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<document id="538F97C3A8C7E94A2BD7014593FC3A21" ID-CLB-Dataset="3320" ID-DOI="10.1093/zoolinnean/zlaa052" ID-GBIF-Dataset="fd4b2931-7bad-4644-b485-f365040ae01e" ID-ISSN="0024-4082" ID-Zenodo-Dep="5724192" ID-ZooBank="0810A91E-7D8D-4F56-B468-9D5B3A6115E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628947432" checkinUser="carolina" docAuthor="Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš &amp; Smith, Aaron D." docDate="2021" docId="BA5587AF637B4A0391A16E032934F9BC" docLanguage="en" docName="ZoolJLinnSoc.191.883-901.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Toktokkus waclawae Kamiński &amp; Gearner &amp; Kanda &amp; Swichtenberg &amp; Purchart &amp; Smith 2021" docType="treatment" docVersion="6" lastPageNumber="899" masterDocId="466CFFD763744A13920F6C4F2B04FFCB" masterDocTitle="First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group" masterLastPageNumber="901" masterPageNumber="883" pageNumber="898" updateTime="1698937967960" updateUser="ExternalLinkService">
<mods:mods id="176B4C16EA5C8203C2E08EA03AA977B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE43A9F23E64AF08EE6F8BBD21177312">
<mods:title id="8164FDF71D06F51EBD42A18E25AF5B3B">First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group</mods:title>
</mods:titleInfo>
<mods:name id="63C2FAF2DD99F814DDCD39E885D8CE44" type="personal">
<mods:role id="52BB8FFFE4AAA8C128AA0CC311FF062F">
<mods:roleTerm id="25F38A38867BD56B02C6D1D4E0785305">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8C51C846541B01944BDEBF0A3D01C696">Kamiński, Marcin J.</mods:namePart>
</mods:name>
<mods:name id="AB668C4A726F3A96B9516676CFD48A31" type="personal">
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<mods:roleTerm id="D4270C32A5966B942E28E9CD862E8EBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C4042A24660F4CF6F88FD727BA0CB89">Gearner, Olivia M.</mods:namePart>
</mods:name>
<mods:name id="67C20007DE7F0C98D4E7C05E4B819852" type="personal">
<mods:role id="9CECF9CF9B1E9956B305929265B5CF9B">
<mods:roleTerm id="6182E1A62131CC7DDB9D77197C6688CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E037871570D91A7D6CBF04CD0C81F593">Kanda, Kojun</mods:namePart>
</mods:name>
<mods:name id="FFD71C845A90E30BE0C8FCA7529245E1" type="personal">
<mods:role id="EAAE0C6C20A9F958CC11BC741641E923">
<mods:roleTerm id="CC02E1B3D907DBF470D4FF1003E6E7BD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8A46480041DD48734DE9D8A00EEBBEE">Swichtenberg, Kali</mods:namePart>
</mods:name>
<mods:name id="0EE532FB9797679F10054ED305110598" type="personal">
<mods:role id="5979CBC36C4775353D8BB5E08789B08A">
<mods:roleTerm id="680D361F982B2AADD5D068557CCBA960">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFA43221BEA76F6C7902FA24367D0905">Purchart, Luboš</mods:namePart>
</mods:name>
<mods:name id="7BF62147CA4EC54FDEB784C0349DB362" type="personal">
<mods:role id="1A6CA65CD4DBAEA1F19992C51C17C1EC">
<mods:roleTerm id="EE77518D3B258B52198CD3399444B6F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E518C0CFC34878D79F779C5C20C62E0">Smith, Aaron D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1E6C5CAA8BCEA5FA3529BB8974A29913">text</mods:typeOfResource>
<mods:relatedItem id="612B8D3D3213284038E76A6FDD1A4D9D" type="host">
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<mods:title id="6F0578B77A1F7F82152163E65FB8349B">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="6F8685880F2D60EF67FE2D6BCF93B411">
<mods:date id="FC9F7DE96B4AF473E60A3FABDCDDA21D">2021</mods:date>
<mods:detail id="03275906ED8EAC4FE316D0CD2A55F3FA" type="pubDate">
<mods:number id="4CAD777550B6876B8E631CF167D9B8D7">2020-06-16</mods:number>
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<mods:detail id="C8EC042F2C12DA83955E98A4A124BC0F" type="volume">
<mods:number id="1CC735647B5B352B65F0A1E95E44970A">191</mods:number>
</mods:detail>
<mods:extent id="ADA03D1C8A35C775A9C03DF2A625D2C4" unit="page">
<mods:start id="AF0CF7FAA197B541970F53E4C8FD052A">883</mods:start>
<mods:end id="B0392E98BE5594A0DF1EBF5515846AB7">901</mods:end>
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<mods:classification id="C57E8C34697DECBF5547875E4144590E">journal article</mods:classification>
<mods:identifier id="B1F143D18454EB45E5328162596B0B7B" type="CLB-Dataset">3320</mods:identifier>
<mods:identifier id="6409BAE17B0AAE2B11CAA8F286E1B693" type="DOI">10.1093/zoolinnean/zlaa052</mods:identifier>
<mods:identifier id="AF867725DA92705B01ED8068CD9A8464" type="GBIF-Dataset">fd4b2931-7bad-4644-b485-f365040ae01e</mods:identifier>
<mods:identifier id="6E47FB65B7209FB912E9E026346016D8" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="18B73517AA36634F0446B4906F8D725F" type="Zenodo-Dep">5724192</mods:identifier>
<mods:identifier id="75F49B25705C60CA137C4D273E69BEEB" type="ZooBank">0810A91E-7D8D-4F56-B468-9D5B3A6115E9</mods:identifier>
</mods:mods>
<treatment id="BA5587AF637B4A0391A16E032934F9BC" ID-DOI="http://doi.org/10.5281/zenodo.5724214" ID-GBIF-Taxon="190683656" ID-Zenodo-Dep="5724214" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC" httpUri="http://treatment.plazi.org/id/BA5587AF637B4A0391A16E032934F9BC" lastPageId="16" lastPageNumber="899" pageId="15" pageNumber="898">
<subSubSection id="7AE66532637B4A1C91A16E032F2FFD63" pageId="15" pageNumber="898" type="nomenclature">
<paragraph id="324336B9637B4A1C91A16E032E2BFDA8" blockId="15.[942,1327,587,612]" box="[942,1327,587,612]" pageId="15" pageNumber="898">
<heading id="690B81D5637B4A1C91A16E032E2BFDA8" box="[942,1327,587,612]" centered="true" fontSize="9" level="2" pageId="15" pageNumber="898" reason="2">
<taxonomicName id="F5FC4D3A637B4A1C91A16E032E2BFDA8" authority="Kaminski" authorityName="Kamiński &amp; Gearner &amp; Kanda &amp; Swichtenberg &amp; Purchart &amp; Smith" authorityYear="2021" box="[942,1327,587,612]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="901" phylum="Arthropoda" rank="genus" status="sp. nov.">
TOKTOKKUS
<emphasis id="0088EAAB637B4A1C96306E002FB2FDA8" box="[1087,1206,591,611]" italics="true" pageId="15" pageNumber="898">WACLAWAE</emphasis>
KAMIŃSKI
</taxonomicName>
</heading>
</paragraph>
<paragraph id="324336B9637B4A1C91346E3B2F2FFD63" blockId="15.[827,1443,628,1048]" pageId="15" pageNumber="898">
lsid
<uri id="466D3ABB637B4A1C91646E3B2F2FFD63" pageId="15" pageNumber="898">
urn:lsid:zoobank.org:act:
<uuid id="465A0C6C637B4A1C96776E3B2F2FFD63" pageId="15" pageNumber="898">CCA6EFFB-401D-40AF- 9341-AB40107E2698</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="7AE66532637B4A0391346EFD2934F9BC" lastPageId="16" lastPageNumber="899" pageId="15" pageNumber="898" type="materials_examined">
<paragraph id="324336B9637B4A1C91346EFD2FB9FBD3" blockId="15.[827,1443,628,1048]" pageId="15" pageNumber="898">
<emphasis id="0088EAAB637B4A1C91346EFD28D9FD0D" box="[827,989,689,711]" italics="true" pageId="15" pageNumber="898">Type material:</emphasis>
<materialsCitation id="82943CE4637B4A1C91E56EFE2896FCCF" ID-GBIF-Occurrence="3411244306" collectingDate="1942-01-08" collectionCode="MIZ, PAS" collectorName="Dr W. Eichler' &amp; Amyot'" country="Zimbabwe" location="Rhodesia" pageId="15" pageNumber="898" specimenCount="1" typeStatus="holotype">
<typeStatus id="ED47881B637B4A1C91E56EFE2F54FD0C" box="[1002,1104,689,711]" pageId="15" pageNumber="898" type="holotype">Holotype</typeStatus>
(
<collectionCode id="54EDAE7C637B4A1C96526EFE2F8BFD0D" box="[1117,1167,689,710]" pageId="15" pageNumber="898">MIZ</collectionCode>
<collectionCode id="54EDAE7C637B4A1C969B6EFE2FC3FD0C" box="[1172,1223,689,711]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="15" pageNumber="898" type="Herbarium">PAS</collectionCode>
), male: ‘N
<collectingCountry id="4AEB7629637B4A1C97356EFE2EA7FD0C" box="[1338,1443,689,711]" name="Zimbabwe" pageId="15" pageNumber="898">Rhodesia</collectingCountry>
/
<date id="46421079637B4A1C91456E9F28ADFD2E" box="[842,937,720,741]" pageId="15" pageNumber="898" value="1942-01-08">
<collectingDate id="5606E991637B4A1C91456E9F28ADFD2E" box="[842,937,720,741]" pageId="15" pageNumber="898" value="1942-01-08">8 I 1942</collectingDate>
</date>
/
<collectorName id="9F09536F637B4A1C91CF6E9F2F61FD2D" box="[960,1125,720,742]" pageId="15" pageNumber="898">Dr W. Eichler’</collectorName>
, ‘42’, ‘
<taxonomicName id="F5FC4D3A637B4A1C96A76E9F2E37FD2D" box="[1192,1331,720,742]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="901" phylum="Arthropoda" rank="genus">Psammodes</taxonomicName>
/ pierreti
<collectorName id="9F09536F637B4A1C91346EA02896FCCF" box="[827,914,751,772]" pageId="15" pageNumber="898">Amyot’</collectorName>
</materialsCitation>
.
<materialsCitation id="82943CE4637B4A1C91906EA02F2BFCE9" ID-GBIF-Occurrence="3411244308" collectingDate="1942-01-10" collectionCode="MIZ, PAS" country="Zimbabwe" location="Rhodesia" pageId="15" pageNumber="898" specimenCount="1" typeStatus="paratype">
<typeStatus id="ED47881B637B4A1C91906EA02F1CFCCF" box="[927,1048,751,772]" pageId="15" pageNumber="898" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="54EDAE7C637B4A1C96276EA02F5FFCCF" box="[1064,1115,751,772]" pageId="15" pageNumber="898">MIZ</collectionCode>
<collectionCode id="54EDAE7C637B4A1C966A6EA02F9FFCCE" box="[1125,1179,751,773]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="15" pageNumber="898" type="Herbarium">PAS</collectionCode>
): male and female: ‘N
<collectingCountry id="4AEB7629637B4A1C91346F4228A3FCE8" box="[827,935,781,803]" name="Zimbabwe" pageId="15" pageNumber="898">Rhodesia</collectingCountry>
/
<date id="46421079637B4A1C91B06F422F2BFCE9" box="[959,1071,781,803]" pageId="15" pageNumber="898" value="1942-01-10">
<collectingDate id="5606E991637B4A1C91B06F422F2BFCE9" box="[959,1071,781,803]" pageId="15" pageNumber="898" value="1942-01-10">10 I 1942</collectingDate>
</date>
</materialsCitation>
/
<materialsCitation id="82943CE4637B4A1C96476F412FD9FC77" ID-GBIF-Occurrence="3411244307" collectingDate="1995-12-19" collectingDateMax="1995-12-22" collectingDateMin="1995-12-19" collectorName="Dr W. Eichler'" country="Zimbabwe" county="Falls" latitude="-17.933332" location="Victoria Falls" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" pageId="15" pageNumber="898" specimenCount="1" typeStatus="paratype">
<collectorName id="9F09536F637B4A1C96476F412FEBFCE8" box="[1096,1263,781,803]" pageId="15" pageNumber="898">Dr W. Eichler’</collectorName>
; male: ‘KafueR / Rhodesia / UnivFilmEx’, ‘HCRaven / Dec`19 col’; male: ‘
<location id="37236062637B4A1C91896F052F23FCAB" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C91896F052F23FCAB" box="[902,1063,842,864]" country="Zimbabwe" county="Falls" latitude="-17.933332" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" name="Victoria Falls" pageId="15" pageNumber="898">Victoria Falls</location>
/ Zimbabwe /??? 11–89’; female: ‘
<taxonomicName id="F5FC4D3A637B4A1C91306F262F2FFCB4" authority="Pierreti" authorityName="Pierreti" box="[831,1067,873,895]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="genus">Psammodes Pierreti</taxonomicName>
/
<location id="37236062637B4A1C964D6F262F82FCB4" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C964D6F262F82FCB4" box="[1090,1158,873,895]" country="Zimbabwe" county="Falls" latitude="-17.933332" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" name="Solier" pageId="15" pageNumber="898">Solier</location>
/
<location id="37236062637B4A1C96936F262E10FCB4" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C96936F262E10FCB4" box="[1180,1300,873,895]" country="Zimbabwe" county="Falls" latitude="-17.933332" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" name="Zamberer'" pageId="15" pageNumber="898">Zamberer’</location>
, ‘
<location id="37236062637B4A1C972C6F262EA6FCB4" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C972C6F262EA6FCB4" box="[1315,1442,873,895]" country="Zimbabwe" county="Falls" latitude="-17.933332" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" name="Sammlung" pageId="15" pageNumber="898">Sammlung</location>
/
<location id="37236062637B4A1C91436FC728CBFC55" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C91436FC728CBFC55" box="[844,975,904,926]" country="Zimbabwe" county="Falls" latitude="-17.933332" longLatPrecision="1277" longitude="25.833334" municipality="Victoria" name="Schroeder'" pageId="15" pageNumber="898">Schroeder’</location>
; female: ‘
<collectingCountry id="4AEB7629637B4A1C964C6FC72FDFFC56" box="[1091,1243,904,925]" name="Zimbabwe" pageId="15" pageNumber="898">ZIMBABWE</collectingCountry>
<collectingMunicipality id="D227ACC3637B4A1C96EB6FC72E41FC55" box="[1252,1349,904,926]" pageId="15" pageNumber="898">Victoria</collectingMunicipality>
/
<collectingCounty id="DB224E35637B4A1C976E6FC72E99FC55" box="[1377,1437,904,926]" pageId="15" pageNumber="898">Falls</collectingCounty>
,
<geoCoordinate id="57C8507E637B4A1C91346FE92891FC77" box="[827,917,934,956]" degrees="17" direction="south" minutes="56" orientation="latitude" pageId="15" pageNumber="898" precision="925" value="-17.933332">17°56’S</geoCoordinate>
<geoCoordinate id="57C8507E637B4A1C91AF6FE928F9FC77" box="[928,1021,934,956]" degrees="25" direction="east" minutes="50" orientation="longitude" pageId="15" pageNumber="898" precision="925" value="25.833334">25°50’E</geoCoordinate>
/
<date id="46421079637B4A1C96166FE82FD9FC77" box="[1049,1245,934,956]" pageId="15" pageNumber="898" value="1995-12-19" valueMax="1995-12-22" valueMin="1995-12-19">
<collectingDate id="5606E991637B4A1C96166FE82FD9FC77" box="[1049,1245,934,956]" pageId="15" pageNumber="898" value="1995-12-19" valueMax="1995-12-22" valueMin="1995-12-19">19–22 Dec 1995</collectingDate>
</date>
</materialsCitation>
/
<materialsCitation id="82943CE4637B4A1C96F86FE82FBDFBD3" ID-GBIF-Occurrence="3411244304" collectingDate="1993-12-26" collectingDateMax="1993-12-30" collectingDateMin="1993-12-26" collectionCode="AFRICA" collectorName="W. J. Pulawski &amp; J. Moravec'" country="Zambia" location="Victoria falls" pageId="15" pageNumber="898" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectorName id="9F09536F637B4A1C96F86FE82EA6FC77" box="[1271,1442,934,956]" pageId="15" pageNumber="898">W.J. Pulawski</collectorName>
collector’; male and
<specimenCount id="24FAFD30637B4A1C963F6F8A2FC6FC10" box="[1072,1218,965,987]" pageId="15" pageNumber="898" type="female">five females</specimenCount>
(LuboŠ Purchart): ‘
<collectionCode id="54EDAE7C637B4A1C91316FAB28A5FC32" box="[830,929,996,1017]" pageId="15" pageNumber="898">AFRICA</collectionCode>
, S
<collectingCountry id="4AEB7629637B4A1C91CB6FAB2F2EFC32" box="[964,1066,996,1017]" name="Zambia" pageId="15" pageNumber="898">ZAMBIA</collectingCountry>
/
<location id="37236062637B4A1C96316FAB2FD5FC31" LSID="urn:lsid:plazi:treatment:BA5587AF637B4A0391A16E032934F9BC:37236062637B4A1C96316FAB2FD5FC31" box="[1086,1233,996,1018]" country="Zambia" name="Victoria falls" pageId="15" pageNumber="898">Victoria falls</location>
env. (Livingstone) /
<date id="46421079637B4A1C9147684D28F1FBD3" box="[840,1013,1026,1048]" pageId="15" pageNumber="898" value="1993-12-26" valueMax="1993-12-30" valueMin="1993-12-26">
<collectingDate id="5606E991637B4A1C9147684D28F1FBD3" box="[840,1013,1026,1048]" pageId="15" pageNumber="898" value="1993-12-26" valueMax="1993-12-30" valueMin="1993-12-26">26.–30.xii.1993</collectingDate>
</date>
/ leg.
<collectorName id="9F09536F637B4A1C9638684C2FBDFBD3" box="[1079,1209,1027,1048]" pageId="15" pageNumber="898">J. Moravec’</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="324336B9637B4A1C9134680D2E8BFAE6" blockId="15.[827,1442,1089,1326]" pageId="15" pageNumber="898">
<emphasis id="0088EAAB637B4A1C9134680D28B8FB9C" box="[827,956,1090,1111]" italics="true" pageId="15" pageNumber="898">Diagnosis:</emphasis>
Similar to
<taxonomicName id="F5FC4D3A637B4A1C965B680D2FBDFB9D" box="[1108,1209,1089,1111]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="vialis">
<emphasis id="0088EAAB637B4A1C965B680D2FBDFB9D" box="[1108,1209,1089,1111]" italics="true" pageId="15" pageNumber="898">T. vialis</emphasis>
</taxonomicName>
due to the lack of microtubercles between the lateral tubercles, and presence of longitudinal ridges on elytral sides and slope. Both species can be distinguished by different morphology (larger and often confluent in
<taxonomicName id="F5FC4D3A637B4A1C971D68F32E9FFB1A" authorityName="Kaminski" authorityYear="2021" box="[1298,1435,1212,1233]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="waclawae">
<emphasis id="0088EAAB637B4A1C971D68F32E9FFB1A" box="[1298,1435,1212,1233]" italics="true" pageId="15" pageNumber="898">T. waclawae</emphasis>
</taxonomicName>
) and arrangement (rows extend over humeri to scutellum in
<taxonomicName id="F5FC4D3A637B4A1C91C168B52F53FAC5" authorityName="Kaminski" authorityYear="2021" box="[974,1111,1273,1295]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="waclawae">
<emphasis id="0088EAAB637B4A1C91C168B52F53FAC5" box="[974,1111,1273,1295]" italics="true" pageId="15" pageNumber="898">T. waclawae</emphasis>
</taxonomicName>
) of tubercles; and elytral disc (almost flat in lateral view in
<taxonomicName id="F5FC4D3A637B4A1C968169572E1CFAE6" authorityName="Kaminski" authorityYear="2021" box="[1166,1304,1304,1325]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="waclawae">
<emphasis id="0088EAAB637B4A1C968169572E1CFAE6" box="[1166,1304,1304,1325]" italics="true" pageId="15" pageNumber="898">T. waclawae</emphasis>
</taxonomicName>
) (
<figureCitation id="AAC72A3C637B4A1C973F69572E7BFAE6" box="[1328,1407,1304,1326]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński &amp; Gearner sp. nov.; B, holotype of T. congolensis Kamiński &amp; Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński &amp; Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="15" pageNumber="898">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="324336B9637B4A03913469182AE3FA4D" blockId="15.[827,1443,1366,1910]" lastBlockId="16.[145,762,197,1415]" lastPageId="16" lastPageNumber="899" pageId="15" pageNumber="898">
<emphasis id="0088EAAB637B4A1C9134691828C7FAA7" box="[827,963,1367,1388]" italics="true" pageId="15" pageNumber="898">Description:</emphasis>
Length 29.0–30.0 mm, width of pronotum 9.0–10.0 mm and elytra 19.0–20.0 mm.
<emphasis id="0088EAAB637B4A1C9758693A2EA7FA41" box="[1367,1443,1397,1418]" italics="true" pageId="15" pageNumber="898">Head:</emphasis>
Hypognathous. Frons finely punctate (3–4 diameters apart); frontoclypeal suture course, with deep groove in middle; apical clypeal margin broadly shallowly emarginate; clypeus projected toward front of body; apical margin of labrum sharply emarginate medially, densely punctate (although punctures fine) in apical half, apical side of labrum densely covered with yellowish, acuminate setae. Eye comma-shaped, with reduced ventral part, strongly emarginate around epistomal base; with deep groove on temporal side. Mentum trapezoidal, with straight base, not fully filling buccal cavity; anterior margin not emarginate; covered with fine setae. Submentum semicircular, concave basally. Antenna slender, moderately covered in recumbent acuminate goldish setae; antennomere 2 short, equal to 0.2 of antennomere 3 length; antennomere 4 about half of antennomere 3 length; length of antenna equal to 0.85 of pronotal length.
<emphasis id="0088EAAB63644A03929E6D4D2A16FEDC" box="[145,274,258,279]" italics="true" pageId="16" pageNumber="899">Prothorax:</emphasis>
Pronotal lateral margin rounded, well visible. Pronotum widest above middle. Disc dull, impunctate; anterior and basal margins, anterior apices strongly produced. Hypomeron convex, without submarginal groove, impunctate. Prosternal process rounded in lateral view, longitudinally depressed in middle (ventral view). Anterior margin of prosternum labiate, strongly projecting ventrally (lateral view).
<emphasis id="0088EAAB63644A0393396DB72AC6FDC6" box="[310,450,504,525]" italics="true" pageId="16" pageNumber="899">Pterothorax:</emphasis>
Scutellum densely covered with microtubercles. Elytra widest in basal third, rounded laterally; disc impunctate, not covered by tubercles; lateral part (below humerus) covered with tubercles (organized in more or less regular rows) and microtubercles (2–4 diameters apart); remaining lateral part of elytra visible only ventrally, impunctate, without tubercles and microtubercles. Elytral slope steep, densely covered with microtubercles (1–4 diameters apart), with sparsely distributed tubercles, elytral apex flattened. Epipleura, impunctate, not tuberculate, clearly distinguishable from neighbouring portion of elytra, widely enfolding fifth ventrite. Mesoventrite with deep median groove and elevated sides. Metaventrite impunctate, densely setose. Lateral regions of metaventrite (between coxae) extremely short. Metaepisternal suture abbreviated posteriorly.
<emphasis id="0088EAAB63644A03929E684D2BD4FBDC" box="[145,208,1026,1047]" italics="true" pageId="16" pageNumber="899">Legs:</emphasis>
Covered with dense, goldish setae. Procoxa exposed basally. Apex of protibia with prominent denticle on outer margin, lateral carina terminating in basal third; median spur reduced, reaching 0.5 of outer lateral spur length. Spurs on meso- and metatibiae of equal length. Tarsi narrowed laterally.
<emphasis id="0088EAAB63644A03908C68D529FCFB64" box="[643,760,1178,1199]" italics="true" pageId="16" pageNumber="899">Abdomen:</emphasis>
Ventrites 1–4 medially densely covered with goldish setae moderately punctate and weakly rugulose; ventrite 5 densely punctate and setose; ventrite 5 without submarginal sulcus, densely punctured (~0.5 diameters apart), each puncture with single goldish setae.
<emphasis id="0088EAAB63644A039339691D2ABAFAAC" box="[310,446,1362,1383]" italics="true" pageId="16" pageNumber="899">Terminalia:</emphasis>
Aedeagus as in
<figureCitation id="AAC72A3C63644A039071691D29F6FAAC" box="[638,754,1362,1384]" captionStart="Figure 1" captionStartId="2.[145,227,1253,1275]" captionTargetBox="[146,1421,201,1212]" captionTargetId="figure-279@2.[145,1425,195,1213]" captionTargetPageId="2" captionText="Figure 1. Phylogeny of Sepidiini imposed with new generic classification. Presented topology is the majority rule consensus of post-burn-in trees obtained in Bayesian analysis of the concatenated CAD, wg, COI JP, COI BC, COII, and 28S matrix. Posterior probabilities are displayed above branches, while the ultrafast IQTree bootstrap values are shown below.Morphological details shown on the right side of the figure corresponds to clades a–e on the tree. Habitus photographs presented on the left illustrates morphological incoherence of the genus Psammodes." figureDoi="http://doi.org/10.5281/zenodo.5724194" httpUri="https://zenodo.org/record/5724194/files/figure.png" pageId="16" pageNumber="899">Figure 1E</figureCitation>
. Ovipositor as in
<figureCitation id="AAC72A3C63644A039343693E2AE4FA4C" box="[332,480,1393,1415]" captionStart="Figure 6" captionStartId="10.[145,225,1641,1663]" captionTargetBox="[154,1423,208,1588]" captionTargetId="figure-81@10.[145,1425,195,1601]" captionTargetPageId="10" captionText="Figure 6. Female terminalia morphology of selected Sepidiini and Adelostomini: A, B, Toktokkus tuberculipennis; C–H, T. waclawae; I, Psammodes pinguis; J, K, Lepidochora kahani; L, M, Eurychora dilatata. Abbreviations: bc, vacuous of valvifer; bp, vacuous of paraproct;c, coxites; c2–c4, coxite plates 2–4; ip, inner plate of paraproct; op, outer plate of paraproct; pa, paraproct; pr, proctiger; ss, sensory setae; v, valvifer." figureDoi="http://doi.org/10.5281/zenodo.5724206" httpUri="https://zenodo.org/record/5724206/files/figure.png" pageId="16" pageNumber="899">Figure 6C–H</figureCitation>
.
</paragraph>
<paragraph id="324336B963644A03929E69FF29FEF9E9" blockId="16.[145,762,1456,1570]" pageId="16" pageNumber="899">
<emphasis id="0088EAAB63644A03929E69FF2A1FFA0E" box="[145,283,1456,1477]" italics="true" pageId="16" pageNumber="899">Etymology:</emphasis>
This new species is dedicated to the memory of the first author’s grandmother, Wacława Kamińska (born on
<date id="4642107963644A03938969A2295AF9C9" box="[390,606,1517,1539]" pageId="16" pageNumber="899" value="1927-11-05">
5 November
<quantity id="F5049B5C63644A03902B69A2295AF9C9" box="[548,606,1517,1538]" metricMagnitude="1" metricUnit="m" metricValue="4.8945799999999995" pageId="16" pageNumber="899" unit="in" value="1927.0">1927</quantity>
</date>
in Bartniki,
<collectingCountry id="4AEB762963644A03929E6A432BE5F9E9" box="[145,225,1548,1570]" name="Poland" pageId="16" pageNumber="899">Poland</collectingCountry>
, died on
<date id="4642107963644A0393306A432913F9EA" box="[319,535,1548,1570]" pageId="16" pageNumber="899" value="2010-09-29">29 September 2010</date>
, in Warsaw,
<collectingCountry id="4AEB762963644A0390936A4329EBF9E9" box="[668,751,1548,1570]" name="Poland" pageId="16" pageNumber="899">Poland</collectingCountry>
).
</paragraph>
<paragraph id="324336B963644A0393556A112934F9BC" blockId="16.[346,560,1630,1655]" box="[346,560,1630,1655]" pageId="16" pageNumber="899">
<collectionCode id="54EDAE7C63644A0393556A11292DF9BD" box="[346,553,1630,1655]" pageId="16" pageNumber="899">SYNONYMY NOTES</collectionCode>
:
</paragraph>
</subSubSection>
</treatment>
</document>
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