195 lines
18 KiB
XML
195 lines
18 KiB
XML
<document id="9A440E2361192D546B46B982D7156A6C" ID-CLB-Dataset="28198" ID-DOI="http://dx.doi.org/10.3897/mycokeys.40.26025" ID-GBIF-Dataset="06667daa-16c0-4b90-9c75-9517f8fc2777" ID-PMC="PMC6170524" ID-Pensoft-Pub="1314-4049--69" ID-PubMed="30294209" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-4049--69" ModsDocOrigin="MycoKeys " ModsDocTitle="Three new species of Krogia (Ramalinaceae, lichenised Ascomycota) from the Paleotropics" checkinTime="1553125612105" checkinUser="pensoft" docAuthor="Kistenich, Sonja, Rikkinen, Jouko K., Thues, Holger, Vairappan, Charles S., Wolseley, Patricia A. & Timdal, Einar" docDate="2018" docId="B287686381EA7EBE1377636F50AF6A12" docLanguage="en" docName="MycoKeys 40: 69-88" docOrigin="MycoKeys 40" docSource="http://dx.doi.org/10.3897/mycokeys.40.26025" docTitle="Krogia borneensis Kistenich & Timdal, sp. nov." docType="treatment" docVersion="6" lastPageNumber="75" masterDocId="FF9E9222FFBF81525B1BFF8CFFF3FF97" masterDocTitle="Three new species of Krogia (Ramalinaceae, lichenised Ascomycota) from the Paleotropics" masterLastPageNumber="88" masterPageNumber="69" pageNumber="74" updateTime="1732831433047" updateUser="ExternalLinkService">
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<mods:title id="FA09CBBDB43484BD824B3CF96BDD2D0E">Three new species of Krogia (Ramalinaceae, lichenised Ascomycota) from the Paleotropics</mods:title>
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<mods:namePart id="64B71F387B4D2B1A10462CCAA93C2D54">Kistenich, Sonja</mods:namePart>
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<mods:namePart id="85DFE64296ACC9EC03AAAD761AEBF729">Rikkinen, Jouko K.</mods:namePart>
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<mods:namePart id="7C98F6AAE11A3D780D928E905C67704F">Thues, Holger</mods:namePart>
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<mods:namePart id="F104C25760D8F168069CBE77E676A9F3">Vairappan, Charles S.</mods:namePart>
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<mods:namePart id="F983D9973D25351EB53E67439AD2E0A2">Wolseley, Patricia A.</mods:namePart>
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<mods:namePart id="80E45A659CC0C5132514C8ACC8C30140">Timdal, Einar</mods:namePart>
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<mods:typeOfResource id="F69720478FD94205F996DA4801ABF039">text</mods:typeOfResource>
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<mods:title id="AD9ACCB4ED6A09310F354EF72AC8330C">MycoKeys</mods:title>
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<mods:date id="DB76218028B36D253DBA14ACE7CF40BD">2018</mods:date>
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<mods:detail id="1D0C3EF935FFCF734B074455FA0A5649" type="volume">
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<mods:number id="C9EAEBBC14668829D38466E05D32E68E">40</mods:number>
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<mods:start id="50BA7DDAEAA517A1C6F1142329B61C6B">69</mods:start>
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<mods:url id="6B96C6276609A867B852403047997E8F">http://dx.doi.org/10.3897/mycokeys.40.26025</mods:url>
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<mods:classification id="8E10E9E9E6E7800177A21859DBB870A4">journal article</mods:classification>
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<mods:identifier id="B38419E88D383610355DB2CD3C590248" type="DOI">http://dx.doi.org/10.3897/mycokeys.40.26025</mods:identifier>
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<mods:identifier id="A5B74FD451FF127D7B21233A247D0047" type="Pensoft-Pub">1314-4049--69</mods:identifier>
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<treatment id="B287686381EA7EBE1377636F50AF6A12" ID-GBIF-Taxon="154473967" LSID="urn:lsid:plazi:treatment:B287686381EA7EBE1377636F50AF6A12" httpUri="http://treatment.plazi.org/id/B287686381EA7EBE1377636F50AF6A12" lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74" scope_family="Ramalinaceae" scope_phylum="Ascomycota">
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<subSubSection id="5A8D93FCFE18E9BD0634B00ADFF62A9B" pageId="5" pageNumber="74" type="nomenclature">
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<paragraph id="F33053AAAD7108BC33139897F5DAB9F7" pageId="5" pageNumber="74">
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<taxonomicName id="9E9350F6FCA48BD4D7218D914D237A03" ID-CoL="6NNQ6" LSID="MB825078" authority="Kistenich & Timdal" class="Lecanoromycetes" family="Ramalinaceae" genus="Krogia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Krogia borneensis" order="Lecanorales" pageId="5" pageNumber="74" phylum="Ascomycota" rank="species" species="borneensis">Krogia borneensis Kistenich & Timdal</taxonomicName>
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<taxonomicNameLabel id="62C7A9682F2D878E233ADAB5ABD1FA7C" pageId="5" pageNumber="74">sp. nov.</taxonomicNameLabel>
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Fig. 2
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</paragraph>
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</subSubSection>
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<subSubSection id="AC5485C34E664F8AAE12C9A491670F2E" pageId="5" pageNumber="74" type="diagnosis">
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<paragraph id="1AFC180261CE6E9968101629BB1CB6C5" pageId="5" pageNumber="74">Diagnosis.</paragraph>
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<paragraph id="59A3E3C768489D4EE892736690B6CFA0" pageId="5" pageNumber="74">
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The species differs from
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<taxonomicName id="1C5A73A4DD50B84AEFFB43B38F49F56B" lsidName="K. isidiata" pageId="5" pageNumber="74" rank="species" species="isidiata">K. isidiata</taxonomicName>
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in forming lacinules as vegetative dispersal units, not isidia, and from the other species in the genus in producing sekikaic and homosekikaic acid.
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</paragraph>
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</subSubSection>
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<subSubSection id="A455AF8AE7573D0A346483A009F202F9" pageId="5" pageNumber="74" type="materials_examined">
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<paragraph id="02736867245BC608493021EA29F435B0" pageId="5" pageNumber="74">Type.</paragraph>
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<paragraph id="472585C3904CF18A6C0FCFADEAAE517F" pageId="5" pageNumber="74">
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Malaysia, Borneo, Sabah, Maliau conservation area, trail between Nepenthes Camp and waterfall Takob Akob,
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<geoCoordinate id="91C9083EA930B821630096875B775988" direction="north" orientation="latitude" precision="92" value="4.7233334">4°43.4'N</geoCoordinate>
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,
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<geoCoordinate id="15D10DEFF3F346A7D2DC40B374D4B3F2" direction="east" orientation="longitude" precision="92" value="116.87">116°52.2'E</geoCoordinate>
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, 900-1000 m alt., in low (few metres) and open pristine montane "Kerangas" (heath) forest with higher trees mostly along a small stream, on smooth barked tree in the vicinity of the stream, 2013-02-23, C. Vairappan & H.
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<normalizedToken id="0FE342C4758DF19BB5DC6CB546F1EC36" originalValue="Thüs">Thues</normalizedToken>
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L291 (BORH, holotype) [TLC: sekikaic and homosekikaic acid; GenBank: MH174275 (mtSSU)].
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</paragraph>
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</subSubSection>
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<subSubSection id="61A45A9A07E6C8573F83B7C0002C758F" pageId="5" pageNumber="74" type="description">
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<paragraph id="7469C91FB105DBE3E41B290230861B33" pageId="5" pageNumber="74">Description.</paragraph>
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<paragraph id="85B612C7D8C161517BE5E76389DB6A01" pageId="5" pageNumber="74">
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Thallus effuse, squamulose; squamules up to 1 mm wide, deeply divided into 0.1-0.2 wide lobes, ascending, imbricate, flattened, elongated to partly linear, often slightly laterally constricted, greyish-green with patches of red (K+ purple) spots, epruinose, glabrous; margin concolorous with upper side, not fibrillose; lower side white; lacinules formed by tips of the lobes. Upper cortex composed of a single layer of thick-walled cells with angular to shortly cylindrical lumina (resembling
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<taxonomicName id="A48B1C7BA764E7677EB4817E834CCADE" class="Lecanoromycetes" family="Ramalinaceae" genus="Eschatogonia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Eschatogonia" order="Lecanorales" pageId="5" pageNumber="74" phylum="Ascomycota" rank="genus">Eschatogonia</taxonomicName>
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-type), not containing crystals (polarised light!); algal layer 30-40
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<normalizedToken id="F1718677E5F197350E684965F77D85D0" originalValue="µm">µm</normalizedToken>
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thick, filled with crystals dissolving in K; medulla composed of loosely interwoven hyphae, not containing crystals dissolving in K; lower cortex resembling upper cortex, both continuing over the edge of the squamule; prothallus brownish-black, often well developed. Apothecia (present in the holotype only) up to 0.6 mm diam. when simple, forming aggregates up to 1.5 mm diam., medium brown with red patches or entirely reddish-brown, more or less plane, with an indistinct, slightly paler, often flexuose margin; excipulum pale brown to colourless, composed of radiating, closely conglutinated hyphae, in inner part containing colourless crystals dissolving in K; hypothecium partly to entirely stained by a blood red pigment which dissolves in K with a purple effusion; epithecium colourless, not containing crystals. Ascospores filiform, curved, non-septate, spirally arranged in ascus, 20-31
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<normalizedToken id="13CE9A9243AD61E79BF8CC622289B537" originalValue="×">x</normalizedToken>
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ca. 1.0
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<normalizedToken id="5C33B57005EC4FC895835391161CE47A" originalValue="µm">µm</normalizedToken>
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(n=10, from holotype). Conidiomata not seen.
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</paragraph>
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</subSubSection>
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<subSubSection id="CA5DBE496422A977C6B111D57A396307" pageId="5" pageNumber="74" type="chemistry">
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<paragraph id="5FF6A61A85E269E8D3F8C2B247EE8553" pageId="5" pageNumber="74">Chemistry.</paragraph>
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<paragraph id="C19C1E7EBA5E5BD0FDD645D0480A5703" pageId="5" pageNumber="74">Sekikaic acid (major), homosekikaic acid (major). Spot tests: all negative, except for red patches being K+ purple.</paragraph>
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</subSubSection>
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<subSubSection id="CCD32F863486967FB2B6951703389A2F" pageId="5" pageNumber="74" type="distribution">
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<paragraph id="607974AAD1E1141C7AD81FB2280CC0A1" pageId="5" pageNumber="74">Distribution.</paragraph>
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<paragraph id="1A545BA8B21A3B4BECAD65D5BBF885F8" pageId="5" pageNumber="74">The species is known from five localities in Borneo.</paragraph>
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<paragraph id="D1731E5C049A5F03A47B39E3B0D59276" pageId="5" pageNumber="74">
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Ecology. The species occurred in rather low "Kerangas" (heath) forest vegetation or on transition vegetation between the heath and oak/conifer (particularly
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<taxonomicName id="AA611C89B93B29A2BC61EC182BFEBB90" class="Insecta" family="Braconidae" genus="Agathis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agathis" order="Hymenoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Agathis</taxonomicName>
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) forest at higher elevations (ca. 1000 m) on very poor soils on sandstone (Fig. 2B). The species always grew on the rather smooth barked, middle-sized trees together with various
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<taxonomicName id="AD582A36CC41EAC3646A780E79E0566B" class="Eurotiomycetes" family="Pyrenulaceae" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Pyrenulales" pageId="5" pageNumber="74" phylum="Ascomycota" rank="family">Pyrenulaceae</taxonomicName>
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and
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<taxonomicName id="9066C9A6EF37CB624F0CED13782B4D5D" class="Lecanoromycetes" family="Graphidaceae" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Ostropales" pageId="5" pageNumber="74" phylum="Ascomycota" rank="family">Graphidaceae</taxonomicName>
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.
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</paragraph>
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<caption id="A0E350A6E6D9742C02D518DF09A3D260" pageId="5" pageNumber="74">
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<paragraph id="AED93550D7A6B3E813FFE429E76F467B" pageId="5" pageNumber="74">
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Figure 2.
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<taxonomicName id="64A9E6C47C8C8F586C34E7A72A7D5F5E" class="Lecanoromycetes" family="Ramalinaceae" genus="Krogia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Krogia borneensis" order="Lecanorales" pageId="5" pageNumber="74" phylum="Ascomycota" rank="species" species="borneensis">Krogia borneensis</taxonomicName>
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. A Field photograph of the holotype B habitat at type locality C herbarium photograph of holotype. Scale bar: 1 mm. Photo: H.
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<normalizedToken id="19EB1C2E489648CAC1A45B97A9CE9D07" originalValue="Thüs">Thues</normalizedToken>
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(A, B), E. Timdal (C).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="16C5A60653D70E193C20BFFC52AC01B8" pageId="5" pageNumber="74" type="etymology">
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<paragraph id="3D2727C579DB098E1B7CFB9244A8E86E" pageId="5" pageNumber="74">Etymology.</paragraph>
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<paragraph id="7110561FC2C5EB8881161F67086C4C58" pageId="5" pageNumber="74">The specific epithet refers to its occurrence in Borneo.</paragraph>
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</subSubSection>
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<subSubSection id="795158BC35564C8174179D290E982928" pageId="6" pageNumber="75" type="remarks">
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<paragraph id="BCAF0257EC8B6A92B0CB214EEB50241B" pageId="6" pageNumber="75">
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<pageBreakToken id="5B5AD1C8E1926FBC8DE40DF99E97CFC2" pageId="6" pageNumber="75" start="start">Remarks</pageBreakToken>
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.
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</paragraph>
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<paragraph id="CF3587FC55CB286BEF0AB54CEB3FB27F" pageId="6" pageNumber="75">
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The medium-sized, flattened squamules make the species morphologically most similar to the neotropical
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<taxonomicName id="6FF477EC09E59F2E41C3183A896C24B5" lsidName="K. antillarum" pageId="6" pageNumber="75" rank="species" species="antillarum">K. antillarum</taxonomicName>
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. The squamules are more elongated, often linear and with more lateral constrictions in
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<taxonomicName id="E143C7E3293C112AF2BD05841B589FB7" lsidName="K. borneensis" pageId="6" pageNumber="75" rank="species" species="borneensis">K. borneensis</taxonomicName>
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than in
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<taxonomicName id="EB9B5AF19E6D86CAE2851D38EBF0FF3E" lsidName="K. antillarum" pageId="6" pageNumber="75" rank="species" species="antillarum">K. antillarum</taxonomicName>
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, which has more fan-shaped squamules. The former species has a thin, unicellular cortex on both upper and lower side, whereas the latter has a multicellular (20-30
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<normalizedToken id="8BAFC3A44DF203F0E600A344E333D25F" originalValue="µm">µm</normalizedToken>
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thick) upper cortex and lacks a lower cortex (
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<bibRefCitation id="51FDAE9D464C214E775E84A7B49F1857" author="Timdal, E" journalOrPublisher="Bryologist" pageId="15" pageNumber="84" pagination="387 - 389" title="Krogiaantillarum (Ramalinaceae), a new lichen species from the West Indies." volume="112" year="2009">Timdal 2009</bibRefCitation>
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). Chemically, the latter species differs in forming 4-O-methylcryptochlorophaeic acid.
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</paragraph>
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<paragraph id="0F3911BABAAF76C8B2E8DE880F779AC2" pageId="6" pageNumber="75">
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<taxonomicName id="4CCD576CD9B6A84BDB4D0735A8AF9CE8" class="Lecanoromycetes" family="Ramalinaceae" genus="Krogia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Krogia isidiata" order="Lecanorales" pageId="6" pageNumber="75" phylum="Ascomycota" rank="species" species="isidiata">Krogia isidiata</taxonomicName>
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shares the secondary chemistry (sekikaic and homosekikaic acid) with
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<taxonomicName id="9229BB4C6E12D3752940D1C3C17D0DC6" lsidName="K. borneensis" pageId="6" pageNumber="75" rank="species" species="borneensis">K. borneensis</taxonomicName>
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, but they differ in their vegetative dispersal units, the former producing cylindrical isidia, the latter flat lacinules (fragmenting squamules). The upper cortex of
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<taxonomicName id="12004A21F82B507FAA74B0AF8287610D" lsidName="K. isidiata" pageId="6" pageNumber="75" rank="species" species="isidiata">K. isidiata</taxonomicName>
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is multicellular (15-30
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<normalizedToken id="468D5DBCF64669B32AAAF5FD1CCAC633" originalValue="µm">µm</normalizedToken>
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thick) and the lower cortex is absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="98CC4AD4A518860138FEB7345A515AED" pageId="6" pageNumber="75" type="materials_examined">
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<paragraph id="36DBD46C9BD89F4652A3F6041F841134" pageId="6" pageNumber="75">Additional specimens examined.</paragraph>
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<paragraph id="53518F9EF84BBFA187518E2B018970C2" pageId="6" pageNumber="75">
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Malaysia, Borneo. Sabah: Danum, plot 88, dipterocarp forest logged in 1988,
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<geoCoordinate id="6F8B852EEC6AC344858D627C97529627" direction="north" orientation="latitude" precision="925" value="4.9666667">4°58'N</geoCoordinate>
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,
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<geoCoordinate id="BE7EBC1A02D9BF2787241073FE92BABC" direction="east" orientation="longitude" precision="925" value="117.833336">117°50'E</geoCoordinate>
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, 131 m alt., 1997-04-30, P. Wolseley Q21 p.p. (BM 001104020); Danum valley, pristine lowland dipterocarp forest
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<geoCoordinate id="AED03177515EDF92B0C7D9B3B35BA539" direction="north" orientation="latitude" precision="9" value="4.966">4°57.96'N</geoCoordinate>
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,
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<geoCoordinate id="86C9A39083763786ED0A339C0C960C38" direction="east" orientation="longitude" precision="9" value="117.788666">117°47.32'E</geoCoordinate>
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, 200-400 m alt., 2012, P. Wolseley, H.
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<normalizedToken id="435F761936D615BC53661A519B858392" originalValue="Thüs">Thues</normalizedToken>
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& C. Vairappan D-3-10-2 (BORH); Maliau conservation area, trail between Nepenthes Camp and waterfall Takob Akob, transition between pristine montane "Kerangas" (heath forest) and montane oak-conifer (
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<taxonomicName id="940A6384E710D084D541E70EECBF4390" class="Insecta" family="Braconidae" genus="Agathis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agathis" order="Hymenoptera" pageId="6" pageNumber="75" phylum="Arthropoda" rank="genus">Agathis</taxonomicName>
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) forest,
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<geoCoordinate id="F13FDA10F3A197997F998AFE94D14C3A" direction="north" orientation="latitude" precision="92" value="4.71">4°42.6'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="61C2CFC5AC0FC905CD393FC2A15B7830" direction="east" orientation="longitude" precision="92" value="116.875">116°52.5'E</geoCoordinate>
|
||
, 900-1000 m alt., 2013, C. Vairappan & H.
|
||
<normalizedToken id="513F7EA5A61B5D12423735797EF35AEF" originalValue="Thüs">Thues</normalizedToken>
|
||
L229 (BM); Ranau district, Kinabalu park, Musang camp on the Tambuyukon trail (loc. T98),
|
||
<geoCoordinate id="2FA921613767E8EB61160FFC27F6DCB6" direction="north" orientation="latitude" precision="1" value="6.212">6°12.720'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C4EEE62B6EA358C92C6FB77BA01B0B02" direction="east" orientation="longitude" precision="1" value="116.68152">116°40.891' E</geoCoordinate>
|
||
, 1429 m alt., epiphytic, 2014-12-09, A. Paukov 2234 (B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |