treatments-xml/data/C0/60/87/C06087AC0E065501FF68C250FE1A0AB0.xml

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<mods:title id="875E8D7ACBC660437F9DE9A4D7325A14">Description of Rhinobatos ranongensis sp. nov. (Rhinopristiformes: Rhinobatidae) from the Andaman Sea and Bay of Bengal with a review of its northern Indian Ocean congeners</mods:title>
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<taxonomicName id="8FC94D390E065504FF68C250FD9C0D2B" authority="Norman 1926" authorityName="Norman" authorityYear="1926" box="[151,572,1340,1366]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="265" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E065504FF68C250FD9C0D2B" bold="true" box="[151,572,1340,1366]" pageId="8" pageNumber="265">
<emphasis id="7ABDEAA80E065504FF68C250FE2A0D2B" bold="true" box="[151,394,1340,1366]" italics="true" pageId="8" pageNumber="265">Rhinobatos lionotus</emphasis>
<bibRefCitation id="2C584B4B0E065504FE6DC250FD9C0D2B" author="Norman, J. R." box="[402,572,1340,1366]" pageId="8" pageNumber="265" pagination="941 - 982" refId="ref12964" refString="Norman, J. R. (1926) A synopsis of the rays of the family Rhinobatidae, with a revision of the genus Rhinobatus. Proceedings of the Zoological Society of London, 1926 (4), 941 - 982. https: // doi. org / 10.1111 / j. 1096 - 3642.1926. tb 02228. x" type="journal article" year="1926">Norman 1926</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="487636BA0E065504FF68C232FE3C0D0A" blockId="8.[151,572,1340,1435]" box="[151,412,1374,1399]" pageId="8" pageNumber="265">Smoothback Guitarfish</paragraph>
</subSubSection>
<subSubSection id="00D365310E065504FF68C2EEFBF60D9D" pageId="8" pageNumber="265" type="description">
<paragraph id="487636BA0E065504FF68C2EEFE610DE6" blockId="8.[151,572,1340,1435]" box="[151,449,1410,1435]" pageId="8" pageNumber="265">
(
<figureCitation id="D0F22A3F0E065504FF60C2EEFF470DE6" box="[159,231,1410,1435]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[177,1416,398,1898]" captionTargetId="figure@11.[151,1436,331,1929]" captionTargetPageId="11" captionText="FIGURE 4. Rhinobatos lionotus: A. female holotype, 495 mm TL (BMNH 1909.7.12.3), India, dorsal view, preserved; female non-type, 760 mm TL (CSIRO H 7867-01), Myanmar, fresh, B. dorsal view, C. ventral view." httpUri="https://zenodo.org/record/2624902/files/figure.png" pageId="8" pageNumber="265">Figs 4</figureCitation>
, 
<figureCitation id="D0F22A3F0E065504FF0CC2EEFEA30DE6" box="[243,259,1410,1435]" captionStart="FIGURE 5" captionStartId="12.[151,250,1976,1998]" captionTargetBox="[330,1252,204,1907]" captionTargetId="figure@12.[151,1436,163,1977]" captionTargetPageId="12" captionText="FIGURE 5. Lateral views of the unpaired fins of Rhinobatos lionotus, female non-type, 760 mm TL (CSIRO H 7867-01), Myanmar, preserved: A. first dorsal fin; B. second dorsal fin; and C. caudal fin." httpUri="https://zenodo.org/record/2624904/files/figure.png" pageId="8" pageNumber="265">5</figureCitation>
, 
<figureCitation id="D0F22A3F0E065504FEF0C2EFFEF80DE6" box="[271,344,1411,1435]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="22.[151,250,1947,1969]" captionStartId-1="23.[151,250,1146,1168]" captionStartId-2="24.[151,250,1492,1514]" captionTargetBox-0="[177,1416,181,1892]" captionTargetBox-1="[172,1396,233,1104]" captionTargetBox-2="[156,1412,525,1459]" captionTargetId-0="figure@22.[151,1436,161,1975]" captionTargetId-1="figure@23.[167,1420,223,1124]" captionTargetId-2="figure@24.[151,1436,517,1471]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionText-0="FIGURE 10. View of dorsal head (preserved): A. Rhinobatos ranongensis sp. nov., adult male holotype 645 mm TL (CSIRO H 7861-02); B. Rhinobatos punctifer, female 570 mm TL (MNHN 2014-0156); C. Rhinobatos annandalei, female 870 mm TL (CSIRO H 7866-01); and D. Rhinobatos lionotus, female 760 mm TL (CSIRO H 7867-01)." captionText-1="FIGURE 11. Views of oronasal region (preserved): A. Rhinobatos ranongensis sp. nov., adult male holotype 645 mm TL (CSIRO H 7861-02); B. Rhinobatos punctifer, female 570 mm TL (MNHN 2014-0156); C. Rhinobatos annandalei, female 870 mm TL (CSIRO H 7866-01); and D. Rhinobatos lionotus, female 760 mm TL (CSIRO H 7867-01)." captionText-2="FIGURE 12. Dorsal view of skin at scapulocoracoids of: A. Rhinobatos ranongensis sp. nov., adult male holotype 645 mm TL (CSIRO H 7861-02); B. Rhinobatos punctifer, adult male 643 mm TL (MNHN 2014-0157); C. Rhinobatos annandalei female 870 mm TL (CSIRO H 7867-01); and D. Rhinobatos lionotus female 760 mm TL (CSIRO H 7866-01)." httpUri-0="https://zenodo.org/record/2624916/files/figure.svg" httpUri-1="https://zenodo.org/record/2624918/files/figure.png" httpUri-2="https://zenodo.org/record/2624920/files/figure.png" pageId="8" pageNumber="265">10–12</figureCitation>
; 
<tableCitation id="054B03010E065504FE9BC2EEFE180DE6" box="[356,440,1410,1435]" captionStart="TABLE 2" captionStartId="9.[151,239,652,675]" captionText="TABLE 2. Morphometric data for the female holotype (BMNH 1909.7.12.3) and newly acquired specimen (CSIRO H 7867-01) of Rhinobatos lionotus, and adult male holotype (CSIRO H 7861-02) and the three paratypes (CSIRO H 8403- 01, 8404-01, 8404-02) of R. ranongensis sp. nov." pageId="8" pageNumber="265">Table 2</tableCitation>
)
</paragraph>
<paragraph id="487636BA0E065504FF68C2A6FBF60D9D" blockId="8.[151,1297,1482,1566]" box="[151,1110,1482,1504]" pageId="8" pageNumber="265">
<taxonomicName id="8FC94D390E065504FF68C2A6FECC0D9D" box="[151,364,1482,1504]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="265" phylum="Chordata" rank="species" species="schlegelii">
<emphasis id="7ABDEAA80E065504FF68C2A6FECC0D9D" box="[151,364,1482,1504]" italics="true" pageId="8" pageNumber="265">Rhinobatis schlegelii</emphasis>
</taxonomicName>
(
<emphasis id="7ABDEAA80E065504FE84C2A7FE3E0D9D" box="[379,414,1483,1504]" italics="true" pageId="8" pageNumber="265">nec</emphasis>
Muller &amp; Henle): Annandale, 1909, Memoirs Indian Museum 2, 15.
</paragraph>
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<subSubSection id="00D365310E065504FF68C285FAB10E63" pageId="8" pageNumber="265" type="reference_group">
<paragraph id="487636BA0E065504FF68C285FC520D82" blockId="8.[151,1297,1482,1566]" box="[151,1010,1513,1535]" pageId="8" pageNumber="265">
<taxonomicName id="8FC94D390E065504FF68C285FE5D0D82" authority="Norman, 1926" authorityName="Norman" authorityYear="1926" box="[151,509,1513,1535]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="265" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E065504FF68C285FEC20D82" box="[151,354,1513,1535]" italics="true" pageId="8" pageNumber="265">Rhinobatus lionotus</emphasis>
<bibRefCitation id="2C584B4B0E065504FE97C286FE5D0D82" author="Norman, J. R." box="[360,509,1513,1535]" pageId="8" pageNumber="265" pagination="941 - 982" refId="ref12964" refString="Norman, J. R. (1926) A synopsis of the rays of the family Rhinobatidae, with a revision of the genus Rhinobatus. Proceedings of the Zoological Society of London, 1926 (4), 941 - 982. https: // doi. org / 10.1111 / j. 1096 - 3642.1926. tb 02228. x" type="journal article" year="1926">Norman, 1926</bibRefCitation>
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, Proc. Zool. Soc. London 1926 (62): 961, fig. 14.
</paragraph>
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<taxonomicName id="8FC94D390E065504FF68C164FE580E63" authority=": Fowler, 1941" authorityName=": Fowler" authorityYear="1941" box="[151,504,1544,1566]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="265" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E065504FF68C164FEC20E63" box="[151,354,1544,1566]" italics="true" pageId="8" pageNumber="265">Rhinobatos lionotus</emphasis>
: Fowler, 1941
</taxonomicName>
, 310; 
<bibRefCitation id="2C584B4B0E065504FDCAC164FCA50E63" author="Akhilesh, K. V. &amp; Bineesh, K. K. &amp; Gopalakrishnan, A. &amp; Jean, J. K. &amp; Basherre, V. S. &amp; Pillai, N. G. K." box="[565,773,1544,1566]" pageId="8" pageNumber="265" pagination="109 - 120" refId="ref11919" refString="Akhilesh, K. V., Bineesh, K. K., Gopalakrishnan, A., Jean, J. K., Basherre, V. S. &amp; Pillai, N. G. K. (2014) Checklist of Chondrichthyans in Indian waters. Journal of Marine Biology Association India, 56 (1), 109 - 120." type="journal article" year="2014">
Akhilesh 
<emphasis id="7ABDEAA80E065504FD68C165FD6A0E63" box="[663,714,1544,1566]" italics="true" pageId="8" pageNumber="265">et al.</emphasis>
2014
</bibRefCitation>
, Checklist Chondrichthyans 
<collectingCountry id="30DE762A0E065504FBD6C164FBC10E63" box="[1065,1121,1544,1566]" name="India" pageId="8" pageNumber="265">India</collectingCountry>
: 115-118 (listed).
</paragraph>
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<subSubSection id="00D365310E065504FF68C126FE860EB2" pageId="8" pageNumber="265" type="materials_examined">
<paragraph id="487636BA0E065504FF68C126FE860EB2" blockId="8.[151,1437,1610,2031]" pageId="8" pageNumber="265">
<emphasis id="7ABDEAA80E065504FF68C126FE210E1E" bold="true" box="[151,385,1610,1635]" pageId="8" pageNumber="265">Material examined.</emphasis>
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, 
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.12.3 [ex 
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], female 
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TL, mouth of 
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, 
<collectingCountry id="30DE762A0E065504FAA4C126FA370E1E" box="[1371,1431,1610,1635]" name="India" pageId="8" pageNumber="265">India</collectingCountry>
, ~ 
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depth; 
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<specimenCode id="186F9EC10E065504FE6EC103FDAD0EFA" box="[401,525,1646,1671]" collectionCode="CSIRO" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="8" pageNumber="265" type="Museum">H 7867-01</specimenCode>
, female 
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TL, 
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, 
<locationDeviation id="59C42F980E065504FC46C103FB210EFB" box="[953,1153,1646,1671]" location="Andaman Is" pageId="8" pageNumber="265">N of Andaman Is</locationDeviation>
, 
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, 
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, 
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, bottom trawl, 
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depth, 
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station
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60, specimen 247, collected 
<collectorName id="E53C536C0E065504FB0AC1FFFA380ED6" box="[1269,1432,1682,1707]" pageId="8" pageNumber="265">P. Psomadakis</collectorName>
, 
<date id="3C77107A0E065504FF68C1DBFE820EB2" box="[151,290,1718,1743]" pageId="8" pageNumber="265" value="2015-05-09">
<collectingDate id="2C33E9920E065504FF68C1DBFE820EB2" box="[151,290,1718,1743]" pageId="8" pageNumber="265" value="2015-05-09">9 May 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="00D365310E065505FF38C1B6FE310989" lastPageId="9" lastPageNumber="266" pageId="8" pageNumber="265" type="diagnosis">
<paragraph id="487636BA0E065505FF38C1B6FE310989" blockId="8.[151,1437,1610,2031]" lastBlockId="9.[151,1437,151,608]" lastPageId="9" lastPageNumber="266" pageId="8" pageNumber="265">
<emphasis id="7ABDEAA80E065504FF38C1B6FEE10E8E" bold="true" box="[199,321,1754,1779]" pageId="8" pageNumber="265">Diagnosis.</emphasis>
A medium-sized species of the genus 
<taxonomicName id="8FC94D390E065504FD08C1B7FCD60E8E" box="[759,886,1755,1779]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="265" phylum="Chordata" rank="genus">
<emphasis id="7ABDEAA80E065504FD08C1B7FCD60E8E" box="[759,886,1755,1779]" italics="true" pageId="8" pageNumber="265">Rhinobatos</emphasis>
</taxonomicName>
(attaining at least 
<quantity id="8F319B5F0E065504FBB5C1B6FB080E8F" box="[1098,1192,1754,1779]" metricMagnitude="-1" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="265" unit="mm" value="760.0">760 mm</quantity>
TL) distinguished by the following combination of characters: disc wedge-shaped, rather large, width 35–37% TL, length 1.2–1.3 times width; snout relatively short, length 2.5–2.7 times interspiracular distance, ~3.4 times interorbital width; orbit diameter 1.2–1.4 times spiracle length; nostrils moderately oblique, their length 1.3–1.5 times internarial distance; mouth width 6.3–6.9% TL; preoral length 5.5–5.9 times internarial distance; posterior nasal flaps broad; two spiracular folds, outermost fold subequal to marginally taller than inner fold; ridges of rostral cartilage almost parallel, widely separated, converging slightly anteriorly but not constricted medially; anterior cartilage weakly sickle shaped, almost truncate posteriorly; distance between fifth gill slits 2.5–2.7 times in ventral head length; prebranchial sensory pore system obvious, extending posteriorly to margin of first gill slit; postscapular sensory canal notched, with exposed lateral pores; weak thorn patches on supraorbit and scapular region, and row of rudimentary thorns (not conspicuous) along dorsal midline; denticles on dorsal fins densest anteriorly, more sparse posteriorly; dorsal fins small, height of first 7.7–8.1% TL; pelvic-fin inner margin subequal to or shorter than its base length; interdorsal distance 2.3–2.9 times first dorsal-fin base; dorsal caudal margin 1.9–2.0 times preventral margin; upper jaw with ~118 tooth rows; snout angle ~61°; 68–72 pectoral radials; ~164 post-synarcual centra; ~49 nasal lamellae; dorsal fins brownish anteriorly, dusky posteriorly and apically; dorsal disc of adults uniformly brown to greenish, with or without large, regularly spaced cloudy blotches (faint orange to dusky, more or less symmetrically arranged and of irregular sizes); ventral disc and tail pale; no large, black, teardrop marking on undersurface of snout.
</paragraph>
</subSubSection>
<subSubSection id="00D365310E075505FF38C693FE6C0A1D" pageId="9" pageNumber="266" type="distribution">
<paragraph id="487636BA0E075505FF38C693FE6C0A1D" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="266">
<emphasis id="7ABDEAA80E075505FF38C693FEC00A65" bold="true" box="[199,352,511,536]" pageId="9" pageNumber="266">Distribution.</emphasis>
Off West Bengal in the Bay of Bengal (Hoogly River mouth) to the northwestern Andaman Sea (off Pathein Peninsula, 
<collectingCountry id="30DE762A0E075505FE62C548FDAF0A41" box="[413,527,548,572]" name="Myanmar" pageId="9" pageNumber="266">Myanmar</collectingCountry>
). Possibly more widespread in the Bay of Bengal. Benthic inshore on continental shelf to at least 
<quantity id="8F319B5F0E075505FEB9C52BFE200A22" box="[326,384,583,608]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="266" unit="m" value="76.0">76 m</quantity>
depth.
</paragraph>
</subSubSection>
<caption id="1CB666320E075505FF68C5E0FD070A9E" pageId="9" pageNumber="266">
<paragraph id="487636BA0E075505FF68C5E0FD070A9E" blockId="9.[151,1435,652,739]" pageId="9" pageNumber="266">
<emphasis id="7ABDEAA80E075505FF68C5E0FEAA0ADE" bold="true" box="[151,266,652,675]" pageId="9" pageNumber="266">TABLE 2.</emphasis>
Morphometric data for the female holotype (BMNH 1909.7.12.3) and newly acquired specimen (CSIRO H 7867-01) of 
<taxonomicName id="8FC94D390E075505FEE3C5C7FE520ABE" authorityName=": Fowler" authorityYear="1941" box="[284,498,683,707]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="266" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E075505FEE3C5C7FE520ABE" box="[284,498,683,707]" italics="true" pageId="9" pageNumber="266">Rhinobatos lionotus</emphasis>
</taxonomicName>
, and adult male holotype (CSIRO H 7861-02) and the three paratypes (CSIRO H 8403- 0 1, 8404-01, 8404-02) of 
<taxonomicName id="8FC94D390E075505FE55C5A0FDEB0A9E" authority="Last &amp; Séret &amp; Naylor, 2019" authorityName="Last &amp; Séret &amp; Naylor" authorityYear="2019" box="[426,587,716,739]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="266" phylum="Chordata" rank="species" species="ranongensis" status="sp. nov.">
<emphasis id="7ABDEAA80E075505FE55C5A0FDEB0A9E" box="[426,587,716,739]" italics="true" pageId="9" pageNumber="266">R. ranongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="618E57D30E075505FDAEC5A0FD070A9E" box="[593,679,716,739]" pageId="9" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="487636BA0E075505FD88C46CFAD70FBE" pageId="9" pageNumber="266">
<table id="3AC9C41A0E07AAF3FF60C592FA250FBE" box="[159,1413,766,1987]" colsContinueIn="10.[159,1398,201,1794]" gridcols="6" gridrows="30" pageId="9" pageNumber="266">
<tr id="F6F934F80E07AAF3FF60C592FA250B68" box="[159,1413,766,789]" gridrow="0" pageId="9" pageNumber="266" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="B5285D840E07AAF3FD88C592FD450B68" box="[631,741,766,789]" gridcol="1" gridrow="0" pageId="9" pageNumber="266">
<taxonomicName id="8FC94D390E075505FD88C46CFD450B69" authorityName=": Fowler" authorityYear="1941" box="[631,741,766,789]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="266" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E075505FD88C46CFD450B69" box="[631,741,766,789]" italics="true" pageId="9" pageNumber="266">R. lionotus</emphasis>
</taxonomicName>
</th>
<th id="B5285D840E07AAF3FC05C592FB320B68" box="[1018,1170,766,789]" gridcol="3" gridrow="0" pageId="9" pageNumber="266">
<taxonomicName id="8FC94D390E075505FC05C46CFB320B68" authorityName="Last &amp; Séret &amp; Naylor" authorityYear="2019" box="[1018,1170,768,789]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="266" phylum="Chordata" rank="species" species="ranongensis">
<emphasis id="7ABDEAA80E075505FC05C46CFB320B68" box="[1018,1170,768,789]" italics="true" pageId="9" pageNumber="266">R. ranongensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="F6F934F80E07AAF3FF60C444FA250B43" box="[159,1413,808,830]" gridrow="1" pageId="9" pageNumber="266" rowspan-0="1">
<td id="B5285D840E07AAF3FD88C444FD450B43" box="[631,741,808,830]" gridcol="1" gridrow="1" pageId="9" pageNumber="266">Holotype</td>
<td id="B5285D840E07AAF3FCF4C444FC680B43" box="[779,968,808,830]" gridcol="2" gridrow="1" pageId="9" pageNumber="266">CSIRO H 7867-01</td>
<td id="B5285D840E07AAF3FC05C444FB320B43" box="[1018,1170,808,830]" gridcol="3" gridrow="1" pageId="9" pageNumber="266">Holotype</td>
<td id="B5285D840E07AAF3FB28C444FA250B43" box="[1239,1413,808,830]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="9" pageNumber="266">Paratypes</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C43EFA250B1A" box="[159,1413,850,871]" gridrow="2" pageId="9" pageNumber="266" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="B5285D840E07AAF3FB28C43EFAA80B1A" box="[1239,1288,850,871]" gridcol="4" gridrow="2" pageId="9" pageNumber="266">MIN</td>
<td id="B5285D840E07AAF3FAB6C43EFA250B1A" box="[1353,1413,850,871]" gridcol="5" gridrow="2" pageId="9" pageNumber="266">MAX</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C416FA250BED" box="[159,1413,890,912]" gridrow="3" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C416FD8F0BED" box="[159,559,890,912]" gridcol="0" gridrow="3" pageId="9" pageNumber="266">Total length (mm)</th>
<td id="B5285D840E07AAF3FD88C416FD450BED" box="[631,741,890,912]" gridcol="1" gridrow="3" pageId="9" pageNumber="266">495</td>
<td id="B5285D840E07AAF3FCF4C416FC680BED" box="[779,968,890,912]" gridcol="2" gridrow="3" pageId="9" pageNumber="266">760</td>
<td id="B5285D840E07AAF3FC05C416FB320BED" box="[1018,1170,890,912]" gridcol="3" gridrow="3" pageId="9" pageNumber="266">645</td>
<td id="B5285D840E07AAF3FB28C416FAA80BED" box="[1239,1288,890,912]" gridcol="4" gridrow="3" pageId="9" pageNumber="266">392</td>
<td id="B5285D840E07AAF3FAB6C416FA250BED" box="[1353,1413,890,912]" gridcol="5" gridrow="3" pageId="9" pageNumber="266">494</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C4C8FA250BC7" box="[159,1413,932,954]" gridrow="4" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C4C8FD8F0BC7" box="[159,559,932,954]" gridcol="0" gridrow="4" pageId="9" pageNumber="266">Disc width—maximum</th>
<td id="B5285D840E07AAF3FD88C4C8FD450BC7" box="[631,741,932,954]" gridcol="1" gridrow="4" pageId="9" pageNumber="266">35.4</td>
<td id="B5285D840E07AAF3FCF4C4C8FC680BC7" box="[779,968,932,954]" gridcol="2" gridrow="4" pageId="9" pageNumber="266">36.9</td>
<td id="B5285D840E07AAF3FC05C4C8FB320BC7" box="[1018,1170,932,954]" gridcol="3" gridrow="4" pageId="9" pageNumber="266">30.5</td>
<td id="B5285D840E07AAF3FB28C4C8FAA80BC7" box="[1239,1288,932,954]" gridcol="4" gridrow="4" pageId="9" pageNumber="266">31.4</td>
<td id="B5285D840E07AAF3FAB6C4C8FA250BC7" box="[1353,1413,932,954]" gridcol="5" gridrow="4" pageId="9" pageNumber="266">33.3</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C4A1FA250B9E" box="[159,1413,973,995]" gridrow="5" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C4A1FD8F0B9E" box="[159,559,973,995]" gridcol="0" gridrow="5" pageId="9" pageNumber="266">Disc length</th>
<td id="B5285D840E07AAF3FD88C4A1FD450B9E" box="[631,741,973,995]" gridcol="1" gridrow="5" pageId="9" pageNumber="266">44.4</td>
<td id="B5285D840E07AAF3FCF4C4A1FC680B9E" box="[779,968,973,995]" gridcol="2" gridrow="5" pageId="9" pageNumber="266">44.1</td>
<td id="B5285D840E07AAF3FC05C4A1FB320B9E" box="[1018,1170,973,995]" gridcol="3" gridrow="5" pageId="9" pageNumber="266">41.0</td>
<td id="B5285D840E07AAF3FB28C4A1FAA80B9E" box="[1239,1288,973,995]" gridcol="4" gridrow="5" pageId="9" pageNumber="266">40.1</td>
<td id="B5285D840E07AAF3FAB6C4A1FA250B9E" box="[1353,1413,973,995]" gridcol="5" gridrow="5" pageId="9" pageNumber="266">42.7</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C49AFA250C71" box="[159,1413,1014,1036]" gridrow="6" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C49AFD8F0C71" box="[159,559,1014,1036]" gridcol="0" gridrow="6" pageId="9" pageNumber="266">Head length—dorsal</th>
<td id="B5285D840E07AAF3FD88C49AFD450C71" box="[631,741,1014,1036]" gridcol="1" gridrow="6" pageId="9" pageNumber="266">29.5</td>
<td id="B5285D840E07AAF3FCF4C49AFC680C71" box="[779,968,1014,1036]" gridcol="2" gridrow="6" pageId="9" pageNumber="266">19.3</td>
<td id="B5285D840E07AAF3FC05C49AFB320C71" box="[1018,1170,1014,1036]" gridcol="3" gridrow="6" pageId="9" pageNumber="266">21.2</td>
<td id="B5285D840E07AAF3FB28C49AFAA80C71" box="[1239,1288,1014,1036]" gridcol="4" gridrow="6" pageId="9" pageNumber="266">21.0</td>
<td id="B5285D840E07AAF3FAB6C49AFA250C71" box="[1353,1413,1014,1036]" gridcol="5" gridrow="6" pageId="9" pageNumber="266">21.9</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C34CFA250C4B" box="[159,1413,1056,1078]" gridrow="7" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C34CFD8F0C4B" box="[159,559,1056,1078]" gridcol="0" gridrow="7" pageId="9" pageNumber="266">Head length—ventral</th>
<td id="B5285D840E07AAF3FD88C34CFD450C4B" box="[631,741,1056,1078]" gridcol="1" gridrow="7" pageId="9" pageNumber="266">28.7</td>
<td id="B5285D840E07AAF3FCF4C34CFC680C4B" box="[779,968,1056,1078]" gridcol="2" gridrow="7" pageId="9" pageNumber="266">26.9</td>
<td id="B5285D840E07AAF3FC05C34CFB320C4B" box="[1018,1170,1056,1078]" gridcol="3" gridrow="7" pageId="9" pageNumber="266">27.3</td>
<td id="B5285D840E07AAF3FB28C34CFAA80C4B" box="[1239,1288,1056,1078]" gridcol="4" gridrow="7" pageId="9" pageNumber="266">26.6</td>
<td id="B5285D840E07AAF3FAB6C34CFA250C4B" box="[1353,1413,1056,1078]" gridcol="5" gridrow="7" pageId="9" pageNumber="266">28.3</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C325FA250C22" box="[159,1413,1097,1119]" gridrow="8" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C325FD8F0C22" box="[159,559,1097,1119]" gridcol="0" gridrow="8" pageId="9" pageNumber="266">Snout length (presocket)</th>
<td id="B5285D840E07AAF3FD88C325FD450C22" box="[631,741,1097,1119]" gridcol="1" gridrow="8" pageId="9" pageNumber="266">13.5</td>
<td id="B5285D840E07AAF3FCF4C325FC680C22" box="[779,968,1097,1119]" gridcol="2" gridrow="8" pageId="9" pageNumber="266">14.0</td>
<td id="B5285D840E07AAF3FC05C325FB320C22" box="[1018,1170,1097,1119]" gridcol="3" gridrow="8" pageId="9" pageNumber="266">15.4</td>
<td id="B5285D840E07AAF3FB28C325FAA80C22" box="[1239,1288,1097,1119]" gridcol="4" gridrow="8" pageId="9" pageNumber="266">15.2</td>
<td id="B5285D840E07AAF3FAB6C325FA250C22" box="[1353,1413,1097,1119]" gridcol="5" gridrow="8" pageId="9" pageNumber="266">15.8</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C31EFA250CF5" box="[159,1413,1138,1160]" gridrow="9" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C31EFD8F0CF5" box="[159,559,1138,1160]" gridcol="0" gridrow="9" pageId="9" pageNumber="266">Orbit diameter</th>
<td id="B5285D840E07AAF3FD88C31EFD450CF5" box="[631,741,1138,1160]" gridcol="1" gridrow="9" pageId="9" pageNumber="266">4.2</td>
<td id="B5285D840E07AAF3FCF4C31EFC680CF5" box="[779,968,1138,1160]" gridcol="2" gridrow="9" pageId="9" pageNumber="266">3.7</td>
<td id="B5285D840E07AAF3FC05C31EFB320CF5" box="[1018,1170,1138,1160]" gridcol="3" gridrow="9" pageId="9" pageNumber="266">3.5</td>
<td id="B5285D840E07AAF3FB28C31EFAA80CF5" box="[1239,1288,1138,1160]" gridcol="4" gridrow="9" pageId="9" pageNumber="266">3.7</td>
<td id="B5285D840E07AAF3FAB6C31EFA250CF5" box="[1353,1413,1138,1160]" gridcol="5" gridrow="9" pageId="9" pageNumber="266">3.9</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C3F0FA250CCF" box="[159,1413,1180,1202]" gridrow="10" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C3F0FD8F0CCF" box="[159,559,1180,1202]" gridcol="0" gridrow="10" pageId="9" pageNumber="266">Spiracle length</th>
<td id="B5285D840E07AAF3FD88C3F0FD450CCF" box="[631,741,1180,1202]" gridcol="1" gridrow="10" pageId="9" pageNumber="266">3.1</td>
<td id="B5285D840E07AAF3FCF4C3F0FC680CCF" box="[779,968,1180,1202]" gridcol="2" gridrow="10" pageId="9" pageNumber="266">3.0</td>
<td id="B5285D840E07AAF3FC05C3F0FB320CCF" box="[1018,1170,1180,1202]" gridcol="3" gridrow="10" pageId="9" pageNumber="266">2.4</td>
<td id="B5285D840E07AAF3FB28C3F0FAA80CCF" box="[1239,1288,1180,1202]" gridcol="4" gridrow="10" pageId="9" pageNumber="266">2.4</td>
<td id="B5285D840E07AAF3FAB6C3F0FA250CCF" box="[1353,1413,1180,1202]" gridcol="5" gridrow="10" pageId="9" pageNumber="266">2.9</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C3A9FA250CA6" box="[159,1413,1221,1243]" gridrow="11" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C3A9FD8F0CA6" box="[159,559,1221,1243]" gridcol="0" gridrow="11" pageId="9" pageNumber="266">Orbit and spiracle length</th>
<td id="B5285D840E07AAF3FD88C3A9FD450CA6" box="[631,741,1221,1243]" gridcol="1" gridrow="11" pageId="9" pageNumber="266">5.4</td>
<td id="B5285D840E07AAF3FCF4C3A9FC680CA6" box="[779,968,1221,1243]" gridcol="2" gridrow="11" pageId="9" pageNumber="266">5.1</td>
<td id="B5285D840E07AAF3FC05C3A9FB320CA6" box="[1018,1170,1221,1243]" gridcol="3" gridrow="11" pageId="9" pageNumber="266">4.6</td>
<td id="B5285D840E07AAF3FB28C3A9FAA80CA6" box="[1239,1288,1221,1243]" gridcol="4" gridrow="11" pageId="9" pageNumber="266">4.7</td>
<td id="B5285D840E07AAF3FAB6C3A9FA250CA6" box="[1353,1413,1221,1243]" gridcol="5" gridrow="11" pageId="9" pageNumber="266">5.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C382FA250D79" box="[159,1413,1262,1284]" gridrow="12" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C382FD8F0D79" box="[159,559,1262,1284]" gridcol="0" gridrow="12" pageId="9" pageNumber="266">Interorbital width</th>
<td id="B5285D840E07AAF3FD88C382FD450D79" box="[631,741,1262,1284]" gridcol="1" gridrow="12" pageId="9" pageNumber="266">4.0</td>
<td id="B5285D840E07AAF3FCF4C382FC680D79" box="[779,968,1262,1284]" gridcol="2" gridrow="12" pageId="9" pageNumber="266">4.1</td>
<td id="B5285D840E07AAF3FC05C382FB320D79" box="[1018,1170,1262,1284]" gridcol="3" gridrow="12" pageId="9" pageNumber="266">3.5</td>
<td id="B5285D840E07AAF3FB28C382FAA80D79" box="[1239,1288,1262,1284]" gridcol="4" gridrow="12" pageId="9" pageNumber="266">3.5</td>
<td id="B5285D840E07AAF3FAB6C382FA250D79" box="[1353,1413,1262,1284]" gridcol="5" gridrow="12" pageId="9" pageNumber="266">3.9</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C274FA250D53" box="[159,1413,1304,1326]" gridrow="13" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C274FD8F0D53" box="[159,559,1304,1326]" gridcol="0" gridrow="13" pageId="9" pageNumber="266">Interspiracular width</th>
<td id="B5285D840E07AAF3FD88C274FD450D53" box="[631,741,1304,1326]" gridcol="1" gridrow="13" pageId="9" pageNumber="266">5.5</td>
<td id="B5285D840E07AAF3FCF4C274FC680D53" box="[779,968,1304,1326]" gridcol="2" gridrow="13" pageId="9" pageNumber="266">5.1</td>
<td id="B5285D840E07AAF3FC05C274FB320D53" box="[1018,1170,1304,1326]" gridcol="3" gridrow="13" pageId="9" pageNumber="266">4.9</td>
<td id="B5285D840E07AAF3FB28C274FAA80D53" box="[1239,1288,1304,1326]" gridcol="4" gridrow="13" pageId="9" pageNumber="266">5.1</td>
<td id="B5285D840E07AAF3FAB6C274FA250D53" box="[1353,1413,1304,1326]" gridcol="5" gridrow="13" pageId="9" pageNumber="266">5.1</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C22DFA250D2A" box="[159,1413,1345,1367]" gridrow="14" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C22DFD8F0D2A" box="[159,559,1345,1367]" gridcol="0" gridrow="14" pageId="9" pageNumber="266">Preoral length</th>
<td id="B5285D840E07AAF3FD88C22DFD450D2A" box="[631,741,1345,1367]" gridcol="1" gridrow="14" pageId="9" pageNumber="266">16.4</td>
<td id="B5285D840E07AAF3FCF4C22DFC680D2A" box="[779,968,1345,1367]" gridcol="2" gridrow="14" pageId="9" pageNumber="266">15.7</td>
<td id="B5285D840E07AAF3FC05C22DFB320D2A" box="[1018,1170,1345,1367]" gridcol="3" gridrow="14" pageId="9" pageNumber="266">18.2</td>
<td id="B5285D840E07AAF3FB28C22DFAA80D2A" box="[1239,1288,1345,1367]" gridcol="4" gridrow="14" pageId="9" pageNumber="266">17.6</td>
<td id="B5285D840E07AAF3FAB6C22DFA250D2A" box="[1353,1413,1345,1367]" gridcol="5" gridrow="14" pageId="9" pageNumber="266">19.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C206FA250DFD" box="[159,1413,1386,1408]" gridrow="15" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C206FD8F0DFD" box="[159,559,1386,1408]" gridcol="0" gridrow="15" pageId="9" pageNumber="266">Mouth width</th>
<td id="B5285D840E07AAF3FD88C206FD450DFD" box="[631,741,1386,1408]" gridcol="1" gridrow="15" pageId="9" pageNumber="266">6.9</td>
<td id="B5285D840E07AAF3FCF4C206FC680DFD" box="[779,968,1386,1408]" gridcol="2" gridrow="15" pageId="9" pageNumber="266">6.3</td>
<td id="B5285D840E07AAF3FC05C206FB320DFD" box="[1018,1170,1386,1408]" gridcol="3" gridrow="15" pageId="9" pageNumber="266">5.4</td>
<td id="B5285D840E07AAF3FB28C206FAA80DFD" box="[1239,1288,1386,1408]" gridcol="4" gridrow="15" pageId="9" pageNumber="266">5.5</td>
<td id="B5285D840E07AAF3FAB6C206FA250DFD" box="[1353,1413,1386,1408]" gridcol="5" gridrow="15" pageId="9" pageNumber="266">6.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C2F8FA250DD7" box="[159,1413,1428,1450]" gridrow="16" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C2F8FD8F0DD7" box="[159,559,1428,1450]" gridcol="0" gridrow="16" pageId="9" pageNumber="266">Prenarial distance</th>
<td id="B5285D840E07AAF3FD88C2F8FD450DD7" box="[631,741,1428,1450]" gridcol="1" gridrow="16" pageId="9" pageNumber="266">12.2</td>
<td id="B5285D840E07AAF3FCF4C2F8FC680DD7" box="[779,968,1428,1450]" gridcol="2" gridrow="16" pageId="9" pageNumber="266">12.1</td>
<td id="B5285D840E07AAF3FC05C2F8FB320DD7" box="[1018,1170,1428,1450]" gridcol="3" gridrow="16" pageId="9" pageNumber="266">14.3</td>
<td id="B5285D840E07AAF3FB28C2F8FAA80DD7" box="[1239,1288,1428,1450]" gridcol="4" gridrow="16" pageId="9" pageNumber="266">13.9</td>
<td id="B5285D840E07AAF3FAB6C2F8FA250DD7" box="[1353,1413,1428,1450]" gridcol="5" gridrow="16" pageId="9" pageNumber="266">14.8</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C2D1FA250DAE" box="[159,1413,1469,1491]" gridrow="17" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C2D1FD8F0DAE" box="[159,559,1469,1491]" gridcol="0" gridrow="17" pageId="9" pageNumber="266">Nostril length</th>
<td id="B5285D840E07AAF3FD88C2D1FD450DAE" box="[631,741,1469,1491]" gridcol="1" gridrow="17" pageId="9" pageNumber="266">4.0</td>
<td id="B5285D840E07AAF3FCF4C2D1FC680DAE" box="[779,968,1469,1491]" gridcol="2" gridrow="17" pageId="9" pageNumber="266">3.9</td>
<td id="B5285D840E07AAF3FC05C2D1FB320DAE" box="[1018,1170,1469,1491]" gridcol="3" gridrow="17" pageId="9" pageNumber="266">3.8</td>
<td id="B5285D840E07AAF3FB28C2D1FAA80DAE" box="[1239,1288,1469,1491]" gridcol="4" gridrow="17" pageId="9" pageNumber="266">3.6</td>
<td id="B5285D840E07AAF3FAB6C2D1FA250DAE" box="[1353,1413,1469,1491]" gridcol="5" gridrow="17" pageId="9" pageNumber="266">3.7</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C28AFA250D81" box="[159,1413,1510,1532]" gridrow="18" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C28AFD8F0D81" box="[159,559,1510,1532]" gridcol="0" gridrow="18" pageId="9" pageNumber="266">Anterior aperture—width</th>
<td id="B5285D840E07AAF3FD88C28AFD450D81" box="[631,741,1510,1532]" gridcol="1" gridrow="18" pageId="9" pageNumber="266">1.4</td>
<td id="B5285D840E07AAF3FCF4C28AFC680D81" box="[779,968,1510,1532]" gridcol="2" gridrow="18" pageId="9" pageNumber="266">1.2</td>
<td id="B5285D840E07AAF3FC05C28AFB320D81" box="[1018,1170,1510,1532]" gridcol="3" gridrow="18" pageId="9" pageNumber="266">1.2</td>
<td id="B5285D840E07AAF3FB28C28AFAA80D81" box="[1239,1288,1510,1532]" gridcol="4" gridrow="18" pageId="9" pageNumber="266">1.0</td>
<td id="B5285D840E07AAF3FAB6C28AFA250D81" box="[1353,1413,1510,1532]" gridcol="5" gridrow="18" pageId="9" pageNumber="266">1.2</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C17CFA250E5B" box="[159,1413,1552,1574]" gridrow="19" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C17CFD8F0E5B" box="[159,559,1552,1574]" gridcol="0" gridrow="19" pageId="9" pageNumber="266">Anterior nasal flap—base length</th>
<td id="B5285D840E07AAF3FD88C17CFD450E5B" box="[631,741,1552,1574]" gridcol="1" gridrow="19" pageId="9" pageNumber="266">2.4</td>
<td id="B5285D840E07AAF3FCF4C17CFC680E5B" box="[779,968,1552,1574]" gridcol="2" gridrow="19" pageId="9" pageNumber="266">2.6</td>
<td id="B5285D840E07AAF3FC05C17CFB320E5B" box="[1018,1170,1552,1574]" gridcol="3" gridrow="19" pageId="9" pageNumber="266">2.5</td>
<td id="B5285D840E07AAF3FB28C17CFAA80E5B" box="[1239,1288,1552,1574]" gridcol="4" gridrow="19" pageId="9" pageNumber="266">2.2</td>
<td id="B5285D840E07AAF3FAB6C17CFA250E5B" box="[1353,1413,1552,1574]" gridcol="5" gridrow="19" pageId="9" pageNumber="266">2.5</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C155FA250E32" box="[159,1413,1593,1615]" gridrow="20" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C155FD8F0E32" box="[159,559,1593,1615]" gridcol="0" gridrow="20" pageId="9" pageNumber="266">Anterior nasal flap—width</th>
<td id="B5285D840E07AAF3FD88C155FD450E32" box="[631,741,1593,1615]" gridcol="1" gridrow="20" pageId="9" pageNumber="266">1.7</td>
<td id="B5285D840E07AAF3FCF4C155FC680E32" box="[779,968,1593,1615]" gridcol="2" gridrow="20" pageId="9" pageNumber="266">1.8</td>
<td id="B5285D840E07AAF3FC05C155FB320E32" box="[1018,1170,1593,1615]" gridcol="3" gridrow="20" pageId="9" pageNumber="266">1.6</td>
<td id="B5285D840E07AAF3FB28C155FAA80E32" box="[1239,1288,1593,1615]" gridcol="4" gridrow="20" pageId="9" pageNumber="266">1.2</td>
<td id="B5285D840E07AAF3FAB6C155FA250E32" box="[1353,1413,1593,1615]" gridcol="5" gridrow="20" pageId="9" pageNumber="266">1.5</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C10EFA250E05" box="[159,1413,1634,1656]" gridrow="21" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C10EFD8F0E05" box="[159,559,1634,1656]" gridcol="0" gridrow="21" pageId="9" pageNumber="266">Posterolateral nasal flap—total length</th>
<td id="B5285D840E07AAF3FD88C10EFD450E05" box="[631,741,1634,1656]" gridcol="1" gridrow="21" pageId="9" pageNumber="266">3.3</td>
<td id="B5285D840E07AAF3FCF4C10EFC680E05" box="[779,968,1634,1656]" gridcol="2" gridrow="21" pageId="9" pageNumber="266">3.1</td>
<td id="B5285D840E07AAF3FC05C10EFB320E05" box="[1018,1170,1634,1656]" gridcol="3" gridrow="21" pageId="9" pageNumber="266">3.3</td>
<td id="B5285D840E07AAF3FB28C10EFAA80E05" box="[1239,1288,1634,1656]" gridcol="4" gridrow="21" pageId="9" pageNumber="266">3.1</td>
<td id="B5285D840E07AAF3FAB6C10EFA250E05" box="[1353,1413,1634,1656]" gridcol="5" gridrow="21" pageId="9" pageNumber="266">3.5</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C1E0FA250EDF" box="[159,1413,1676,1698]" gridrow="22" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C1E0FD8F0EDF" box="[159,559,1676,1698]" gridcol="0" gridrow="22" pageId="9" pageNumber="266">Posterolateral nasal flap—width</th>
<td id="B5285D840E07AAF3FD88C1E0FD450EDF" box="[631,741,1676,1698]" gridcol="1" gridrow="22" pageId="9" pageNumber="266">0.6</td>
<td id="B5285D840E07AAF3FCF4C1E0FC680EDF" box="[779,968,1676,1698]" gridcol="2" gridrow="22" pageId="9" pageNumber="266">0.7</td>
<td id="B5285D840E07AAF3FC05C1E0FB320EDF" box="[1018,1170,1676,1698]" gridcol="3" gridrow="22" pageId="9" pageNumber="266">0.7</td>
<td id="B5285D840E07AAF3FB28C1E0FAA80EDF" box="[1239,1288,1676,1698]" gridcol="4" gridrow="22" pageId="9" pageNumber="266">0.6</td>
<td id="B5285D840E07AAF3FAB6C1E0FA250EDF" box="[1353,1413,1676,1698]" gridcol="5" gridrow="22" pageId="9" pageNumber="266">0.7</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C1D9FA250EB6" box="[159,1413,1717,1739]" gridrow="23" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C1D9FD8F0EB6" box="[159,559,1717,1739]" gridcol="0" gridrow="23" pageId="9" pageNumber="266">Posterior nasal flap—base length</th>
<td id="B5285D840E07AAF3FD88C1D9FD450EB6" box="[631,741,1717,1739]" gridcol="1" gridrow="23" pageId="9" pageNumber="266">2.6</td>
<td id="B5285D840E07AAF3FCF4C1D9FC680EB6" box="[779,968,1717,1739]" gridcol="2" gridrow="23" pageId="9" pageNumber="266">2.0</td>
<td id="B5285D840E07AAF3FC05C1D9FB320EB6" box="[1018,1170,1717,1739]" gridcol="3" gridrow="23" pageId="9" pageNumber="266">2.2</td>
<td id="B5285D840E07AAF3FB28C1D9FAA80EB6" box="[1239,1288,1717,1739]" gridcol="4" gridrow="23" pageId="9" pageNumber="266">2.0</td>
<td id="B5285D840E07AAF3FAB6C1D9FA250EB6" box="[1353,1413,1717,1739]" gridcol="5" gridrow="23" pageId="9" pageNumber="266">2.2</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C1B2FA250E89" box="[159,1413,1758,1780]" gridrow="24" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C1B2FD8F0E89" box="[159,559,1758,1780]" gridcol="0" gridrow="24" pageId="9" pageNumber="266">Posterior nasal flap—width</th>
<td id="B5285D840E07AAF3FD88C1B2FD450E89" box="[631,741,1758,1780]" gridcol="1" gridrow="24" pageId="9" pageNumber="266">1.2</td>
<td id="B5285D840E07AAF3FCF4C1B2FC680E89" box="[779,968,1758,1780]" gridcol="2" gridrow="24" pageId="9" pageNumber="266">1.1</td>
<td id="B5285D840E07AAF3FC05C1B2FB320E89" box="[1018,1170,1758,1780]" gridcol="3" gridrow="24" pageId="9" pageNumber="266">1.0</td>
<td id="B5285D840E07AAF3FB28C1B2FAA80E89" box="[1239,1288,1758,1780]" gridcol="4" gridrow="24" pageId="9" pageNumber="266">0.8</td>
<td id="B5285D840E07AAF3FAB6C1B2FA250E89" box="[1353,1413,1758,1780]" gridcol="5" gridrow="24" pageId="9" pageNumber="266">1.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C064FA250F63" box="[159,1413,1800,1822]" gridrow="25" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C064FD8F0F63" box="[159,559,1800,1822]" gridcol="0" gridrow="25" pageId="9" pageNumber="266">Distance across anterior nasal apertures</th>
<td id="B5285D840E07AAF3FD88C064FD450F63" box="[631,741,1800,1822]" gridcol="1" gridrow="25" pageId="9" pageNumber="266">3.7</td>
<td id="B5285D840E07AAF3FCF4C064FC680F63" box="[779,968,1800,1822]" gridcol="2" gridrow="25" pageId="9" pageNumber="266">9.2</td>
<td id="B5285D840E07AAF3FC05C064FB320F63" box="[1018,1170,1800,1822]" gridcol="3" gridrow="25" pageId="9" pageNumber="266">9.0</td>
<td id="B5285D840E07AAF3FB28C064FAA80F63" box="[1239,1288,1800,1822]" gridcol="4" gridrow="25" pageId="9" pageNumber="266">8.5</td>
<td id="B5285D840E07AAF3FAB6C064FA250F63" box="[1353,1413,1800,1822]" gridcol="5" gridrow="25" pageId="9" pageNumber="266">9.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C05DFA250F3A" box="[159,1413,1841,1863]" gridrow="26" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C05DFD8F0F3A" box="[159,559,1841,1863]" gridcol="0" gridrow="26" pageId="9" pageNumber="266">Internarial distance (minimum)</th>
<td id="B5285D840E07AAF3FD88C05DFD450F3A" box="[631,741,1841,1863]" gridcol="1" gridrow="26" pageId="9" pageNumber="266">3.0</td>
<td id="B5285D840E07AAF3FCF4C05DFC680F3A" box="[779,968,1841,1863]" gridcol="2" gridrow="26" pageId="9" pageNumber="266">2.7</td>
<td id="B5285D840E07AAF3FC05C05DFB320F3A" box="[1018,1170,1841,1863]" gridcol="3" gridrow="26" pageId="9" pageNumber="266">2.6</td>
<td id="B5285D840E07AAF3FB28C05DFAA80F3A" box="[1239,1288,1841,1863]" gridcol="4" gridrow="26" pageId="9" pageNumber="266">2.5</td>
<td id="B5285D840E07AAF3FAB6C05DFA250F3A" box="[1353,1413,1841,1863]" gridcol="5" gridrow="26" pageId="9" pageNumber="266">3.0</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C036FA250F0D" box="[159,1413,1882,1904]" gridrow="27" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C036FD8F0F0D" box="[159,559,1882,1904]" gridcol="0" gridrow="27" pageId="9" pageNumber="266">Distance between anterior nasal flaps</th>
<td id="B5285D840E07AAF3FD88C036FD450F0D" box="[631,741,1882,1904]" gridcol="1" gridrow="27" pageId="9" pageNumber="266">10.0</td>
<td id="B5285D840E07AAF3FCF4C036FC680F0D" box="[779,968,1882,1904]" gridcol="2" gridrow="27" pageId="9" pageNumber="266">2.6</td>
<td id="B5285D840E07AAF3FC05C036FB320F0D" box="[1018,1170,1882,1904]" gridcol="3" gridrow="27" pageId="9" pageNumber="266">2.3</td>
<td id="B5285D840E07AAF3FB28C036FAA80F0D" box="[1239,1288,1882,1904]" gridcol="4" gridrow="27" pageId="9" pageNumber="266">2.3</td>
<td id="B5285D840E07AAF3FAB6C036FA250F0D" box="[1353,1413,1882,1904]" gridcol="5" gridrow="27" pageId="9" pageNumber="266">2.7</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C0E8FA250FE7" box="[159,1413,1924,1946]" gridrow="28" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C0E8FD8F0FE7" box="[159,559,1924,1946]" gridcol="0" gridrow="28" pageId="9" pageNumber="266">Distance from nostril to disc margin</th>
<td id="B5285D840E07AAF3FD88C0E8FD450FE7" box="[631,741,1924,1946]" gridcol="1" gridrow="28" pageId="9" pageNumber="266">2.8</td>
<td id="B5285D840E07AAF3FCF4C0E8FC680FE7" box="[779,968,1924,1946]" gridcol="2" gridrow="28" pageId="9" pageNumber="266">3.5</td>
<td id="B5285D840E07AAF3FC05C0E8FB320FE7" box="[1018,1170,1924,1946]" gridcol="3" gridrow="28" pageId="9" pageNumber="266">3.5</td>
<td id="B5285D840E07AAF3FB28C0E8FAA80FE7" box="[1239,1288,1924,1946]" gridcol="4" gridrow="28" pageId="9" pageNumber="266">3.3</td>
<td id="B5285D840E07AAF3FAB6C0E8FA250FE7" box="[1353,1413,1924,1946]" gridcol="5" gridrow="28" pageId="9" pageNumber="266">3.7</td>
</tr>
<tr id="F6F934F80E07AAF3FF60C0C1FA250FBE" box="[159,1413,1965,1987]" gridrow="29" pageId="9" pageNumber="266">
<th id="B5285D840E07AAF3FF60C0C1FD8F0FBE" box="[159,559,1965,1987]" gridcol="0" gridrow="29" pageId="9" pageNumber="266">Third gill opening—width</th>
<td id="B5285D840E07AAF3FD88C0C1FD450FBE" box="[631,741,1965,1987]" gridcol="1" gridrow="29" pageId="9" pageNumber="266">2.9</td>
<td id="B5285D840E07AAF3FCF4C0C1FC680FBE" box="[779,968,1965,1987]" gridcol="2" gridrow="29" pageId="9" pageNumber="266">2.4</td>
<td id="B5285D840E07AAF3FC05C0C1FB320FBE" box="[1018,1170,1965,1987]" gridcol="3" gridrow="29" pageId="9" pageNumber="266">1.7</td>
<td id="B5285D840E07AAF3FB28C0C1FAA80FBE" box="[1239,1288,1965,1987]" gridcol="4" gridrow="29" pageId="9" pageNumber="266">1.8</td>
<td id="B5285D840E07AAF3FAB6C0C1FA250FBE" box="[1353,1413,1965,1987]" gridcol="5" gridrow="29" pageId="9" pageNumber="266">1.9</td>
</tr>
</table>
</paragraph>
<paragraph id="487636BA0E075505FBA3C0BAFA330F90" blockId="9.[1116,1427,2006,2029]" box="[1116,1427,2006,2029]" pageId="9" pageNumber="266">
<tableNote id="B52F37340E075505FBA3C0BAFA330F90" box="[1116,1427,2006,2029]" pageId="9" pageNumber="266" targetBox="[159,1413,766,1987]" targetPageId="9">......continued on the next page</tableNote>
</paragraph>
<caption id="1CB666320E045506FF60C7CCFE2D08CB" box="[159,397,160,182]" pageId="10" pageNumber="267">
<paragraph id="487636BA0E045506FF60C7CCFE2D08CB" blockId="10.[159,397,160,182]" box="[159,397,160,182]" pageId="10" pageNumber="267">
<emphasis id="7ABDEAA80E045506FF60C7CCFEAC08CB" bold="true" box="[159,268,160,182]" pageId="10" pageNumber="267">TABLE 2.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="487636BA0E045506FD90C7A6FAFB0F7F" pageId="10" pageNumber="267">
<table id="3AC9C41A0E04AAF3FF60C7A5FAD60F7F" box="[159,1398,201,1794]" colsContinueFrom="9.[159,1413,766,1987]" gridcols="6" gridrows="39" pageId="10" pageNumber="267">
<tr id="F6F934F80E04AAF3FF60C7A5FAD608A2" box="[159,1398,201,223]" gridrow="0" pageId="10" pageNumber="267" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="B5285D840E04AAF3FD90C7A5FD7C08A2" box="[623,732,201,223]" gridcol="1" gridrow="0" pageId="10" pageNumber="267">
<taxonomicName id="8FC94D390E045506FD90C7A6FD7C08A2" authorityName=": Fowler" authorityYear="1941" box="[623,732,201,223]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="267" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E045506FD90C7A6FD7C08A2" box="[623,732,201,223]" italics="true" pageId="10" pageNumber="267">R. lionotus</emphasis>
</taxonomicName>
</th>
<th id="B5285D840E04AAF3FC19C7A5FBDE08A2" box="[998,1150,201,223]" gridcol="3" gridrow="0" pageId="10" pageNumber="267">
<taxonomicName id="8FC94D390E045506FC19C7A6FBDE08A2" authorityName="Last &amp; Séret &amp; Naylor" authorityYear="2019" box="[998,1150,202,223]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="267" phylum="Chordata" rank="species" species="ranongensis">
<emphasis id="7ABDEAA80E045506FC19C7A6FBDE08A2" box="[998,1150,202,223]" italics="true" pageId="10" pageNumber="267">R. ranongensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="F6F934F80E04AAF3FF60C79EFAD60975" box="[159,1398,242,264]" gridrow="1" pageId="10" pageNumber="267" rowspan-0="1" rowspan-5="1">
<td id="B5285D840E04AAF3FD90C79EFD7C0975" box="[623,732,242,264]" gridcol="1" gridrow="1" pageId="10" pageNumber="267">Holotype</td>
<td id="B5285D840E04AAF3FD02C79EFC1B0975" box="[765,955,242,264]" gridcol="2" gridrow="1" pageId="10" pageNumber="267">CSIRO H 7867-01</td>
<td id="B5285D840E04AAF3FC19C79EFBDE0975" box="[998,1150,242,264]" gridcol="3" gridrow="1" pageId="10" pageNumber="267">Holotype</td>
<td id="B5285D840E04AAF3FB4AC79EFAB80975" box="[1205,1304,242,264]" gridcol="4" gridrow="1" pageId="10" pageNumber="267">Paratypes</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C670FAD6094F" box="[159,1398,284,306]" gridrow="2" pageId="10" pageNumber="267" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="B5285D840E04AAF3FB4AC670FAB8094F" box="[1205,1304,284,306]" gridcol="4" gridrow="2" pageId="10" pageNumber="267">MIN</td>
<td id="B5285D840E04AAF3FAC5C670FAD6094F" box="[1338,1398,284,306]" gridcol="5" gridrow="2" pageId="10" pageNumber="267">MAX</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C629FAD60926" box="[159,1398,325,347]" gridrow="3" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C629FDEB0926" box="[159,587,325,347]" gridcol="0" gridrow="3" pageId="10" pageNumber="267">Distance between first gill openings</th>
<td id="B5285D840E04AAF3FD90C629FD7C0926" box="[623,732,325,347]" gridcol="1" gridrow="3" pageId="10" pageNumber="267">14.9</td>
<td id="B5285D840E04AAF3FD02C629FC1B0926" box="[765,955,325,347]" gridcol="2" gridrow="3" pageId="10" pageNumber="267">14.3</td>
<td id="B5285D840E04AAF3FC19C629FBDE0926" box="[998,1150,325,347]" gridcol="3" gridrow="3" pageId="10" pageNumber="267">12.4</td>
<td id="B5285D840E04AAF3FB4AC629FAB80926" box="[1205,1304,325,347]" gridcol="4" gridrow="3" pageId="10" pageNumber="267">12.4</td>
<td id="B5285D840E04AAF3FAC5C629FAD60926" box="[1338,1398,325,347]" gridcol="5" gridrow="3" pageId="10" pageNumber="267">13.4</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C602FAD609F9" box="[159,1398,366,388]" gridrow="4" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C602FDEB09F9" box="[159,587,366,388]" gridcol="0" gridrow="4" pageId="10" pageNumber="267">Distance between fifth gill openings</th>
<td id="B5285D840E04AAF3FD90C602FD7C09F9" box="[623,732,366,388]" gridcol="1" gridrow="4" pageId="10" pageNumber="267">10.5</td>
<td id="B5285D840E04AAF3FD02C602FC1B09F9" box="[765,955,366,388]" gridcol="2" gridrow="4" pageId="10" pageNumber="267">10.9</td>
<td id="B5285D840E04AAF3FC19C602FBDE09F9" box="[998,1150,366,388]" gridcol="3" gridrow="4" pageId="10" pageNumber="267">8.6</td>
<td id="B5285D840E04AAF3FB4AC602FAB809F9" box="[1205,1304,366,388]" gridcol="4" gridrow="4" pageId="10" pageNumber="267">8.8</td>
<td id="B5285D840E04AAF3FAC5C602FAD609F9" box="[1338,1398,366,388]" gridcol="5" gridrow="4" pageId="10" pageNumber="267">9.5</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C6F4FAD609D3" box="[159,1398,408,430]" gridrow="5" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C6F4FDEB09D3" box="[159,587,408,430]" gridcol="0" gridrow="5" pageId="10" pageNumber="267">Pelvic fin—length</th>
<td id="B5285D840E04AAF3FD90C6F4FD7C09D3" box="[623,732,408,430]" gridcol="1" gridrow="5" pageId="10" pageNumber="267">15.8</td>
<td id="B5285D840E04AAF3FD02C6F4FC1B09D3" box="[765,955,408,430]" gridcol="2" gridrow="5" pageId="10" pageNumber="267">16.3</td>
<td id="B5285D840E04AAF3FC19C6F4FBDE09D3" box="[998,1150,408,430]" gridcol="3" gridrow="5" pageId="10" pageNumber="267">14.4</td>
<td id="B5285D840E04AAF3FB4AC6F4FAB809D3" box="[1205,1304,408,430]" gridcol="4" gridrow="5" pageId="10" pageNumber="267">14.5</td>
<td id="B5285D840E04AAF3FAC5C6F4FAD609D3" box="[1338,1398,408,430]" gridcol="5" gridrow="5" pageId="10" pageNumber="267">15.4</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C6ADFAD609AA" box="[159,1398,449,471]" gridrow="6" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C6ADFDEB09AA" box="[159,587,449,471]" gridcol="0" gridrow="6" pageId="10" pageNumber="267">Pelvic fin—anterior margin length</th>
<td id="B5285D840E04AAF3FD90C6ADFD7C09AA" box="[623,732,449,471]" gridcol="1" gridrow="6" pageId="10" pageNumber="267">6.5</td>
<td id="B5285D840E04AAF3FD02C6ADFC1B09AA" box="[765,955,449,471]" gridcol="2" gridrow="6" pageId="10" pageNumber="267">8.5</td>
<td id="B5285D840E04AAF3FC19C6ADFBDE09AA" box="[998,1150,449,471]" gridcol="3" gridrow="6" pageId="10" pageNumber="267">8.4</td>
<td id="B5285D840E04AAF3FB4AC6ADFAB809AA" box="[1205,1304,449,471]" gridcol="4" gridrow="6" pageId="10" pageNumber="267">8.4</td>
<td id="B5285D840E04AAF3FAC5C6ADFAD609AA" box="[1338,1398,449,471]" gridcol="5" gridrow="6" pageId="10" pageNumber="267">9.0</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C686FAD60A7D" box="[159,1398,490,512]" gridrow="7" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C686FDEB0A7D" box="[159,587,490,512]" gridcol="0" gridrow="7" pageId="10" pageNumber="267">Pelvic fin—width</th>
<td id="B5285D840E04AAF3FD90C686FD7C0A7D" box="[623,732,490,512]" gridcol="1" gridrow="7" pageId="10" pageNumber="267">6.3</td>
<td id="B5285D840E04AAF3FD02C686FC1B0A7D" box="[765,955,490,512]" gridcol="2" gridrow="7" pageId="10" pageNumber="267">6.7</td>
<td id="B5285D840E04AAF3FC19C686FBDE0A7D" box="[998,1150,490,512]" gridcol="3" gridrow="7" pageId="10" pageNumber="267">5.5</td>
<td id="B5285D840E04AAF3FB4AC686FAB80A7D" box="[1205,1304,490,512]" gridcol="4" gridrow="7" pageId="10" pageNumber="267">5.1</td>
<td id="B5285D840E04AAF3FAC5C686FAD60A7D" box="[1338,1398,490,512]" gridcol="5" gridrow="7" pageId="10" pageNumber="267">5.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C578FAD60A57" box="[159,1398,532,554]" gridrow="8" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C578FDEB0A57" box="[159,587,532,554]" gridcol="0" gridrow="8" pageId="10" pageNumber="267">Pelvic fin—base length</th>
<td id="B5285D840E04AAF3FD90C578FD7C0A57" box="[623,732,532,554]" gridcol="1" gridrow="8" pageId="10" pageNumber="267">9.3</td>
<td id="B5285D840E04AAF3FD02C578FC1B0A57" box="[765,955,532,554]" gridcol="2" gridrow="8" pageId="10" pageNumber="267">9.1</td>
<td id="B5285D840E04AAF3FC19C578FBDE0A57" box="[998,1150,532,554]" gridcol="3" gridrow="8" pageId="10" pageNumber="267">7.8</td>
<td id="B5285D840E04AAF3FB4AC578FAB80A57" box="[1205,1304,532,554]" gridcol="4" gridrow="8" pageId="10" pageNumber="267">7.2</td>
<td id="B5285D840E04AAF3FAC5C578FAD60A57" box="[1338,1398,532,554]" gridcol="5" gridrow="8" pageId="10" pageNumber="267">8.1</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C551FAD60A2E" box="[159,1398,573,595]" gridrow="9" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C551FDEB0A2E" box="[159,587,573,595]" gridcol="0" gridrow="9" pageId="10" pageNumber="267">Pelvic fin—inner margin length</th>
<td id="B5285D840E04AAF3FD90C551FD7C0A2E" box="[623,732,573,595]" gridcol="1" gridrow="9" pageId="10" pageNumber="267">7.1</td>
<td id="B5285D840E04AAF3FD02C551FC1B0A2E" box="[765,955,573,595]" gridcol="2" gridrow="9" pageId="10" pageNumber="267">8.9</td>
<td id="B5285D840E04AAF3FC19C551FBDE0A2E" box="[998,1150,573,595]" gridcol="3" gridrow="9" pageId="10" pageNumber="267">7.7</td>
<td id="B5285D840E04AAF3FB4AC551FAB80A2E" box="[1205,1304,573,595]" gridcol="4" gridrow="9" pageId="10" pageNumber="267">6.9</td>
<td id="B5285D840E04AAF3FAC5C551FAD60A2E" box="[1338,1398,573,595]" gridcol="5" gridrow="9" pageId="10" pageNumber="267">7.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C50AFAD60A01" box="[159,1398,614,636]" gridrow="10" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C50AFDEB0A01" box="[159,587,614,636]" gridcol="0" gridrow="10" pageId="10" pageNumber="267">First dorsal fin—length</th>
<td id="B5285D840E04AAF3FD90C50AFD7C0A01" box="[623,732,614,636]" gridcol="1" gridrow="10" pageId="10" pageNumber="267">6.1</td>
<td id="B5285D840E04AAF3FD02C50AFC1B0A01" box="[765,955,614,636]" gridcol="2" gridrow="10" pageId="10" pageNumber="267">7.2</td>
<td id="B5285D840E04AAF3FC19C50AFBDE0A01" box="[998,1150,614,636]" gridcol="3" gridrow="10" pageId="10" pageNumber="267">6.5</td>
<td id="B5285D840E04AAF3FB4AC50AFAB80A01" box="[1205,1304,614,636]" gridcol="4" gridrow="10" pageId="10" pageNumber="267">6.3</td>
<td id="B5285D840E04AAF3FAC5C50AFAD60A01" box="[1338,1398,614,636]" gridcol="5" gridrow="10" pageId="10" pageNumber="267">6.7</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C5FCFAD60ADB" box="[159,1398,656,678]" gridrow="11" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C5FCFDEB0ADB" box="[159,587,656,678]" gridcol="0" gridrow="11" pageId="10" pageNumber="267">First dorsal fin—anterior margin length</th>
<td id="B5285D840E04AAF3FD90C5FCFD7C0ADB" box="[623,732,656,678]" gridcol="1" gridrow="11" pageId="10" pageNumber="267">9.9</td>
<td id="B5285D840E04AAF3FD02C5FCFC1B0ADB" box="[765,955,656,678]" gridcol="2" gridrow="11" pageId="10" pageNumber="267">10.2</td>
<td id="B5285D840E04AAF3FC19C5FCFBDE0ADB" box="[998,1150,656,678]" gridcol="3" gridrow="11" pageId="10" pageNumber="267">8.9</td>
<td id="B5285D840E04AAF3FB4AC5FCFAB80ADB" box="[1205,1304,656,678]" gridcol="4" gridrow="11" pageId="10" pageNumber="267">9.2</td>
<td id="B5285D840E04AAF3FAC5C5FCFAD60ADB" box="[1338,1398,656,678]" gridcol="5" gridrow="11" pageId="10" pageNumber="267">9.3</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C5D5FAD60AB2" box="[159,1398,697,719]" gridrow="12" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C5D5FDEB0AB2" box="[159,587,697,719]" gridcol="0" gridrow="12" pageId="10" pageNumber="267">First dorsal fin—height</th>
<td id="B5285D840E04AAF3FD90C5D5FD7C0AB2" box="[623,732,697,719]" gridcol="1" gridrow="12" pageId="10" pageNumber="267">8.1</td>
<td id="B5285D840E04AAF3FD02C5D5FC1B0AB2" box="[765,955,697,719]" gridcol="2" gridrow="12" pageId="10" pageNumber="267">7.7</td>
<td id="B5285D840E04AAF3FC19C5D5FBDE0AB2" box="[998,1150,697,719]" gridcol="3" gridrow="12" pageId="10" pageNumber="267">5.9</td>
<td id="B5285D840E04AAF3FB4AC5D5FAB80AB2" box="[1205,1304,697,719]" gridcol="4" gridrow="12" pageId="10" pageNumber="267">5.4</td>
<td id="B5285D840E04AAF3FAC5C5D5FAD60AB2" box="[1338,1398,697,719]" gridcol="5" gridrow="12" pageId="10" pageNumber="267">6.2</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C58EFAD60A85" box="[159,1398,738,760]" gridrow="13" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C58EFDEB0A85" box="[159,587,738,760]" gridcol="0" gridrow="13" pageId="10" pageNumber="267">First dorsal fin—base length</th>
<td id="B5285D840E04AAF3FD90C58EFD7C0A85" box="[623,732,738,760]" gridcol="1" gridrow="13" pageId="10" pageNumber="267">4.1</td>
<td id="B5285D840E04AAF3FD02C58EFC1B0A85" box="[765,955,738,760]" gridcol="2" gridrow="13" pageId="10" pageNumber="267">4.8</td>
<td id="B5285D840E04AAF3FC19C58EFBDE0A85" box="[998,1150,738,760]" gridcol="3" gridrow="13" pageId="10" pageNumber="267">4.2</td>
<td id="B5285D840E04AAF3FB4AC58EFAB80A85" box="[1205,1304,738,760]" gridcol="4" gridrow="13" pageId="10" pageNumber="267">4.0</td>
<td id="B5285D840E04AAF3FAC5C58EFAD60A85" box="[1338,1398,738,760]" gridcol="5" gridrow="13" pageId="10" pageNumber="267">4.3</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C460FAD60B5F" box="[159,1398,780,802]" gridrow="14" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C460FDEB0B5F" box="[159,587,780,802]" gridcol="0" gridrow="14" pageId="10" pageNumber="267">First dorsal fin—inner margin length</th>
<td id="B5285D840E04AAF3FD90C460FD7C0B5F" box="[623,732,780,802]" gridcol="1" gridrow="14" pageId="10" pageNumber="267">2.8</td>
<td id="B5285D840E04AAF3FD02C460FC1B0B5F" box="[765,955,780,802]" gridcol="2" gridrow="14" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FC19C460FBDE0B5F" box="[998,1150,780,802]" gridcol="3" gridrow="14" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FB4AC460FAB80B5F" box="[1205,1304,780,802]" gridcol="4" gridrow="14" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FAC5C460FAD60B5F" box="[1338,1398,780,802]" gridcol="5" gridrow="14" pageId="10" pageNumber="267">2.9</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C459FAD60B36" box="[159,1398,821,843]" gridrow="15" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C459FDEB0B36" box="[159,587,821,843]" gridcol="0" gridrow="15" pageId="10" pageNumber="267">Second dorsal fin—length</th>
<td id="B5285D840E04AAF3FD90C459FD7C0B36" box="[623,732,821,843]" gridcol="1" gridrow="15" pageId="10" pageNumber="267">5.9</td>
<td id="B5285D840E04AAF3FD02C459FC1B0B36" box="[765,955,821,843]" gridcol="2" gridrow="15" pageId="10" pageNumber="267">7.2</td>
<td id="B5285D840E04AAF3FC19C459FBDE0B36" box="[998,1150,821,843]" gridcol="3" gridrow="15" pageId="10" pageNumber="267">6.6</td>
<td id="B5285D840E04AAF3FB4AC459FAB80B36" box="[1205,1304,821,843]" gridcol="4" gridrow="15" pageId="10" pageNumber="267">6.7</td>
<td id="B5285D840E04AAF3FAC5C459FAD60B36" box="[1338,1398,821,843]" gridcol="5" gridrow="15" pageId="10" pageNumber="267">7.0</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C432FAD60B09" box="[159,1398,862,884]" gridrow="16" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C432FDEB0B09" box="[159,587,862,884]" gridcol="0" gridrow="16" pageId="10" pageNumber="267">Second dorsal fin—anterior margin length</th>
<td id="B5285D840E04AAF3FD90C432FD7C0B09" box="[623,732,862,884]" gridcol="1" gridrow="16" pageId="10" pageNumber="267">8.7</td>
<td id="B5285D840E04AAF3FD02C432FC1B0B09" box="[765,955,862,884]" gridcol="2" gridrow="16" pageId="10" pageNumber="267">9.7</td>
<td id="B5285D840E04AAF3FC19C432FBDE0B09" box="[998,1150,862,884]" gridcol="3" gridrow="16" pageId="10" pageNumber="267">8.5</td>
<td id="B5285D840E04AAF3FB4AC432FAB80B09" box="[1205,1304,862,884]" gridcol="4" gridrow="16" pageId="10" pageNumber="267">8.8</td>
<td id="B5285D840E04AAF3FAC5C432FAD60B09" box="[1338,1398,862,884]" gridcol="5" gridrow="16" pageId="10" pageNumber="267">9.2</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C4E4FAD60BE3" box="[159,1398,904,926]" gridrow="17" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C4E4FDEB0BE3" box="[159,587,904,926]" gridcol="0" gridrow="17" pageId="10" pageNumber="267">Second dorsal fin—height</th>
<td id="B5285D840E04AAF3FD90C4E4FD7C0BE3" box="[623,732,904,926]" gridcol="1" gridrow="17" pageId="10" pageNumber="267">6.2</td>
<td id="B5285D840E04AAF3FD02C4E4FC1B0BE3" box="[765,955,904,926]" gridcol="2" gridrow="17" pageId="10" pageNumber="267">6.4</td>
<td id="B5285D840E04AAF3FC19C4E4FBDE0BE3" box="[998,1150,904,926]" gridcol="3" gridrow="17" pageId="10" pageNumber="267">5.5</td>
<td id="B5285D840E04AAF3FB4AC4E4FAB80BE3" box="[1205,1304,904,926]" gridcol="4" gridrow="17" pageId="10" pageNumber="267">4.8</td>
<td id="B5285D840E04AAF3FAC5C4E4FAD60BE3" box="[1338,1398,904,926]" gridcol="5" gridrow="17" pageId="10" pageNumber="267">5.2</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C4DDFAD60BBA" box="[159,1398,945,967]" gridrow="18" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C4DDFDEB0BBA" box="[159,587,945,967]" gridcol="0" gridrow="18" pageId="10" pageNumber="267">Second dorsal fin—base length</th>
<td id="B5285D840E04AAF3FD90C4DDFD7C0BBA" box="[623,732,945,967]" gridcol="1" gridrow="18" pageId="10" pageNumber="267">4.3</td>
<td id="B5285D840E04AAF3FD02C4DDFC1B0BBA" box="[765,955,945,967]" gridcol="2" gridrow="18" pageId="10" pageNumber="267">4.9</td>
<td id="B5285D840E04AAF3FC19C4DDFBDE0BBA" box="[998,1150,945,967]" gridcol="3" gridrow="18" pageId="10" pageNumber="267">4.6</td>
<td id="B5285D840E04AAF3FB4AC4DDFAB80BBA" box="[1205,1304,945,967]" gridcol="4" gridrow="18" pageId="10" pageNumber="267">4.5</td>
<td id="B5285D840E04AAF3FAC5C4DDFAD60BBA" box="[1338,1398,945,967]" gridcol="5" gridrow="18" pageId="10" pageNumber="267">4.7</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C4B6FAD60B8D" box="[159,1398,986,1008]" gridrow="19" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C4B6FDEB0B8D" box="[159,587,986,1008]" gridcol="0" gridrow="19" pageId="10" pageNumber="267">Second dorsal fin—inner margin length</th>
<td id="B5285D840E04AAF3FD90C4B6FD7C0B8D" box="[623,732,986,1008]" gridcol="1" gridrow="19" pageId="10" pageNumber="267">2.0</td>
<td id="B5285D840E04AAF3FD02C4B6FC1B0B8D" box="[765,955,986,1008]" gridcol="2" gridrow="19" pageId="10" pageNumber="267">2.3</td>
<td id="B5285D840E04AAF3FC19C4B6FBDE0B8D" box="[998,1150,986,1008]" gridcol="3" gridrow="19" pageId="10" pageNumber="267">2.2</td>
<td id="B5285D840E04AAF3FB4AC4B6FAB80B8D" box="[1205,1304,986,1008]" gridcol="4" gridrow="19" pageId="10" pageNumber="267">2.2</td>
<td id="B5285D840E04AAF3FAC5C4B6FAD60B8D" box="[1338,1398,986,1008]" gridcol="5" gridrow="19" pageId="10" pageNumber="267">2.4</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C368FAD60C67" box="[159,1398,1028,1050]" gridrow="20" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C368FDEB0C67" box="[159,587,1028,1050]" gridcol="0" gridrow="20" pageId="10" pageNumber="267">Caudal fin—dorsal margin</th>
<td id="B5285D840E04AAF3FD90C368FD7C0C67" box="[623,732,1028,1050]" gridcol="1" gridrow="20" pageId="10" pageNumber="267">12.6</td>
<td id="B5285D840E04AAF3FD02C368FC1B0C67" box="[765,955,1028,1050]" gridcol="2" gridrow="20" pageId="10" pageNumber="267">13.2</td>
<td id="B5285D840E04AAF3FC19C368FBDE0C67" box="[998,1150,1028,1050]" gridcol="3" gridrow="20" pageId="10" pageNumber="267">13.1</td>
<td id="B5285D840E04AAF3FB4AC368FAB80C67" box="[1205,1304,1028,1050]" gridcol="4" gridrow="20" pageId="10" pageNumber="267">12.5</td>
<td id="B5285D840E04AAF3FAC5C368FAD60C67" box="[1338,1398,1028,1050]" gridcol="5" gridrow="20" pageId="10" pageNumber="267">13.0</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C341FAD60C3E" box="[159,1398,1069,1091]" gridrow="21" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C341FDEB0C3E" box="[159,587,1069,1091]" gridcol="0" gridrow="21" pageId="10" pageNumber="267">Caudal fin—preventral margin</th>
<td id="B5285D840E04AAF3FD90C341FD7C0C3E" box="[623,732,1069,1091]" gridcol="1" gridrow="21" pageId="10" pageNumber="267">6.4</td>
<td id="B5285D840E04AAF3FD02C341FC1B0C3E" box="[765,955,1069,1091]" gridcol="2" gridrow="21" pageId="10" pageNumber="267">6.8</td>
<td id="B5285D840E04AAF3FC19C341FBDE0C3E" box="[998,1150,1069,1091]" gridcol="3" gridrow="21" pageId="10" pageNumber="267">5.4</td>
<td id="B5285D840E04AAF3FB4AC341FAB80C3E" box="[1205,1304,1069,1091]" gridcol="4" gridrow="21" pageId="10" pageNumber="267">5.2</td>
<td id="B5285D840E04AAF3FAC5C341FAD60C3E" box="[1338,1398,1069,1091]" gridcol="5" gridrow="21" pageId="10" pageNumber="267">6.1</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C33AFAD60C11" box="[159,1398,1110,1132]" gridrow="22" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C33AFDEB0C11" box="[159,587,1110,1132]" gridcol="0" gridrow="22" pageId="10" pageNumber="267">Snout to first dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C33AFD7C0C11" box="[623,732,1110,1132]" gridcol="1" gridrow="22" pageId="10" pageNumber="267">59.6</td>
<td id="B5285D840E04AAF3FD02C33AFC1B0C11" box="[765,955,1110,1132]" gridcol="2" gridrow="22" pageId="10" pageNumber="267">57.9</td>
<td id="B5285D840E04AAF3FC19C33AFBDE0C11" box="[998,1150,1110,1132]" gridcol="3" gridrow="22" pageId="10" pageNumber="267">57.1</td>
<td id="B5285D840E04AAF3FB4AC33AFAB80C11" box="[1205,1304,1110,1132]" gridcol="4" gridrow="22" pageId="10" pageNumber="267">6.8</td>
<td id="B5285D840E04AAF3FAC5C33AFAD60C11" box="[1338,1398,1110,1132]" gridcol="5" gridrow="22" pageId="10" pageNumber="267">7.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C3ECFAD60CEB" box="[159,1398,1152,1174]" gridrow="23" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C3ECFDEB0CEB" box="[159,587,1152,1174]" gridcol="0" gridrow="23" pageId="10" pageNumber="267">Snout to second dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C3ECFD7C0CEB" box="[623,732,1152,1174]" gridcol="1" gridrow="23" pageId="10" pageNumber="267">75.4</td>
<td id="B5285D840E04AAF3FD02C3ECFC1B0CEB" box="[765,955,1152,1174]" gridcol="2" gridrow="23" pageId="10" pageNumber="267">73.0</td>
<td id="B5285D840E04AAF3FC19C3ECFBDE0CEB" box="[998,1150,1152,1174]" gridcol="3" gridrow="23" pageId="10" pageNumber="267">74.4</td>
<td id="B5285D840E04AAF3FB4AC3ECFAB80CEB" box="[1205,1304,1152,1174]" gridcol="4" gridrow="23" pageId="10" pageNumber="267">73.6</td>
<td id="B5285D840E04AAF3FAC5C3ECFAD60CEB" box="[1338,1398,1152,1174]" gridcol="5" gridrow="23" pageId="10" pageNumber="267">74.0</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C3C5FAD60CC2" box="[159,1398,1193,1215]" gridrow="24" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C3C5FDEB0CC2" box="[159,587,1193,1215]" gridcol="0" gridrow="24" pageId="10" pageNumber="267">Snout to upper caudal-fin origin</th>
<td id="B5285D840E04AAF3FD90C3C5FD7C0CC2" box="[623,732,1193,1215]" gridcol="1" gridrow="24" pageId="10" pageNumber="267">87.2</td>
<td id="B5285D840E04AAF3FD02C3C5FC1B0CC2" box="[765,955,1193,1215]" gridcol="2" gridrow="24" pageId="10" pageNumber="267">86.8</td>
<td id="B5285D840E04AAF3FC19C3C5FBDE0CC2" box="[998,1150,1193,1215]" gridcol="3" gridrow="24" pageId="10" pageNumber="267">86.9</td>
<td id="B5285D840E04AAF3FB4AC3C5FAB80CC2" box="[1205,1304,1193,1215]" gridcol="4" gridrow="24" pageId="10" pageNumber="267">87.0</td>
<td id="B5285D840E04AAF3FAC5C3C5FAD60CC2" box="[1338,1398,1193,1215]" gridcol="5" gridrow="24" pageId="10" pageNumber="267">87.5</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C3BEFAD60C95" box="[159,1398,1234,1256]" gridrow="25" pageId="10" pageNumber="267" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="B5285D840E04AAF3FF60C3BEFDEB0C95" box="[159,587,1234,1256]" gridcol="0" gridrow="25" pageId="10" pageNumber="267">Snout to lower caudal-fin origin</th>
<td id="B5285D840E04AAF3FD90C3BEFD7C0C95" box="[623,732,1234,1256]" gridcol="1" gridrow="25" pageId="10" pageNumber="267">88.9</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C390FAD60D6F" box="[159,1398,1276,1298]" gridrow="26" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C390FDEB0D6F" box="[159,587,1276,1298]" gridcol="0" gridrow="26" pageId="10" pageNumber="267">Snout to pelvic-fin origin</th>
<td id="B5285D840E04AAF3FD90C390FD7C0D6F" box="[623,732,1276,1298]" gridcol="1" gridrow="26" pageId="10" pageNumber="267">40.2</td>
<td id="B5285D840E04AAF3FD02C390FC1B0D6F" box="[765,955,1276,1298]" gridcol="2" gridrow="26" pageId="10" pageNumber="267">38.8</td>
<td id="B5285D840E04AAF3FC19C390FBDE0D6F" box="[998,1150,1276,1298]" gridcol="3" gridrow="26" pageId="10" pageNumber="267">37.5</td>
<td id="B5285D840E04AAF3FB4AC390FAB80D6F" box="[1205,1304,1276,1298]" gridcol="4" gridrow="26" pageId="10" pageNumber="267">36.8</td>
<td id="B5285D840E04AAF3FAC5C390FAD60D6F" box="[1338,1398,1276,1298]" gridcol="5" gridrow="26" pageId="10" pageNumber="267">38.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C249FAD60D46" box="[159,1398,1317,1339]" gridrow="27" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C249FDEB0D46" box="[159,587,1317,1339]" gridcol="0" gridrow="27" pageId="10" pageNumber="267">Snout to anterior vent</th>
<td id="B5285D840E04AAF3FD90C249FD7C0D46" box="[623,732,1317,1339]" gridcol="1" gridrow="27" pageId="10" pageNumber="267">43.8</td>
<td id="B5285D840E04AAF3FD02C249FC1B0D46" box="[765,955,1317,1339]" gridcol="2" gridrow="27" pageId="10" pageNumber="267">41.6</td>
<td id="B5285D840E04AAF3FC19C249FBDE0D46" box="[998,1150,1317,1339]" gridcol="3" gridrow="27" pageId="10" pageNumber="267">40.0</td>
<td id="B5285D840E04AAF3FB4AC249FAB80D46" box="[1205,1304,1317,1339]" gridcol="4" gridrow="27" pageId="10" pageNumber="267">39.8</td>
<td id="B5285D840E04AAF3FAC5C249FAD60D46" box="[1338,1398,1317,1339]" gridcol="5" gridrow="27" pageId="10" pageNumber="267">42.3</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C222FAD60D19" box="[159,1398,1358,1380]" gridrow="28" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C222FDEB0D19" box="[159,587,1358,1380]" gridcol="0" gridrow="28" pageId="10" pageNumber="267">Pelvic-fin insertion to dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C222FD7C0D19" box="[623,732,1358,1380]" gridcol="1" gridrow="28" pageId="10" pageNumber="267">12.2</td>
<td id="B5285D840E04AAF3FD02C222FC1B0D19" box="[765,955,1358,1380]" gridcol="2" gridrow="28" pageId="10" pageNumber="267">10.9</td>
<td id="B5285D840E04AAF3FC19C222FBDE0D19" box="[998,1150,1358,1380]" gridcol="3" gridrow="28" pageId="10" pageNumber="267">10.2</td>
<td id="B5285D840E04AAF3FB4AC222FAB80D19" box="[1205,1304,1358,1380]" gridcol="4" gridrow="28" pageId="10" pageNumber="267">11.6</td>
<td id="B5285D840E04AAF3FAC5C222FAD60D19" box="[1338,1398,1358,1380]" gridcol="5" gridrow="28" pageId="10" pageNumber="267">12.3</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C214FAD60DF3" box="[159,1398,1400,1422]" gridrow="29" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C214FDEB0DF3" box="[159,587,1400,1422]" gridcol="0" gridrow="29" pageId="10" pageNumber="267">Interdorsal distance</th>
<td id="B5285D840E04AAF3FD90C214FD7C0DF3" box="[623,732,1400,1422]" gridcol="1" gridrow="29" pageId="10" pageNumber="267">11.7</td>
<td id="B5285D840E04AAF3FD02C214FC1B0DF3" box="[765,955,1400,1422]" gridcol="2" gridrow="29" pageId="10" pageNumber="267">11.2</td>
<td id="B5285D840E04AAF3FC19C214FBDE0DF3" box="[998,1150,1400,1422]" gridcol="3" gridrow="29" pageId="10" pageNumber="267">13.8</td>
<td id="B5285D840E04AAF3FB4AC214FAB80DF3" box="[1205,1304,1400,1422]" gridcol="4" gridrow="29" pageId="10" pageNumber="267">12.3</td>
<td id="B5285D840E04AAF3FAC5C214FAD60DF3" box="[1338,1398,1400,1422]" gridcol="5" gridrow="29" pageId="10" pageNumber="267">12.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C2CDFAD60DCA" box="[159,1398,1441,1463]" gridrow="30" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C2CDFDEB0DCA" box="[159,587,1441,1463]" gridcol="0" gridrow="30" pageId="10" pageNumber="267">Caudal peduncle length (dorsal)</th>
<td id="B5285D840E04AAF3FD90C2CDFD7C0DCA" box="[623,732,1441,1463]" gridcol="1" gridrow="30" pageId="10" pageNumber="267">6.9</td>
<td id="B5285D840E04AAF3FD02C2CDFC1B0DCA" box="[765,955,1441,1463]" gridcol="2" gridrow="30" pageId="10" pageNumber="267">6.8</td>
<td id="B5285D840E04AAF3FC19C2CDFBDE0DCA" box="[998,1150,1441,1463]" gridcol="3" gridrow="30" pageId="10" pageNumber="267">7.8</td>
<td id="B5285D840E04AAF3FB4AC2CDFAB80DCA" box="[1205,1304,1441,1463]" gridcol="4" gridrow="30" pageId="10" pageNumber="267">7.9</td>
<td id="B5285D840E04AAF3FAC5C2CDFAD60DCA" box="[1338,1398,1441,1463]" gridcol="5" gridrow="30" pageId="10" pageNumber="267">8.7</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C2A6FAD60D9D" box="[159,1398,1482,1504]" gridrow="31" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C2A6FDEB0D9D" box="[159,587,1482,1504]" gridcol="0" gridrow="31" pageId="10" pageNumber="267">Body width—pelvic insertion (tail)</th>
<td id="B5285D840E04AAF3FD90C2A6FD7C0D9D" box="[623,732,1482,1504]" gridcol="1" gridrow="31" pageId="10" pageNumber="267">16.2</td>
<td id="B5285D840E04AAF3FD02C2A6FC1B0D9D" box="[765,955,1482,1504]" gridcol="2" gridrow="31" pageId="10" pageNumber="267">11.7</td>
<td id="B5285D840E04AAF3FC19C2A6FBDE0D9D" box="[998,1150,1482,1504]" gridcol="3" gridrow="31" pageId="10" pageNumber="267">10.2</td>
<td id="B5285D840E04AAF3FB4AC2A6FAB80D9D" box="[1205,1304,1482,1504]" gridcol="4" gridrow="31" pageId="10" pageNumber="267">9.1</td>
<td id="B5285D840E04AAF3FAC5C2A6FAD60D9D" box="[1338,1398,1482,1504]" gridcol="5" gridrow="31" pageId="10" pageNumber="267">10.1</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C298FAD60E77" box="[159,1398,1524,1546]" gridrow="32" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C298FDEB0E77" box="[159,587,1524,1546]" gridcol="0" gridrow="32" pageId="10" pageNumber="267">Disc width—anterior orbit</th>
<td id="B5285D840E04AAF3FD90C298FD7C0E77" box="[623,732,1524,1546]" gridcol="1" gridrow="32" pageId="10" pageNumber="267">17.4</td>
<td id="B5285D840E04AAF3FD02C298FC1B0E77" box="[765,955,1524,1546]" gridcol="2" gridrow="32" pageId="10" pageNumber="267">19.9</td>
<td id="B5285D840E04AAF3FC19C298FBDE0E77" box="[998,1150,1524,1546]" gridcol="3" gridrow="32" pageId="10" pageNumber="267">17.6</td>
<td id="B5285D840E04AAF3FB4AC298FAB80E77" box="[1205,1304,1524,1546]" gridcol="4" gridrow="32" pageId="10" pageNumber="267">17.6</td>
<td id="B5285D840E04AAF3FAC5C298FAD60E77" box="[1338,1398,1524,1546]" gridcol="5" gridrow="32" pageId="10" pageNumber="267">18.2</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C171FAD60E4E" box="[159,1398,1565,1587]" gridrow="33" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C171FDEB0E4E" box="[159,587,1565,1587]" gridcol="0" gridrow="33" pageId="10" pageNumber="267">Body width—first dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C171FD7C0E4E" box="[623,732,1565,1587]" gridcol="1" gridrow="33" pageId="10" pageNumber="267">9.7</td>
<td id="B5285D840E04AAF3FD02C171FC1B0E4E" box="[765,955,1565,1587]" gridcol="2" gridrow="33" pageId="10" pageNumber="267">10.6</td>
<td id="B5285D840E04AAF3FC19C171FBDE0E4E" box="[998,1150,1565,1587]" gridcol="3" gridrow="33" pageId="10" pageNumber="267">9.8</td>
<td id="B5285D840E04AAF3FB4AC171FAB80E4E" box="[1205,1304,1565,1587]" gridcol="4" gridrow="33" pageId="10" pageNumber="267">8.4</td>
<td id="B5285D840E04AAF3FAC5C171FAD60E4E" box="[1338,1398,1565,1587]" gridcol="5" gridrow="33" pageId="10" pageNumber="267">9.8</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C12AFAD60E21" box="[159,1398,1606,1628]" gridrow="34" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C12AFDEB0E21" box="[159,587,1606,1628]" gridcol="0" gridrow="34" pageId="10" pageNumber="267">Body width—second dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C12AFD7C0E21" box="[623,732,1606,1628]" gridcol="1" gridrow="34" pageId="10" pageNumber="267">5.0</td>
<td id="B5285D840E04AAF3FD02C12AFC1B0E21" box="[765,955,1606,1628]" gridcol="2" gridrow="34" pageId="10" pageNumber="267">5.2</td>
<td id="B5285D840E04AAF3FC19C12AFBDE0E21" box="[998,1150,1606,1628]" gridcol="3" gridrow="34" pageId="10" pageNumber="267">4.9</td>
<td id="B5285D840E04AAF3FB4AC12AFAB80E21" box="[1205,1304,1606,1628]" gridcol="4" gridrow="34" pageId="10" pageNumber="267">4.2</td>
<td id="B5285D840E04AAF3FAC5C12AFAD60E21" box="[1338,1398,1606,1628]" gridcol="5" gridrow="34" pageId="10" pageNumber="267">4.9</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C11CFAD60EFB" box="[159,1398,1648,1670]" gridrow="35" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C11CFDEB0EFB" box="[159,587,1648,1670]" gridcol="0" gridrow="35" pageId="10" pageNumber="267">Body depth—maximum (scap)</th>
<td id="B5285D840E04AAF3FD90C11CFD7C0EFB" box="[623,732,1648,1670]" gridcol="1" gridrow="35" pageId="10" pageNumber="267">5.1</td>
<td id="B5285D840E04AAF3FD02C11CFC1B0EFB" box="[765,955,1648,1670]" gridcol="2" gridrow="35" pageId="10" pageNumber="267">5.8</td>
<td id="B5285D840E04AAF3FC19C11CFBDE0EFB" box="[998,1150,1648,1670]" gridcol="3" gridrow="35" pageId="10" pageNumber="267">5.0</td>
<td id="B5285D840E04AAF3FB4AC11CFAB80EFB" box="[1205,1304,1648,1670]" gridcol="4" gridrow="35" pageId="10" pageNumber="267">3.9</td>
<td id="B5285D840E04AAF3FAC5C11CFAD60EFB" box="[1338,1398,1648,1670]" gridcol="5" gridrow="35" pageId="10" pageNumber="267">4.2</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C1F5FAD60ED2" box="[159,1398,1689,1711]" gridrow="36" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C1F5FDEB0ED2" box="[159,587,1689,1711]" gridcol="0" gridrow="36" pageId="10" pageNumber="267">Body depth—pelvic-fin insertion</th>
<td id="B5285D840E04AAF3FD90C1F5FD7C0ED2" box="[623,732,1689,1711]" gridcol="1" gridrow="36" pageId="10" pageNumber="267">5.1</td>
<td id="B5285D840E04AAF3FD02C1F5FC1B0ED2" box="[765,955,1689,1711]" gridcol="2" gridrow="36" pageId="10" pageNumber="267">5.1</td>
<td id="B5285D840E04AAF3FC19C1F5FBDE0ED2" box="[998,1150,1689,1711]" gridcol="3" gridrow="36" pageId="10" pageNumber="267">4.7</td>
<td id="B5285D840E04AAF3FB4AC1F5FAB80ED2" box="[1205,1304,1689,1711]" gridcol="4" gridrow="36" pageId="10" pageNumber="267">3.6</td>
<td id="B5285D840E04AAF3FAC5C1F5FAD60ED2" box="[1338,1398,1689,1711]" gridcol="5" gridrow="36" pageId="10" pageNumber="267">3.9</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C1AEFAD60EA5" box="[159,1398,1730,1752]" gridrow="37" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C1AEFDEB0EA5" box="[159,587,1730,1752]" gridcol="0" gridrow="37" pageId="10" pageNumber="267">Body depth—first dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C1AEFD7C0EA5" box="[623,732,1730,1752]" gridcol="1" gridrow="37" pageId="10" pageNumber="267">3.8</td>
<td id="B5285D840E04AAF3FD02C1AEFC1B0EA5" box="[765,955,1730,1752]" gridcol="2" gridrow="37" pageId="10" pageNumber="267">4.0</td>
<td id="B5285D840E04AAF3FC19C1AEFBDE0EA5" box="[998,1150,1730,1752]" gridcol="3" gridrow="37" pageId="10" pageNumber="267">4.0</td>
<td id="B5285D840E04AAF3FB4AC1AEFAB80EA5" box="[1205,1304,1730,1752]" gridcol="4" gridrow="37" pageId="10" pageNumber="267">3.2</td>
<td id="B5285D840E04AAF3FAC5C1AEFAD60EA5" box="[1338,1398,1730,1752]" gridcol="5" gridrow="37" pageId="10" pageNumber="267">3.5</td>
</tr>
<tr id="F6F934F80E04AAF3FF60C180FAD60F7F" box="[159,1398,1772,1794]" gridrow="38" pageId="10" pageNumber="267">
<th id="B5285D840E04AAF3FF60C180FDEB0F7F" box="[159,587,1772,1794]" gridcol="0" gridrow="38" pageId="10" pageNumber="267">Body depth—second dorsal-fin origin</th>
<td id="B5285D840E04AAF3FD90C180FD7C0F7F" box="[623,732,1772,1794]" gridcol="1" gridrow="38" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FD02C180FC1B0F7F" box="[765,955,1772,1794]" gridcol="2" gridrow="38" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FC19C180FBDE0F7F" box="[998,1150,1772,1794]" gridcol="3" gridrow="38" pageId="10" pageNumber="267">2.5</td>
<td id="B5285D840E04AAF3FB4AC180FAB80F7F" box="[1205,1304,1772,1794]" gridcol="4" gridrow="38" pageId="10" pageNumber="267">2.1</td>
<td id="B5285D840E04AAF3FAC5C180FAD60F7F" box="[1338,1398,1772,1794]" gridcol="5" gridrow="38" pageId="10" pageNumber="267">2.3</td>
</tr>
</table>
</paragraph>
<subSubSection id="00D365310E045501FF38C051FE1A0AB0" lastPageId="13" lastPageNumber="270" pageId="10" pageNumber="267" type="discussion">
<paragraph id="487636BA0E045507FF38C051FBFF093D" blockId="10.[151,1436,1853,2022]" lastBlockId="11.[151,1436,151,320]" lastPageId="11" lastPageNumber="268" pageId="10" pageNumber="267">
<emphasis id="7ABDEAA80E045506FF38C051FE9B0F2B" bold="true" box="[199,315,1853,1878]" pageId="10" pageNumber="267">Remarks.</emphasis>
<taxonomicName id="8FC94D390E045506FEBBC051FD870F28" authorityName=": Fowler" authorityYear="1941" box="[324,551,1853,1877]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="267" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E045506FEBBC051FD870F28" box="[324,551,1853,1877]" italics="true" pageId="10" pageNumber="267">Rhinobatos lionotus</emphasis>
</taxonomicName>
remains poorly known and more material is needed to adequately define the species and determine its geographic range. However, data and images from the recently acquired specimen from 
<collectingCountry id="30DE762A0E045506FF68C0EAFEA40FE3" box="[151,260,1926,1950]" name="Myanmar" pageId="10" pageNumber="267">Myanmar</collectingCountry>
(kindly made available for study by P. Psomadakis), are congruent with Norman’s 
<typeStatus id="977288180E045506FB36C0E9FA8B0FE3" box="[1225,1323,1925,1950]" pageId="10" pageNumber="267" type="holotype">holotype</typeStatus>
and have helped immensely. Sequence, meristic and morphometric data suggest this species is most closely allied to 
<taxonomicName id="8FC94D390E045506FA7BC0C6FEB50F98" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="267" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E045506FA7BC0C6FEB50F98" italics="true" pageId="10" pageNumber="267">R. annandalei</emphasis>
</taxonomicName>
and 
<taxonomicName id="8FC94D390E045506FEB5C0A2FDEE0F9B" authority="(Indonesia)" baseAuthorityName="Indonesia" box="[330,590,1997,2022]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="267" phylum="Chordata" rank="species" species="pengalli">
<emphasis id="7ABDEAA80E045506FEB5C0A2FE640F98" box="[330,452,1997,2021]" italics="true" pageId="10" pageNumber="267">R. pengalli</emphasis>
(
<collectingCountry id="30DE762A0E045506FE2DC0A1FDE60F9B" box="[466,582,1997,2022]" name="Indonesia" pageId="10" pageNumber="267">Indonesia</collectingCountry>
)
</taxonomicName>
; all three species are characterised by a large, broad disc (width 38–44% TL in 
<taxonomicName id="8FC94D390E055507FF4AC7F5FEF308CD" box="[181,339,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E055507FF4AC7F5FEF308CD" box="[181,339,152,176]" italics="true" pageId="11" pageNumber="268">R. annandalei</emphasis>
</taxonomicName>
, 35–37% TL in 
<taxonomicName id="8FC94D390E055507FDEEC7F5FD2B08CD" authorityName=": Fowler" authorityYear="1941" box="[529,651,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E055507FDEEC7F5FD2B08CD" box="[529,651,152,176]" italics="true" pageId="11" pageNumber="268">R. lionotus</emphasis>
</taxonomicName>
, 36–37% TL in 
<taxonomicName id="8FC94D390E055507FCB6C7F5FC6608CD" box="[841,966,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="pengalli">
<emphasis id="7ABDEAA80E055507FCB6C7F5FC6608CD" box="[841,966,152,176]" italics="true" pageId="11" pageNumber="268">R. pengalli</emphasis>
</taxonomicName>
), broad mouth (width 7.1–8.0% TL in 
<taxonomicName id="8FC94D390E055507FA7BC7F5FEB508A9" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E055507FA7BC7F5FEB508A9" italics="true" pageId="11" pageNumber="268">R. annandalei</emphasis>
</taxonomicName>
, 6.3–6.9% TL in 
<taxonomicName id="8FC94D390E055507FE20C7D1FDFA08A9" authorityName=": Fowler" authorityYear="1941" box="[479,602,188,212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E055507FE20C7D1FDFA08A9" box="[479,602,188,212]" italics="true" pageId="11" pageNumber="268">R. lionotus</emphasis>
</taxonomicName>
, 6.0–6.7% TL in 
<taxonomicName id="8FC94D390E055507FCDBC7D1FC0008A9" box="[804,928,188,212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="pengalli">
<emphasis id="7ABDEAA80E055507FCDBC7D1FC0008A9" box="[804,928,188,212]" italics="true" pageId="11" pageNumber="268">R. pengalli</emphasis>
</taxonomicName>
), and relatively low post-synarcual vertebral counts (~158 centra in 
<taxonomicName id="8FC94D390E055507FE64C78DFD980885" box="[411,568,224,248]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E055507FE64C78DFD980885" box="[411,568,224,248]" italics="true" pageId="11" pageNumber="268">R. annandalei</emphasis>
</taxonomicName>
, ~ 
<quantity id="8F319B5F0E055507FDAAC7B3FD3D0885" box="[597,669,223,248]" metricMagnitude="0" metricUnit="m" metricValue="4.1655999999999995" pageId="11" pageNumber="268" unit="in" value="164.0">164 in</quantity>
<taxonomicName id="8FC94D390E055507FD5BC78DFCBE0885" authorityName=": Fowler" authorityYear="1941" box="[676,798,224,248]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E055507FD5BC78DFCBE0885" box="[676,798,224,248]" italics="true" pageId="11" pageNumber="268">R. lionotus</emphasis>
</taxonomicName>
, and 
<quantity id="8F319B5F0E055507FCA2C7B3FC7C0885" box="[861,988,223,248]" metricMagnitude="0" metricUnit="m" metricValue="4.2672" metricValueMax="4.318" metricValueMin="4.2164" pageId="11" pageNumber="268" unit="in" value="168.0" valueMax="170.0" valueMin="166.0">166–170 in</quantity>
<taxonomicName id="8FC94D390E055507FC1BC78DFBFF0885" box="[996,1119,224,248]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="pengalli">
<emphasis id="7ABDEAA80E055507FC1BC78DFBFF0885" box="[996,1119,224,248]" italics="true" pageId="11" pageNumber="268">R. pengalli</emphasis>
</taxonomicName>
). The two other 
<taxonomicName id="8FC94D390E055507FAE3C78CFA3B0885" box="[1308,1435,224,248]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="7ABDEAA80E055507FAE3C78CFA3B0885" box="[1308,1435,224,248]" italics="true" pageId="11" pageNumber="268">Rhinobatos</emphasis>
</taxonomicName>
species in the Northern Indian Ocean, 
<taxonomicName id="8FC94D390E055507FDB1C669FD740961" authorityName="Compagno &amp; Randall" authorityYear="1987" box="[590,724,260,284]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="punctifer">
<emphasis id="7ABDEAA80E055507FDB1C669FD740961" box="[590,724,260,284]" italics="true" pageId="11" pageNumber="268">R. punctifer</emphasis>
</taxonomicName>
and 
<taxonomicName id="8FC94D390E055507FCF2C669FC170961" authority="Last &amp; Séret &amp; Naylor, 2019" authorityName="Last &amp; Séret &amp; Naylor" authorityYear="2019" box="[781,951,261,284]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="ranongensis" status="sp. nov.">
<emphasis id="7ABDEAA80E055507FCF2C669FC170961" box="[781,951,261,284]" italics="true" pageId="11" pageNumber="268">R. ranongensis</emphasis>
</taxonomicName>
<emphasis id="7ABDEAA80E055507FC3FC668FBBB0960" bold="true" box="[960,1051,260,285]" pageId="11" pageNumber="268">
<taxonomicNameLabel id="618E57D30E055507FC3FC668FBBB0960" box="[960,1051,260,285]" pageId="11" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, have a narrower disc (width 29– 34% TL) and mouth (width 5.4–6.0% TL), and more post-synarcual centra (172–184).
</paragraph>
<caption id="1CB666320E055507FF68C0F2FBE40FAE" httpUri="https://zenodo.org/record/2624902/files/figure.png" pageId="11" pageNumber="268" startId="11.[151,250,1950,1972]" targetBox="[177,1416,398,1898]" targetPageId="11">
<paragraph id="487636BA0E055507FF68C0F2FBE40FAE" blockId="11.[151,1435,1950,2003]" pageId="11" pageNumber="268">
<emphasis id="7ABDEAA80E055507FF68C0F2FEB20FC9" bold="true" box="[151,274,1950,1972]" pageId="11" pageNumber="268">FIGURE 4.</emphasis>
<taxonomicName id="8FC94D390E055507FEE6C0F2FE430FC9" authorityName=": Fowler" authorityYear="1941" box="[281,483,1950,1972]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="268" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E055507FEE6C0F2FE430FC9" box="[281,483,1950,1972]" italics="true" pageId="11" pageNumber="268">Rhinobatos lionotus</emphasis>
</taxonomicName>
: A. female holotype, 495 mm TL (BMNH 1909.7.12.3), India, dorsal view, preserved; female non-type, 760 mm TL (CSIRO H 7867-01), Myanmar, fresh, B. dorsal view, C. ventral view.
</paragraph>
</caption>
<caption id="1CB666320E025500FF68C0D4FC1C0F90" httpUri="https://zenodo.org/record/2624904/files/figure.png" pageId="12" pageNumber="269" startId="12.[151,250,1976,1998]" targetBox="[330,1252,204,1907]" targetPageId="12">
<paragraph id="487636BA0E025500FF68C0D4FC1C0F90" blockId="12.[151,1436,1976,2029]" pageId="12" pageNumber="269">
<emphasis id="7ABDEAA80E025500FF68C0D4FEB50FB3" bold="true" box="[151,277,1976,1998]" pageId="12" pageNumber="269">FIGURE 5.</emphasis>
Lateral views of the unpaired fins of 
<taxonomicName id="8FC94D390E025500FD58C0D4FCD50FB3" authorityName=": Fowler" authorityYear="1941" box="[679,885,1976,1998]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="269" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E025500FD58C0D4FCD50FB3" box="[679,885,1976,1998]" italics="true" pageId="12" pageNumber="269">Rhinobatos lionotus</emphasis>
</taxonomicName>
, female non-type, 760 mm TL (CSIRO H 7867-01), Myanmar, preserved: A. first dorsal fin; B. second dorsal fin; and C. caudal fin.
</paragraph>
</caption>
<paragraph id="487636BA0E035501FF38C7FBFC2709F5" blockId="13.[151,1436,151,717]" pageId="13" pageNumber="270">
The newly examined material fit Norman’s descriptions of 
<taxonomicName id="8FC94D390E035501FC82C7F5FC5A08CD" authorityName=": Fowler" authorityYear="1941" box="[893,1018,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FC82C7F5FC5A08CD" box="[893,1018,152,176]" italics="true" pageId="13" pageNumber="270">R. lionotus</emphasis>
</taxonomicName>
and 
<taxonomicName id="8FC94D390E035501FBC7C7F5FB7908CD" box="[1080,1241,152,176]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E035501FBC7C7F5FB7908CD" box="[1080,1241,152,176]" italics="true" pageId="13" pageNumber="270">R. annandalei</emphasis>
</taxonomicName>
well despite the absence of information on colour in preservation in the original descriptions. 
<taxonomicName id="8FC94D390E035501FBF6C7D0FB4B08A9" authorityName=": Fowler" authorityYear="1941" box="[1033,1259,188,212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FBF6C7D0FB4B08A9" box="[1033,1259,188,212]" italics="true" pageId="13" pageNumber="270">Rhinobatos lionotus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="8FC94D390E035501FA7BC7D1FEB50885" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E035501FA7BC7D1FEB50885" italics="true" pageId="13" pageNumber="270">R. annandalei</emphasis>
</taxonomicName>
, in lacking symmetrically arranged white spots (being uniformly coloured or with faint or well-defined dark blotches) on the dorsal surface. However, squamation is particularly useful for distinguishing these species; Norman noticed the absence of a median row of short thorns in 
<taxonomicName id="8FC94D390E035501FC81C645FC59093D" authorityName=": Fowler" authorityYear="1941" box="[894,1017,296,320]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FC81C645FC59093D" box="[894,1017,296,320]" italics="true" pageId="13" pageNumber="270">R. lionotus</emphasis>
</taxonomicName>
and their presence in 
<taxonomicName id="8FC94D390E035501FB03C645FA3B093D" box="[1276,1435,296,320]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E035501FB03C645FA3B093D" box="[1276,1435,296,320]" italics="true" pageId="13" pageNumber="270">R. annandalei</emphasis>
</taxonomicName>
(evident in his text figs. 13 and 14 and in our material). These species occur sympatrically at their 
<typeStatus id="977288180E035501FAF0C621FA9F0918" box="[1295,1343,333,357]" pageId="13" pageNumber="270">type</typeStatus>
locality (Hooghly River mouth, Bay of Bengal) and presumably elsewhere.
</paragraph>
<paragraph id="487636BA0E035501FF38C6F8FE1A0AB0" blockId="13.[151,1436,151,717]" pageId="13" pageNumber="270">
Annandale (1909) confused 
<taxonomicName id="8FC94D390E035501FDF5C6F8FD4C09D1" authorityName=": Fowler" authorityYear="1941" box="[522,748,404,428]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FDF5C6F8FD4C09D1" box="[522,748,404,428]" italics="true" pageId="13" pageNumber="270">Rhinobatos lionotus</emphasis>
</taxonomicName>
with 
<taxonomicName id="8FC94D390E035501FCD0C6F8FC0E09D1" box="[815,942,404,428]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="genus">
<emphasis id="7ABDEAA80E035501FCD0C6F8FC0E09D1" box="[815,942,404,428]" italics="true" pageId="13" pageNumber="270">Rhinobatos</emphasis>
</taxonomicName>
(as 
<taxonomicName id="8FC94D390E035501FC1FC6F8FBF909D1" authorityName="Blainville" authorityYear="1825" box="[992,1113,404,428]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="genus">
<emphasis id="7ABDEAA80E035501FC1FC6F8FBF909D1" box="[992,1113,404,428]" italics="true" pageId="13" pageNumber="270">Rhinobatis</emphasis>
</taxonomicName>
) 
<taxonomicName id="8FC94D390E035501FB93C6F8FB7709D1" box="[1132,1239,404,428]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="schlegelii">
<emphasis id="7ABDEAA80E035501FB93C6F8FB7709D1" box="[1132,1239,404,428]" italics="true" pageId="13" pageNumber="270">schlegelii</emphasis>
</taxonomicName>
based on several adult specimens taken from the 
<typeStatus id="977288180E035501FDFEC6D4FD9109AD" box="[513,561,440,464]" pageId="13" pageNumber="270">type</typeStatus>
locality. A somewhat similar guitarfish, 
<taxonomicName id="8FC94D390E035501FBFFC6D4FB5209AD" authorityName="Muller &amp; Henle" authorityYear="1841" box="[1024,1266,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="schlegelii">
<emphasis id="7ABDEAA80E035501FBFFC6D4FB5209AD" box="[1024,1266,440,464]" italics="true" pageId="13" pageNumber="270">Rhinobatos schlegelii</emphasis>
</taxonomicName>
, is confined to the western North Pacific from 
<collectingCountry id="30DE762A0E035501FDFBC6B1FDE40989" box="[516,580,477,500]" name="Japan" pageId="13" pageNumber="270">Japan</collectingCountry>
to 
<collectingCountry id="30DE762A0E035501FD93C6B0FD1E0988" box="[620,702,476,501]" name="Taiwan" pageId="13" pageNumber="270">Taiwan</collectingCountry>
(
<bibRefCitation id="2C584B4B0E035501FD30C6B0FC390988" author="Seret, B. &amp; Last, P. R. &amp; Naylor, G. J. P." box="[719,921,476,501]" pageId="13" pageNumber="270" pagination="77 - 109" refId="ref13286" refString="Seret, B., Last, P. R. &amp; Naylor, G. J. P. (2016) Guitarfishes. Family Rhinobatidae. In: Last, P. R., White, W. T., de Carvalho, M., Seret, B. Stehmann, M. &amp; Naylor, G. J. P. (Eds.), Rays of the World. CSIRO Publishing, Comstock Publishing Associates, pp. 77 - 109." type="book chapter" year="2016">
Séret 
<emphasis id="7ABDEAA80E035501FCECC6B1FCE70989" box="[787,839,476,500]" italics="true" pageId="13" pageNumber="270">et al</emphasis>
., 2016
</bibRefCitation>
, Séret &amp; Last, unpublished data). Annandale described the colour of these adult 
<taxonomicName id="8FC94D390E035501FDCCC56DFD0F0A65" authorityName=": Fowler" authorityYear="1941" box="[563,687,512,536]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FDCCC56DFD0F0A65" box="[563,687,512,536]" italics="true" pageId="13" pageNumber="270">R. lionotus</emphasis>
</taxonomicName>
as uniform brownish grey to olive green, but noted that unborn young had ‘numerous faint white spots all over the dorsal surface ...’. His reference to ‘all over the dorsal surface’ suggests the spots were densely arranged (unlike the regular pattern of sparse spots in 
<taxonomicName id="8FC94D390E035501FB93C525FAAA0A1D" box="[1132,1290,584,608]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7ABDEAA80E035501FB93C525FAAA0A1D" box="[1132,1290,584,608]" italics="true" pageId="13" pageNumber="270">R. annandalei</emphasis>
</taxonomicName>
). Annandale also made no reference to white spots in adults, so if these unborn were young of 
<taxonomicName id="8FC94D390E035501FBDBC501FB3D0AF9" authorityName=": Fowler" authorityYear="1941" box="[1060,1181,620,644]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="270" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7ABDEAA80E035501FBDBC501FB3D0AF9" box="[1060,1181,620,644]" italics="true" pageId="13" pageNumber="270">R. lionotus</emphasis>
</taxonomicName>
, presumably the white spots are an ontogenetic feature that is lost as the fish grows. More material of the young of both species is needed to shed light on this issue.
</paragraph>
</subSubSection>
</treatment>
</document>