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<document id="C8C4B2896E6182A3D3A53B15666644BB" ID-DOI="10.11646/zootaxa.5492.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13234851" ID-ZooBank="8F81F053-DE78-4684-BCC9-875AE0C41615" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722932859467" checkinUser="plazi" docAuthor="Schmidt, Joachim &amp; Merene, Yeshitla" docDate="2024" docId="B36D0606FFD2AC1DFF28FB45FF28F9A9" docLanguage="en" docName="zootaxa.5492.3.3.pdf" docOrigin="Zootaxa 5492 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus (Abyssinotus) yitbareki Schmidt &amp; Merene 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="349" masterDocId="4F547E7EFFD6AC1BFFBFFFDAFFC3FFF8" masterDocTitle="Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)" masterLastPageNumber="355" masterPageNumber="343" pageNumber="347" updateTime="1722933277969" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="55A518D722F311C4DAA1EB61BEEE0DC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F06EA92400F3812858F7EF85453400E7">
<mods:title id="07C8676D61726F3238773882CE805066">Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)</mods:title>
</mods:titleInfo>
<mods:name id="25F23F771CA9F50298FE0F01CA83441E" type="personal">
<mods:role id="277FCA8B19C52D1BD9C83480F25E3583">
<mods:roleTerm id="48281BB66B234DC2ACF4E2F92F9C3E93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EEFF966BDD9B161321B3C3B129ED1076">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="3BA0A6BB53C67201EDC4BCD140999FA1">General and Systematic Zoology, University of Rostock, Germany</mods:affiliation>
</mods:name>
<mods:name id="36272D8AC05DC6C4E34CE4FB93007495" type="personal">
<mods:role id="2BFD966415074E656C150305218F799A">
<mods:roleTerm id="33DBD7E90553466617CEBF75FF020F09">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B8612EA0FA07AC4C6DD1487FFE908FB">Merene, Yeshitla</mods:namePart>
<mods:affiliation id="821C1FB7D82BE3C5BFCC98BC54B6FA89">Department of Zoological Sciences, Addis Ababa University, Ethiopia &amp; Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6A081C2A84A3A2530975F74C00D9DBE2">text</mods:typeOfResource>
<mods:relatedItem id="9DE8F87A70DAA3DDB86948F72FA2A9E0" type="host">
<mods:titleInfo id="FBF713E3477C9BEE7A37070BFDFA16E9">
<mods:title id="5BDA35A243F2CF098D96019A6E21C649">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6ACE91E524CA5F6E8295193BE10CD446">
<mods:date id="0A951598204E69CA924D7BD19F0F9EA1">2024</mods:date>
<mods:detail id="47BAE73BFF25451A19BFCB33AFDCDA4B" type="pubDate">
<mods:number id="3BEBE6375B28A61BC7D9E3A68023046E">2024-08-06</mods:number>
</mods:detail>
<mods:detail id="398AF53FE2C43C285AC9EF929947262F" type="volume">
<mods:number id="6DCB07C86B939A5D5D1C14DBFDEBB3B1">5492</mods:number>
</mods:detail>
<mods:detail id="BED93DAA9109E5532F524FCB946A0839" type="issue">
<mods:number id="0BF6D3639657CADBCEDAFE736664A29B">3</mods:number>
</mods:detail>
<mods:extent id="1D88509744EEBE8537B136EE62F2CE2F" unit="page">
<mods:start id="837729CB8B3452DAF20841F5BF40D608">343</mods:start>
<mods:end id="5872FD72B15DCC9359AC7AA2078C3925">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="60D8AFC01483BA3F7122635F70CD97D6">
<mods:url id="446DEBEA7A5F70119EA6F75301843B99">http://dx.doi.org/10.11646/zootaxa.5492.3.3</mods:url>
</mods:location>
<mods:classification id="B224FCC13A43C4D285801ED0B67D2C0F">journal article</mods:classification>
<mods:identifier id="158981C8AA3E52374A5FB494DE7ABCEA" type="DOI">10.11646/zootaxa.5492.3.3</mods:identifier>
<mods:identifier id="25AC41417985C807F5B2B8E1861361E3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FD923AC2E38396DA532C26B6F3455F1C" type="Zenodo-Dep">13234851</mods:identifier>
<mods:identifier id="9E882957F13C5464A1E35EB9DE789D7C" type="ZooBank">8F81F053-DE78-4684-BCC9-875AE0C41615</mods:identifier>
</mods:mods>
<treatment id="B36D0606FFD2AC1DFF28FB45FF28F9A9" ID-DOI="http://doi.org/10.5281/zenodo.13234867" ID-Zenodo-Dep="13234867" LSID="urn:lsid:plazi:treatment:B36D0606FFD2AC1DFF28FB45FF28F9A9" httpUri="http://treatment.plazi.org/id/B36D0606FFD2AC1DFF28FB45FF28F9A9" lastPageId="6" lastPageNumber="349" pageId="4" pageNumber="347">
<subSubSection id="73DEE49BFFD2AC1FFF28FB45FD14FB42" box="[151,727,1183,1210]" pageId="4" pageNumber="347" type="nomenclature">
<paragraph id="3B7BB710FFD2AC1FFF28FB45FD14FB42" blockId="4.[151,727,1183,1247]" box="[151,727,1183,1210]" pageId="4" pageNumber="347">
<heading id="6033007CFFD2AC1FFF28FB45FD14FB42" bold="true" box="[151,727,1183,1210]" fontSize="11" level="1" pageId="4" pageNumber="347" reason="1">
<emphasis id="09B06B02FFD2AC1FFF28FB45FD14FB42" bold="true" box="[151,727,1183,1210]" pageId="4" pageNumber="347">
<taxonomicName id="FCC4CC93FFD2AC1FFF28FB45FD4BFB42" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[151,648,1183,1210]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. nov." subGenus="Abyssinotus">
<emphasis id="09B06B02FFD2AC1FFF28FB45FF3AFB42" bold="true" box="[151,249,1183,1210]" italics="true" pageId="4" pageNumber="347">Trechus</emphasis>
(
<emphasis id="09B06B02FFD2AC1FFEB4FB45FE5FFB42" bold="true" box="[267,412,1183,1210]" italics="true" pageId="4" pageNumber="347">Abyssinotus</emphasis>
)
<emphasis id="09B06B02FFD2AC1FFE12FB45FDD4FB42" bold="true" box="[429,535,1183,1210]" italics="true" pageId="4" pageNumber="347">yitbareki</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD2AC1FFD2AFB7AFD14FB42" box="[661,727,1184,1210]" pageId="4" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD2AC1FFF28FB1EFE94FB27" box="[151,343,1220,1247]" pageId="4" pageNumber="347" type="description">
<paragraph id="3B7BB710FFD2AC1FFF28FB1EFE94FB27" blockId="4.[151,727,1183,1247]" box="[151,343,1220,1247]" pageId="4" pageNumber="347">
<figureCitation id="A3FFAB95FFD2AC1FFF28FB1EFF2DFB26" box="[151,238,1220,1246]" captionStart="FIGURE 2" captionStartId="5.[152,255,1889,1913]" captionTargetBox="[280,1308,182,1866]" captionTargetId="figure-24@5.[280,1308,181,1866]" captionTargetPageId="5" captionText="FIGURE 2. Habitus photographs of Trechus (Abyssinotus) species; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. A, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, T. yitbareki Schmidt, sp. n., paratype. C, T. kniphofia Schmidt, sp. n., paratype. D, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234855" httpUri="https://zenodo.org/record/13234855/files/figure.png" pageId="4" pageNumber="347">Fig. 2B</figureCitation>
,
<figureCitation id="A3FFAB95FFD2AC1FFF46FB1EFE94FB27" box="[249,343,1220,1247]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="4" pageNumber="347">3C, F, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD2AC1FFF28FAD6FB60FA4E" pageId="4" pageNumber="347" type="materials_examined">
<paragraph id="3B7BB710FFD2AC1FFF28FAD6FB60FA4E" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">
<emphasis id="09B06B02FFD2AC1FFF28FAD6FE81FADE" bold="true" box="[151,322,1292,1318]" pageId="4" pageNumber="347">Type material.</emphasis>
<materialsCitation id="8BACBD4DFFD2AC1FFEF9FAD6FBD5FAB2" collectingDate="2019-02-23" collectionCode="CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="4" pageNumber="347" specimenCount="1" specimenCount-male="1" stateProvince="Amara" typeStatus="holotype">
<typeStatus id="E47F09B2FFD2AC1FFEF9FAD6FE70FADE" box="[326,435,1292,1318]" pageId="4" pageNumber="347" type="holotype">Holotype</typeStatus>
:
<specimenCount id="2DC27C99FFD2AC1FFE04FAD7FE11FADF" box="[443,466,1293,1319]" count="1" pageId="4" pageNumber="347" type="male">♂</specimenCount>
, with label data:
<collectingCountry id="43D3F780FFD2AC1FFD39FAD6FD2BFADE" box="[646,744,1292,1318]" name="Ethiopia" pageId="4" pageNumber="347">Ethiopia</collectingCountry>
,
<collectingRegion id="F90079F2FFD2AC1FFD50FAD6FC8FFADE" box="[751,844,1292,1318]" country="Ethiopia" name="Amara" pageId="4" pageNumber="347">Amhara</collectingRegion>
, W-slope
<location id="3E1BE1CBFFD2AC1FFC0AFAD7FBEEFADE" LSID="urn:lsid:plazi:treatment:B36D0606FFD2AC1DFF28FB45FF28F9A9:3E1BE1CBFFD2AC1FFC0AFAD7FBEEFADE" box="[949,1069,1292,1319]" country="Ethiopia" latitude="10.635278" longLatPrecision="21" longitude="37.764168" name="Mt. Choke" pageId="4" pageNumber="347" stateProvince="Amara">Mt.Choke</location>
,
<quantity id="FC3C1AF5FFD2AC1FFB89FAD6FB49FADF" box="[1078,1162,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="4" pageNumber="347" unit="m" value="3370.0">
<elevation id="B0E95023FFD2AC1FFB89FAD6FB49FADF" box="[1078,1162,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="4" pageNumber="347" unit="m" value="3370.0">3370 m</elevation>
</quantity>
,
<date id="4F7A91D0FFD2AC1FFB2CFAD6FAC8FADE" box="[1171,1291,1292,1318]" pageId="4" pageNumber="347" value="2019-02-23">
<collectingDate id="5F3E6838FFD2AC1FFB2CFAD6FAC8FADE" box="[1171,1291,1292,1318]" pageId="4" pageNumber="347" value="2019-02-23">23.II.2019</collectingDate>
</date>
,
<geoCoordinate id="5EF0D1D7FFD2AC1FFAACFAD6FA5FFADE" box="[1299,1436,1292,1318]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="347" precision="15" seconds="07" value="10.635278">10°38‘07“N</geoCoordinate>
<geoCoordinate id="5EF0D1D7FFD2AC1FFF28FAEAFEE3FAB3" box="[151,288,1328,1355]" degrees="37" direction="east" minutes="45" orientation="longitude" pageId="4" pageNumber="347" precision="15" seconds="51" value="37.764168">37°45‘51“E</geoCoordinate>
, leg.
<collectorName id="9631D2C6FFD2AC1FFEDBFAEBFE12FAB2" box="[356,465,1328,1355]" pageId="4" pageNumber="347">D. Hauth</collectorName>
, J. Schmidt,
<collectorName id="9631D2C6FFD2AC1FFDD3FAEAFD30FAB3" box="[620,755,1328,1355]" pageId="4" pageNumber="347">Yeshitla M.</collectorName>
,
<collectorName id="9631D2C6FFD2AC1FFCBDFAEAFC57FAB3" box="[770,916,1328,1355]" pageId="4" pageNumber="347">Yitbarek, W.</collectorName>
(
<collectionCode id="5DD52FD5FFD2AC1FFC19FAEAFBCCFAB2" box="[934,1039,1328,1354]" pageId="4" pageNumber="347">CSCHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8BACBD4DFFD2AC1FFB9AFAEBFCD2FA96" collectingDate="2019-02-23" collectionCode="NHMAA, ZSM, CAF, CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="4" pageNumber="347" specimenCount="50" specimenCount-female="17" specimenCount-male="33" stateProvince="Amara" typeStatus="paratype">
<typeStatus id="E47F09B2FFD2AC1FFB9AFAEBFB55FAB3" box="[1061,1174,1329,1355]" pageId="4" pageNumber="347" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2DC27C99FFD2AC1FFB1BFAEAFB34FAB3" box="[1188,1271,1328,1355]" count="33" pageId="4" pageNumber="347" type="male">33 ♂♂</specimenCount>
,
<specimenCount id="2DC27C99FFD2AC1FFAB8FAEAFA9AFAB3" box="[1287,1369,1328,1355]" count="17" pageId="4" pageNumber="347" type="female">17 ♀♀</specimenCount>
, with same data as holotype (
<collectionCode id="5DD52FD5FFD2AC1FFE22FA8EFDC8FA96" box="[413,523,1364,1390]" pageId="4" pageNumber="347">NHMAA</collectionCode>
,
<collectionCode id="5DD52FD5FFD2AC1FFDA8FA8EFD90FA96" box="[535,595,1364,1390]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="347" type="Museum">ZSM</collectionCode>
,
<collectionCode id="5DD52FD5FFD2AC1FFDE0FA8EFD55FA96" box="[607,662,1364,1390]" country="China" lsid="urn:lsid:biocol.org:col:12802" name="Chinese Academy of Forestry" pageId="4" pageNumber="347" type="Herbarium">CAF</collectionCode>
,
<collectionCode id="5DD52FD5FFD2AC1FFD1DFA8EFCCAFA96" box="[674,777,1364,1390]" pageId="4" pageNumber="347">CSCHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8BACBD4DFFD2AC1FFCA4FA8EFAF7FA6A" collectingDate="2019-02-22" collectionCode="NHMAA, ZSM, CSCHM" collectorName="D. Hauth &amp; J. Schmidt &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3450" latitude="10.635833" location="Mt. Choke" longLatPrecision="21" longitude="37.768333" pageId="4" pageNumber="347" specimenCount="79" specimenCount-female="31" specimenCount-male="48" stateProvince="Amara" typeStatus="paratype">
<specimenCount id="2DC27C99FFD2AC1FFCA4FA8EFCA8FA97" box="[795,875,1364,1391]" count="48" pageId="4" pageNumber="347" type="male">48 ♂♂</specimenCount>
,
<specimenCount id="2DC27C99FFD2AC1FFCC8FA8EFC06FA97" box="[887,965,1364,1391]" count="31" pageId="4" pageNumber="347" type="female">31 ♀♀</specimenCount>
, W-slope
<location id="3E1BE1CBFFD2AC1FFB88FA8FFB71FA96" LSID="urn:lsid:plazi:treatment:B36D0606FFD2AC1DFF28FB45FF28F9A9:3E1BE1CBFFD2AC1FFB88FA8FFB71FA96" box="[1079,1202,1364,1391]" country="Ethiopia" latitude="10.635833" longLatPrecision="21" longitude="37.768333" name="Mt. Choke" pageId="4" pageNumber="347" stateProvince="Amara">Mt. Choke</location>
,
<quantity id="FC3C1AF5FFD2AC1FFB02FA8EFAD7FA97" box="[1213,1300,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="4" pageNumber="347" unit="m" value="3450.0">
<elevation id="B0E95023FFD2AC1FFB02FA8EFAD7FA97" box="[1213,1300,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="4" pageNumber="347" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,
<date id="4F7A91D0FFD2AC1FFA9EFA8EFA5BFA96" box="[1313,1432,1364,1390]" pageId="4" pageNumber="347" value="2019-02-22">
<collectingDate id="5F3E6838FFD2AC1FFA9EFA8EFA5BFA96" box="[1313,1432,1364,1390]" pageId="4" pageNumber="347" value="2019-02-22">22.II.2019</collectingDate>
</date>
,
<geoCoordinate id="5EF0D1D7FFD2AC1FFF28FAA2FEE3FA6A" box="[151,288,1400,1426]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="347" precision="15" seconds="09" value="10.635833">10°38‘09“N</geoCoordinate>
<geoCoordinate id="5EF0D1D7FFD2AC1FFE98FAA2FE6CFA6A" box="[295,431,1400,1426]" degrees="37" direction="east" minutes="46" orientation="longitude" pageId="4" pageNumber="347" precision="15" seconds="06" value="37.768333">37°46‘06“E</geoCoordinate>
, leg.
<collectorName id="9631D2C6FFD2AC1FFE56FAA3FD91FA6A" box="[489,594,1400,1427]" pageId="4" pageNumber="347">D. Hauth</collectorName>
,
<collectorName id="9631D2C6FFD2AC1FFDE2FAA3FD16FA6A" box="[605,725,1400,1427]" pageId="4" pageNumber="347">J. Schmidt</collectorName>
,
<collectorName id="9631D2C6FFD2AC1FFD61FAA2FCA2FA6B" box="[734,865,1400,1427]" pageId="4" pageNumber="347">Yeshitla M.</collectorName>
,
<collectorName id="9631D2C6FFD2AC1FFCD4FAA2FC3BFA6B" box="[875,1016,1400,1427]" pageId="4" pageNumber="347">Yitbarek, W.</collectorName>
(
<collectionCode id="5DD52FD5FFD2AC1FFBBAFAA2FBB0FA6A" box="[1029,1139,1400,1426]" pageId="4" pageNumber="347">NHMAA</collectionCode>
,
<collectionCode id="5DD52FD5FFD2AC1FFBC1FAA2FB79FA6A" box="[1150,1210,1400,1426]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="347" type="Museum">ZSM</collectionCode>
,
<collectionCode id="5DD52FD5FFD2AC1FFB7AFAA2FAEFFA6A" box="[1221,1324,1400,1426]" pageId="4" pageNumber="347">CSCHM</collectionCode>
)
</materialsCitation>
; of these,
<materialsCitation id="8BACBD4DFFD2AC1FFF28FA46FDDEFA4E" box="[151,541,1436,1463]" country="Ethiopia" pageId="4" pageNumber="347" specimenCode="SMNS-L1821" specimenCount="1" specimenCount-male="1" stateProvince="Amara" typeStatus="paratype">
<specimenCount id="2DC27C99FFD2AC1FFF28FA46FF02FA4F" box="[151,193,1436,1463]" count="1" pageId="4" pageNumber="347" type="male">1 ♂</specimenCount>
(DNA voucher
<specimenCode id="6B621F6BFFD2AC1FFECAFA46FDD6FA4E" box="[373,533,1436,1462]" collectionCode="SMNS-L" pageId="4" pageNumber="347">SMNS-L1821</specimenCode>
)
</materialsCitation>
was used for molecular genetic study (
<bibRefCitation id="5F55CAE1FFD2AC1FFC6DFA46FB54FA4E" author="Faille, A. &amp; Hofmann, S. &amp; Merene, Y. &amp; Hauth, D. &amp; Opgenoorth, L. &amp; Woldehawariat, Y. &amp; Schmidt, J." box="[978,1175,1436,1462]" pageId="4" pageNumber="347" pagination="311 - 335" refId="ref9248" refString="Faille, A., Hofmann, S., Merene, Y., Hauth, D., Opgenoorth, L., Woldehawariat, Y. &amp; Schmidt, J. (2023) Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift, 70 (2), 311 - 335. https: // doi. org / 10.3897 / dez. 70.107425" type="journal article" year="2023">
Faille
<emphasis id="09B06B02FFD2AC1FFBA4FA47FB90FA4E" box="[1051,1107,1436,1462]" italics="true" pageId="4" pageNumber="347">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD2AC1FFF78FA1AFBF4FA06" pageId="4" pageNumber="347" type="etymology">
<paragraph id="3B7BB710FFD2AC1FFF78FA1AFBF4FA06" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">
<emphasis id="09B06B02FFD2AC1FFF78FA1AFE8BFA22" bold="true" box="[199,328,1472,1498]" pageId="4" pageNumber="347">Etymology</emphasis>
. The new species is dedicated to one of its collectors, Yitbarek Woldehawariat, professor and entomologist at the Department of Zoological Science, University of
<collectingRegion id="F90079F2FFD2AC1FFC21FA3EFBF0FA06" box="[926,1075,1508,1534]" country="Ethiopia" name="Adis Abeba" pageId="4" pageNumber="347">Addis Ababa</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD2AC1DFF78F9D2FEE2FB3D" lastPageId="6" lastPageNumber="349" pageId="4" pageNumber="347" type="description">
<paragraph id="3B7BB710FFD2AC1FFF78F9D2FD45F9DB" blockId="4.[151,1437,1292,1967]" box="[199,646,1544,1571]" pageId="4" pageNumber="347">
<emphasis id="09B06B02FFD2AC1FFF78F9D2FE92F9DA" bold="true" box="[199,337,1544,1570]" pageId="4" pageNumber="347">Description</emphasis>
. Body length:
<quantity id="FC3C1AF5FFD2AC1FFE47F9D2FD41F9DB" box="[504,642,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="4" pageNumber="347" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
.
</paragraph>
<paragraph id="3B7BB710FFD2AC1FFF78F9F6FA4CF992" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">Proportions (n = 10): PW/HW = 1.39–1.47 (Ø = 1.43); PW/PL = 1.33–1.42 (Ø = 1.37); PW/PBW = 1.18–1.22 (Ø = 1.20); PBW/PAW = 1.21–1.29 (Ø = 1.24); EW/PW = 1.41–1.51 (Ø = 1.46); EL/EW = 1.28–1.35 (Ø = 1.31).</paragraph>
<paragraph id="3B7BB710FFD2AC1FFF78F9AEFD55F92E" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">
Colour: Head, pronotum and elytra blackish brown, pronotum sometimes more reddish; mandibles, palps, antennal base and legs yellowish brown; antennae darkened beginning from second or third antennomere; in most specimens 2
<superScript id="CCB11A58FFD2AC1FFE9EF966FEF2F932" attach="left" box="[289,305,1724,1738]" fontSize="6" pageId="4" pageNumber="347">nd</superScript>
maxillary palpomere darkened.
</paragraph>
<paragraph id="3B7BB710FFD2AC1FFF78F93AFEDEF89E" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric, on clypeus very finely impressed, slightly transverse. Sculpticells on pronotal disc moderately impressed, slightly transverse. Elytra with large and deeply impressed, almost isodiametric or somewhat irregularly formed sculpticells.</paragraph>
<paragraph id="3B7BB710FFD2AC1FFF78F8AAFDACF857" blockId="4.[151,1437,1292,1967]" pageId="4" pageNumber="347">
Head: Compound eyes rather small, moderately convex; tempora about 4/5 of length of eyes. In all other characters as described in
<taxonomicName id="FCC4CC93FFD2AC1FFE02F84FFDEEF856" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[445,557,1941,1966]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD2AC1FFE02F84FFDEEF856" box="[445,557,1941,1966]" italics="true" pageId="4" pageNumber="347">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD2AC1FFD8BF84FFDACF857" box="[564,623,1941,1967]" pageId="4" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<caption id="6FBBE798FFD3AC1EFF27F8BBFDAEF81E" ID-DOI="http://doi.org/10.5281/zenodo.13234855" ID-Zenodo-Dep="13234855" httpUri="https://zenodo.org/record/13234855/files/figure.png" pageId="5" pageNumber="348" startId="5.[152,255,1889,1913]" targetBox="[280,1308,182,1866]" targetPageId="5" targetType="figure">
<paragraph id="3B7BB710FFD3AC1EFF27F8BBFDAEF81E" blockId="5.[151,1437,1889,2022]" pageId="5" pageNumber="348">
<emphasis id="09B06B02FFD3AC1EFF27F8BBFEDAF881" bold="true" box="[152,281,1889,1913]" pageId="5" pageNumber="348">FIGURE 2.</emphasis>
Habitus photographs of
<taxonomicName id="FCC4CC93FFD3AC1EFDABF8B8FD30F882" box="[532,755,1890,1914]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="subGenus" subGenus="Abyssinotus">
<emphasis id="09B06B02FFD3AC1EFDABF8B8FDA0F882" box="[532,611,1890,1914]" italics="true" pageId="5" pageNumber="348">Trechus</emphasis>
(
<emphasis id="09B06B02FFD3AC1EFDCDF8B8FD28F882" box="[626,747,1890,1914]" italics="true" pageId="5" pageNumber="348">Abyssinotus</emphasis>
)
</taxonomicName>
species; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series.
<emphasis id="09B06B02FFD3AC1EFB29F85FFB6BF865" bold="true" box="[1174,1192,1925,1949]" pageId="5" pageNumber="348">A</emphasis>
,
<taxonomicName id="FCC4CC93FFD3AC1EFB0AF85CFEECF83A" authority="(Queinnec &amp; Ollivier, 2021)" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFD3AC1EFB0AF85CFAD1F866" box="[1205,1298,1926,1950]" italics="true" pageId="5" pageNumber="348">T. reebae</emphasis>
(
<bibRefCitation id="5F55CAE1FFD3AC1EFAA0F85CFEEBF83A" author="Queinnec, E. &amp; Ollivier, E. &amp; Reeb, C." pageId="5" pageNumber="348" pagination="1 - 70" refId="ref9659" refString="Queinnec, E., Ollivier, E. &amp; Reeb, C. (2021) Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity, 27 (1), 1 - 70. https: // doi. org / 10.12976 / jib / 2021.27.1.1" type="journal article" year="2021">Quéinnec &amp; Ollivier, 2021</bibRefCitation>
)
</taxonomicName>
, specimen from western crater valley.
<emphasis id="09B06B02FFD3AC1EFD05F870FD08F83A" bold="true" box="[698,715,1962,1986]" pageId="5" pageNumber="348">B</emphasis>
,
<taxonomicName id="FCC4CC93FFD3AC1EFD67F870FC6AF83A" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[728,937,1962,1986]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD3AC1EFD67F870FC88F83A" box="[728,843,1962,1986]" italics="true" pageId="5" pageNumber="348">T. yitbareki</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD3AC1EFC0BF870FC2EF839" box="[948,1005,1962,1985]" pageId="5" pageNumber="348" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
<emphasis id="09B06B02FFD3AC1EFBE2F873FBACF839" bold="true" box="[1117,1135,1961,1985]" pageId="5" pageNumber="348">C</emphasis>
,
<taxonomicName id="FCC4CC93FFD3AC1EFBC2F870FA97F83A" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1149,1364,1962,1986]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFD3AC1EFBC2F870FB35F83A" box="[1149,1270,1962,1986]" italics="true" pageId="5" pageNumber="348">T. kniphofia</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD3AC1EFAE0F870FA54F839" box="[1375,1431,1962,1985]" pageId="5" pageNumber="348" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
<emphasis id="09B06B02FFD3AC1EFF44F814FECEF81E" bold="true" box="[251,269,1998,2022]" pageId="5" pageNumber="348">D</emphasis>
,
<taxonomicName id="FCC4CC93FFD3AC1EFEA5F814FE02F81E" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[282,449,1998,2022]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFD3AC1EFEA5F814FEA7F81D" box="[282,356,1998,2021]" italics="true" pageId="5" pageNumber="348">T. igori</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD3AC1EFE73F814FDC0F81D" box="[460,515,1998,2021]" pageId="5" pageNumber="348" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
</paragraph>
</caption>
<paragraph id="3B7BB710FFD0AC1DFF78FF4DFB4AFE55" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">Pronotum: Pronotum large and transverse, sub-discoidal, broadest slightly before middle, with base distinctly broader than apex; disc moderately convex.Anterior margin slightly concave, with anterior angles slightly protruded anteriorly, rounded; basal margin convex, with laterobasal angles markedly shifted anteriorly; lateral margin almost evenly rounded throughout; laterobasal angles obtuse, sometimes indistinct. Marginal gutter narrow, extending towards laterobasal foveae, slightly widened between laterobasal angle and foveae. Median longitudinal impression finely incised, disappearing near apex and base; anterior transverse impression indistinct, smooth, posterior transverse impression broad and very shallow, smooth; laterobasal foveae small, moderately deep, smooth. Lateral and laterobasal setae present, with the former situated at the end of anterior pronotal third.</paragraph>
<paragraph id="3B7BB710FFD0AC1DFF78FE6DFC04FE29" blockId="6.[151,1437,151,1617]" box="[199,967,439,465]" pageId="6" pageNumber="349">Pro- and mesothorax: Pro- and mesepisternum glabrous and smooth.</paragraph>
<paragraph id="3B7BB710FFD0AC1DFF78FE01FD16FCC1" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
Pterothorax: Elytra short, ovate, with disc convex, in dorsal view broadest in middle, humerus rounded, subapical sinuation indistinct, apex rounded. Striae 1–3 slightly impressed, indistinct in some specimens, smooth or suggestively punctate, external striae absent apart from the apical portion of stria 8 which is distinctly deepened before apex; parascutellar stria absent. Recurrent preapical stria short, deep, with outer curvature directed towards the fifth stria. Parascutellar seta present; three discal setae located in the 3rd interval, adjoined to the 3rd stria: anterior seta located near the end of the anterior elytral 8th; middle seta located anterior of elytral middle, near the end of elytral 3/7; posterior seta (= subapical seta) located about 1/8 of elytral length from elytral apex; seta of the recurrent stria isolated, removed from this stria by distance of the diameter of the pore. Number and positions of the setae of the marginal umbilicate series as normal for
<taxonomicName id="FCC4CC93FFD0AC1DFD53FD21FC87FCED" authorityName="Clairville" authorityYear="1806" box="[748,836,763,789]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="09B06B02FFD0AC1DFD53FD21FC87FCED" box="[748,836,763,789]" italics="true" pageId="6" pageNumber="349">Trechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFCF4FD21FCB0FCED" box="[843,883,763,789]" pageId="6" pageNumber="349" sensu="lato">s. l.</taxonomicNameLabel>
Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin.
</paragraph>
<paragraph id="3B7BB710FFD0AC1DFF78FC99FC1AFC5D" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
Legs: Moderately short and robust. Protibia dilated towards apex, straight on external margin, its dorsal surface with longitudinal groove developed but very flat, without micro-setae on anterior surface. Two basal protarsomeres of males markedly dilated and dentoid at the inner apical border (
<figureCitation id="A3FFAB95FFD0AC1DFCCCFC51FC0FFC5D" box="[883,972,907,933]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="3B7BB710FFD0AC1DFF78FC75FEE2FB3D" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
Male genitalia: EL/AL = 1.77–1.89 (Ø = 1.82; n = 8). Aedeagal median lobe robust, in lateral view markedly bent in basal half, almost straight towards apex, with moderately long apical lamella hook-like bent dorsally (
<figureCitation id="A3FFAB95FFD0AC1DFAD0FC09FF7BFBE9" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3F</figureCitation>
); in dorsal view with side margin almost parallel, very slightly widened in apical quarter and shortly narrowed towards slender apical lamella (
<figureCitation id="A3FFAB95FFD0AC1DFDBFFBC1FD9BFBCD" box="[512,600,1051,1077]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3H</figureCitation>
). Dorsal opening extends basally near to the apical margin of basal bulb. Sagittal aileron absent. Endophallus symmetrical in dorsal view, without a distinct copulatory piece, densely covered by tiny scales, in the inactive position with a sac-like folding structure near its base and an area covered by large sclerotized scales on its dorsal side (
<figureCitation id="A3FFAB95FFD0AC1DFD86FB5DFD53FB59" box="[569,656,1159,1185]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3F</figureCitation>
). Parameres similar in size and shape, rather slender, each with four apical setae.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD0AC1DFF78FB14FEACFA65" pageId="6" pageNumber="349" type="diagnosis">
<paragraph id="3B7BB710FFD0AC1DFF78FB14FEACFA65" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
<emphasis id="09B06B02FFD0AC1DFF78FB14FE06FB10" bold="true" box="[199,453,1230,1256]" pageId="6" pageNumber="349">Differential diagnosis</emphasis>
. Externally,
<taxonomicName id="FCC4CC93FFD0AC1DFDEEFB15FD0DFB11" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[593,718,1231,1257]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFDEEFB15FD0DFB11" box="[593,718,1231,1257]" italics="true" pageId="6" pageNumber="349">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFD6CFB15FCCEFB11" box="[723,781,1231,1257]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="FCC4CC93FFD0AC1DFC6FFB15FBF6FB11" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[976,1077,1231,1257]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFD0AC1DFC6FFB15FBF6FB11" box="[976,1077,1231,1257]" italics="true" pageId="6" pageNumber="349">T. reebae</emphasis>
</taxonomicName>
, but differs in the proportionally longer median lobe of the aedeagus (EL/AL &lt;
<quantity id="FC3C1AF5FFD0AC1DFD09FB29FD3AFAF5" box="[694,761,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="6" pageNumber="349" unit="in" value="1.9">1.9 in</quantity>
<taxonomicName id="FCC4CC93FFD0AC1DFCBDFB29FC47FAF5" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[770,900,1267,1293]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFCBDFB29FC47FAF5" box="[770,900,1267,1293]" italics="true" pageId="6" pageNumber="349">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFC32FB29FC0FFAF5" box="[909,972,1267,1293]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
,&gt; 2.0 in =
<taxonomicName id="FCC4CC93FFD0AC1DFBE4FB29FB07FAF5" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[1115,1220,1267,1293]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFD0AC1DFBE4FB29FB07FAF5" box="[1115,1220,1267,1293]" italics="true" pageId="6" pageNumber="349">T. reebae</emphasis>
</taxonomicName>
) and in the longer apical lamella (
<figureCitation id="A3FFAB95FFD0AC1DFEF8FACDFE5DFAC9" box="[327,414,1303,1329]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3F</figureCitation>
). The new species differs from the likewise similar
<taxonomicName id="FCC4CC93FFD0AC1DFC4DFACDFB85FAC8" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1010,1094,1303,1328]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFC4DFACDFB85FAC8" box="[1010,1094,1303,1328]" italics="true" pageId="6" pageNumber="349">T. igori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFBF0FACDFB4EFAC9" box="[1103,1165,1303,1329]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="FCC4CC93FFD0AC1DFB78FACDFA8CFAC9" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1223,1359,1303,1329]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFB78FACDFA8CFAC9" box="[1223,1359,1303,1329]" italics="true" pageId="6" pageNumber="349">T. kniphofia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFAE8FACDFA55FAC9" box="[1367,1430,1303,1329]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
, prominently in the more markedly dilated protarsomeres 1 and 2 (
<figureCitation id="A3FFAB95FFD0AC1DFCCFFAE1FC05FAAD" box="[880,966,1339,1365]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="6" pageNumber="349">Fig. 3C</figureCitation>
), and additionally in the more robust body which is, on an average larger, and in the larger aedeagus with longer apical lamella which is more markedly hook-like bent upwardly.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD0AC1DFF78FA7CFF28F9A9" pageId="6" pageNumber="349" type="distribution">
<paragraph id="3B7BB710FFD0AC1DFF78FA7CFEB1F9F1" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
<emphasis id="09B06B02FFD0AC1DFF78FA7CFE9AFA38" bold="true" box="[199,345,1446,1472]" pageId="6" pageNumber="349">Distribution</emphasis>
. Endemic to Mt. Choke in northern
<collectingCountry id="43D3F780FFD0AC1DFD46FA7DFC9FFA39" box="[761,860,1447,1473]" name="Ethiopia" pageId="6" pageNumber="349">Ethiopia</collectingCountry>
. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west were it occurs sympatric with
<taxonomicName id="FCC4CC93FFD0AC1DFB27FA11FACAFA1C" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1176,1289,1483,1508]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFB27FA11FACAFA1C" box="[1176,1289,1483,1508]" italics="true" pageId="6" pageNumber="349">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFAADFA11FA8CFA1D" box="[1298,1359,1483,1509]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="FCC4CC93FFD0AC1DFA37FA11FF0EF9F0" authorityName="Schmidt &amp; Merene" authorityYear="2024" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFA37FA11FF0EF9F0" italics="true" pageId="6" pageNumber="349">T. igori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFF6AFA35FED2F9F1" box="[213,273,1519,1545]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="A3FFAB95FFD0AC1DFE9FFA35FEA6F9F1" box="[288,357,1519,1545]" captionStart="FIGURE 4" captionStartId="9.[152,255,1559,1583]" captionTargetBox="[151,1436,561,1535]" captionTargetId="figure-219@9.[151,1436,561,1535]" captionTargetPageId="9" captionText="FIGURE 4. Topographic map of Mt. Choke (highest point marked by a white triangle), showing sampling localities of six Trechus (Abyssinotus) species of the niloticus group (white circles; the numbers refer to the different species). 1, T. apertus Schmidt, sp. n. 2, T. reebae (Quéinnec &amp; Ollivier). 3, T. yitbareki Schmidt, sp. n. 4, T. kniphofia Schmidt, sp. n. 5, T. igori Schmidt, sp. n. 6, T. niloticus (Quéinnec &amp; Ollivier). Sampling data of T. reebae and T. niloticus published by Quéinnec et al. (2021) are considered. Base map was downloaded from Topographic-map.com." figureDoi="http://doi.org/10.5281/zenodo.13234861" httpUri="https://zenodo.org/record/13234861/files/figure.png" pageId="6" pageNumber="349">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="3B7BB710FFD0AC1DFF78F9C8FF28F9A9" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="349">
<emphasis id="09B06B02FFD0AC1DFF78F9C8FE77F9D4" bold="true" box="[199,436,1554,1580]" pageId="6" pageNumber="349">Habitat preferences</emphasis>
. The new species was found syntopic with
<taxonomicName id="FCC4CC93FFD0AC1DFC1BF9C9FBD6F9D4" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[932,1045,1555,1580]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD0AC1DFC1BF9C9FBD6F9D4" box="[932,1045,1555,1580]" italics="true" pageId="6" pageNumber="349">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD0AC1DFBA2F9C9FB9FF9D5" box="[1053,1116,1555,1581]" pageId="6" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
; see above for details of the habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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