248 lines
30 KiB
XML
248 lines
30 KiB
XML
<document id="7AF58441EBA81D6B8A19FBBA588997E2" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="10114201" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630351968237" checkinUser="diego" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="BA2A87DCFF9AFFFBF128FA2BFC4EFA70" docLanguage="en" docName="0010-065X-72.mo4.1.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:868A6B4378E6626262F3806A93742F8C.2:TheColeopBull.2017-.monograph" docStyleId="868A6B4378E6626262F3806A93742F8C" docStyleName="TheColeopBull.2017-.monograph" docStyleVersion="2" docTitle="Phlaeopterus frosti Hatch 1957" docType="treatment" docVersion="4" masterDocId="4613FFA4FF85FFDBF221FFDFFFB0FFA2" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1700036274245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="01115ED2EEF4F3BC35C760763795895F">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="92E204D6E00FE70E034920A678D426D2">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="03813B727C6F0DEA8556EA6C139AF960">Campbell, J. M.</mods:namePart>
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<mods:roleTerm id="5B4F361286699406A8E5E4024B93467F">Author</mods:roleTerm>
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<mods:namePart id="A3BDE9202E247565C2F9F575B40BA5DF">Sikes, Derek S.</mods:namePart>
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<mods:typeOfResource id="9072A2E6AEA150350D67CD750292713E">text</mods:typeOfResource>
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<mods:title id="2B3D71E9777EB16DB5A4C9BD14057F62">The Coleopterists Bulletin</mods:title>
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<mods:part id="B6299C9CC6020A97CF100314827B12E5">
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<mods:date id="914E180454BAF3170A0ECB7D3DC5A1D8">2018</mods:date>
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<mods:number id="8BFF91BCC75EEFB392D8258E82979EC8">2018-12-28</mods:number>
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<mods:number id="BB3AE9EFD97D553B23A35CE4FAEE4556">72</mods:number>
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<mods:start id="AEDF6BCCDE8AC7978EC1687A0C6D8746">1</mods:start>
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<mods:url id="76DF024E48459B10A79368F711027C26">http://dx.doi.org/10.1649/0010-065x-72.mo4.1</mods:url>
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<mods:classification id="99154EE645FE471097DD0352C548D7F0">journal article</mods:classification>
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<mods:identifier id="96E5583A38C582E39996449DAA1B624E" type="DOI">10.1649/0010-065X-72.mo4.1</mods:identifier>
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<mods:identifier id="360DED9AE46480EF68BFEBAF3806ED19" type="ISSN">1938-4394</mods:identifier>
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<treatment id="BA2A87DCFF9AFFFBF128FA2BFC4EFA70" LSID="urn:lsid:plazi:treatment:BA2A87DCFF9AFFFBF128FA2BFC4EFA70" httpUri="http://treatment.plazi.org/id/BA2A87DCFF9AFFFBF128FA2BFC4EFA70" lastPageId="32" pageId="31">
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<subSubSection id="7A996541FF9AFFC4F128FA2BFBD3F9AB" box="[777,1123,1524,1545]" pageId="31" type="nomenclature">
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<paragraph id="323C36CAFF9AFFC4F128FA2BFBD3F9AB" blockId="31.[743,1156,1524,1572]" box="[777,1123,1524,1545]" pageId="31">
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<heading id="697481A6FF9AFFC4F128FA2BFBD3F9AB" box="[777,1123,1524,1545]" centered="true" fontSize="10" level="2" pageId="31" reason="2">
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<emphasis id="00F7EAD8FF9AFFC4F128FA2BFBD3F9AB" bold="true" box="[777,1123,1524,1545]" pageId="31">
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7.
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<taxonomicName id="F5834D49FF9AFFC4F103FA2BFBD3F9AB" ID-CoL="76ZZX" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[802,1123,1524,1545]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="31" phylum="Arthropoda" rank="species" species="frosti">
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<emphasis id="00F7EAD8FF9AFFC4F103FA2BFC50F9AB" bold="true" box="[802,992,1524,1545]" italics="true" pageId="31">Phlaeopterus frosti</emphasis>
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<bibRefCitation id="56124B3BFF9AFFC4F1C9FA2BFBD3F9AB" author="Hatch" box="[1000,1123,1524,1545]" pageId="31" refId="ref29117" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch, 1957</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7A996541FF9AFFC4F0C6F9D0FB34F986" box="[743,1156,1551,1572]" pageId="31" type="description">
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<paragraph id="323C36CAFF9AFFC4F0C6F9D0FB34F986" blockId="31.[743,1156,1524,1572]" box="[743,1156,1551,1572]" pageId="31">
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(
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||
<figureCitation id="AAB82A4FFF9AFFC4F0CFF9D0FCF3F986" box="[750,835,1551,1572]" captionStart="Fig" captionStartId="12.[175,206,1726,1745]" captionTargetBox="[156,1167,196,1708]" captionTargetId="figure-1@12.[154,1169,194,1710]" captionTargetPageId="12" captionText="Fig. 2. Phlaeopterus species, habitus. A) P. elongatus, male (UAMObs:Ento:232543), B) P. filicornis, female" figureDoi="http://doi.org/10.5281/zenodo.10130804" httpUri="https://zenodo.org/record/10130804/files/figure.png" pageId="31">Figs. 2C</figureCitation>
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,
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||
<figureCitation id="AAB82A4FFF9AFFC4F16EF9D0FCDEF986" box="[847,878,1551,1572]" captionStart="Fig" captionStartId="18.[175,206,1027,1046]" captionText="Fig. 9. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. frosti, B) P. fusconiger. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="31">9A</figureCitation>
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,
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<figureCitation id="AAB82A4FFF9AFFC4F15BF9D0FC14F986" box="[890,932,1551,1572]" captionStart="Fig" captionStartId="24.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@24.[164,1159,194,1532]" captionTargetPageId="24" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130828" httpUri="https://zenodo.org/record/10130828/files/figure.png" pageId="31">17B</figureCitation>
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,
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<figureCitation id="AAB82A4FFF9AFFC4F18EF9D0FC6BF986" box="[943,987,1551,1572]" captionStart="Fig" captionStartId="25.[197,228,482,501]" captionTargetBox="[177,1188,196,465]" captionTargetId="figure-576@25.[175,1190,194,467]" captionTargetPageId="25" captionText="Fig. 18. Mesotibiae, dorsal view, of A) Phlaeopterus frosti (UAMObs:Ento:235497), and B) Phlaeopterus fusconiger (UAMObs:Ento:235751). Red highlights are added to emphasize the difference in the length of the subglabrous region (lacking dense pubescence but with longer spine-like setae) at the apex of the tibia." figureDoi="http://doi.org/10.5281/zenodo.10130830" httpUri="https://zenodo.org/record/10130830/files/figure.png" pageId="31">18A</figureCitation>
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,
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<figureCitation id="AAB82A4FFF9AFFC4F1C7F9D0FBA1F986" box="[998,1041,1551,1572]" captionStart="Fig" captionStartId="26.[709,740,1185,1204]" captionTargetBox="[689,1169,195,1168]" captionTargetId="figure-245@26.[688,1170,194,1169]" captionTargetPageId="26" captionText="Fig. 20. Phlaeopterus species, mesosterna, ventral oblique view. A) P. longipennis, B) P. obsoletus, C) P. olympicus, D) P. frosti, E) P. occidentalis, F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130832" httpUri="https://zenodo.org/record/10130832/files/figure.png" pageId="31">20D</figureCitation>
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,
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<figureCitation id="AAB82A4FFF9AFFC4F63CF9D0FBF8F986" box="[1053,1096,1551,1572]" captionStart="Fig" captionStartId="31.[197,228,1342,1361]" captionTargetBox="[177,1188,196,1324]" captionTargetId="figure-145@31.[175,1190,194,1326]" captionTargetPageId="31" captionText="Fig. 25. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) Phlaeopterus frosti, B) P. hatchi, C) P. filicornis, D) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130844" httpUri="https://zenodo.org/record/10130844/files/figure.png" pageId="31">25A</figureCitation>
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,
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<figureCitation id="AAB82A4FFF9AFFC4F672F9D0FBCEF986" box="[1107,1150,1551,1572]" captionStart="Fig" captionStartId="43.[731,762,1272,1291]" captionTargetBox="[711,1189,196,1255]" captionTargetId="figure-251@43.[709,1191,194,1257]" captionTargetPageId="43" captionText="Fig. 36. Phlaeopterus species, eyes, dorsal view. A) P. lagrandeuri, B) P. houkae, C) P. longipennis, D) P. frosti, E) P. bakerensis, F) P. olympicus." figureDoi="http://doi.org/10.5281/zenodo.10130869" httpUri="https://zenodo.org/record/10130869/files/figure.png" pageId="31">36D</figureCitation>
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)
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||
</paragraph>
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</subSubSection>
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<subSubSection id="7A996541FF9AFFC4F0E2F99CFC64F92F" pageId="31" type="reference_group">
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<paragraph id="323C36CAFF9AFFC4F0E2F99CFC64F92F" blockId="31.[707,1193,1603,1677]" pageId="31">
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<treatmentCitationGroup id="129311E4FF9AFFC4F0E2F99CFC64F92F" pageId="31">
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<treatmentCitation id="B32210DBFF9AFFC4F0E2F99CFB93F9FA" author="Hatch" box="[707,1059,1603,1624]" page="59" pageId="31" year="1957">
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<taxonomicName id="F5834D49FF9AFFC4F0E2F99CFB93F9FA" ID-CoL="76ZZX" authority="Hatch 1957: 59" authorityName="Hatch" authorityPageNumber="59" authorityYear="1957" box="[707,1059,1603,1624]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="31" phylum="Arthropoda" rank="species" species="frosti">
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<emphasis id="00F7EAD8FF9AFFC4F0E2F99CFCCDF9FA" box="[707,893,1603,1624]" italics="true" pageId="31">Phlaeopterus frosti</emphasis>
|
||
<bibRefCitation id="56124B3BFF9AFFC4F1A6F99CFB93F9FA" author="Hatch" box="[903,1059,1603,1624]" pageId="31" refId="ref29117" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 59</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
[original description]. Campbell and Davies 1991: 5;
|
||
<treatmentCitation id="B32210DBFF9AFFC4F0FCF9A7FC60F92F" author="Bousquet" box="[733,976,1656,1677]" page="89" pageId="31" year="2013">
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||
<bibRefCitation id="56124B3BFF9AFFC4F0FCF9A7FC60F92F" author="Bousquet" box="[733,976,1656,1677]" pageId="31" refId="ref28595" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition. Pensoft, Sofia, Bulgaria." type="book" year="2013">
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||
Bousquet
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<emphasis id="00F7EAD8FF9AFFC4F11EF9A7FCC1F92F" box="[831,881,1656,1677]" italics="true" pageId="31">et al.</emphasis>
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2013: 89
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7A996541FF9AFFC4F0FAF972FB14F960" box="[731,1188,1709,1730]" pageId="31" type="materials_examined">
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<paragraph id="323C36CAFF9AFFC4F0FAF972FB14F960" blockId="31.[731,1188,1709,1730]" box="[731,1188,1709,1730]" pageId="31">
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<materialsCitation id="82EB3C97FF9AFFC4F0FAF972FB11F960" box="[731,1185,1709,1730]" country="United States of America" location="Mt. Rainier" pageId="31" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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||
<emphasis id="00F7EAD8FF9AFFC4F0FAF972FCDCF960" bold="true" box="[731,876,1709,1730]" pageId="31">
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||
<typeStatus id="ED388868FF9AFFC4F0FAF972FCBDF960" box="[731,781,1709,1730]" pageId="31">Type</typeStatus>
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Locality.
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||
</emphasis>
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||
<location id="375C6011FF9AFFC4F152F972FC54F960" LSID="urn:lsid:plazi:treatment:BA2A87DCFF9AFFFBF128FA2BFC4EFA70:375C6011FF9AFFC4F152F972FC54F960" box="[883,996,1709,1730]" country="United States of America" name="Mt. Rainier" pageId="31" stateProvince="Washington">Mt. Rainier</location>
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||
,
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<collectingRegion id="F047F828FF9AFFC4F1CEF972FBD2F960" box="[1007,1122,1709,1730]" country="United States of America" name="Washington" pageId="31">Washington</collectingRegion>
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||
,
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<collectingCountry id="4A94765AFF9AFFC4F64FF972FB11F960" box="[1134,1185,1709,1730]" name="United States of America" pageId="31">USA</collectingCountry>
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="66FC6642FFA5FFFBF28EFD5AFDB5FD6A" ID-DOI="http://doi.org/10.5281/zenodo.10130846" ID-Zenodo-Dep="10130846" httpUri="https://zenodo.org/record/10130846/files/figure.png" pageId="32" startId="32.[175,206,645,664]" targetBox="[155,633,196,621]" targetPageId="32">
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<paragraph id="323C36CAFFA5FFFBF28EFD5AFDB5FD6A" blockId="32.[151,639,645,712]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF28EFD5AFEB3FD3A" bold="true" box="[175,259,645,664]" pageId="32">Fig. 26.</emphasis>
|
||
<taxonomicName id="F5834D49FFA5FFFBF339FD5AFE52FD3A" baseAuthorityName="Shavrin" baseAuthorityYear="2001" box="[280,482,645,664]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="czerskyi">
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||
<emphasis id="00F7EAD8FFA5FFFBF339FD5AFE52FD3A" box="[280,482,645,664]" italics="true" pageId="32">Phlaeopterus czerskyi</emphasis>
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||
</taxonomicName>
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, aedeagus. A) Ventral view, B) Lateral view. Scale bar = 0.1 mm. Adapted from Shavrin and Mullen (2015).
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||
</paragraph>
|
||
</caption>
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||
<subSubSection id="7A996541FFA5FFFBF28EFCD4FD6AFE42" pageId="32" type="description">
|
||
<paragraph id="323C36CAFFA5FFFBF28EFCD4FD6AFE42" blockId="32.[151,639,779,1730]" lastBlockId="32.[685,1173,194,1490]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF28EFCD4FE15FC82" bold="true" box="[175,421,779,800]" pageId="32">Redescription. Habitus:</emphasis>
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||
Length
|
||
<quantity id="F57B9B2FFFA5FFFBF3DEFCD4FDCBFC82" box="[511,635,779,801]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.2" metricValueMin="5.8" pageId="32" unit="mm" value="6.5" valueMax="7.2" valueMin="5.8">5.8–7.2 mm</quantity>
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||
. Reddish brown to black; legs, elytral epipleura, tarsi, and apices of tibiae lighter reddish brown to yellowish brown (
|
||
<figureCitation id="AAB82A4FFFA5FFFBF365FC84FE20FCD2" box="[324,400,859,880]" captionStart="Fig" captionStartId="12.[175,206,1726,1745]" captionTargetBox="[156,1167,196,1708]" captionTargetId="figure-1@12.[154,1169,194,1710]" captionTargetPageId="12" captionText="Fig. 2. Phlaeopterus species, habitus. A) P. elongatus, male (UAMObs:Ento:232543), B) P. filicornis, female" figureDoi="http://doi.org/10.5281/zenodo.10130804" httpUri="https://zenodo.org/record/10130804/files/figure.png" pageId="32">Fig. 2C</figureCitation>
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||
).
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<emphasis id="00F7EAD8FFA5FFFBF381FC84FE6EFCD2" bold="true" box="[416,478,859,880]" pageId="32">Head:</emphasis>
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||
Slightly broader than long, ratio of width across eyes to head length = 4:3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered hairs near ventral margin (
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<figureCitation id="AAB82A4FFFA5FFFBF032FC3FFDC4FC57" box="[531,628,992,1013]" captionStart="Fig" captionStartId="43.[731,762,1272,1291]" captionTargetBox="[711,1189,196,1255]" captionTargetId="figure-251@43.[709,1191,194,1257]" captionTargetPageId="43" captionText="Fig. 36. Phlaeopterus species, eyes, dorsal view. A) P. lagrandeuri, B) P. houkae, C) P. longipennis, D) P. frosti, E) P. bakerensis, F) P. olympicus." figureDoi="http://doi.org/10.5281/zenodo.10130869" httpUri="https://zenodo.org/record/10130869/files/figure.png" pageId="32">Fig. 36D</figureCitation>
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). Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
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<superScript id="C5F69B82FFA5FFFBF04EFBA5FDCEFB2A" attach="none" box="[623,638,1146,1160]" fontSize="6" pageId="32">rd</superScript>
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palpomere 1.6–1.8 times longer than palpomere 2.
|
||
<emphasis id="00F7EAD8FFA5FFFBF2B9FB6BFF5AFB6B" bold="true" box="[152,234,1204,1225]" pageId="32">Thorax:</emphasis>
|
||
Pronotum moderately wide (
|
||
<figureCitation id="AAB82A4FFFA5FFFBF037FB6BFDC2FB6B" box="[534,626,1204,1225]" captionStart="Fig" captionStartId="24.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@24.[164,1159,194,1532]" captionTargetPageId="24" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130828" httpUri="https://zenodo.org/record/10130828/files/figure.png" pageId="32">Fig. 17B</figureCitation>
|
||
), length to width ratio = 0.68–0.72; ratio of pronotal width to head width = 1.54–1.67; maximum width subequal to elytral width at humeral angles; punctures separated by distance equal to slightly greater than diameter of a puncture; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1–2.2 times longer than pronotum; apical margins convex or subtruncate. Wings fully developed in most indivuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum distinct on anterior half, distinct or reduced on posterior half (
|
||
<figureCitation id="AAB82A4FFFA5FFFBF03CF9F7FDC3F99F" box="[541,627,1576,1597]" captionStart="Fig" captionStartId="26.[709,740,1185,1204]" captionTargetBox="[689,1169,195,1168]" captionTargetId="figure-245@26.[688,1170,194,1169]" captionTargetPageId="26" captionText="Fig. 20. Phlaeopterus species, mesosterna, ventral oblique view. A) P. longipennis, B) P. obsoletus, C) P. olympicus, D) P. frosti, E) P. occidentalis, F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130832" httpUri="https://zenodo.org/record/10130832/files/figure.png" pageId="32">Fig. 20D</figureCitation>
|
||
).
|
||
<emphasis id="00F7EAD8FFA5FFFBF2B6F99CFF7EF9FA" bold="true" box="[151,206,1603,1624]" pageId="32">Legs:</emphasis>
|
||
Apices of all tibiae glabrous, subglabrous apex of mesotibia subequal to length of basal 4 mesotarsomeres (
|
||
<figureCitation id="AAB82A4FFFA5FFFBF31CF9A7FE27F92F" box="[317,407,1656,1677]" captionStart="Fig" captionStartId="25.[197,228,482,501]" captionTargetBox="[177,1188,196,465]" captionTargetId="figure-576@25.[175,1190,194,467]" captionTargetPageId="25" captionText="Fig. 18. Mesotibiae, dorsal view, of A) Phlaeopterus frosti (UAMObs:Ento:235497), and B) Phlaeopterus fusconiger (UAMObs:Ento:235751). Red highlights are added to emphasize the difference in the length of the subglabrous region (lacking dense pubescence but with longer spine-like setae) at the apex of the tibia." figureDoi="http://doi.org/10.5281/zenodo.10130830" httpUri="https://zenodo.org/record/10130830/files/figure.png" pageId="32">Fig. 18A</figureCitation>
|
||
), length of subglabrous apical region of metatibia as ratio to metatibial length in males = 4.9–8.4, in females = 4.2–5.5. Metatrochanter without tooth on apical margin.
|
||
<emphasis id="00F7EAD8FFA5FFFBF08CFF03FCA6FF53" bold="true" box="[685,790,220,241]" pageId="32">Abdomen:</emphasis>
|
||
Wing-folding spicules on tergites IV and V; shape of wing-folding patches on tergite V broadly oval and narrowly separate.
|
||
<emphasis id="00F7EAD8FFA5FFFBF1FAFECEFBF2FE84" bold="true" box="[987,1090,273,294]" pageId="32">Aedeagus:</emphasis>
|
||
Length
|
||
<quantity id="F57B9B2FFFA5FFFBF08CFEF3FC8EFEE3" box="[685,830,299,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.4" metricValueMin="1.23" pageId="32" unit="mm" value="1.315" valueMax="1.4" valueMin="1.23">1.23–1.40 mm</quantity>
|
||
. Median lobe somewhat triangular with sides narrowing from base to apex; apex not carinate (
|
||
<figureCitation id="AAB82A4FFFA5FFFBF128FEBEFCD6FED4" box="[777,870,353,374]" captionStart="Fig" captionStartId="31.[197,228,1342,1361]" captionTargetBox="[177,1188,196,1324]" captionTargetId="figure-145@31.[175,1190,194,1326]" captionTargetPageId="31" captionText="Fig. 25. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) Phlaeopterus frosti, B) P. hatchi, C) P. filicornis, D) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.10130844" httpUri="https://zenodo.org/record/10130844/files/figure.png" pageId="32">Fig. 25A</figureCitation>
|
||
). Parameres extending slightly beyond apex of median lobe. Internal sac rectangular and elongate; heavily sclerotized, uniformly covered with microspinules; lacking subapical transverse fold.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A996541FFA5FFFBF0E4FE39FD49FD38" pageId="32" type="materials_examined">
|
||
<paragraph id="323C36CAFFA5FFFBF0E4FE39FD49FD38" blockId="32.[685,1173,194,1490]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF0E4FE39FCC6FE59" bold="true" box="[709,886,486,507]" pageId="32">Type Specimens.</emphasis>
|
||
<materialsCitation id="82EB3C97FFA5FFFBF1A5FE39FD49FD38" collectingDate="1930-08-23" collectorName="M. H. Hatch & M. H. Hatch. Both" location="Mt. Rainier" municipality="Sluskin Falls" pageId="32" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<typeStatus id="ED388868FFA5FFFBF1A5FE39FC6CFE59" box="[900,988,486,507]" pageId="32" type="holotype">Holotype</typeStatus>
|
||
male (UAMObs: Ento:235778) and
|
||
<typeStatus id="ED388868FFA5FFFBF17CFDDFFC19FDB7" box="[861,937,512,533]" pageId="32" type="allotype">allotype</typeStatus>
|
||
female (UAMObs:Ento: 235779) labeled as follows:
|
||
<location id="375C6011FFA5FFFBF1E7FDC4FB88FD92" LSID="urn:lsid:plazi:treatment:BA2A87DCFF9AFFFBF128FA2BFC4EFA70:375C6011FFA5FFFBF1E7FDC4FB88FD92" box="[966,1080,539,560]" municipality="Sluskin Falls" name="Mt. Rainier" pageId="32">Mt. Rainier</location>
|
||
, WASH., below
|
||
<collectingMunicipality id="D258ACB0FFA5FFFBF0D3FDEAFCC4FDE8" box="[754,884,565,586]" pageId="32">Sluskin Falls</collectingMunicipality>
|
||
,
|
||
<date id="463D100AFFA5FFFBF1A3FDEAFBA4FDE8" box="[898,1044,565,586]" pageId="32" value="1930-08-23">
|
||
<collectingDate id="5679E9E2FFA5FFFBF1A3FDEAFBA4FDE8" box="[898,1044,565,586]" pageId="32" value="1930-08-23">Aug. 23, 1930</collectingDate>
|
||
</date>
|
||
<collectorName id="9F76531CFFA5FFFBF63FFDEAFB23FDE8" box="[1054,1171,565,586]" pageId="32">M.H. Hatch</collectorName>
|
||
TYPE ³ (or
|
||
<typeStatus id="ED388868FFA5FFFBF111FD8FFC1AFDC7" box="[816,938,592,613]" pageId="32" type="allotype">ALLOTYPE</typeStatus>
|
||
<specimenCount id="2485FD43FFA5FFFBF195FD8FFC71FDC4" box="[948,961,592,614]" pageId="32" type="female">♀</specimenCount>
|
||
),
|
||
<taxonomicName id="F5834D49FFA5FFFBF1F9FD8FFD6DFD22" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="frosti">Phlaeopterus frosti, 1951</taxonomicName>
|
||
–
|
||
<collectorName id="9F76531CFFA5FFFBF0C8FDB4FC17FD22" box="[745,935,619,640]" pageId="32">M.H. Hatch. Both</collectorName>
|
||
specimens are in the USNM.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A996541FFA5FFFBF0E4FD7FFBE1FB0F" pageId="32" type="distribution">
|
||
<paragraph id="323C36CAFFA5FFFBF0E4FD7FFC5CFCF6" blockId="32.[685,1173,194,1490]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF0E4FD7FFCF6FD17" bold="true" box="[709,838,672,693]" pageId="32">Distribution.</emphasis>
|
||
<taxonomicName id="F5834D49FFA5FFFBF16CFD7FFBB3FD17" authorityName="Hatch" authorityYear="1957" box="[845,1027,672,693]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="frosti">
|
||
<emphasis id="00F7EAD8FFA5FFFBF16CFD7FFBB3FD17" box="[845,1027,672,693]" italics="true" pageId="32">Phlaeopterus frosti</emphasis>
|
||
</taxonomicName>
|
||
is found in the Cascade Range from
|
||
<collectingRegion id="F047F828FFA5FFFBF1ADFD65FC63FD6D" box="[908,979,698,719]" country="United States of America" name="Oregon" pageId="32">Oregon</collectingRegion>
|
||
north to Manning Provincial Park,
|
||
<collectingRegion id="F047F828FFA5FFFBF178FD0AFBBAFD48" box="[857,1034,725,746]" country="Canada" name="British Columbia" pageId="32">British Columbia</collectingRegion>
|
||
, the Olympic Mountains of
|
||
<collectingRegion id="F047F828FFA5FFFBF16DFD2FFC74FCA7" box="[844,964,752,773]" country="United States of America" name="Washington" pageId="32">Washington</collectingRegion>
|
||
, the Pacific Coast Range, Selkirk Mountains, and southern Rocky Mountains of
|
||
<collectingRegion id="F047F828FFA5FFFBF160FCFAFC43FC98" box="[833,1011,805,826]" country="Canada" name="British Columbia" pageId="32">British Columbia</collectingRegion>
|
||
, and the Kenai Mountains of
|
||
<collectingRegion id="F047F828FFA5FFFBF11AFCE0FCCFFCF6" box="[827,895,831,852]" country="United States of America" name="Alaska" pageId="32">Alaska</collectingRegion>
|
||
(
|
||
<figureCitation id="AAB82A4FFFA5FFFBF1AEFCE0FC51FCF6" box="[911,993,831,852]" captionStart="Fig" captionStartId="18.[175,206,1027,1046]" captionText="Fig. 9. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. frosti, B) P. fusconiger. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="32">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="323C36CAFFA5FFFBF0E4FC85FBE1FB0F" blockId="32.[685,1173,194,1490]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF0E4FC85FC88FCCD" bold="true" box="[709,824,858,879]" pageId="32">Bionomics.</emphasis>
|
||
Adults have been collected at
|
||
<quantity id="F57B9B2FFFA5FFFBF08CFCABFC95FC2B" box="[685,805,884,906]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="2.05" metricValueMin="0.05" pageId="32" unit="m" value="1050.0" valueMax="2050.0" valueMin="50.0">50–2,050 m</quantity>
|
||
elevation during June–August. This species has been collected on the surface of permanent or long-lasting snowfields at night in
|
||
<collectingRegion id="F047F828FFA5FFFBF621FC76FBDDFC1C" box="[1024,1133,937,958]" country="United States of America" name="Washington" pageId="32">Washington</collectingRegion>
|
||
and
|
||
<collectingRegion id="F047F828FFA5FFFBF08CFC1BFD44FC7B" box="[685,756,964,985]" country="United States of America" name="Oregon" pageId="32">Oregon</collectingRegion>
|
||
, where they forage immediately after dusk, and during the day in
|
||
<collectingRegion id="F047F828FFA5FFFBF1A2FC00FB96FC56" box="[899,1062,991,1012]" country="Canada" name="British Columbia" pageId="32">British Columbia</collectingRegion>
|
||
. They have also been collected under rocks at the edges of snowfields and cold, fast-flowing streams, in moss growing in the splash-zones of waterfalls, and occasionally in the gravelly, marshy edges of high elevation lakes. On the surface of snowfields, they have been observed feeding on small insects (mostly Diptera) stranded on the surface of the snow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A996541FFA5FFFBF0E4FB6CFC4EFA70" pageId="32" type="discussion">
|
||
<paragraph id="323C36CAFFA5FFFBF0E4FB6CFC4EFA70" blockId="32.[685,1173,194,1490]" pageId="32">
|
||
<emphasis id="00F7EAD8FFA5FFFBF0E4FB6CFC97FB6A" bold="true" box="[709,807,1203,1224]" pageId="32">Remarks.</emphasis>
|
||
<taxonomicName id="F5834D49FFA5FFFBF117FB6CFC44FB6A" authorityName="Hatch" authorityYear="1957" box="[822,1012,1203,1224]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="frosti">
|
||
<emphasis id="00F7EAD8FFA5FFFBF117FB6CFC44FB6A" box="[822,1012,1203,1224]" italics="true" pageId="32">Phlaeopterus frosti</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="F5834D49FFA5FFFBF1A9FB11FBE1FB41" box="[904,1105,1230,1251]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="00F7EAD8FFA5FFFBF1A9FB11FBB6FB41" box="[904,1030,1230,1251]" italics="true" pageId="32">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the shape of the pronotum (length:width ratio, lateral margins, and lateral foveae), the glabrous region at the apex of the tibiae, and the form of the aedeagus. This species is very similar to
|
||
<taxonomicName id="F5834D49FFA5FFFBF1ECFAE7FBF8FAEF" box="[973,1096,1336,1357]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="32" phylum="Arthropoda" rank="species" species="fusconiger">
|
||
<emphasis id="00F7EAD8FFA5FFFBF1ECFAE7FBF8FAEF" box="[973,1096,1336,1357]" italics="true" pageId="32">P. fusconiger</emphasis>
|
||
</taxonomicName>
|
||
but can be distinguished by the characters discussed in the Remarks section of the latter species. These two species are sympatric in the Pacific Coast Range from the Olympic Mountains of
|
||
<collectingRegion id="F047F828FFA5FFFBF1C8FA7DFBEAFA15" box="[1001,1114,1442,1463]" country="United States of America" name="Washington" pageId="32">Washington</collectingRegion>
|
||
north to the Kenai Mountains of
|
||
<collectingRegion id="F047F828FFA5FFFBF194FA62FC4AFA70" box="[949,1018,1469,1490]" country="United States of America" name="Alaska" pageId="32">Alaska</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |