionStart="FIGURES 1  5" captionStartId="2.[264,383,1788,1812]" captionTargetBox="[332,1234,835,1756]" captionTargetId="figure@2.[329,1237,830,1756]" captionTargetPageId="2" captionText="FIGURES 1  5. Mouthparts of Microlia meticola (Casey) (Arizona). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155782/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
). Mandibles (
<figureCitation id="D48B5791FFB86274CEE1A5D3B534F922" captionStart="FIGURES 1  5" captionStartId="2.[264,383,1788,1812]" captionTargetBox="[332,1234,835,1756]" captionTargetId="figure@2.[329,1237,830,1756]" captionTargetPageId="2" captionText="FIGURES 1  5. Mouthparts of Microlia meticola (Casey) (Arizona). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155782/files/figure.png" pageId="3" pageNumber="4">Figs. 35</figureCitation>
) broad, right mandible with a small medial tooth; dorsal molar area with velvety patch consisting of tiny denticles (visible at x400;
<figureCitation id="D48B5791FFB86274C91DA5A3B749F8FA" box="[780,850,1772,1798]" captionStart="FIGURES 1  5" captionStartId="2.[264,383,1788,1812]" captionTargetBox="[332,1234,835,1756]" captionTargetId="figure@2.[329,1237,830,1756]" captionTargetPageId="2" captionText="FIGURES 1  5. Mouthparts of Microlia meticola (Casey) (Arizona). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155782/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). Maxilla (
<figureCitation id="D48B5791FFB86274C9CBA5A3B058F8FA" box="[986,1091,1772,1798]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="3" pageNumber="4">Figs. 68</figureCitation>
) with galea extending beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; apical half of lacinia with row of closely spaced spines, middle portion produced medially and covered with numerous setae. Maxillary palpus with four segments and pseudosegment (
<figureCitation id="D48B5791FFBF6273CBC1A20BB600FEA2" box="[464,539,324,350]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="4" pageNumber="5">Fig. 6</figureCitation>
). Labium as in
<figureCitation id="D48B5791FFBF6273C8F8A20BB77EFEA2" box="[745,869,324,350]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="4" pageNumber="5">Figs. 911</figureCitation>
; labial palpi with three segments and pseudosegment (
<figureCitation id="D48B5791FFBF6273CBDEA223B602FE7A" box="[463,537,364,390]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="4" pageNumber="5">Fig. 9</figureCitation>
); apical half of ligula divided into two lobes; medial area of prementum without pores or pseudopores, lateral areas with 46 pores and single spinose pore. Hypopharyngeal lobes as in
<figureCitation id="D48B5791FFBF6273C8A4A2F3B714FE2A" box="[693,783,444,470]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="4" pageNumber="5">Fig. 10</figureCitation>
. Mentum (
<figureCitation id="D48B5791FFBF6273C98EA2F3B7EDFE2A" box="[927,1014,444,470]" captionStart="FIGURES 6  12" captionStartId="3.[264,383,1352,1376]" captionText="FIGURES 6  12. Mouthparts and antenna of Microlia meticola (Casey) (Arizona). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum; 12 – right antenna. Scale bar 0.14 mm (6), 0.1 mm (7  11), 0.28 mm (12)." pageId="4" pageNumber="5">Fig. 11</figureCitation>
) with protruding anterior angles, straight anterior margin, medial area without pores.
</paragraph>
<paragraph id="4C0F4B14FFBF6273CB29A143B61BFBFA" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">
Pronotum (
<figureCitation id="D48B5791FFBF6273CBACA143B608FDDA" box="[445,531,524,550]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="4" pageNumber="5">Fig. 13</figureCitation>
) strongly transverse, broadest near middle, sides broadly rounded; anterior margin straight, posterior margin convex; surface covered with microsetae directed posteriorly in midline, laterally and obliquely posteriorly in lateral areas (
<typeStatus id="930BF5B6FFBF6273CEDEA113B110FD8A" box="[1231,1291,604,630]" pageId="4" pageNumber="5">Type</typeStatus>
V,
<bibRefCitation id="282136E5FFBF6273CB19A1CBB602FD62" author="Benick" box="[264,537,644,670]" pageId="4" pageNumber="24" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); macrosetae short; hypomera invisible in lateral view. Meso  metasternum as in
<figureCitation id="D48B5791FFBF6273CBF9A1E3B624FD3A" box="[488,575,684,710]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="4" pageNumber="5">Fig. 14</figureCitation>
, mesosternal process long and wide, extended about 1/2 length of mesocoxal cavities, metasternal process almost nonexistent, anterior margin of metasternum only slightly convex medially; mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 11:10:2; mesocoxal cavities margined posteriorly; mesocoxae contiguous. Medial microseta of mesotibia inconspicuous, shorter than tibial width. Tarsal segmentation 45 5 or 445 (
<figureCitation id="D48B5791FFBF6273CB8AA0D3B631FC4A" box="[411,554,924,950]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="4" pageNumber="5">Figs. 1520</figureCitation>
); metatarsal segment 1 not longer than segment 2. One short empodial seta. Wings fully developed. Posterior margin of elytra slightly concave near posterolateral angle.
</paragraph>
<paragraph id="4C0F4B14FFBF6273CB29A75BB763FB82" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">Abdominal terga 35 with moderately transverse basal impressions. Tergum 7 1.41.6 times longer than tergum 6. Puncturation on terga 67 not significantly sparser than on terga 35. Tergum 7 with wide white palisade fringe.</paragraph>
<paragraph id="4C0F4B14FFBF6273CB29A7C3B647FA42" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">
Male secondary characters include some of the following: lateral triangular lobes or medial knob at posterior margin of tergum 3, longitudinal tubercles on terga 7 and 8, crenulation of posterior margin of tergum 8. Posterior margin of male abdominal sternum 8 extended posteriorly as pointed triangular lobe (
<figureCitation id="D48B5791FFBF6273C95FA64BB7ABFAE2" box="[846,944,1284,1310]" captionStart="FIGURES 21  25" captionStartId="9.[264,383,1766,1790]" captionTargetBox="[324,1235,612,1747]" captionTargetId="figure@9.[320,1236,612,1749]" captionTargetPageId="9" captionText="FIGURES 21  25. Abdominal segment 8 and female accessory sclerites of Microlia silacea (Erichson) (21 – paralectotype of Homalota silacea; 22, 25 – North Carolina; 23  24 – lectotype of Homalota silacea). 21 – male sternum 8; 22 – male tergum 8; 23 – female sternum 8; 24 – female tergum 8; 25 – female accessory sclerites. Scale bar 0.25 mm (21  24), 0.1 mm (25)." httpUri="https://zenodo.org/record/155783/files/figure.png" pageId="4" pageNumber="5">Figs. 21</figureCitation>
,
<figureCitation id="D48B5791FFBF6273C9ADA64BB7C7FAE2" box="[956,988,1284,1310]" captionStart="FIGURES 34  37" captionStartId="12.[264,383,1107,1131]" captionTargetBox="[339,1231,452,1085]" captionTargetId="figure@12.[337,1231,452,1086]" captionTargetPageId="12" captionText="FIGURES 34  37. Details of Microlia meticola (Casey) (Arizona). 34 – male sternum 8; 35 – male tergum 8; 36 – female sternum 8; 37 – female tergum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155785/files/figure.png" pageId="4" pageNumber="5">34</figureCitation>
,
<figureCitation id="D48B5791FFBF6273C9F9A64BB012FAE2" box="[1000,1033,1284,1310]" captionStart="FIGURES 62  67" captionStartId="19.[264,383,1428,1452]" captionTargetBox="[360,1260,320,1405]" captionTargetId="figure@19.[354,1262,320,1409]" captionTargetPageId="19" captionText="FIGURES 62  67. Details of Microlia tetramera Gusarov, sp. nov. (62  63 – holotype; 64  67 – possible female of M. tetramera from Costa Rica, 64  65 – type A, 66  67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155790/files/figure.png" pageId="4" pageNumber="5">62</figureCitation>
). Median lobe of aedeagus with long and narrow apical process (
<figureCitation id="D48B5791FFBF6273C8EEA663B791FABA" box="[767,906,1324,1350]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="4" pageNumber="5">Figs. 2629</figureCitation>
,
<figureCitation id="D48B5791FFBF6273C989A663B7FAFABA" box="[920,993,1324,1350]" captionStart="FIGURES 68  76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68  76. Details of Microlia tetramera Gusarov, sp. nov. (68  72, 75  76 – holotype; 73  74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73  74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73  74), 0.1 mm (69, 71  72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="4" pageNumber="5">6872</figureCitation>
). Parameres with two long and two short macrosetae (
<figureCitation id="D48B5791FFBF6273C852A61BB6BEFA92" box="[579,677,1364,1390]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="4" pageNumber="5">Figs. 31</figureCitation>
,
<figureCitation id="D48B5791FFBF6273C8A3A61BB6CFFA92" box="[690,724,1364,1390]" captionStart="FIGURES 38  44" captionStartId="13.[264,383,1768,1792]" captionTargetBox="[362,1223,685,1753]" captionTargetId="figure@13.[358,1230,683,1753]" captionTargetPageId="13" captionText="FIGURES 38  44. Details of Microlia meticola (Casey) (Arizona). 38 – median lobe of aedeagus, lateral view; 39 – apex of median lobe of aedeagus, lateral view; 40 – median lobe of aedeagus, ventral view; 41 – apex of median lobe of aedeagus, ventral view; 42  43 – spermatheca; 44 – apical lobe of right paramere, internal view. Scale bar 0.2 mm (38, 40, 42  43), 0.1 mm (39, 41, 44)." httpUri="https://zenodo.org/record/155786/files/figure.png" pageId="4" pageNumber="5">44</figureCitation>
). Copulatory piece of internal sac with long apical process (
<figureCitation id="D48B5791FFBF6273CB60A633B5D2FA6A" box="[369,457,1404,1430]" captionStart="FIGURES 53  61" captionStartId="17.[264,383,1492,1516]" captionTargetBox="[350,1231,590,1462]" captionTargetId="figure@17.[350,1231,585,1464]" captionTargetPageId="17" captionText="FIGURES 53  61. Details of Microlia pentamera Gusarov, sp. nov. (53  58, 61 – holotype; 59  60 – paratypes). 53 – median lobe of aedeagus, lateral view; 54 – apex of median lobe of aedeagus, lateral view; 55 – median lobe of aedeagus, ventral view; 56 – apex of median lobe of aedeagus, ventral view; 57 – copulatory piece of internal sac; 58 – apical lobe of right paramere, internal view; 59  60 – spermatheca; 61 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (53, 55, 59  60), 0.1 mm (54, 56  58), 0.55 mm (61)." httpUri="https://zenodo.org/record/155789/files/figure.png" pageId="4" pageNumber="5">Fig. 57</figureCitation>
). Spermatheca forming two to many coils (
<figureCitation id="D48B5791FFBF6273C9C0A633B040FA6A" box="[977,1115,1404,1430]" captionStart="FIGURES 26  33" captionStartId="10.[264,383,1674,1698]" captionTargetBox="[363,1225,458,1639]" captionTargetId="figure@10.[359,1226,452,1639]" captionTargetPageId="10" captionText="FIGURES 26  33. Details of Microlia silacea (Erichson) (26  29, 31 – paralectotype of Homalota silacea; 30, 33 – North Carolina; 32 – lectotype of Homalota silacea). 26 – median lobe of aedeagus, lateral view; 27 – apex of median lobe of aedeagus, lateral view; 28 – median lobe of aedeagus, ventral view; 29 – apex of median lobe of aedeagus, ventral view; 30 – antenna; 31 – apical lobe of right paramere, internal view; 32  33 – spermatheca. Scale bar 0.2 mm (25, 27, 29, 31  33), 0.1 mm (26, 28, 30)." httpUri="https://zenodo.org/record/155784/files/figure.png" pageId="4" pageNumber="5">Figs. 3233</figureCitation>
,
<figureCitation id="D48B5791FFBF6273CE79A633B09CFA6A" box="[1128,1159,1404,1430]" captionStart="FIGURES 45  48" captionStartId="15.[264,383,1232,1256]" captionTargetBox="[335,1229,335,1210]" captionTargetId="figure@15.[329,1237,329,1216]" captionTargetPageId="15" captionText="FIGURES 45  48. Details of Microlia azteca (Casey) (paralectotype). 45 – female sternum 8; 46 – female tergum 8; 47 – spermatheca; 48 – right antenna. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155787/files/figure.png" pageId="4" pageNumber="5">47</figureCitation>
) or numerous irregular loops (
<figureCitation id="D48B5791FFBF6273CBD5A6EBB64BFA42" box="[452,592,1444,1470]" captionStart="FIGURES 38  44" captionStartId="13.[264,383,1768,1792]" captionTargetBox="[362,1223,685,1753]" captionTargetId="figure@13.[358,1230,683,1753]" captionTargetPageId="13" captionText="FIGURES 38  44. Details of Microlia meticola (Casey) (Arizona). 38 – median lobe of aedeagus, lateral view; 39 – apex of median lobe of aedeagus, lateral view; 40 – median lobe of aedeagus, ventral view; 41 – apex of median lobe of aedeagus, ventral view; 42  43 – spermatheca; 44 – apical lobe of right paramere, internal view. Scale bar 0.2 mm (38, 40, 42  43), 0.1 mm (39, 41, 44)." httpUri="https://zenodo.org/record/155786/files/figure.png" pageId="4" pageNumber="5">Figs. 4243</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBF6273CB29A684B707F9A2" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="4C0F4B14FFBF6273CB29A684B079FA1A" blockId="4.[264,1324,284,1870]" box="[312,1122,1483,1510]" pageId="4" pageNumber="5">
<emphasis id="7EC49706FFBF6273CB29A684B5C1FA19" bold="true" box="[312,474,1483,1509]" pageId="4" pageNumber="5">
<typeStatus id="930BF5B6FFBF6273CB29A684B56EFA19" box="[312,373,1483,1509]" pageId="4" pageNumber="5">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8BB03097FFBF6273CBF8A683B724FA1A" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[489,831,1484,1510]" class="Insecta" family="Staphylinidae" genus="Dolosota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pernix">
<emphasis id="7EC49706FFBF6273CBF8A683B6BDFA19" box="[489,678,1484,1509]" italics="true" pageId="4" pageNumber="5">Dolosota pernix</emphasis>
<bibRefCitation id="282136E5FFBF6273C8BFA683B724FA1A" author="Casey" box="[686,831,1484,1510]" pageId="4" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="4C0F4B14FFBF6273CB29A6BCB707F9A2" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">
<emphasis id="7EC49706FFBF6273CB29A6BCB5DBF9F1" bold="true" box="[312,448,1523,1549]" pageId="4" pageNumber="5">Synonyms.</emphasis>
<taxonomicName id="8BB03097FFBF6273CBC1A6BBB708F9F2" authority="Casey, 1911" authorityName="Casey" authorityYear="1911" box="[464,787,1524,1550]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="7EC49706FFBF6273CBC1A6BBB661F9F1" box="[464,634,1524,1549]" italics="true" pageId="4" pageNumber="5">Nosora azteca</emphasis>
<bibRefCitation id="282136E5FFBF6273C890A6BBB708F9F2" author="Casey" box="[641,787,1524,1550]" pageId="4" pageNumber="24" refString="Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. Lancaster: The New Era Printing Company, 245 pp." type="book" year="1911">Casey, 1911</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="930BF5B6FFBF6273C95DA6BBB764F9F2" box="[844,895,1524,1550]" pageId="4" pageNumber="5">type</typeStatus>
species of the genus
<taxonomicName id="8BB03097FFBF6273CE6DA6BBB0C9F9F1" box="[1148,1234,1524,1549]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBF6273CE6DA6BBB0C9F9F1" box="[1148,1234,1524,1549]" italics="true" pageId="4" pageNumber="5">Nosora</emphasis>
</taxonomicName>
by original designation is similar to
<taxonomicName id="8BB03097FFBF6273C879A553B6C2F9C9" box="[616,729,1564,1589]" class="Insecta" family="Staphylinidae" genus="Dolosota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pernix">
<emphasis id="7EC49706FFBF6273C879A553B6C2F9C9" box="[616,729,1564,1589]" italics="true" pageId="4" pageNumber="5">M. pernix</emphasis>
</taxonomicName>
in all characters listed in the Diagnosis. Therefore
<taxonomicName id="8BB03097FFBF6273CB19A50BB545F9A1" box="[264,350,1604,1629]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBF6273CB19A50BB545F9A1" box="[264,350,1604,1629]" italics="true" pageId="4" pageNumber="5">Nosora</emphasis>
</taxonomicName>
is placed in synynymy with
<taxonomicName id="8BB03097FFBF6273C8A1A50BB70EF9A1" box="[688,789,1604,1629]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBF6273C8A1A50BB70EF9A1" box="[688,789,1604,1629]" italics="true" pageId="4" pageNumber="5">Microlia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="04AA189FFFBF6270CB29A524B130FE52" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="discussion">
<paragraph id="4C0F4B14FFBF6272CB29A524B5B3FE7A" blockId="4.[264,1324,284,1870]" lastBlockId="5.[264,1323,284,390]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="7EC49706FFBF6273CB29A524B5D8F979" bold="true" box="[312,451,1643,1669]" pageId="4" pageNumber="5">Discussion.</emphasis>
My examination of the
<typeStatus id="930BF5B6FFBF6273C912A523B775F97A" box="[771,878,1644,1670]" pageId="4" pageNumber="5" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="8BB03097FFBF6273C9B0A523B131F97A" authority="Casey, 1911" authorityName="Casey" authorityYear="1911" box="[929,1322,1644,1670]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFBF6273C9B0A523B06CF979" box="[929,1143,1644,1669]" italics="true" pageId="4" pageNumber="5">Microlia meticola</emphasis>
(
<bibRefCitation id="282136E5FFBF6273CE9DA523B139F97A" author="Casey" box="[1164,1314,1644,1670]" pageId="4" pageNumber="24" refString="Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. Lancaster: The New Era Printing Company, 245 pp." type="book" year="1911">Casey, 1911</bibRefCitation>
)
</taxonomicName>
(original combination:
<taxonomicName id="8BB03097FFBF6273C808A5DBB6C6F951" box="[537,733,1684,1709]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFBF6273C808A5DBB6C6F951" box="[537,733,1684,1709]" italics="true" pageId="4" pageNumber="5">Nosora meticola</emphasis>
</taxonomicName>
) revealed that in this species the tarsal formula is 445 (
<figureCitation id="D48B5791FFBF6273CB49A5F3B5B5F92A" box="[344,430,1724,1750]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="4" pageNumber="5">Fig. 15</figureCitation>
). Examination of additional material demonstrated that in
<taxonomicName id="8BB03097FFBF6273CE72A5F3B0F4F929" box="[1123,1263,1724,1749]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFBF6273CE72A5F3B0F4F929" box="[1123,1263,1724,1749]" italics="true" pageId="4" pageNumber="5">M. meticola</emphasis>
</taxonomicName>
both sexes have this tarsal formula and there is no intraspecific variation in this character (in total more than
<specimenCount id="5AB6809DFFBF6273CBD7A443B67DF8DA" box="[454,614,1804,1830]" pageId="4" pageNumber="5" type="generic">80 specimens</specimenCount>
were examined). Up to now this fact has been overlooked (despite the
<typeStatus id="930BF5B6FFBF6273CB88A47BB5D0F8B2" box="[409,459,1844,1870]" pageId="4" pageNumber="5">type</typeStatus>
of
<taxonomicName id="8BB03097FFBF6273CBE4A47BB666F8B1" box="[501,637,1844,1869]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFBF6273CBE4A47BB666F8B1" box="[501,637,1844,1869]" italics="true" pageId="4" pageNumber="5">N. meticola</emphasis>
</taxonomicName>
carrying a small label “445”, apparently by Casey), and all authors who wrote about the group believed the formula was 455. One of the new species of
<taxonomicName id="8BB03097FFBE6272CB94A20BB5F1FEA1" box="[389,490,324,349]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBE6272CB94A20BB5F1FEA1" box="[389,490,324,349]" italics="true" pageId="5" pageNumber="6">Microlia</emphasis>
</taxonomicName>
described below (
<taxonomicName id="8BB03097FFBE6272C8D5A20BB77BFEA1" box="[708,864,324,349]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tetramera">
<emphasis id="7EC49706FFBE6272C8D5A20BB77BFEA1" box="[708,864,324,349]" italics="true" pageId="5" pageNumber="6">M. tetramera</emphasis>
</taxonomicName>
) also has mesotarsus with four articles (
<figureCitation id="D48B5791FFBE6272CB00A223B580FE7A" box="[273,411,364,390]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="5" pageNumber="6">Figs. 1718</figureCitation>
).
</paragraph>
<caption id="18CF1B9CFFBE6272CB19A5D2B783F89D" pageId="5" pageNumber="6">
<paragraph id="4C0F4B14FFBE6272CB19A5D2B783F89D" blockId="5.[264,1323,1693,1889]" pageId="5" pageNumber="6">
<emphasis id="7EC49706FFBE6272CB19A5D2B5DEF949" bold="true" box="[264,453,1693,1717]" pageId="5" pageNumber="6">FIGURES 1320</emphasis>
. Details of
<taxonomicName id="8BB03097FFBE6272C85FA5D2B772F948" authority="Casey" authorityName="Casey" box="[590,873,1693,1716]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="meticola">
<emphasis id="7EC49706FFBE6272C85FA5D2B715F948" box="[590,782,1693,1716]" italics="true" pageId="5" pageNumber="6">Microlia meticola</emphasis>
(Casey)
</taxonomicName>
(1315, Arizona),
<taxonomicName id="8BB03097FFBE6272CE25A5D2B0E3F948" authority="Casey" authorityName="Casey" box="[1076,1272,1693,1716]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="7EC49706FFBE6272CE25A5D2B085F948" box="[1076,1182,1693,1716]" italics="true" pageId="5" pageNumber="6">M. azteca</emphasis>
(Casey)
</taxonomicName>
(16, paralectotype),
<taxonomicName id="8BB03097FFBE6272CBBEA58FB6D9F92B" authority="Gusarov" authorityName="Gusarov" box="[431,706,1728,1751]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tetramera" status="sp. nov.">
<emphasis id="7EC49706FFBE6272CBBEA58FB5D7F92B" box="[431,460,1728,1751]" italics="true" pageId="5" pageNumber="6">M.</emphasis>
cf.
<emphasis id="7EC49706FFBE6272CBE4A58FB645F92B" box="[501,606,1728,1751]" italics="true" pageId="5" pageNumber="6">tetramera</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="65F72A7DFFBE6272C8DDA5F0B739F92B" box="[716,802,1727,1751]" pageId="5" pageNumber="6" rank="species">
<emphasis id="7EC49706FFBE6272C8DDA5F0B739F92B" bold="true" box="[716,802,1727,1751]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
(1718, female from Costa Rica) and
<taxonomicName id="8BB03097FFBE6272CEAAA58FB563F905" authority="Erichson" authorityName="Erichson" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silacea">
<emphasis id="7EC49706FFBE6272CEAAA58FB0C2F92B" box="[1211,1241,1728,1751]" italics="true" pageId="5" pageNumber="6">M.</emphasis>
<emphasis id="7EC49706FFBE6272CECEA58FB131F92B" box="[1247,1322,1728,1751]" italics="true" pageId="5" pageNumber="6">silacea</emphasis>
(Erichson)
</taxonomicName>
(1920, paralectotype of
<taxonomicName id="8BB03097FFBE6272C897A5ACB759F906" box="[646,834,1763,1786]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silacea">
<emphasis id="7EC49706FFBE6272C897A5ACB759F906" box="[646,834,1763,1786]" italics="true" pageId="5" pageNumber="6">Homalota silacea</emphasis>
</taxonomicName>
). 13 – pronotum; 14 – mesometathorax, ventral view; 1516 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (1314), 0.28 mm (1516), 0.1 mm (1720).
</paragraph>
</caption>
<paragraph id="4C0F4B14FFBD6271CB29A253B0A7FE52" blockId="6.[264,1324,284,1030]" pageId="6" pageNumber="7">
Traditionally in the aleocharine taxonomy the tarsal formula was considered an important character, often stable at the level of tribe. However in a few groups the tarsal formula was found to vary within the genus (as in
<taxonomicName id="8BB03097FFBD6271C912A223B0FCFE7A" authority="Casey, 1893 (Seevers 1978)" authorityName="Casey, 1893 (Seevers" authorityYear="1978" box="[771,1255,364,390]" class="Insecta" family="Staphylinidae" genus="Gyronycha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C912A223B79FFE79" box="[771,900,364,389]" italics="true" pageId="6" pageNumber="7">Gyronycha</emphasis>
Casey, 1893
<bibRefCitation id="282136E5FFBD6271CE39A223B0FCFE7A" author="Seevers" box="[1064,1255,364,390]" pageId="6" pageNumber="24" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">(Seevers 1978))</bibRefCitation>
</taxonomicName>
. The genus
<taxonomicName id="8BB03097FFBD6271CB45A2DBB5A2FE51" box="[340,441,404,429]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CB45A2DBB5A2FE51" box="[340,441,404,429]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
is a new example of such intrageneric variation in tarsal formula.
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB29A2F3B0BCFC9A" blockId="6.[264,1324,284,1030]" pageId="6" pageNumber="7">
To establish homologies of tarsal segments between the species with different tarsal formulas the setation was examined. In 455 species every segment of mesotarsus carries a pair of lateral setae (
<figureCitation id="D48B5791FFBD6271C80BA143B6BDFDDA" box="[538,678,524,550]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Figs. 1920</figureCitation>
, l); on ventral side the first segment has two pairs of setae, secondfourth have one pair of ventral setae each, fifth has no ventral setae (
<figureCitation id="D48B5791FFBD6271CEDEA17BB13DFDB2" box="[1231,1318,564,590]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Fig. 20</figureCitation>
, v); on dorsal side the first segment has no setae, secondfourth segments have one or two dorsal setae each, fifth has two pairs of dorsal setae (
<figureCitation id="D48B5791FFBD6271C97FA1CBB7D5FD62" box="[878,974,644,670]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Figs. 19</figureCitation>
, d). Comparison between the 455 (
<figureCitation id="D48B5791FFBD6271CB48A1E3B5FAFD3A" box="[345,481,684,710]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Figs. 1920</figureCitation>
) and 445 (
<figureCitation id="D48B5791FFBD6271C864A1E3B6E9FD3A" box="[629,754,684,710]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Fig. 1718</figureCitation>
) species shows that in the 445 species the last segment of mesotarsus has the setae found on both fourth and fifth segments of the 455 species: a pair of ventral setae (
<figureCitation id="D48B5791FFBD6271C86AA1B3B6CAFCEA" box="[635,721,764,790]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Fig. 18</figureCitation>
, v), 1+1+2 dorsal setae (
<figureCitation id="D48B5791FFBD6271C9FEA1B3B05FFCEA" box="[1007,1092,764,790]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Fig. 17</figureCitation>
, d) and two pairs of lateral setae (
<figureCitation id="D48B5791FFBD6271CBB4A06BB62BFCC2" box="[421,560,804,830]" captionStart="FIGURES 13  20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13  20. Details of Microlia meticola (Casey) (13  15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17  18, female from Costa Rica) and M. silacea (Erichson) (19  20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15  16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13  14), 0.28 mm (15  16), 0.1 mm (17  20)." pageId="6" pageNumber="7">Figs. 1718</figureCitation>
, l). Apparently the last segment of mesotarsus in the 445 species is homologous to the two last segments of mesotarsus in the 455 species.
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB29A03BB7D6FBFA" blockId="6.[264,1324,284,1030]" pageId="6" pageNumber="7">
The key for aleocharine genera by
<bibRefCitation id="282136E5FFBD6271C8C7A03BB009FC72" box="[726,1042,884,910]" pageId="6" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter">
Ashe (Newton
<emphasis id="7EC49706FFBD6271C998A03BB7A5FC71" box="[905,958,884,909]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2000)
</bibRefCitation>
does not allow to identify the species of
<taxonomicName id="8BB03097FFBD6271CBCCA0D3B659FC49" box="[477,578,924,949]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CBCCA0D3B659FC49" box="[477,578,924,949]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
(=
<taxonomicName id="8BB03097FFBD6271C872A0D3B6A2FC49" box="[611,697,924,949]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C872A0D3B6A2FC49" box="[611,697,924,949]" italics="true" pageId="6" pageNumber="7">Nosora</emphasis>
</taxonomicName>
) with tarsal formula 445. These species run to couplet
<quantity id="8B48E6F1FFBD6271CB53A08BB598FC22" box="[322,387,964,990]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="6" pageNumber="7" unit="in" value="24.0">24 in</quantity>
Key E. To incorporate
<taxonomicName id="8BB03097FFBD6271C8D3A08BB73CFC21" box="[706,807,964,989]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C8D3A08BB73CFC21" box="[706,807,964,989]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
the key should be modified by inserting additional couplet as follows (p.
<quantity id="8B48E6F1FFBD6271C899A0A3B6C8FBFA" box="[648,723,1004,1030]" metricMagnitude="0" metricUnit="m" metricValue="7.8232" pageId="6" pageNumber="7" unit="in" value="308.0">308 in</quantity>
<bibRefCitation id="282136E5FFBD6271C8CBA0A3B7DAFBFA" author="Newton" box="[730,961,1004,1030]" pageId="6" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter" year="2000">
Newton
<emphasis id="7EC49706FFBD6271C92FA0A3B76AFBF9" box="[830,881,1004,1029]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2000
</bibRefCitation>
):
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB19A773B137FBAA" blockId="6.[264,1324,1084,1230]" box="[264,1324,1084,1110]" pageId="6" pageNumber="7">24(22) Maxillary (Fig. 155.22, arrow) and labial (Fig. 159.22) palpi with apical pseudo</paragraph>
<paragraph id="4C0F4B14FFBD6271CB79A72BB7FBFB32" blockId="6.[264,1324,1084,1230]" pageId="6" pageNumber="7">
segment, appearing to have 5 and 4 palpomeres, respectively........
<taxonomicName id="8BB03097FFBD6271CEAAA72BB13BFB81" box="[1211,1312,1124,1149]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CEAAA72BB13BFB81" box="[1211,1312,1124,1149]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
. – Maxillary and labial palpi without apical pseudosegments............... 24a 24a(24) Body distinctly … (from here follow the original key)
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB29A64BB561FABA" blockId="6.[264,1324,1284,1870]" pageId="6" pageNumber="7">
The 455 species of
<taxonomicName id="8BB03097FFBD6271C83EA64BB68EFAE1" box="[559,661,1284,1309]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C83EA64BB68EFAE1" box="[559,661,1284,1309]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
reach
<taxonomicName id="8BB03097FFBD6271C8F3A64BB723FAE1" box="[738,824,1284,1309]" class="Insecta" family="Staphylinidae" genus="Nosora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C8F3A64BB723FAE1" box="[738,824,1284,1309]" italics="true" pageId="6" pageNumber="7">Nosora</emphasis>
</taxonomicName>
at couplet
<quantity id="8B48E6F1FFBD6271C9A8A64BB7FDFAE2" box="[953,998,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="7" unit="in" value="3.0">3 in</quantity>
Key C by
<bibRefCitation id="282136E5FFBD6271CE75A64BB56DFABA" pageId="6" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter">
Ashe (Newton
<emphasis id="7EC49706FFBD6271CF04A64BB505FAB9" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2000)
</bibRefCitation>
.
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB29A61BB636FA42" blockId="6.[264,1324,1284,1870]" pageId="6" pageNumber="7">
Two species were originally included in subgenus
<taxonomicName id="8BB03097FFBD6271C99AA61BB7EBFA91" box="[907,1008,1364,1389]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C99AA61BB7EBFA91" box="[907,1008,1364,1389]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
by
<bibRefCitation id="282136E5FFBD6271CE0FA61BB0DBFA92" author="Casey" box="[1054,1216,1364,1390]" pageId="6" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey (1910)</bibRefCitation>
. The second species,
<taxonomicName id="8BB03097FFBD6271CBA1A633B745FA6A" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[432,862,1404,1430]" class="Insecta" family="Staphylinidae" genus="Dolosota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="petulans" subGenus="Microlia">
<emphasis id="7EC49706FFBD6271CBA1A633B607FA69" box="[432,540,1404,1429]" italics="true" pageId="6" pageNumber="7">Dolosota</emphasis>
(
<emphasis id="7EC49706FFBD6271C83FA633B655FA69" box="[558,590,1404,1429]" italics="true" pageId="6" pageNumber="7">M.</emphasis>
)
<emphasis id="7EC49706FFBD6271C871A633B6DFFA69" box="[608,708,1404,1429]" italics="true" pageId="6" pageNumber="7">petulans</emphasis>
<bibRefCitation id="282136E5FFBD6271C8DCA633B745FA6A" author="Casey" box="[717,862,1404,1430]" pageId="6" pageNumber="24" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
belongs to the genus
<taxonomicName id="8BB03097FFBD6271CE77A633B566FA42" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CE77A633B0C9FA69" box="[1126,1234,1404,1429]" italics="true" pageId="6" pageNumber="7">Acrotona</emphasis>
Thomson, 1859
</taxonomicName>
, tribe
<taxonomicName id="8BB03097FFBD6271CBD4A6EBB632FA42" box="[453,553,1444,1470]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
.
</paragraph>
<paragraph id="4C0F4B14FFBD6271CB29A683B0CBF8DA" blockId="6.[264,1324,1284,1870]" pageId="6" pageNumber="7">
All species of
<taxonomicName id="8BB03097FFBD6271CBE0A683B64CFA19" box="[497,599,1484,1509]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CBE0A683B64CFA19" box="[497,599,1484,1509]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
mentioned in this paper are similar in external characters including proportions of antennal segments. The species represented in my material by many specimens show considerable variation in body coloration and sometimes in puncturation of pronotum. Some species are known from only few specimens which does not allow to assess variability of body coloration. Considering the observed intraspecific variation and the possibility that undescribed species of
<taxonomicName id="8BB03097FFBD6271C969A5DBB7C6F951" box="[888,989,1684,1709]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271C969A5DBB7C6F951" box="[888,989,1684,1709]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
exist in Central
<collectingCountry id="34A70B84FFBD6271CEB7A5DBB115F952" box="[1190,1294,1684,1710]" name="United States of America" pageId="6" pageNumber="7">America</collectingCountry>
, it seems reasonable that a key to known species based on coloration of specimens will be of limited value. That is why the key below is based mostly on the characters of genitalia and secondary sexual characters even though some species are known by one sex only.
</paragraph>
<paragraph id="4C0F4B14FFBD6270CB29A47BB130FE52" blockId="6.[264,1324,1284,1870]" lastBlockId="7.[264,1323,284,430]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
There can be little doubt that many undescribed species of
<taxonomicName id="8BB03097FFBD6271CE12A47BB072F8B1" box="[1027,1129,1844,1869]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBD6271CE12A47BB072F8B1" box="[1027,1129,1844,1869]" italics="true" pageId="6" pageNumber="7">Microlia</emphasis>
</taxonomicName>
exist in Central and possibly South
<collectingCountry id="34A70B84FFBC6270CBE5A253B646FECA" box="[500,605,284,310]" name="United States of America" pageId="7" pageNumber="8">America</collectingCountry>
. The high diversity of
<taxonomicName id="8BB03097FFBC6270C966A253B7C6FEC9" box="[887,989,284,309]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBC6270C966A253B7C6FEC9" box="[887,989,284,309]" italics="true" pageId="7" pageNumber="8">Microlia</emphasis>
</taxonomicName>
in Central
<collectingCountry id="34A70B84FFBC6270CE75A253B0D2FECA" box="[1124,1225,284,310]" name="United States of America" pageId="7" pageNumber="8">America</collectingCountry>
is illustrated by the fact that in a single locality in
<collectingCountry id="34A70B84FFBC6270C93AA20BB7B4FEA2" box="[811,943,324,350]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
three species of
<taxonomicName id="8BB03097FFBC6270CE90A20BB0FCFEA1" box="[1153,1255,324,349]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="7EC49706FFBC6270CE90A20BB0FCFEA1" box="[1153,1255,324,349]" italics="true" pageId="7" pageNumber="8">Microlia</emphasis>
</taxonomicName>
were found (
<taxonomicName id="8BB03097FFBC6270CB4FA223B674FE7A" authority="Gusarov" authorityName="Gusarov" box="[350,623,364,390]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pentamera" status="sp. nov.">
<emphasis id="7EC49706FFBC6270CB4FA223B619FE79" box="[350,514,364,389]" italics="true" pageId="7" pageNumber="8">M. pentamera</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="65F72A7DFFBC6270C86AA224B6C2FE79" box="[635,729,363,389]" pageId="7" pageNumber="8" rank="species">
<emphasis id="7EC49706FFBC6270C86AA224B6C2FE79" bold="true" box="[635,729,363,389]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="8BB03097FFBC6270C8F9A223B7F5FE7A" authority="Gusarov" authorityName="Gusarov" box="[744,1006,364,390]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tetramera" status="sp. nov.">
<emphasis id="7EC49706FFBC6270C8F9A223B799FE79" box="[744,898,364,389]" italics="true" pageId="7" pageNumber="8">M. tetramera</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="65F72A7DFFBC6270C9EBA224B043FE79" box="[1018,1112,363,389]" pageId="7" pageNumber="8" rank="species">
<emphasis id="7EC49706FFBC6270C9EBA224B043FE79" bold="true" box="[1018,1112,363,389]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
and an additional species represented by females only; see Discussion after the description of
<taxonomicName id="8BB03097FFBC6270CE6EA2DBB101FE51" box="[1151,1306,404,429]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tetramera">
<emphasis id="7EC49706FFBC6270CE6EA2DBB101FE51" box="[1151,1306,404,429]" italics="true" pageId="7" pageNumber="8">M. tetramera</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>