275 lines
37 KiB
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275 lines
37 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. & Branham, Marc A." docDate="2018" docId="B279B3F17F82958E13D9037481B01922" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Cyclocephala tridentata" docType="treatment" docVersion="6" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="220" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="219" updateTime="1668165599527" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moore, Matthew R.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cyclocephala@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cave, Ronald D.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branham, Marc A.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>745</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>378</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.745.23685</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.745.23685</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-745-101</mods:identifier>
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<mods:identifier type="ZooBank">8785DC6BC2A244FD94B6243EB07C717F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">047DFFCAFFA5F32EA97C873F4708943F</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1222435</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="142480307" LSID="urn:lsid:plazi:treatment:B279B3F17F82958E13D9037481B01922" httpUri="http://treatment.plazi.org/id/B279B3F17F82958E13D9037481B01922" lastPageId="119" lastPageNumber="220" pageId="118" pageNumber="219">
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<subSubSection pageId="118" pageNumber="219" type="nomenclature">
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName LSID="B279B3F1-7F82-958E-13D9-037481B01922" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Cyclocephala tridentata (Fabricius, 1801)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="118" pageNumber="219" type="reference_group">
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Melolontha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melolontha tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Melolontha tridentata</taxonomicName>
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Fabricius, 1801: 170-171 [original combination].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Cyclocephala tridentata</taxonomicName>
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(Fabricius) [new combination by
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<bibRefCitation author="Burmeister, H" pageId="168" pageNumber="269" refId="B140" refString="Burmeister, H, 1847. Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." title="Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." year="1847">Burmeister 1847</bibRefCitation>
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: 47].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<emphasis bold="true" pageId="118" pageNumber="219">syn.</emphasis>
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala annamariae" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="annamariae">Cyclocephala annamariae</taxonomicName>
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Dutrillaux & Chalumeau, 2013: 64-65 [original combination in
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<bibRefCitation author="Dutrillaux, B" pageId="177" pageNumber="278" refId="B292" refString="Dutrillaux, B, Chalumeau, F, Dutrillaux, A-M, Giannoulis, T, Mamuris, Z, 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61-67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61 - 67." year="2013">Dutrillaux et al. 2013</bibRefCitation>
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].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Cyclocephala tridentata</taxonomicName>
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Fabricius [synonymy by
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<bibRefCitation author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B983" refString="Ratcliffe, BC, Cave, RD, 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1-346." title="The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 346." year="2015">Ratcliffe and Cave 2015</bibRefCitation>
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: 113].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<emphasis bold="true" pageId="118" pageNumber="219">syn.</emphasis>
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata subsp. dominicensis" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="dominicensis">Cyclocephala tridentata dominicensis</taxonomicName>
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Cartwright & Chalumeau, 1978: 25 [original combination].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala dominicensis" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="dominicensis">Cyclocephala dominicensis</taxonomicName>
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Cartwright & Chalumeau [new species status by
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<bibRefCitation author="Dutrillaux, B" pageId="177" pageNumber="278" refId="B292" refString="Dutrillaux, B, Chalumeau, F, Dutrillaux, A-M, Giannoulis, T, Mamuris, Z, 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61-67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61 - 67." year="2013">Dutrillaux et al. 2013</bibRefCitation>
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: 64].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Cyclocephala tridentata</taxonomicName>
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(Fabricius) [synonymy by
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<bibRefCitation author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B983" refString="Ratcliffe, BC, Cave, RD, 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1-346." title="The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 346." year="2015">Ratcliffe and Cave 2015</bibRefCitation>
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: 113].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<emphasis bold="true" pageId="118" pageNumber="219">syn.</emphasis>
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<taxonomicName authorityName="Gyllenhal" authorityYear="1817" class="Insecta" family="Scarabaeidae" genus="Melolontha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melolontha biliturata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="biliturata">Melolontha biliturata</taxonomicName>
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Gyllenhal, 1817b: 105-106 [original combination].
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</paragraph>
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<paragraph pageId="118" pageNumber="219">
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala tridentata" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="species" species="tridentata">Cyclocephala tridentata</taxonomicName>
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(Fabricius) [synonymy by
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<bibRefCitation author="Burmeister, H" pageId="168" pageNumber="269" refId="B140" refString="Burmeister, H, 1847. Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." title="Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." year="1847">Burmeister 1847</bibRefCitation>
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: 48].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="118" pageNumber="219" type="types">
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<paragraph pageId="118" pageNumber="219">Types.</paragraph>
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<paragraph pageId="118" pageNumber="219">
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Lectotype ♀ of
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<taxonomicName lsidName="M. tridentata" pageId="118" pageNumber="219" rank="species" species="tridentata">
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<emphasis italics="true" pageId="118" pageNumber="219">M. tridentata</emphasis>
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</taxonomicName>
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deposited at ZMUK, now housed at ZMUC (
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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).
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<bibRefCitation author="Chalumeau, F" pageId="170" pageNumber="271" refId="B166" refString="Chalumeau, F, 1982. Contribution a l'etude des Scarabaeoidea des Antilles (III). Nouveau Review d'Entomologie 12: 321-345." title="Contribution a l'etude des Scarabaeoidea des Antilles (III). Nouveau Review d'Entomologie 12: 321 - 345." year="1982">Chalumeau (1982)</bibRefCitation>
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did not consider the lectotype valid because
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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selected a specimen from Martinique rather than Guadeloupe, the type locality. Therefore, Chalumeau designated a neotype deposited at ZMUC.Holotype ♂ of
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<taxonomicName family="Scarabaeidae" kingdom="Animalia" lsidName="C. annamariae" order="Coleoptera" pageId="118" pageNumber="219" rank="species" species="annamariae">
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<emphasis italics="true" pageId="118" pageNumber="219">C. annamariae</emphasis>
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</taxonomicName>
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at IREC (
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<bibRefCitation author="Dutrillaux, B" pageId="177" pageNumber="278" refId="B292" refString="Dutrillaux, B, Chalumeau, F, Dutrillaux, A-M, Giannoulis, T, Mamuris, Z, 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61-67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetoques, chromosomiques et geographiques. Annales de la Societe Entomologique de France (n. s.) 49: 61 - 67." year="2013">Dutrillaux et al. 2013</bibRefCitation>
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). Holotype ♂ of
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<taxonomicName family="Scarabaeidae" kingdom="Animalia" lsidName="C. tridentata subsp. dominicensis" order="Coleoptera" pageId="118" pageNumber="219" rank="subSpecies" species="tridentata" subSpecies="dominicensis">
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<emphasis italics="true" pageId="118" pageNumber="219">C. tridentata dominicensis</emphasis>
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</taxonomicName>
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in the
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<normalizedToken originalValue="Fortuné">Fortune</normalizedToken>
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Chalumeau Collection (
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<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.279" author="Cartwright, OL" pageId="169" pageNumber="270" refId="B156" refString="Cartwright, OL, Chalumeau, FE, 1978. Bredin-Archbold-Smithsonian biological survey of Dominica. The superfamily Scarabaeoidea (Coleoptera). Smithsonian Contributions to Zoology 279: 1-32. https://doi.org/10.5479/si.00810282.279" title="Bredin-Archbold-Smithsonian biological survey of Dominica. The superfamily Scarabaeoidea (Coleoptera). Smithsonian Contributions to Zoology 279: 1 - 32." url="https://doi.org/10.5479/si.00810282.279" year="1978">Cartwright and Chalumeau 1978</bibRefCitation>
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). The type material of
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<taxonomicName lsidName="M. biliturata" pageId="118" pageNumber="219" rank="species" species="biliturata">
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<emphasis italics="true" pageId="118" pageNumber="219">M. biliturata</emphasis>
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</taxonomicName>
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was not reported by
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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(1966)
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</bibRefCitation>
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, and it is not with the other Gyllenhal
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Melolontha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melolontha" order="Coleoptera" pageId="118" pageNumber="219" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="118" pageNumber="219">Melolontha</emphasis>
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</taxonomicName>
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types at UUZM (
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<bibRefCitation author="Wallin, L" pageId="230" pageNumber="331" refId="B1179" refString="Wallin, L, 2001. Catalogue of type specimens. 3. Entomology. Version 4. Uppsala University, Museum of Evolution, Zoology Section, Uppsala, 55 pp." title="Catalogue of type specimens. 3. Entomology. Version 4. Uppsala University, Museum of Evolution, Zoology Section, Uppsala, 55 pp." year="2001">Wallin 2001</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="118" pageNumber="219" type="distribution">
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<paragraph pageId="118" pageNumber="219">Distribution.</paragraph>
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<paragraph pageId="118" pageNumber="219">
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BARBADOS. COLOMBIA: Cauca. DOMINICA: St. David, St. George, St. John, St. Joseph, St. Luke, St. Mark, St. Patrick, St. Paul. GRENADA: St. Andrew. GUADELOUPE: Basse-Terre, Grande-Terre. MARTINIQUE: Fort-de-France, La
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<normalizedToken originalValue="Trinité">Trinite</normalizedToken>
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, Le Marin, Saint-Pierre. MONTSERRAT: Saint Georges. ST. LUCIA: Anse la Raye, Castries, Choiseul, Dauphin, Dennery, Micoud, Praslin,
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<normalizedToken originalValue="Soufrière">Soufriere</normalizedToken>
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, Vieux Fort. SURINAME.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="119" lastPageNumber="220" pageId="118" pageNumber="219" type="references">
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<paragraph pageId="118" pageNumber="219">References.</paragraph>
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<paragraph lastPageId="119" lastPageNumber="220" pageId="118" pageNumber="219">
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<bibRefCitation author="Voet, JE" pageId="230" pageNumber="331" refId="B1176" refString="Voet, JE, 1769. Catalogus systematicus Coleopterorum. Volume 1. G. Bakhuysen, The Hague, sections paginated separately + 54 pls." title="Catalogus systematicus Coleopterorum. Volume 1. G. Bakhuysen, The Hague, sections paginated separately + 54 pls." year="1769">Voet 1769</bibRefCitation>
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,
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<bibRefCitation author="Fabricius, JC" pageId="179" pageNumber="280" refId="B338" refString="Fabricius, JC, 1801. Systema eleutheratorum secundum ordines, genera, species: Adiectis synonymis, ocis, oberservationibus, descriptionibus. Volume 2. Impensis Bibliopolii Academici Novi, Kiel, 687 pp." title="Systema eleutheratorum secundum ordines, genera, species: Adiectis synonymis, ocis, oberservationibus, descriptionibus. Volume 2. Impensis Bibliopolii Academici Novi, Kiel, 687 pp." year="1801">Fabricius 1801</bibRefCitation>
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,
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<bibRefCitation author="Gyllenhal, L" pageId="186" pageNumber="287" refId="B449" refString="Gyllenhal, L, 1817b. In: Schoenherr CJ (Ed.) Appendix ad C. J. Schoenherr synonymiam insectorum sistens descriptions novarum specierum. Volume 1, Part 3. Scaris, in officina Lewerentziana, Stockholm, 266 pp. [+ 2 pls]" title="In: Schoenherr CJ (Ed.) Appendix ad C. J. Schoenherr synonymiam insectorum sistens descriptions novarum specierum. Volume 1, Part 3. Scaris, in officina Lewerentziana, Stockholm, 266 pp. [+ 2 pls]" year="1817 b">Gyllenhal 1817b</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11259" author="Dejean, PFMA" pageId="174" pageNumber="275" refId="B251" refString="Dejean, PFMA, 1821. Catalogue de la Collection de Coleopteres de M. le Baron Dejean. Crevot Libraire, Paris, 136 pp. https://doi.org/10.5962/bhl.title.11259" title="Catalogue de la Collection de Coleopteres de M. le Baron Dejean. Crevot Libraire, Paris, 136 pp." url="https://doi.org/10.5962/bhl.title.11259" year="1821">Dejean 1821</bibRefCitation>
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,
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</paragraph>
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</subSubSection>
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<subSubSection pageId="119" pageNumber="220" type="remarks">
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<paragraph pageId="119" pageNumber="220">Remarks.</paragraph>
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<paragraph pageId="119" pageNumber="220">
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The identification of the specimens from Colombia and Suriname need to be critically evaluated (
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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,
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<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
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,
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<bibRefCitation author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B983" refString="Ratcliffe, BC, Cave, RD, 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1-346." title="The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 346." year="2015">Ratcliffe and Cave 2015</bibRefCitation>
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).
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<bibRefCitation author="Chalumeau, F" pageId="170" pageNumber="271" refId="B166" refString="Chalumeau, F, 1982. Contribution a l'etude des Scarabaeoidea des Antilles (III). Nouveau Review d'Entomologie 12: 321-345." title="Contribution a l'etude des Scarabaeoidea des Antilles (III). Nouveau Review d'Entomologie 12: 321 - 345." year="1982">Chalumeau (1982)</bibRefCitation>
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reported four specimens of
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<taxonomicName family="Scarabaeidae" kingdom="Animalia" lsidName="C. tridentata" order="Coleoptera" pageId="119" pageNumber="220" rank="species" species="tridentata">
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<emphasis italics="true" pageId="119" pageNumber="220">C. tridentata</emphasis>
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</taxonomicName>
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from Puerto Rico, though major faunistic studies did not find additional specimens from Puerto Rico (
|
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<bibRefCitation author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B983" refString="Ratcliffe, BC, Cave, RD, 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1-346." title="The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 346." year="2015">Ratcliffe and Cave 2015</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |