242 lines
24 KiB
XML
242 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.232.110110" ID-Pensoft-Pub="1314-2003-232-145" ID-Pensoft-UUID="B24E305A09B9582DBFDBAD8120894678" ModsDocID="1314-2003-232-145" checkinTime="1695263963467" checkinUser="pensoft" docAuthor="Goyder, David J., Davies, Nina, Finckh, Manfred, Gomes, Amandio, Goncalves, Francisco Maiato P., Meller, Paulina & Paton, Alan J." docDate="2023" docId="B3859CC85EBA53908D5AB828A10A4B47" docLanguage="en" docName="PhytoKeys 232: 145-166" docOrigin="PhytoKeys 232" docPubDate="2023-09-20" docSource="http://dx.doi.org/10.3897/phytokeys.232.110110" docTitle="Endostemon palustris A. J. Paton & Goyder 2023, sp. nov." docType="treatment" docVersion="1" id="B24E305A09B9582DBFDBAD8120894678" lastPageNumber="145" masterDocId="B24E305A09B9582DBFDBAD8120894678" masterDocTitle="New species of Asclepias (Apocynaceae), Baphia (Leguminosae), Cochlospermum (Bixaceae) and Endostemon (Lamiaceae) from the Kalahari sands of Angola and NW Zambia, with one new combination in Vangueria (Rubiaceae)" masterLastPageNumber="166" masterPageNumber="145" pageNumber="145" updateTime="1695263963467" updateUser="pensoft">
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<mods:titleInfo id="46137701561D51AB11C8D9B8072A86E7">
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<mods:title id="D9ABB1457FA437ABF995B1AA5AFD85E8">New species of Asclepias (Apocynaceae), Baphia (Leguminosae), Cochlospermum (Bixaceae) and Endostemon (Lamiaceae) from the Kalahari sands of Angola and NW Zambia, with one new combination in Vangueria (Rubiaceae)</mods:title>
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<mods:namePart id="733F0DAB16EF0A885BC75B0B25987C70">Goyder, David J.</mods:namePart>
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<mods:nameIdentifier id="88543E8A7DC9A567A1374D5ABC0F94A6" type="ORCID">https://orcid.org/0000-0002-3449-7313</mods:nameIdentifier>
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<mods:affiliation id="F9CA7C86FA934732D18CBA9238FE45E5">Royal Botanic Gardens, Kew, TW 9 3 AE, London, UK & National Geographic Okavango Wilderness Project, Wild Bird Trust, Hogsback, South Africa</mods:affiliation>
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<mods:nameIdentifier id="1DD2BA2B875D02A2497FBB2FC5B68ED0" type="email">d.goyder@kew.org</mods:nameIdentifier>
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<mods:roleTerm id="A3EBB53A45F4E839A3113BF0EC37425E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A85FF1A98DDED6FA0347AEA882343BE0">Davies, Nina</mods:namePart>
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<mods:affiliation id="768B5BE1236637E8E15F0713C9849A35">Royal Botanic Gardens, Kew, TW 9 3 AE, London, UK</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="3ADEE6CCC7838C377DC8E11D71463DED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="126A91775B773F924DD1A91EA0168844">Finckh, Manfred</mods:namePart>
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<mods:nameIdentifier id="51E1E105BE157AA95F7544AC3E5D2517" type="ORCID">https://orcid.org/0000-0003-2186-0854</mods:nameIdentifier>
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<mods:affiliation id="2F7FD5927CCEED19082FBAB8FC36732F">Institute for Plant Science and Microbiology, University of Hamburg, Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="136B13FCFCE4E37095D3550FEE37D280" type="personal">
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<mods:roleTerm id="2369A05E328FDE8194A4E4E5FF4AED61">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8F3EBD8955974C03E508E546F89D9A65">Gomes, Amandio</mods:namePart>
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<mods:nameIdentifier id="6F252F2332DD7094DB6073DCE85B6D4F" type="ORCID">https://orcid.org/0000-0002-7812-5490</mods:nameIdentifier>
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<mods:affiliation id="9EF01262451BFC3BED9AFEC54B516826">National Geographic Okavango Wilderness Project, Wild Bird Trust, Hogsback, South Africa & Natural Science Faculty of Agostinho Neto University, Luanda, Angola</mods:affiliation>
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</mods:name>
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<mods:name id="ACEBB74FEB1D2CC5D49E85F2B360A076" type="personal">
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<mods:roleTerm id="578A74F76872B30D9A8C7C2C559678FD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="393CC227AD235EAEE7EA5CE90F860BEC">Goncalves, Francisco Maiato P.</mods:namePart>
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<mods:nameIdentifier id="C2A3EA87551E0CA90D7D57DE79CD3289" type="ORCID">https://orcid.org/0000-0002-8859-7491</mods:nameIdentifier>
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<mods:affiliation id="312D75AD0CE4742F9006523D2A32C29F">National Geographic Okavango Wilderness Project, Wild Bird Trust, Hogsback, South Africa & Herbarium of Lubango, ISCED-Huila, Lubango, Angola</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="B493BE60CF5BF0ABC8576B894483B8C6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BD51F8B8F580E27F52225AF13425F473">Meller, Paulina</mods:namePart>
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<mods:nameIdentifier id="3537D0834FC6D1CC3BB86C270BF123CC" type="ORCID">https://orcid.org/0000-0001-6711-4385</mods:nameIdentifier>
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<mods:affiliation id="B301CE95DA0ADC46A967614AF177F98C">Institute for Plant Science and Microbiology, University of Hamburg, Hamburg, Germany</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="5773E99AC2E2CA0E428EC6B7963EEF9D">Paton, Alan J.</mods:namePart>
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<mods:nameIdentifier id="581ACB9DCA378B52D50B8687AB878FA5" type="ORCID">https://orcid.org/0000-0002-6052-6675</mods:nameIdentifier>
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<mods:affiliation id="36669552CE11960F3974070CAFB57782">Royal Botanic Gardens, Kew, TW 9 3 AE, London, UK</mods:affiliation>
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<mods:typeOfResource id="760971CAE9FD0F819D0BC82AF73746A1">text</mods:typeOfResource>
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<mods:title id="73C8988F4DDA123D916D0E66AF7039A5">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="D97F80A3367FCFB4D510A691A9CFAC39">
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<mods:date id="0C65E7AD86C00CE0C4535CC985C0C704">2023</mods:date>
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<mods:detail id="764A0159D6E3B2BC6C02ABB267661622" type="pubDate">
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<mods:number id="9D1062D8B858998025470ABC2BC4395D">2023-09-20</mods:number>
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</mods:detail>
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<mods:detail id="C7ED0D77934D841C2AE7CCA490E9DCAF" type="volume">
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<mods:number id="0F04F2488B35449F6501A0C8923983F9">232</mods:number>
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</mods:detail>
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<mods:start id="B32F3793B0285E2F14B4F98775827FBF">145</mods:start>
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<mods:end id="BB35D5A8A028822CAEF7369C9B7A0C70">166</mods:end>
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</mods:extent>
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<mods:location id="E8121C699CBCF757190733BC5D11227D">
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<mods:url id="A8B5D918699BC23614A061C82D9EC4F7">http://dx.doi.org/10.3897/phytokeys.232.110110</mods:url>
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</mods:location>
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<mods:classification id="34F5CE1BAF3224A613A6996C85825CD7">journal article</mods:classification>
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<mods:identifier id="D869DBF47A48D3E4299BCD8A2754B0E7" type="DOI">http://dx.doi.org/10.3897/phytokeys.232.110110</mods:identifier>
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<mods:identifier id="D769DEDC7EED65A09068C500CA6F8FD9" type="Pensoft-Pub">1314-2003-232-145</mods:identifier>
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<mods:identifier id="2DC7B18038CBD3FF1FAB83807DC7E89E" type="Pensoft-UUID">B24E305A09B9582DBFDBAD8120894678</mods:identifier>
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</mods:mods>
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<treatment id="B3859CC85EBA53908D5AB828A10A4B47" LSID="urn:lsid:plazi:treatment:B3859CC85EBA53908D5AB828A10A4B47" httpUri="http://treatment.plazi.org/id/B3859CC85EBA53908D5AB828A10A4B47" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection id="A6478A00FE38E5F8D87BA8C13843B467" pageId="0" pageNumber="145" type="nomenclature">
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<paragraph id="2DD506881902C0BF6E587185FCA9933A" pageId="0" pageNumber="145">
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<taxonomicName id="565BF150F07484F34302F48D7FE7EFE9" LSID="B3859CC8-5EBA-5390-8D5A-B828A10A4B47" authority="A. J. Paton & Goyder" authorityName="A. J. Paton & Goyder" authorityYear="2023" class="Magnoliopsida" family="Lamiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon palustris" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="palustris" status="sp. nov.">Endostemon palustris A.J.Paton & Goyder</taxonomicName>
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<taxonomicNameLabel id="60A977E89FA705481909FF6A5EFD2920" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="A411BDCA8902E5E745E2652823B033FD" pageId="0" pageNumber="145" type="diagnosis">
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<paragraph id="F23EF13496CD5A877D1C7C977D25DA39" pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph id="061E48842FF9F738AE1913057FB17C29" pageId="0" pageNumber="145">
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Differs from
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<taxonomicName id="2707DC92B8B2A4B292A393180D88D38A" lsidName="E. tubulascens" pageId="0" pageNumber="145" rank="species" species="tubulascens">
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<emphasis id="EFE7148005D1DA42897D52ABB79F403E" italics="true" pageId="0" pageNumber="145">E. tubulascens</emphasis>
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</taxonomicName>
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(Briq.) M.Ashby in the little-branched habit, the sessile rather than petiolate and linear rather than elliptic leaves, the fewer-flowered (4-6-flowered rather than 6-10-flowered) verticils in the inflorescence, the pale violet rather than pinkish white flowers and the longer calyx (fruiting calyx 8-9 mm long rather than 5.5-6 mm).
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</paragraph>
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</subSubSection>
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<subSubSection id="1AD26F86C279E8714D19316800DCD43A" pageId="0" pageNumber="145" type="materials_examined">
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<paragraph id="76FA4E045BC4A84FA960970225EE0960" pageId="0" pageNumber="145">
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<typeStatus id="075AE803517E6C85E155C55C5817CD39">Type</typeStatus>
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.
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</paragraph>
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<paragraph id="96C8B4872D3615AFDFC2C831DF80F6B9" pageId="0" pageNumber="145">
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<materialsCitation id="C2A86B50D16B34EFBBC049E77DFC10D7" collectingDate="2016-10-20" collectorName="D. Goyder, F. Maiato" country="Angola" latitude="-12.748611" location="Kalua River" longLatPrecision="21" longitude="18.354445" municipality="Confluence of Cuito River" specimenCount="1" stateProvince="Moxico Province" typeStatus="holotype">
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<collectingCountry id="1C825456499360557C83D732E90C884E" name="Angola">Angola</collectingCountry>
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.
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<collectingRegion id="14E7D8B9518B419D21EB5BEE2C3EABFE" country="Angola" name="Moxico">Moxico Province</collectingRegion>
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:
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<collectingMunicipality id="08E8A0B5DB577159C1667D98516DB2D4">Confluence of Cuito River</collectingMunicipality>
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and its 1st tributary, the
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<location id="E8FC47F2EA86EDAD72374A501F8F4701" LSID="urn:lsid:plazi:treatment:B3859CC85EBA53908D5AB828A10A4B47:E8FC47F2EA86EDAD72374A501F8F4701" country="Angola" latitude="-12.748611" longLatPrecision="21" longitude="18.354445" municipality="Confluence of Cuito River" name="Kalua River" stateProvince="Moxico Province">Kalua River</location>
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, c.
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<locationDeviation id="5D28A117D499A545CAE868D15D92E7C4" location="Munhango">
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<quantity id="87193D8D5D20004BF421C6F99A8F6BAA" metricMagnitude="4" metricUnit="m" metricValue="6.5" unit="km" value="65.0">65 km</quantity>
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SSW of Munhango
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</locationDeviation>
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,
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<geoCoordinate id="38828A5CB53D42A70549A7AE70C66306" degrees="12" direction="south" minutes="44" orientation="latitude" precision="15" seconds="55" value="-12.748611">12°44'55"S</geoCoordinate>
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,
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<geoCoordinate id="A413DC75956376BC46D09C6E134DB09E" degrees="018" direction="east" minutes="21" orientation="longitude" precision="15" seconds="16" value="18.354445">018°21'16"E</geoCoordinate>
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, fl.
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<collectingDate id="71C24AD4B9FC5638B811EAAE4517D544" value="2016-10-20">20 Oct. 2016</collectingDate>
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,
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<emphasis id="3A83FC78B5AE01A9C40824638F1F378F" italics="true" pageId="0" pageNumber="145">
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<collectorName id="F5F9F3BEE94AFCA0004114F6115C98A6">D.Goyder</collectorName>
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&
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<collectorName id="95F0E60D2BF27D996146744DB68196BD">F.Maiato</collectorName>
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</emphasis>
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8762 (
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<typeStatus id="AA593505041B666DAC7C03D5042717FC">holotype</typeStatus>
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: K (K001333409); isotypes: INBAC, LUBA)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BF2D820C1856DCB9B898A4C196A5AEF0" pageId="0" pageNumber="145" type="description">
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<paragraph id="6C06DFC407BCEDE729CAF623C48D122E" pageId="0" pageNumber="145">Description.</paragraph>
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<paragraph id="C15D78FB692B9ACA21AD6B361D81B1AF" pageId="0" pageNumber="145">
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Aromatic perennial suffrutex with few stems arising from a thick woody rootstock; stems erect, 15-30 cm tall, branched near the base, square in section above, more rounded below, pubescent with both glandular and eglandular hairs. Leaves verticillate, ascending, sessile, linear and folded along the midvein, 2.5-5 cm long, 0.1-0.2 cm wide, pubescent. Inflorescence lax with 4-6-flowered verticils 1.5-2.5 cm apart; bracts lanceolate or narrowly ovate, 3.5-4.5 mm long; pedicels 1-5 mm long, longer in lower verticils. Calyx 5 mm long at anthesis, pubescent with spreading hairs and sessile glands, posterior lip purplish; fruiting calyx 8-9 mm long. Corolla pale violet, 8-10 mm long; tube 6-7 mm long, straight, parallel-sided, dilating at the throat. Filaments c. 0.5 mm long, glabrous or pilose. Ovaries pubescent at apex. (Fig.
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<figureCitation id="03D3FE6D705D7C5620557C783CA8453A" captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Endostemon palustris A habit B flower C opened corolla showing androecium and gynoecium D base of gynoecium showing three of the four nutlets and the gynobasic style E individual nutlet F calyx G indumentum. Drawn by Margaret Tebbs from Goyder & Maiato 8762." figureDoi="10.3897/phytokeys.232.110110.figure4" httpUri="https://binary.pensoft.net/fig/908502" pageId="0" pageNumber="145">4</figureCitation>
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).
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</paragraph>
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<caption id="F306E9FE1A6AEE8ADFEE26079614378A" doi="10.3897/phytokeys.232.110110.figure4" httpUri="https://binary.pensoft.net/fig/908502" pageId="0" pageNumber="145" start="Figure 4" startId="F6">
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<paragraph id="E9E287144166A4E20D0FBD3A3CAB91D3" pageId="0" pageNumber="145">
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<emphasis id="94C6B09413794A1E6A92C2194CF1929B" bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
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<taxonomicName id="B15D474737EB8EBEE97B3DBBC0F516FE" authorityName="A. J. Paton & Goyder" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon palustris" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="palustris">
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<emphasis id="E928A715342EFF4BDA5D1FBC4D06D7AB" italics="true" pageId="0" pageNumber="145">Endostemon palustris</emphasis>
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</taxonomicName>
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<emphasis id="293DDEB7E85E9E9AA2358FA19593A0BE" bold="true" pageId="0" pageNumber="145">A</emphasis>
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habit
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<emphasis id="FDF48ED360EF785B7B2FC86396295EEF" bold="true" pageId="0" pageNumber="145">B</emphasis>
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flower
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<emphasis id="BE830F14023ACA338CBDF12D6ECCD015" bold="true" pageId="0" pageNumber="145">C</emphasis>
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opened corolla showing androecium and gynoecium
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<emphasis id="4DB1C6E74F4E9E8AEEE36552B13DB674" bold="true" pageId="0" pageNumber="145">D</emphasis>
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base of gynoecium showing three of the four nutlets and the gynobasic style
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<emphasis id="0AEC353B8399C95E4E13A8196AEDE2A5" bold="true" pageId="0" pageNumber="145">E</emphasis>
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individual nutlet
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<emphasis id="11185B2B30A9A5F06BBF057C5B546AE3" bold="true" pageId="0" pageNumber="145">F</emphasis>
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calyx
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<emphasis id="60E84E3C226A6127C0B3405B11D67C25" bold="true" pageId="0" pageNumber="145">G</emphasis>
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indumentum. Drawn by Margaret Tebbs from
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<emphasis id="6A158499B74AFBD45F2846D75C261367" italics="true" pageId="0" pageNumber="145">Goyder & Maiato</emphasis>
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8762.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="C5F97C226AD3889345BC97B4D36E89DE" pageId="0" pageNumber="145" type="distribution">
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<paragraph id="FD5CF349CA9848EF8FB3BDE423ACC308" pageId="0" pageNumber="145">Distribution and ecology.</paragraph>
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<paragraph id="E27108FD6E90360452290991DE01F7C5" pageId="0" pageNumber="145">
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Known only from the type collection close to the source of the Cuito River. It was found towards the upper edge of a marsh fringing the river. (Map
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<figureCitation id="5F613F2BD73B30D3812F786443EA8DEF" captionStart="Map 3" captionStartId="F7" captionText="Map 3. Known distribution of Endostemon palustris (star), and of the related species E. tubulascens (cross)." figureDoi="10.3897/phytokeys.232.110110.map3" httpUri="https://binary.pensoft.net/fig/908503" pageId="0" pageNumber="145">3</figureCitation>
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).
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</paragraph>
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<caption id="EB8248EFB69AF8173228CE6B23421DAA" doi="10.3897/phytokeys.232.110110.map3" httpUri="https://binary.pensoft.net/fig/908503" pageId="0" pageNumber="145" start="Map 3" startId="F7">
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<paragraph id="98DC899345228EE5EAEB90C87EAA2823" pageId="0" pageNumber="145">
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|
<emphasis id="9A2C50BB39D69290B50D2044B8D9D6B8" bold="true" pageId="0" pageNumber="145">Map 3.</emphasis>
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|
Known distribution of
|
|
<taxonomicName id="8B8AA453C49A99857EBBB38D01E55061" authorityName="A. J. Paton & Goyder" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon palustris" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="palustris">
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|
<emphasis id="FA63DC0CAAAA4A4477D5E2D834621D6F" italics="true" pageId="0" pageNumber="145">Endostemon palustris</emphasis>
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|
</taxonomicName>
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|
(star), and of the related species
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|
<taxonomicName id="5F3D18CDE425A1F70411B143D990D3A2" lsidName="E. tubulascens" pageId="0" pageNumber="145" rank="species" species="tubulascens">
|
|
<emphasis id="BE03BE398088CA2D2C6CDEF892B81C16" italics="true" pageId="0" pageNumber="145">E. tubulascens</emphasis>
|
|
</taxonomicName>
|
|
(cross).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="18D28DFC54CB2A6D12EA775176AA2091" pageId="0" pageNumber="145" type="conservation status">
|
|
<paragraph id="2B4577131A5C60E79F09A851AFC25F05" pageId="0" pageNumber="145">Conservation status.</paragraph>
|
|
<paragraph id="BB5424DFBEA6570A53B68F259A78E2AC" pageId="0" pageNumber="145">
|
|
Although known from a single locality,
|
|
<taxonomicName id="279209ADFF0110534B114318E8BB98BF" authorityName="A. J. Paton & Goyder" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon palustris" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="palustris">
|
|
<emphasis id="A826D81ADE739A50767D938BACE99BDB" italics="true" pageId="0" pageNumber="145">Endostemon palustris</emphasis>
|
|
</taxonomicName>
|
|
occupies a habitat that is extensive within the upper catchment of the Cuito and its tributaries. The area is not threatened with agricultural development being both nutrient poor and many kilometres from any human habitation, but with so little information the species is provisionally assessed as Data Deficient.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A6AE4EA542DF51F9054DC9D3B7C4341E" pageId="0" pageNumber="145" type="taxonomic notes">
|
|
<paragraph id="84BD5F78B5F0B1299E9A5A12C574A560" pageId="0" pageNumber="145">Taxonomic notes.</paragraph>
|
|
<paragraph id="1F9B9695F3C9E89EF3CED5AF6E2EE9BD" pageId="0" pageNumber="145">
|
|
<taxonomicName id="37D400826993E30094DEC37F3EB10B6F" authorityName="N.E.Brown" authorityYear="1910" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="7676913784B4349282DDBC0E56A7AECD" italics="true" pageId="0" pageNumber="145">Endostemon</emphasis>
|
|
</taxonomicName>
|
|
N.E.Br. is an isolated genus of 20 species within the tribe Ocimeae, with two centres of endemism - Angola and the Horn of Africa. It can be recognised within the
|
|
<taxonomicName id="68C543774689083E0117BDCF875D6AD2" class="Magnoliopsida" family="Lamiaceae" genus="Orthosiphon" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="Orthosiphon" order="Lamiales" pageId="0" pageNumber="145" phylum="Magnoliophyta" rank="genus">
|
|
<emphasis id="1DF636461E51E5E1CC9145CDCF1C6B31" italics="true" pageId="0" pageNumber="145">Orthosiphon</emphasis>
|
|
</taxonomicName>
|
|
Benth. group of genera by its short, villous staminal filaments, an expanded shield-like base to the style, and pollen with alternating wide and narrow mesocolpia (
|
|
<bibRefCitation id="412F5AB069BE7B36107AB1B22B6C6A9B" DOI="https://doi.org/10.2307/4118065" author="Paton, A" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="145" pagination="673 - 716" refId="B43" refString="Paton, A, Harley, MM, Harley, RM, Weeks, S, 1994. A Revision of Endostemon (Labiatae). Kew Bulletin 49 (4): 673 - 716, DOI: https://doi.org/10.2307/4118065" title="A Revision of Endostemon (Labiatae)." url="https://doi.org/10.2307/4118065" volume="49" year="1994">Paton et al. 1994</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="B55297B086DD3F3F22B96A228FEDDBED" DOI="https://doi.org/10.2307/4111205" author="Ryding, O" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="145" pagination="919 - 927" refId="B50" refString="Ryding, O, Paton, A, Thulin, M, Springate, D, 2003. [publ. 2004] Reconsideration of the genus Puntia and a new species of the genus Endostemon (Lamiaceae). Kew Bulletin 58 (4): 919 - 927, DOI: https://doi.org/10.2307/4111205" title="[publ. 2004] Reconsideration of the genus Puntia and a new species of the genus Endostemon (Lamiaceae)." url="https://doi.org/10.2307/4111205" volume="58" year="2003">Ryding et al. 2003</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="B891C932B2B433B4A2362E207B28DF95" DOI="https://doi.org/10.1016/j.ympev.2003.08.002" author="Paton, AJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="145" pagination="277 - 299" refId="B44" refString="Paton, AJ, Springate, D, Suddee, S, Otieno, D, Grayer, RJ, Harley, MM, Willis, F, Simmonds, MSJ, Powell, MP, Savolainen, V, 2004. Phylogeny and Evolution of Basils and Allies (Ocimeae, Labiatae) based on three Plastid DNA Regions. Molecular Phylogenetics and Evolution 31 (1): 277 - 299, DOI: https://doi.org/10.1016/j.ympev.2003.08.002" title="Phylogeny and Evolution of Basils and Allies (Ocimeae, Labiatae) based on three Plastid DNA Regions." url="https://doi.org/10.1016/j.ympev.2003.08.002" volume="31" year="2004">Paton et al. 2004</bibRefCitation>
|
|
). With the exception of
|
|
<taxonomicName id="4BEE3160F6A0161F198B958D94D62A68" lsidName="E. tereticaulis" pageId="0" pageNumber="145" rank="species" species="tereticaulis">
|
|
<emphasis id="6E4164A37EDC50C8654E05EEA9A0AD03" italics="true" pageId="0" pageNumber="145">E. tereticaulis</emphasis>
|
|
</taxonomicName>
|
|
(Poir.) M. Ashby which is in
|
|
<taxonomicName id="AD2FE0C5C54F993740379B9BC9B2799D" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon subsp. sect." order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="section" section="Oblongi">Endostemon sect. Oblongi</taxonomicName>
|
|
Ayob. ex A.J.Paton, Harley & M.M.Harley, all of the Angolan species fall within
|
|
<taxonomicName id="A7A0B3FDFF7884048ACC12C746EF71D8" class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon subsp. sect." order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="section" section="Endostemon">Endostemon sect. Endostemon</taxonomicName>
|
|
. This section is characterised by a relatively short calyx tube with an open glabrous throat, and the form of the lateral calyx lobes - the posterior margins of these lobes are not extended towards the upper lip of the calyx.
|
|
<taxonomicName id="91B8022AD447C152A192B72EBC1EE5B6" authorityName="M.R.Ashby" authorityYear="1936" baseAuthorityName="Briq." class="Magnoliopsida" family="Rubiaceae" genus="Endostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endostemon tubulascens" order="Lamiales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="tubulascens">
|
|
<emphasis id="9D3C2FABF1327054E3B4D9B5DBEBE1D7" italics="true" pageId="0" pageNumber="145">Endostemon tubulascens</emphasis>
|
|
</taxonomicName>
|
|
, to which this new species is compared, appears largely restricted to the high escarpment zone around Lubango,
|
|
<normalizedToken id="27DF9A02FBB94A04EFE596D583B9132B" originalValue="Huíla">Huila</normalizedToken>
|
|
Province, Angola.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |