460 lines
44 KiB
XML
460 lines
44 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.72564" ID-GBIF-Dataset="10e84242-30e4-4279-b9ae-6da1d74a4512" ID-GBIF-Taxon="190304341" ID-Pensoft-Pub="2535-0730-2-335" ID-Pensoft-UUID="F54528451556597C8689D7DBC1E0158E" ID-ZooBank="CA45E26222CD463E9B8A13B324319E87" LSID="urn:lsid:zoobank.org:act:E7D48769-300F-43B4-8F8B-45E0A8ED414B" ModsDocID="2535-0730-5-2-335" checkinTime="1635891671126" checkinUser="pensoft" docAuthor="Mirza, Zeeshan A., Bhardwaj, Virender Kumar & Patel, Harshil" docDate="2021" docId="C330BC54C36C5677AE14780365216079" docLanguage="en" docName="EvolutSyst 5(2): 335-345" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-11-02" docSource="http://dx.doi.org/10.3897/evolsyst.5.72564" docTitle="Oligodon churahensis Mirza & Bhardwaj & Patel 2021, sp. nov." docType="treatment" docUuid="E7D48769-300F-43B4-8F8B-45E0A8ED414B" docUuidSource="ZooBank" docVersion="4" id="F54528451556597C8689D7DBC1E0158E" lastPageNumber="335" masterDocId="F54528451556597C8689D7DBC1E0158E" masterDocTitle="A new species of snake of the genus Oligodon Boie in Fitzinger, 1826 (Reptilia, Serpentes) from the Western Himalayas" masterLastPageNumber="345" masterPageNumber="335" pageId="0" pageNumber="335" updateTime="1637334872922" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of snake of the genus Oligodon Boie in Fitzinger, 1826 (Reptilia, Serpentes) from the Western Himalayas</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mirza, Zeeshan A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1685-9816</mods:nameIdentifier>
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<mods:affiliation>National Centre for Biological Sciences, Tata Institute for Fundamental Research, Bangalore, Karnataka 560065, India & The University of Trans-Disciplinary Health Sciences and Technology (TDU), 74 / 2, Post Attur via Yelahanka, Jarakabande Kaval, Bengaluru, Karnataka 560064, India</mods:affiliation>
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<mods:nameIdentifier type="email">snakeszeeshan@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bhardwaj, Virender Kumar</mods:namePart>
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<mods:affiliation>Department of Zoology, Guru Nanak Dev University, Amritsar, Punjab 143005, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Patel, Harshil</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4679-5211</mods:nameIdentifier>
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<mods:affiliation>Department of Biosciences, Veer Narmad South Gujarat University, Surat 395007, Gujarat, India & Voluntary Nature Conservancy, 101 - Radha Darshan, Behind Union Bank, Vallabh Vidyanagar 388120, Gujarat, India</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>335</mods:start>
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<mods:end>345</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.5.72564</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.5.72564</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-335</mods:identifier>
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<mods:identifier type="ZooBank">CA45E26222CD463E9B8A13B324319E87</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F54528451556597C8689D7DBC1E0158E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="190304341" LSID="urn:lsid:zoobank.org:act:E7D48769-300F-43B4-8F8B-45E0A8ED414B" httpUri="http://treatment.plazi.org/id/C330BC54C36C5677AE14780365216079" lastPageNumber="335" pageId="0" pageNumber="335">
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<subSubSection pageId="0" pageNumber="335" type="nomenclature">
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<paragraph pageId="0" pageNumber="335">
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<taxonomicName LSID="http://zoobank.org/E7D48769-300F-43B4-8F8B-45E0A8ED414B" authority="Mirza & Bhardwaj & Patel, 2021" authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis" status="sp. nov.">Oligodon churahensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="335">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="description">
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<paragraph pageId="0" pageNumber="335">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019 (a) dorsal view, (b) ventral view. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/606329" pageId="0" pageNumber="335">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micro CT scan images of the skull, (a-c) Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019; (d-f) Oligodon arnensis NCBS NRC-AA- 020 showing dorsal view (a, d), lateral view (b, e) and ventral view (c, f)." httpUri="https://binary.pensoft.net/fig/606331" pageId="0" pageNumber="335">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hemipenis of Oligodon churahensis sp. nov. paratype male BNHS 3657 (a) sulcal view, (b) asulcal view. Scale bars: 1 mm." httpUri="https://binary.pensoft.net/fig/606333" pageId="0" pageNumber="335">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019 in life (a); a general view of the habitat at the type locality (b), Photos by Virender Kumar." httpUri="https://binary.pensoft.net/fig/606335" pageId="0" pageNumber="335">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="holotype">
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<paragraph pageId="0" pageNumber="335">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="335">
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female NCBS NRC-AA-019 from near Thanei Kothi village, Churah Valley, Chamba District, Himachal Pradesh, India (
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<geoCoordinate degrees="32.835467" direction="north" orientation="latitude" precision="1" value="32.83547">32.835467</geoCoordinate>
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,
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<geoCoordinate degrees="76.119381" direction="east" orientation="longitude" precision="1" value="76.119385">76.119381</geoCoordinate>
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, elevation 1864 m) collected by Virendar Kumar on 22nd June 2020.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="paratype">
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<paragraph pageId="0" pageNumber="335">Paratype.</paragraph>
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<paragraph pageId="0" pageNumber="335">male BNHS 3657 collected on 25 June 2020, same data as for the holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="etymology">
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<paragraph pageId="0" pageNumber="335">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="335">The specific epithet refers to the Churah Valley where the new species was collected.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="suggested common name">
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<paragraph pageId="0" pageNumber="335">Suggested common name.</paragraph>
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<paragraph pageId="0" pageNumber="335">Churah Valley Kukri</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="diagnosis">
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<paragraph pageId="0" pageNumber="335">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="335">
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A medium sized
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<taxonomicName class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="335">Oligodon</emphasis>
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</taxonomicName>
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(SVL 275 mm) with 17 dorsal scale rows at midbody. Seven supralabials, 3rd and 4th in contact with the eye. Loreal present. 170-175 ventrals, 46-47 subcaudals. Palatine and pterygoid teeth absent. Dorsal patterns consist of 1-2 dorsal scales wide black bands edged with yellow. Ventral scales white with brown smear along the width of each scale, the smear is darker on the lateral edges forming a blotch on each side. 48 to 54 bands in total on the body. Hemipenis forked and spinose throughout.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="335" type="comparisons">
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<paragraph pageId="0" pageNumber="335">Comparison.</paragraph>
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<paragraph pageId="0" pageNumber="335">
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Morphologically
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<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
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<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
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</taxonomicName>
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sp. nov. differs from all South Asian congeners, except
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<taxonomicName genus="O." lsidName="O. affinis" pageId="0" pageNumber="335" rank="species" species="affinis">
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<emphasis italics="true" pageId="0" pageNumber="335">O. affinis</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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, 1862,
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<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
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<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="O." lsidName="O. cinereus" pageId="0" pageNumber="335" rank="species" species="cinereus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. cinereus</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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, 1864),
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<taxonomicName genus="O." lsidName="O. erythrogaster" pageId="0" pageNumber="335" rank="species" species="erythrogaster">
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<emphasis italics="true" pageId="0" pageNumber="335">O. erythrogaster</emphasis>
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</taxonomicName>
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Boulenger, 1907,
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<taxonomicName genus="O." lsidName="O. melanozonatus" pageId="0" pageNumber="335" rank="species" species="melanozonatus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. melanozonatus</emphasis>
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</taxonomicName>
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Wall, 1922,
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<taxonomicName genus="O." lsidName="O. theobaldi" pageId="0" pageNumber="335" rank="species" species="theobaldi">
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<emphasis italics="true" pageId="0" pageNumber="335">O. theobaldi</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Günther">Guenther</normalizedToken>
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, 1868),
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<taxonomicName genus="O." lsidName="O. travancoricus" pageId="0" pageNumber="335" rank="species" species="travancoricus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. travancoricus</emphasis>
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</taxonomicName>
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Beddome, 1877,
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<taxonomicName genus="O." lsidName="O. venustus" pageId="0" pageNumber="335" rank="species" species="venustus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. venustus</emphasis>
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</taxonomicName>
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(Jerdon, 1853) and
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<taxonomicName genus="O." lsidName="O. woodmasoni" pageId="0" pageNumber="335" rank="species" species="woodmasoni">
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<emphasis italics="true" pageId="0" pageNumber="335">O. woodmasoni</emphasis>
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</taxonomicName>
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(Sclater, 1891), by having 17 dorsal scale rows at midbody. The new species differs from
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<taxonomicName genus="O." lsidName="O. affinis" pageId="0" pageNumber="335" rank="species" species="affinis">
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<emphasis italics="true" pageId="0" pageNumber="335">O. affinis</emphasis>
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</taxonomicName>
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in having higher (170-175) number of ventral scales (vs. 128-133), higher (46-47) number of subcaudal scales (vs. 23-36) and loreal present (vs. absent) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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;
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<bibRefCitation author="Whitaker, R" journalOrPublisher="Draco Books, Chennai" pageId="0" pageNumber="335" refId="B38" refString="Whitaker, R, Captain, A, 2004. Snakes of India. The field guide. Draco Books, Chennai" title="Snakes of India. The field guide." year="2004">Whitaker and Captain 2004</bibRefCitation>
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). It differs from
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<taxonomicName genus="O." lsidName="O. cinereus" pageId="0" pageNumber="335" rank="species" species="cinereus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. cinereus</emphasis>
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</taxonomicName>
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in having higher (46-47) number of subcaudal scales (vs. 29-42), 7 supralabials (vs. 8), hemipenis forked and spinose (vs. hemipenis not forked, lack spines) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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). It differs from
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<taxonomicName genus="O." lsidName="O. erythrogaster" pageId="0" pageNumber="335" rank="species" species="erythrogaster">
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<emphasis italics="true" pageId="0" pageNumber="335">O. erythrogaster</emphasis>
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</taxonomicName>
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by having a loreal present (vs. absent), hemipenis forked and spinose (vs. hemipenis not forked, lack spines) and dorsum with darker bands (vs. dorsum with stripes, bands absent) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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). The new species differs from
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<taxonomicName genus="O." lsidName="O. melanozonatus" pageId="0" pageNumber="335" rank="species" species="melanozonatus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. melanozonatus</emphasis>
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</taxonomicName>
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in having 7 supralabials (vs. 6), higher (46-47) number of subcaudal scales (vs. 42-45) and loreal present (vs. absent) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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). It differs from
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<taxonomicName genus="O." lsidName="O. theobaldi" pageId="0" pageNumber="335" rank="species" species="theobaldi">
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<emphasis italics="true" pageId="0" pageNumber="335">O. theobaldi</emphasis>
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</taxonomicName>
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in having 7 supralabials (vs. 8), higher (46-47) number of subcaudal scales (vs. 30-42), hemipenis forked and spinose (vs. hemipenis not forked, lack spines) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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). The new species differs from
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<taxonomicName genus="O." lsidName="O. travancoricus" pageId="0" pageNumber="335" rank="species" species="travancoricus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. travancoricus</emphasis>
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</taxonomicName>
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in having higher (170-175) number of ventral scales (vs. 150-155), higher (46-47) number of subcaudal scales (vs. 31-39), loreal present (vs. absent) and hemipenis forked (vs. not forked) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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;
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<bibRefCitation author="Ganesh, SR" journalOrPublisher="Herpetological Bulletin" pageId="0" pageNumber="335" pagination="25 - 28" refId="B14" refString="Ganesh, SR, Asokan, S, Kannan, P, 2009. Record of Oligodon travancoricus Beddome, 1877 (Serpentes, Colubridae) from Grizzled Squirrel Sanctuary, Western Ghats, Tamil Nadu, India. Herpetological Bulletin 109: 25 - 28" title="Record of Oligodon travancoricus Beddome, 1877 (Serpentes, Colubridae) from Grizzled Squirrel Sanctuary, Western Ghats, Tamil Nadu, India." volume="109" year="2009">Ganesh et al. 2009</bibRefCitation>
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). It differs from
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<taxonomicName genus="O." lsidName="O. venustus" pageId="0" pageNumber="335" rank="species" species="venustus">
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<emphasis italics="true" pageId="0" pageNumber="335">O. venustus</emphasis>
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</taxonomicName>
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in having higher (170-175) number of ventral scales (vs. 138-165), higher (46-47) number of subcaudal scales (vs. 27-36), loreal present (vs. absent) and hemipenis forked (vs. not forked) (
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<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
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). It differs from
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<taxonomicName genus="O." lsidName="O. woodmasoni" pageId="0" pageNumber="335" rank="species" species="woodmasoni">
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<emphasis italics="true" pageId="0" pageNumber="335">O. woodmasoni</emphasis>
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</taxonomicName>
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in having 7 supralabials (vs. 6), hemipenis forked and spinose (vs. hemipenis not forked, lack spines) and dorsum with darker bands (vs. dorsum with stripes, bands absent) (
|
||
<bibRefCitation author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="335" refId="B30" refString="Smith, MA, 1943. Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London" title="Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. 3. Serpentes." year="1943">Smith 1943</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Vijayakumar, SP" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="335" pagination="11 - 40" refId="B35" refString="Vijayakumar, SP, David, P, 2006. Taxonomy, natural history and distribution of the snakes of Nicobar Islands (India), based on new materials and with an emphasis on endemic species. Russian Journal of Herpetology 13: 11 - 40" title="Taxonomy, natural history and distribution of the snakes of Nicobar Islands (India), based on new materials and with an emphasis on endemic species." volume="13" year="2006">Vijayakumar and David 2006</bibRefCitation>
|
||
). See Table
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Comparison of the new species with Oligodon species bearing 17 DSR from south Asia. Missing data denoted by N. A." httpUri="http://table.plazi.org/id/C06C68436E088318161AA03E03E51D62" pageId="0" pageNumber="335" tableUuid="C06C68436E088318161AA03E03E51D62">3</tableCitation>
|
||
for a summary of comparison with south Asian species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
The new species is most similar to
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
in bearing a banded dorsum, 17 mid-dorsal scale rows and in shape of the Hemipenis. However, differs from
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis italics="true" pageId="0" pageNumber="335">sensu stricto</emphasis>
|
||
(see Discussion) in bearing a loreal shield (vs. absent in
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
); palatine and pterygoid teeth absent (vs, present in
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micro CT scan images of the skull, (a-c) Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019; (d-f) Oligodon arnensis NCBS NRC-AA- 020 showing dorsal view (a, d), lateral view (b, e) and ventral view (c, f)." httpUri="https://binary.pensoft.net/fig/606331" pageId="0" pageNumber="335">4d-f</figureCitation>
|
||
); maxillary teeth gradually increase in size posteriorly (vs. posterior most teeth enlarged abruptly, in
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
); ventral scales white with brown smear along the width of each scale, the smear is darker on the lateral edges forming a blotch on each side (vs. ventral scales white, lacking markings in
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
); 48 to 54 bands in total on the body (vs. ≤40 in
|
||
<taxonomicName genus="O." lsidName="O. arnensis" pageId="0" pageNumber="335" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">O. arnensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="335" type="description">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
Description of female holotype NCBS NRC-AA-0019 (Figure
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019 (a) dorsal view, (b) ventral view. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/606329" pageId="0" pageNumber="335">2</figureCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
The specimen is in good condition preserved in a coil with its head resting outside the coil. The specimen bears a single longitudinal incision (spanning over nine ventral scales) (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019 (a) dorsal view, (b) ventral view. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/606329" pageId="0" pageNumber="335">2a, b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/606329" pageId="0" pageNumber="335" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype female NCBS NRC-AA-019 (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a</emphasis>
|
||
) dorsal view, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b</emphasis>
|
||
) ventral view. Scale bars: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
Head short, measuring 10.1 mm from snout to the posterior tip to the parietal scale, comprising 3.6% of total length; high, 5 mm, with steeply domed snout in lateral view; upper jaw visible from ventral side. Head of the same width as the neck (width 6.8 mm). Snout gradually tapering to blunt, rounded tip in dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3a</figureCitation>
|
||
). Rostral subtriangular, slightly visible when viewed from top; wider (2.5 mm) than deep (1.6 mm). Nostrils small, elliptical shaped, present in the posterior border of the anterior nasal. The posterior nasal borders the nostrils. Paired internasals, wider (2 mm) than long (1.1 mm); smaller than prefrontals. Prefrontals, wider (2.5 mm) than long (1.7 mm). Frontal roughly hexagonal, bell shaped, 2.7 mm at the widest anterior border, median length 3.8 mm. Parietals 4.2 mm long, 3.2 mm at its widest anterior border. Temporals 1+2, anterior temporal 2.2 mm long and 0.9 mm wide, posterior upper temporal 2.3 mm long and 0.9 mm wide and the lower posterior temporal 1.6 mm and 1 mm long and wide respectively. Six nuchal scales, slightly larger than adjacent dorsal scales, bordering parietals. Supraocular larger than preocular; preocular large, deeper (2.5 mm) than wide (1.5 mm). Loreal roughly pentagonal, longer (1 mm) than high (0.6 mm). Two postoculars, subequal. Eye circular, 1.9 mm diameter of the eye with a circular pupil. Seven supralabials, third and fourth in contact with eye (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3c, d</figureCitation>
|
||
). Supralabial increase in size gradually, fifth and seventh larger than the rest. Second supralabial in contact with posterior nasal, loreal and first + third supralabials. Sixth supralabial deeply embedded between fifth and seventh supralabial, remotely making contact with the upper jawline on the right lateral side of the head (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3c</figureCitation>
|
||
). The sixth supralabial makes broad contact with the jawline on the left side of the head (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype female NCBS NRC-AA-019, view of the head, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a</emphasis>
|
||
) dorsal, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b</emphasis>
|
||
) ventral, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">c</emphasis>
|
||
) lateral right, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">d</emphasis>
|
||
) lateral left.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
Mental short, triangular, wider (1.9) than long (1.2). Infralabials 7, first really long, II to VI infralabials short and thin, fifth onwards larger (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3b</figureCitation>
|
||
). First four infralabials in contact with the genials. Anterior genials almost one and a half times as long (2.5 mm) as wide (1.7 mm); posterior genials as long as wide (1.2 mm) (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019, view of the head, (a) dorsal, (b) ventral, (c) lateral right, (d) lateral left." httpUri="https://binary.pensoft.net/fig/606330" pageId="0" pageNumber="335">3b</figureCitation>
|
||
). The right genials are fused.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">Body rounded, compressed, ventral surface distinctly flattened. Dorsal scales in 17-17-15 rows. First lateral reduction observed after the 10th ventral is at 50% of the ventrals where third and fourth DSR are involved in reduction from 17 DSR to 16 at ventral 85, thereafter from 16 DSR to 15 at the 90th ventral, row 2 +3 involved (52.9%). Dorsal scales imbricate, regularly arranged, vertebral scales not enlarged. All body scales smooth and glossy, without apical pit. Ventral scales 175 in number excluding two preventrals. Anal shield divided, slightly larger than last ventral scale. Subcaudals paired, 46 in number. Tail terminates in a sharp, tapering apical spine. Total length 335 mm, tail length 60 mm, tail/total length ratio 0.18.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="335" type="colouration">
|
||
<paragraph pageId="0" pageNumber="335">Colouration.</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
'In
|
||
<normalizedToken originalValue="preservative’">preservative'</normalizedToken>
|
||
Overall in a shade of pale brown with broad dark brown to black bands throughout the body. Each dorsal scale bears dark brown mottling. The first band is thin that runs across the internasals, the second in shape of a chevron that runs from fourth supralabial along the eye on the either side of the head and joins at the prefrontals. The third one, runs from the posterior of the angle of the jaw along the posterior temporals and further along the parietal and meets the frontal. The fourth broad chevron mark (4-5 scales wide) meets at the posterior portion of the parietal. The dorsal broad black bands are 1-2 scale wide edged with cream colour. The bands in the anterior 1/3rd of the SVL are not connected at the vertebral scale row. Each dorsal band is spaced by 3-4 dorsal scales. Forty-five bands on the dorsum from the nape to the vent and nine bands on the tail. Ventral scales white with brown smear along the width of each scale, the smear is darker on the lateral edges forming a blotch on each side.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
Skull features: A complete and robust skull typical of members of the family
|
||
<taxonomicName authorityName="Oppell" authorityYear="1811" class="Reptilia" family="Colubridae" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="family">Colubridae</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Cundall, D" journalOrPublisher="Taylor & Francis Group, London" pageId="0" pageNumber="335" refId="B6" refString="Cundall, D, Irish, F, 2008. The snake skull. In: Gaunt AS, Adler K (Eds) The skull of Lepidosauria, 349-692." title="The snake skull. In: Gaunt AS, Adler K (Eds) The skull of Lepidosauria, 349 - 692." year="2008">Cundall and Irish 2008</bibRefCitation>
|
||
). Maxilla with 8-10 functional teeth, gradually increasing in size posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micro CT scan images of the skull, (a-c) Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019; (d-f) Oligodon arnensis NCBS NRC-AA- 020 showing dorsal view (a, d), lateral view (b, e) and ventral view (c, f)." httpUri="https://binary.pensoft.net/fig/606331" pageId="0" pageNumber="335">4b</figureCitation>
|
||
). The palatine bone is slender, lacking teeth so is the pterygoid (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micro CT scan images of the skull, (a-c) Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019; (d-f) Oligodon arnensis NCBS NRC-AA- 020 showing dorsal view (a, d), lateral view (b, e) and ventral view (c, f)." httpUri="https://binary.pensoft.net/fig/606331" pageId="0" pageNumber="335">4c</figureCitation>
|
||
). Postorbital bone slender, conical, directed downwards, not making contact with the maxillary bone.
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/606331" pageId="0" pageNumber="335" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Figure 4.</emphasis>
|
||
Micro CT scan images of the skull, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a-c</emphasis>
|
||
)
|
||
<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype female NCBS NRC-AA-019; (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">d-f</emphasis>
|
||
)
|
||
<taxonomicName baseAuthorityName="Shaw" baseAuthorityYear="1802" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon arnensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="arnensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon arnensis</emphasis>
|
||
</taxonomicName>
|
||
NCBS NRC-AA-020 showing dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a, d</emphasis>
|
||
), lateral view (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b, e</emphasis>
|
||
) and ventral view (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">c, f</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="335" type="hemipenial morphology of paratype male bnhs 3657 (everted organs, n=2)">
|
||
<paragraph pageId="0" pageNumber="335">Hemipenial morphology of paratype male BNHS 3657 (everted organs, N=2):</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">The hemipenis is fully everted and expanded. The organ is stout, slightly bilobed, and semicapitate; lobes extend to about 20% of the hemipenis. The organ is spinous throughout; the spines are larger near the base and mid body and gradually decreases in size distally; sulcus spermaticus deep, bifurcating on the terminal fifth of the hemipenial body, with centrolineal orientation; the spine line on either side of the sulcus spermaticus is weak; hemipenial base is nude, with few spinnules.</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/606333" pageId="0" pageNumber="335" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Figure 5.</emphasis>
|
||
Hemipenis of
|
||
<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratype male BNHS 3657 (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a</emphasis>
|
||
) sulcal view, (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b</emphasis>
|
||
) asulcal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="335" type="natural history">
|
||
<paragraph pageId="0" pageNumber="335">Natural history notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
The holotype and the paratype were found actively moving along a mud road around 20:00 hours (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Oligodon churahensis sp. nov. holotype female NCBS NRC-AA- 019 in life (a); a general view of the habitat at the type locality (b), Photos by Virender Kumar." httpUri="https://binary.pensoft.net/fig/606335" pageId="0" pageNumber="335">6b</figureCitation>
|
||
). Additionally, five more individuals (uncollected) of the species were encountered at the same locality between the last weeks of May to late June 2020. All the individuals were found near the village after dusk. Sympatric reptile species recorded at the locality include
|
||
<taxonomicName authorityName="Agarwal, Khandekar & Bauer" authorityYear="2018" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus chamba" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="chamba">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Cyrtodactylus chamba</emphasis>
|
||
</taxonomicName>
|
||
Agarwal, Khandekar & Bauer, 2018,
|
||
<taxonomicName class="Reptilia" family="Scincidae" genus="Asymblepharus" kingdom="Animalia" lsidName="Asymblepharus" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Asymblepharus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName authorityName="Wall" authorityYear="1906" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" lsidName="Lycodon mackinnoni" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="mackinnoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Lycodon mackinnoni</emphasis>
|
||
</taxonomicName>
|
||
Wall, 1906,
|
||
<taxonomicName baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Reptilia" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius himalayanus" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="himalayanus">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Gloydius himalayanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
, 1864),
|
||
<taxonomicName class="Reptilia" family="Colubridae" genus="Orthriophis" kingdom="Animalia" lsidName="Orthriophis hodgsonii" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="hodgsonii">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Orthriophis hodgsonii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
, 1864),
|
||
<taxonomicName authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" lsidName="Laudakia" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Laudakia</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName authorityName="Fitzinger" authorityYear="1843" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" lsidName="Liopeltis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Liopeltis</emphasis>
|
||
</taxonomicName>
|
||
sp. None of the observed individuals showed any sign of aggression nor did they try to bite except for the male paratype, which bit one of us upon capture.
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/606335" pageId="0" pageNumber="335" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="Mirza & Bhardwaj & Patel" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Oligodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligodon churahensis" order="Squamata" pageId="0" pageNumber="335" phylum="Chordata" rank="species" species="churahensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="335">Oligodon churahensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype female NCBS NRC-AA-019 in life (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a</emphasis>
|
||
); a general view of the habitat at the type locality (
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b</emphasis>
|
||
), Photos by Virender Kumar.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="335" type="variation">
|
||
<paragraph pageId="0" pageNumber="335">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
The paratype male agrees with the description of the female holotype except for differences listed here: V 170 (+2 preventrals) Sc 47, first 8 subcaudals undivided thereafter the 9th subcaudal is divided followed by three undivided subcaudals and the rest are divided (13th onwards). The colouration of the male is much darker in comparison with the holotype. The number of dark bands on the dorsum in the paratype male is 37 bands from nape to the vent and 11 on the tail. Variation in the scale reduction formula is presented in Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Scale reduction formula for the type specimens, (a) holotype female NCBS NRC-AA- 019 (b) paratype male BNHS 3657." httpUri="http://table.plazi.org/id/25D23A1E25F8D6197FA6174A7048BE62" pageId="0" pageNumber="335" tableUuid="25D23A1E25F8D6197FA6174A7048BE62">1</tableCitation>
|
||
(Suppl. material 3).
|
||
</paragraph>
|
||
<caption ID-Table-UUID="25D23A1E25F8D6197FA6174A7048BE62" httpUri="http://table.plazi.org/id/25D23A1E25F8D6197FA6174A7048BE62" pageId="0" pageNumber="335" start="Table 1" startId="T1">
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<emphasis bold="true" pageId="0" pageNumber="335">Table 1.</emphasis>
|
||
Scale reduction formula for the type specimens, (a) holotype female NCBS NRC-AA-019 (b) paratype male BNHS 3657.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="335">
|
||
<table pageId="0" pageNumber="335">
|
||
<tr pageId="0" pageNumber="335">
|
||
<td colspan="1" pageId="0" pageNumber="335" rowspan="1">
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="335">a</emphasis>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="335" rowspan="1">17 (10) 2 + 3 (90) 3 + 4 (85) 15 (170)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="335">
|
||
<td colspan="1" pageId="0" pageNumber="335" rowspan="1">
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="335">b</emphasis>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="335" rowspan="1">17 (10) 3 + 4 (88) 3 + 4 (87) 15 (175)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |