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<document ID-CLB-Dataset="23646" ID-DOI="http://dx.doi.org/10.3897/zookeys.917.47326" ID-GBIF-Dataset="292ff02a-f10f-416e-be2e-90c8624f83fe" ID-PMC="PMC7076059" ID-Pensoft-Pub="1313-2970-917-85" ID-Pensoft-UUID="AE848A9099625A9B9D72ED28EB542ED0" ID-PubMed="32206019" ID-ZooBank="DEDB8D24052545B18F3E9A97EF23DA37" ModsDocID="1313-2970-917-85" checkinTime="1583773828879" checkinUser="pensoft" docAuthor="Chang, Zhi-Min, Yang, Lin &amp; Chen, Xiang-Sheng" docDate="2020" docId="B32F475B77145B73A8040D59020FC0C4" docLanguage="en" docName="ZooKeys 917: 85-104" docOrigin="ZooKeys 917" docSource="http://dx.doi.org/10.3897/zookeys.917.47326" docTitle="Kodaianella furcata Chang &amp; Chen 2020, sp. nov." docType="treatment" docUuid="EB9B5313-A0A2-4F2B-821F-FD5E8B5A342E" docUuidSource="ZooBank" docVersion="6" id="AE848A9099625A9B9D72ED28EB542ED0" lastPageNumber="85" masterDocId="AE848A9099625A9B9D72ED28EB542ED0" masterDocTitle="First records of the genera Sivaloka Distant, 1906, with two new species from China, and description of a new species of genus Kodaianella Fennah, 1956 (Hemiptera, Fulgoromorpha, Issidae)" masterLastPageNumber="104" masterPageNumber="85" pageNumber="85" updateTime="1732751919278" updateUser="ExternalLinkService">
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<mods:title id="DEBD0887BA8A4DAF4B1FA76CFC57E81E">First records of the genera Sivaloka Distant, 1906, with two new species from China, and description of a new species of genus Kodaianella Fennah, 1956 (Hemiptera, Fulgoromorpha, Issidae)</mods:title>
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<mods:namePart id="9FD1777AB87C9B9892F59B03EF6D9DAD">Chang, Zhi-Min</mods:namePart>
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<mods:namePart id="BEB1CF272C847D2B8AB7CB501CCB0EA5">Yang, Lin</mods:namePart>
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<mods:namePart id="A35E6D0E17676FE77EB34841D5740EEB">Chen, Xiang-Sheng</mods:namePart>
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<treatment id="B32F475B77145B73A8040D59020FC0C4" ID-GBIF-Taxon="162467315" LSID="urn:lsid:zoobank.org:act:EB9B5313-A0A2-4F2B-821F-FD5E8B5A342E" httpUri="http://treatment.plazi.org/id/B32F475B77145B73A8040D59020FC0C4" lastPageNumber="85" pageId="0" pageNumber="85" scope_family="Issidae" scope_order="Hemiptera" scope_subFamily="Hemisphaeriinae">
<subSubSection id="F4F11100A5870BC8E771C036C4DA9B97" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="DD0874A8802FCC046127656BFE9143FB" pageId="0" pageNumber="85">
<taxonomicName id="4BC3A59E010D7744C34D7B7E4767FF9B" ID-CoL="3RBCX" LSID="B32F475B-7714-5B73-A804-0D59020FC0C4" authority="Chang &amp; Chen" authorityName="Chang &amp; Chen" authorityYear="2020" class="Insecta" family="Issidae" genus="Kodaianella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kodaianella furcata" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="furcata" status="sp. nov.">Kodaianella furcata Chang &amp; Chen</taxonomicName>
<taxonomicNameLabel id="4704B51574C7B8456B6708A94F4F12A3" pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
<figureCitation id="6ACC703C99417A69074DCA9C886A0AA9" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">Figs 1-13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="580B662122C5B4B0268817BA273C9C6C" pageId="0" pageNumber="85" type="type material">
<paragraph id="C501CDDEAC4982273D3C104FFDB9A90F" pageId="0" pageNumber="85">Type material.</paragraph>
<paragraph id="B432B785B559BD0CFDA8A99770132E1B" pageId="0" pageNumber="85">
Holotype: ♂, China: Guangxi, Nonggang National Nature Reserve (
<geoCoordinate id="FDC5019EED4B77926CF4E4A9B8DA0F14" degrees="22" direction="north" minutes="28" orientation="latitude" precision="925" value="22.466667">22°28'N</geoCoordinate>
,
<geoCoordinate id="8A7A74F9491275BE3D1A6CDE8998F3A4" degrees="106" direction="east" minutes="58" orientation="longitude" precision="925" value="106.96667">106°58'E</geoCoordinate>
), 8 May 2011, H Li leg.; paratypes: 5♂♂, same data as holotype (IEGU); 1♂, Guangxi, Nonggang National Nature Reserve (
<geoCoordinate id="D1E40444995AC4C662CBEE4D9D449C59" degrees="22" direction="north" minutes="28" orientation="latitude" precision="925" value="22.466667">22°28'N</geoCoordinate>
,
<geoCoordinate id="5206EEC09F70E329908F1CB828D05ECD" degrees="106" direction="east" minutes="58" orientation="longitude" precision="925" value="106.96667">106°58'E</geoCoordinate>
), 7-8 May 2012, H Li and N-N Yang, leg.
</paragraph>
</subSubSection>
<subSubSection id="59F0EC2C79B3E0DE0B0F428B3B024FBE" pageId="0" pageNumber="85" type="diagnosis">
<paragraph id="2B6B7D156D1A64529E9D1311C4C8B3AE" pageId="0" pageNumber="85">Diagnosis.</paragraph>
<paragraph id="23F511C16784C26B216FBED55C74C1F6" pageId="0" pageNumber="85">
This species is similar to
<taxonomicName id="742FA34AAFC2B6641B24F39EADC956DC" lsidName="K. longispina" pageId="0" pageNumber="85" rank="species" species="longispina">
<emphasis id="4AE567A2E7D6056AF0E4D7835947243E" italics="true" pageId="0" pageNumber="85">K. longispina</emphasis>
</taxonomicName>
Zhang &amp; Chen, 2010 in appearance, but it differs from the latter in having the phallobase with dorsal lobe bearing one rod-like process near its apical part in lateral view (Fig.
<figureCitation id="FAFA9E86C5603EDB06222EC66877FAA0" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12a</figureCitation>
); the phallobase with ventral lobe distinct short, apical part finger-liked in ventral view (Fig.
<figureCitation id="AC90CF5AB16341601AC47422208C265C" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13c</figureCitation>
); and the aedeagus in lateral view, with a forked and hooked process near the apical 1/3 (Fig.
<figureCitation id="9EBB34DDBC37761001D3DE9B4F8A023C" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F0ED383E9666EBA013B8B50DAF979C09" pageId="0" pageNumber="85" type="description">
<paragraph id="1EDD3E7ABE1DC40023F6A3CAC8A79768" pageId="0" pageNumber="85">Description.</paragraph>
<paragraph id="96F9545C15EC45EAB85DCB1BE562BDAC" pageId="0" pageNumber="85">Body length: male 3.94-4.18 mm; forewing: male 3.16-3.45 mm.</paragraph>
</subSubSection>
<subSubSection id="26D6805A10B5C265789D16A5EAB046FE" pageId="0" pageNumber="85" type="coloration">
<paragraph id="A3AF8F635782669543EBB1D45727D95F" pageId="0" pageNumber="85">Coloration.</paragraph>
<paragraph id="1E5901F4F917FC8297C701DF4EF7884A" pageId="0" pageNumber="85">
General color brown (Figs
<figureCitation id="C33D570786808FB1980369F2E00FF5F4" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">1</figureCitation>
,
<figureCitation id="403652A4FAF6E4BA159C4292998D156A" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">2</figureCitation>
). Vertex, pronotum, and mesonotum (Fig.
<figureCitation id="2B700637F0C9B5356714D95028BDF310" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">3</figureCitation>
) black-brown. Frons (Fig.
<figureCitation id="6F9450E6703EAB9CC2430253ED3F574F" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">5</figureCitation>
) black-brown, with pale yellow verrucae along base and lateral margins. Clypeus (Fig.
<figureCitation id="32F4469498802D160CE2C15C5D060D58" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">5</figureCitation>
) yellow-brown. Compound eyes black-brown, ocelli pale green (Fig.
<figureCitation id="08766C4D7E20193CC647D442A0501664" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">4</figureCitation>
). Forewings (Fig.
<figureCitation id="7A6C0133DDB2CA3BC20C750C04DB17CB" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">2</figureCitation>
) yellow-brown, with dark spots. Legs yellow-brown, with tips of spines on hind tibiae and tarsi black.
</paragraph>
</subSubSection>
<subSubSection id="28A39B47AC96CA04E8967E24C6D9E0D7" pageId="0" pageNumber="85" type="head and thorax">
<paragraph id="C457379524F2E35817BD37553575FB3D" pageId="0" pageNumber="85">Head and thorax.</paragraph>
<paragraph id="2DCF7D9AA55E6FE08B0F767D67EFD788" pageId="0" pageNumber="85">
Head (Fig.
<figureCitation id="D8CF5DF39FEB06AEE85F6B2E8FA14CF9" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">3</figureCitation>
) including eyes, slightly narrower than pronotum (0.96: 1.00). Vertex (Fig.
<figureCitation id="662A64AEDF60F8CA2AA660AF726BD29A" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">3</figureCitation>
) shorter in middle than the width at base (0.43: 1.00), with median carina linear; anterior margin slightly convexly arched; posterior margin obviously obtusely concave. Frons (Fig.
<figureCitation id="121A0FC39CE75BAD2A69FE0CC0530D86" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">5</figureCitation>
) shorter in middle than the widest breadth (0.60: 1.00); median carina obvious and straight, reaching to 2/3 level of frons. Clypeus (Fig.
<figureCitation id="30343A963B8FBBC7701C1022C6710EAB" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">5</figureCitation>
) triangular, without median carina. Pronotum (Fig.
<figureCitation id="6F0D37850246F37D5245B5EB785CE08A" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">3</figureCitation>
) with median carina obscure, lateral carina reaching to the posterior margin. Mesonotum (Fig.
<figureCitation id="06F180E0D8400D391E4CDB65E8ADD3AB" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">3</figureCitation>
) with median carina obscure. Forewings (Fig.
<figureCitation id="484602D4DAED896930424BD810644EDB" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">6</figureCitation>
) 1.60 times as long as maximum breadth; with wide &quot;hypocostal plate&quot;; ScP and RP convergent near base, ScP and RP veins long, nearly reaching apical margin; MP bifurcating two branches near basal 1/3, MP1 forked near apical 1/3; CuA forked into two branches near middle; CuP present, Pcu and A1 united near middle of clavus. Hindwings (Fig.
<figureCitation id="376AE0259FECDBA06B08485F3437C65E" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">7</figureCitation>
) with ScP+R and M simple, not forked, CuA forked near apical part, with one vein between R and M, M and CuA1; and Pcu and A11 jointed near apical 1/4, without short transverse vein between Pcu + A11 and A12; A2 simple, reaching to 2/3 of A2 lobe. Spinal formula of hind leg (2)8/8/2.
</paragraph>
</subSubSection>
<subSubSection id="05393BBA18A57429779F7BFA4E0CE3AE" pageId="0" pageNumber="85" type="male genitalia">
<paragraph id="3C820161586E35BA8F8F5A7CA03AD57A" pageId="0" pageNumber="85">Male genitalia.</paragraph>
<paragraph id="6E11E04AF172C91E5BA97429FF4935E9" pageId="0" pageNumber="85">
Anal tube (Fig.
<figureCitation id="FE23AA1137B2DB089D950BDF6E46F36F" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">9</figureCitation>
) longer in midline than the width (2.49: 1.00) in dorsal view; lateral margins nearly parallel and widest in apical part; apical margin nearly truncated, with unobvious small, angular process near lateral margin. Anal style (Fig.
<figureCitation id="776311797CDDF260EFED4A0160F22E9E" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">9</figureCitation>
) stout and long, located at the base 2/5 of anal tube, surpassing the opening of anal pore. Pygofer (Fig.
<figureCitation id="70BF540B42D937B7E5DDBA9E56570DD2" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">8</figureCitation>
) irregularly rectangular; dorsal margin slightly broader than ventral margin; anterior margin arched near dorsal 1/3; posterior margin nearly straight. Genital styles (Figs
<figureCitation id="A4BBB245763138C0387D617B85841DA4" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">8</figureCitation>
,
<figureCitation id="521BBA24FCCF6CEE7D6A92FA0592F36B" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">10</figureCitation>
) relatively triangular; anterior margin without triangular process; posterior margin with triangular process. Capitulum of genital styles irregularly triangular, with small irregular triangular, relatively long and stout neck (Fig.
<figureCitation id="5EF2501248AB02A2174AED0C31176D59" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">11</figureCitation>
). Phallobase (Figs
<figureCitation id="007F12982957467DAE443FA104B819E3" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12</figureCitation>
,
<figureCitation id="DB662D58CA9EC8C48394F047DF870686" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13</figureCitation>
) with dorsal lobe cystiform near apical part, with stout rod-like process (Figs
<figureCitation id="A92BF032A9893EF460CD68EE5B477420" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12a</figureCitation>
,
<figureCitation id="70FA878E4C9101E3B8676DFEE9D6B568" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13a</figureCitation>
) in apical 1/6, directed to posterior, with dorso-lateral lobe with short hook-like process (Fig.
<figureCitation id="4B28033BB6C9F31ACAAFCC348453CA3C" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12b</figureCitation>
), pointed to dorsal margin in lateral view; lateral lobe splitting into two stout branches; ventral lobe membranous, apical part narrow, surface with microvilli in lateral view (Fig.
<figureCitation id="77F17159E14A86454737AAF4FFBB20D8" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12c</figureCitation>
); ventral lobe in ventral view obviously shorter than dorsal lobe, with apical part projecting into finger-like process in middle (Fig.
<figureCitation id="4B69332F012AA5CFF05BF4098A144422" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13c</figureCitation>
). Aedeagus (Figs
<figureCitation id="119BDA0203CC2C2B3B83AE026A6924A0" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12</figureCitation>
,
<figureCitation id="9746CA30764CB546C61EDA945D2417D2" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13</figureCitation>
) with long, hooked process near apical 1/3 in ventral view, tip of process directed to ventro-posterior in lateral view (Fig.
<figureCitation id="74D29B3100539E23BC9966CCCD8B9FED" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">12d</figureCitation>
); in ventral view, hooked process forked into asymmetrical hooks (Fig.
<figureCitation id="9241AB43B5A922AFFDC6FEE2BC748C47" captionStart="Figures 113" captionStartId="F1" captionText="Figures 1 - 13. Kodaianella furcata Chang &amp; Chen, sp. nov. 1 habitus, dorsal view 2 habitus, lateral view 3 head and thorax, dorsal view 4 head and thorax, lateral view 5 head, ventral view 6 forewing 7 hindwing 8 male genitalia, lateral view 9 male anal segment, dorsal view 10 genital styles, lateral view 11 capitulum of genital styles, ventral view 12 phallobase and aedeagus, lateral view 13 phallobase and aedeagus, ventral view. Scale bars: 0.5 mm. Abbreviations: a = rod-like process; b = hook-like process; c = ventral lobe; d = long forked process." figureDoi="10.3897/zookeys.917.47326.figures1-13" httpUri="https://binary.pensoft.net/fig/387722" pageId="0" pageNumber="85">13d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9C55A5E3C537AC3DDA6523FB5B7008E5" pageId="0" pageNumber="85" type="etymology">
<paragraph id="4C2ACC4D775C8D3FCDE0B63C42F7C1FF" pageId="0" pageNumber="85">Etymology.</paragraph>
<paragraph id="73EDA7B100CC9928725641ECB16E5668" pageId="0" pageNumber="85">
The new species is derived from the Latin word &quot;
<taxonomicName id="E16AC153B135243359593AFAD17A10EA" lsidName="furcata" pageId="0" pageNumber="85" rank="species" species="furcata">
<emphasis id="BC8D6CD76A54419A3F89C2856FF9F22E" italics="true" pageId="0" pageNumber="85">furcata</emphasis>
</taxonomicName>
&quot;, in reference to the aedeagus, which bears a forked, hooked process.
</paragraph>
</subSubSection>
<subSubSection id="467527658B25512A37E68236EDF19B7D" pageId="0" pageNumber="85" type="host">
<paragraph id="D7AF14AA537950D371A7F1AA259523BF" pageId="0" pageNumber="85">Host plant.</paragraph>
<paragraph id="5831C677DC9E2F23C525D1B0FC9505F4" pageId="0" pageNumber="85">Unknown.</paragraph>
</subSubSection>
<subSubSection id="DDE37EC5F5FEA4833D833AB687AD5C9A" pageId="0" pageNumber="85" type="distribution">
<paragraph id="69BC4360D0991F274E1ED54245C2F933" pageId="0" pageNumber="85">Distribution.</paragraph>
<paragraph id="3DAED4E3F31471395B6444E3FEE54BAF" pageId="0" pageNumber="85">China (Guangxi).</paragraph>
</subSubSection>
</treatment>
</document>