262 lines
36 KiB
XML
262 lines
36 KiB
XML
<document id="663489E0CB11772B244D5BC8E987131D" ID-CLB-Dataset="3320" ID-DOI="10.1093/zoolinnean/zlaa052" ID-GBIF-Dataset="fd4b2931-7bad-4644-b485-f365040ae01e" ID-ISSN="0024-4082" ID-Zenodo-Dep="5724192" ID-ZooBank="0810A91E-7D8D-4F56-B468-9D5B3A6115E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628947432" checkinUser="carolina" docAuthor="Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D." docDate="2021" docId="BA5587AF637F4A18901F6C912E8DFB7F" docLanguage="en" docName="ZoolJLinnSoc.191.883-901.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Toktokkus Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith 2021" docType="key" docVersion="6" lastPageNumber="894" masterDocId="466CFFD763744A13920F6C4F2B04FFCB" masterDocTitle="First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group" masterLastPageNumber="901" masterPageNumber="883" pageNumber="894" updateTime="1698937967960" updateUser="ExternalLinkService">
|
||
<mods:mods id="D1855324159D4E606B23FE2DBE1B1C38" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="858A0121E563D75AE6FFC5CD2026D82F">
|
||
<mods:title id="A697D4CF00EB2C3BF9E650B43346A40F">First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="18F9563AE67ABE7E8B08344FFF65438B" type="personal">
|
||
<mods:role id="F6F02017BE4610B61BAB7F7CCB56565C">
|
||
<mods:roleTerm id="B6184626BEF383C2DDDA76FD53E150CB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="49CB560F35CBB1FB290F862E50574918">Kamiński, Marcin J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F75168CC3DB7D3324448F80EE33FAFB7" type="personal">
|
||
<mods:role id="9FE0B8DBF0EA6C98401FAF182ACA1491">
|
||
<mods:roleTerm id="FBADB7B58ABA1FD386E4BC72220A6FB6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A0B4AB5D802AC4E2DBD56969692A0042">Gearner, Olivia M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B6D872EC0F0B1A8EBDDC0057581FB84D" type="personal">
|
||
<mods:role id="EA922F8224C94198858C644CF4875936">
|
||
<mods:roleTerm id="865334F9C73EE6D0213D512300B8D79E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9A3BD4345B2C78CD0565EFED5F335AE2">Kanda, Kojun</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C0E447083D2705F8C2BA45C072FC2FEA" type="personal">
|
||
<mods:role id="5F4D43F223236813B69EE562FA845393">
|
||
<mods:roleTerm id="35328633A5E6328375DE365C2030CD1C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B258E504230470EDE4E0967320A8ED4E">Swichtenberg, Kali</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CC966AE9E14AA4026BAE9484F474AB4F" type="personal">
|
||
<mods:role id="01AB3C63029592815783864BDF94C448">
|
||
<mods:roleTerm id="C7230384578C74CD46CBC4A1A71BF764">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E9D804309ACCB9966FBFEC0C71FCB3CC">Purchart, Luboš</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="45E0F2F6F2562281ACFE99E4078BD292" type="personal">
|
||
<mods:role id="CEE326E2D76A3FCA7EDA4043336682BE">
|
||
<mods:roleTerm id="1834C4A9AE21623D9B3BD58A93C261FC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6B2C100F675DF31639425643FC55D886">Smith, Aaron D.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="018E267480EDC1A0011A4A8CE55E3351">text</mods:typeOfResource>
|
||
<mods:relatedItem id="290F03D599003E936B9EC26A137AA39B" type="host">
|
||
<mods:titleInfo id="EC35D9BF9340CD9A68C086ADC2473F68">
|
||
<mods:title id="5C2F6DF3FF2B4D8256D68E3C210B5A7B">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BE566CC70889B7EDB04021F3DDE113E4">
|
||
<mods:date id="AF7D597ABC184196580A7B70D11DC035">2021</mods:date>
|
||
<mods:detail id="C6400D05D3C9B47E1970157B97DFECDF" type="pubDate">
|
||
<mods:number id="5CF082298F1ED589C18156D8121F1571">2020-06-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F2DA03BA5598876C352DBABB459AC791" type="volume">
|
||
<mods:number id="03F4B06B023804EE09891C311C63DEFF">191</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4D44AD9FC07DFCE16F165422D30EB700" unit="page">
|
||
<mods:start id="338250F9546E924AB44DEBD6AFB8AAE7">883</mods:start>
|
||
<mods:end id="68B8FEB9809643AE3EC04018C85803B7">901</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="FCFF791FFE01846F9D9D135834207D00">journal article</mods:classification>
|
||
<mods:identifier id="BD90E354239863AD4354377100DCDF7B" type="CLB-Dataset">3320</mods:identifier>
|
||
<mods:identifier id="24BDC8A9EB7AF42187AEA9F7AA96CC09" type="DOI">10.1093/zoolinnean/zlaa052</mods:identifier>
|
||
<mods:identifier id="24910F1A7E2448DECEA9382EA0EEB609" type="GBIF-Dataset">fd4b2931-7bad-4644-b485-f365040ae01e</mods:identifier>
|
||
<mods:identifier id="C539DA6DCC87C2F313CA82039AE53844" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="E75B0B570B691C52A4D0FDB9171763CF" type="Zenodo-Dep">5724192</mods:identifier>
|
||
<mods:identifier id="7A0E348013A6147A89F3701E6B97649E" type="ZooBank">0810A91E-7D8D-4F56-B468-9D5B3A6115E9</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="BA5587AF637F4A18901F6C912E8DFB7F" ID-DOI="http://doi.org/10.5281/zenodo.5724246" ID-GBIF-Taxon="190683655" ID-Zenodo-Dep="5724246" LSID="urn:lsid:plazi:treatment:BA5587AF637F4A18901F6C912E8DFB7F" httpUri="http://treatment.plazi.org/id/BA5587AF637F4A18901F6C912E8DFB7F" lastPageNumber="894" pageId="11" pageNumber="894">
|
||
<table id="40FCC419637FB5EC92AE6C8C2EA7FB01" box="[161,1443,195,1226]" pageId="11" pageNumber="894">
|
||
<subSubSection id="7AE66532637F4A18901F6C912F32FF3D" box="[528,1078,222,247]" pageId="11" pageNumber="894" type="nomenclature">
|
||
<paragraph id="324336B9637F4A18901F6C912F32FF3D" blockId="11.[528,1078,222,247]" box="[528,1078,222,247]" pageId="11" pageNumber="894">
|
||
<heading id="690B81D5637F4A18901F6C912F32FF3D" box="[528,1078,222,247]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="894" reason="2">
|
||
KEY TO THE SPECIES OF THE GENUS
|
||
<taxonomicName id="F5FC4D3A637F4A1891A36C902F32FF3D" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" box="[940,1078,223,246]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="901" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0088EAAB637F4A1891A36C902F32FF3D" box="[940,1078,223,246]" italics="true" pageId="11" pageNumber="894">TOKTOKKUS</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AE66532637F4A1892C36D482E8DFB7F" pageId="11" pageNumber="894" type="key">
|
||
<keyStep id="89082E1C637F4A1892C36D482E8CFEF0" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36D482E8CFED7" blockId="11.[188,1417,263,1205]" box="[204,1416,263,285]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36D482E8CFED7" box="[204,1416,263,285]" pageId="11" pageNumber="894">
|
||
1. Apex of elytra not sloped (dorsal view) and depressed with clear margin (
|
||
<figureCitation id="AAC72A3C637F4A1896256D482F7FFED7" box="[1066,1147,263,285]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3B</figureCitation>
|
||
)...............
|
||
<taxonomicName id="F5FC4D3A637F4A1896E36D472E8CFED7" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1260,1416,263,285]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="congolensis">
|
||
<emphasis id="0088EAAB637F4A1896E36D472E8CFED7" box="[1260,1416,263,285]" italics="true" pageId="11" pageNumber="894">T. congolensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86D692E8CFEF0" blockId="11.[188,1417,263,1205]" box="[215,1416,294,316]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86D692E8CFEF0" box="[215,1416,294,316]" pageId="11" pageNumber="894">
|
||
- Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g.
|
||
<figureCitation id="AAC72A3C637F4A1891D46D692F2FFEF0" box="[987,1067,294,316]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3A</figureCitation>
|
||
) ...............................................2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36D0B2E8CFE7E" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36D0B2E8CFEB3" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36D0B2E8CFEB3" pageId="11" pageNumber="894">
|
||
2. Margin of prosternal collar expanded and folded out into a large lip (
|
||
<figureCitation id="AAC72A3C637F4A1891FE6D0B2F45FE92" box="[1009,1089,324,346]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3C</figureCitation>
|
||
); elytra round, almost heart- shaped (
|
||
<figureCitation id="AAC72A3C637F4A1893436D2C2A99FEB3" box="[332,413,355,377]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3C</figureCitation>
|
||
); declivous portion of elytra with little to no tubercles.....................................
|
||
<taxonomicName id="F5FC4D3A637F4A1896F06D2B2E8CFEB3" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1279,1416,355,377]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="tschinkeli">
|
||
<emphasis id="0088EAAB637F4A1896F06D2B2E8CFEB3" box="[1279,1416,355,377]" italics="true" pageId="11" pageNumber="894">T. tschinkeli</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86DCD2E8CFE7E" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86DCD2E8CFE7E" pageId="11" pageNumber="894">- Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows .........................................................................................................3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36DF02E8CFE38" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36DF02E8CFE1F" blockId="11.[188,1417,263,1205]" box="[204,1416,447,469]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36DF02E8CFE1F" box="[204,1416,447,469]" pageId="11" pageNumber="894">
|
||
3. Disc of pronotum with prominent punctures (
|
||
<figureCitation id="AAC72A3C637F4A1890FF6DF02847FE1F" box="[752,835,447,469]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3D</figureCitation>
|
||
); gold setae present on elytra (
|
||
<figureCitation id="AAC72A3C637F4A1896AD6DF02FF0FE1F" box="[1186,1268,447,469]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3D</figureCitation>
|
||
).....
|
||
<taxonomicName id="F5FC4D3A637F4A1897296D8F2E8CFE1F" authorityName="Gearner" authorityYear="2021" box="[1318,1416,447,469]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="0088EAAB637F4A1897296D8F2E8CFE1F" box="[1318,1416,447,469]" italics="true" pageId="11" pageNumber="894">T. herero</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86D912E8CFE38" blockId="11.[188,1417,263,1205]" box="[215,1416,478,500]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86D912E8CFE38" box="[215,1416,478,500]" pageId="11" pageNumber="894">- Disc of pronotum only with micropunctures; elytra not covered with setae................................................4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36DB32E8CFDFA" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36DB32E8CFDD9" blockId="11.[188,1417,263,1205]" box="[204,1416,508,530]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36DB32E8CFDD9" box="[204,1416,508,530]" pageId="11" pageNumber="894">
|
||
4. Tubercles on elytral sides round/globular, vertically direct (
|
||
<figureCitation id="AAC72A3C637F4A1891836DB328FEFDDA" box="[908,1018,508,530]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2B, C</figureCitation>
|
||
) ......................................................5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86E542E8CFDFA" blockId="11.[188,1417,263,1205]" box="[215,1416,539,561]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86E542E8CFDFA" box="[215,1416,539,561]" pageId="11" pageNumber="894">
|
||
- Tubercles on elytral sides pointed, directed posteriorly (e.g.
|
||
<figureCitation id="AAC72A3C637F4A18919F6E5428DBFDFB" box="[912,991,539,561]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3F</figureCitation>
|
||
) ..........................................................6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36E762E8CFD60" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36E762E8DFDA6" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36E762E8DFDA6" pageId="11" pageNumber="894">
|
||
5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round (
|
||
<figureCitation id="AAC72A3C637F4A1897236E752E84FD84" box="[1324,1408,570,592]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2B</figureCitation>
|
||
) ...........................................................................................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A1897166E162E8DFDA6" ID-CoL="4NCBY" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1899" box="[1305,1417,600,622]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="mulleri">
|
||
<emphasis id="0088EAAB637F4A1897166E162E8DFDA6" box="[1305,1417,600,622]" italics="true" pageId="11" pageNumber="894">T. mulleri</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86E382E8CFD60" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86E382E8CFD60" pageId="11" pageNumber="894">
|
||
- Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round (
|
||
<figureCitation id="AAC72A3C637F4A1893316ED92A8BFD60" box="[318,399,662,684]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2C</figureCitation>
|
||
)................................................................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A1897056ED92E8CFD60" ID-CoL="6WBRW" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1908" box="[1290,1416,662,683]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="schultzei">
|
||
<emphasis id="0088EAAB637F4A1897056ED92E8CFD60" box="[1290,1416,662,683]" italics="true" pageId="11" pageNumber="894">T. schultzei</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36EFB2E8CFD23" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36EFB2E8CFD01" blockId="11.[188,1417,263,1205]" box="[204,1416,692,714]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36EFB2E8CFD01" box="[204,1416,692,714]" pageId="11" pageNumber="894">6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86E9C2E8CFD23" blockId="11.[188,1417,263,1205]" box="[215,1416,723,745]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86E9C2E8CFD23" box="[215,1416,723,745]" pageId="11" pageNumber="894">
|
||
- Elytral tubercles dense (more than 6 lateral rows), slightly angled (
|
||
<figureCitation id="AAC72A3C637F4A1891E66E9C2F5CFD23" box="[1001,1112,723,745]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2B–F</figureCitation>
|
||
).........................................8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36EBD2E8CFC8F" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36EBD2E8CFCCC" blockId="11.[188,1417,263,1205]" box="[204,1416,754,776]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36EBD2E8CFCCC" box="[204,1416,754,776]" pageId="11" pageNumber="894">
|
||
7. Elytral tubercles distinct, not confluent, nearly reaching humeri (
|
||
<figureCitation id="AAC72A3C637F4A1891DA6EBD2F22FCCC" box="[981,1062,754,776]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3A</figureCitation>
|
||
) ................................
|
||
<taxonomicName id="F5FC4D3A637F4A18971F6EBD2E8CFCCC" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1296,1416,754,775]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="barclayi">
|
||
<emphasis id="0088EAAB637F4A18971F6EBD2E8CFCCC" box="[1296,1416,754,775]" italics="true" pageId="11" pageNumber="894">T. barclayi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86F5E2E8CFC8F" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86F5E2E8CFC8F" pageId="11" pageNumber="894">
|
||
- Elytral tubercles small and short, confluent into rows, terminating well before humeri (
|
||
<figureCitation id="AAC72A3C637F4A1897276F5F2E7BFCEE" box="[1320,1407,784,806]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2A</figureCitation>
|
||
) ......................................................................................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A1896F86F7F2E8CFC8F" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1896" box="[1271,1416,815,837]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="mashunus">
|
||
<emphasis id="0088EAAB637F4A1896F86F7F2E8CFC8F" box="[1271,1416,815,837]" italics="true" pageId="11" pageNumber="894">T. mashunus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36F012E8CFC74" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36F012E8CFC4A" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36F012E8CFC4A" pageId="11" pageNumber="894">
|
||
8. Microtubercles present between tuberculate rows (e.g.
|
||
<figureCitation id="AAC72A3C637F4A1891796F0128CEFCA8" box="[886,970,846,868]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3E</figureCitation>
|
||
), tuberculate rows rarely elevated on ridges ...............................................................................................................................................................9
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D86FC42E8CFC74" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D86FC42E8CFC74" pageId="11" pageNumber="894">- Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges ................................................................................................................................10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892C36F872E8CFBF2" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892C36F872E8CFC37" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892C36F872E8CFC37" pageId="11" pageNumber="894">
|
||
9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short ....................................................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A1896B26FA72E8CFC37" box="[1213,1416,999,1021]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="tuberculipennis">
|
||
<emphasis id="0088EAAB637F4A1896B26FA72E8CFC37" box="[1213,1416,999,1021]" italics="true" pageId="11" pageNumber="894">T. tuberculipennis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D868492E8CFBF2" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D868492E8CFBF2" pageId="11" pageNumber="894">
|
||
- Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller ................................................................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A18971E686A2E8CFBF2" authorityName="Gearner" authorityYear="2021" box="[1297,1416,1060,1082]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="makuya">
|
||
<emphasis id="0088EAAB637F4A18971E686A2E8CFBF2" box="[1297,1416,1060,1082]" italics="true" pageId="11" pageNumber="894">T. makuya</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="89082E1C637F4A1892B3680C2E8DFB7F" pageId="11" pageNumber="894">
|
||
<paragraph id="324336B9637F4A1892B3680C2E8CFBBD" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892B3680C2E8CFBBD" pageId="11" pageNumber="894">
|
||
10. Elytral tubercles large, often confluent (
|
||
<figureCitation id="AAC72A3C637F4A1890C8680C281EFB93" box="[711,794,1091,1113]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3F</figureCitation>
|
||
); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat (
|
||
<figureCitation id="AAC72A3C637F4A18918C682E28D6FBBC" box="[899,978,1121,1143]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="11" pageNumber="894">Fig. 3F</figureCitation>
|
||
) .........................................
|
||
<taxonomicName id="F5FC4D3A637F4A1896F1682D2E8CFBBD" authorityName="Kaminski" authorityYear="2021" box="[1278,1416,1121,1143]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="waclawae">
|
||
<emphasis id="0088EAAB637F4A1896F1682D2E8CFBBD" box="[1278,1416,1121,1143]" italics="true" pageId="11" pageNumber="894">T. waclawae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="324336B9637F4A1892D868CE2E8DFB7F" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="890D958C637F4A1892D868CE2E8DFB7F" pageId="11" pageNumber="894">
|
||
- Tubercles medium sized (
|
||
<figureCitation id="AAC72A3C637F4A18900968CF296BFB5D" box="[518,623,1152,1174]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724196" httpUri="https://zenodo.org/record/5724196/files/figure.png" pageId="11" pageNumber="894">Fig. 2E, F</figureCitation>
|
||
), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex ....................................................................................
|
||
<taxonomicName id="F5FC4D3A637F4A18972168D02E8DFB7F" ID-CoL="4NCFP" baseAuthorityName="Burchell" baseAuthorityYear="1822" box="[1326,1417,1183,1204]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="vialis">T. vialis</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</table>
|
||
</treatment>
|
||
</document> |