222 lines
32 KiB
XML
222 lines
32 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="B1BA5EB37B3D5113BF9C991F6618A2D2" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole sava Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="B7516043-C09C-4573-9A1B-9B649C329C67" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:mods id="3A9C9E208FB5D32A56A78FE3E4241051" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DA6483844CE8874DF6DD2A4203B84F4D">
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<mods:title id="F569155BE3492B4BBD6174A72803A3E5">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="87EAF5722E9B2DAE750FE36F03CA863B" type="personal">
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<mods:role id="0321CCA89E93556718068C5A72AB5128">
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<mods:roleTerm id="623283FA516C571C6CD52D5E290801E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="45FC19C26A316F8F0750E6BFB7DC1CAA">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="C7A4015EF9F5A9B15E6007B72663035D">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="2EB9CFA3B29D052BB9712550F6CE3C8C" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="9D3712E120F1C84BB14C532926596470" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:name id="10EE995E5B08E6A7E18BBE4D9C7FC2FF" type="personal">
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<mods:namePart id="224EED2A7C95956367BB6A127E5F1DD8">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="2E4AB97DFE3A1C9E50A6F3322892E9DD">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="0830679E6C2887A5579FF010D40775B0" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:typeOfResource id="55BFCA7D3485D11F139E64362209E1E9">text</mods:typeOfResource>
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<mods:title id="3205EDBEDF818664BBC6B998D77D606F">ZooKeys</mods:title>
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<mods:part id="35B11E3B0C61C29970C0BE6159A3B63B">
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<mods:date id="456CA3C7887C48848FBF3CC1417D16C3">2020</mods:date>
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<mods:detail id="5A7F139147197C64AD784C14327DEE7A" type="volume">
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<mods:number id="77072F7C139F040822593AF9EAA6D4AB">949</mods:number>
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<mods:start id="85EF99F8FCE49D6BDDF94857C41FCF3C">1</mods:start>
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<mods:url id="8EB5C49BAC32096886C8D85472B6EE8E">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="2F6E142872C7157D5CA6D3D1FADC4E40">journal article</mods:classification>
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<mods:identifier id="EC897E999A30E7ABD90FEE373E0D6988" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier id="CC3EAC2E635A2BF7DA49F298D1DD2457" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<treatment id="B1BA5EB37B3D5113BF9C991F6618A2D2" ID-GBIF-Taxon="165090168" LSID="urn:lsid:zoobank.org:act:B7516043-C09C-4573-9A1B-9B649C329C67" httpUri="http://treatment.plazi.org/id/B1BA5EB37B3D5113BF9C991F6618A2D2" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="8DFF46CEA8E17ED8636DE5B813BF3D75" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="C6D3CFEDE7B6FDA164591AF1A68819C3" pageId="0" pageNumber="1">
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<taxonomicName id="5595D14EFA30C196C858D8ABFCB6B0F3" ID-CoL="CMLVN" LSID="B1BA5EB3-7B3D-5113-BF9C-991F6618A2D2" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava" status="sp. nov.">Pheidole sava</taxonomicName>
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<taxonomicNameLabel id="4492A1DACB22584FAEF663795056B1D0" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="DEFC5D26C9668DCBDA1FEC8FF2AFD2D2" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">Figs 52A-F</figureCitation>
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<figureCitation id="15BA1E12E23008935EB3F6E40A38F24E" captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64J</figureCitation>
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<figureCitation id="F34577272D919694BB2F613BE4CB0CEC" captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="993408FB6F82BBD8400AEAA57DF48CD9" pageId="0" pageNumber="1" type="type material">
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<paragraph id="22342E64902DA421025B00F8E3CAF57A" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="3EAC5410A1186AF937D607D1BF677BCB" pageId="0" pageNumber="1">
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<emphasis id="99DC231CA7BBFC70E3E9DF57F0B48637" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate id="7328906432CD34307E9B24B12499C29E" degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate id="DD9DBDC8D1FB0B82A2EF3D949C2A6C11" degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF41084; CASENT0808425 (CASC).
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<emphasis id="87B0D2CCC60AEB62BE23282F246BB3E9" bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype; CASENT0923296 (CASC).
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</paragraph>
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<caption id="3E1447C212E030010D22B4AFCD14E944" doi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
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<paragraph id="C59D8E25D547F1B20775992E9D7E2C81" pageId="0" pageNumber="1">
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<emphasis id="140652ED47F7F871823B890710239463" bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
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<taxonomicName id="ACD2BEE75436D554F685E3EC12903EC4" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
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<emphasis id="6A47ACE2B509DDB33DD8E280F4496FEE" italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="FE2825C0B623F436003941A8E829BA54" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="014825823C5274C789422E816D23F0E1" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="0AB95BCD36DEBA7118FBC775B1C81085" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923296) and full-face view (
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<emphasis id="C98E1333C84CC22C773CE55525B2EAB3" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="52D3FCD40DE4DA661D6F312C2EC39FB1" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="8792BC59A82437679A0DA01C04E1794A" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0808425).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="3FBF958275B4230823EC52D35A4CA6ED" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="2673BAA4960F87B94FFFB39AA8ED6FC4" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="824A8CAF6192E9F9CE70532BD96AB683" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="FADCD69964A85EA30E2C11D51CDB9A06" bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: • 1w., 1s.; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate id="425D16226ECE7B4DA6AF750974CA6068" degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate id="EE1013C5386E2FFBD13E004B7630D215" degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex root mat; BLF41089 (CASC). • 1w., 1s.; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate id="CEFB43246768D60EC7184482504A2B78" degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate id="650880C959AA858B5BEE1C7BA80336B0" degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF41097 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="73C1D10B0CD9973C8DC6BE3D42A5B258" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="882CE7571358B3A7832AED66D4B63EF7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="53CD5F2AE34C54515E7552CEA04F1E7D" pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis id="0BEA415F09E4104775CB18DFE0B1F347" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with dense, short, suberect pilosity; anterior and medial parts of frons with moderately dense, thick, and longitudinal rugae sometimes interrupted in the posteromedial part, interspaces with sparse and distinct rugulae; posterolateral sides with longitudinal to irregular and thick rugae with distinctly rugulate interspaces; area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity; mesosoma with fine rugofoveolae, dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches; gaster smooth; body brownish orange.
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<emphasis id="12ACDDA3BD455205553EF0E79ADE0925" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate with additional indistinct longitudinal rugulae on medial frons, area posterolateral from eyes smooth; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum moderately high and long, arched; promesonotal groove present; propodeal spines indistinct, triangular; mesosoma smooth and shiny, only promesonotal dorsum with arched rugae and indistinct foveolae, mesonotum and lateral sides of propodeum with indistinct and sparse foveolae; body yellowish brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="9C1B2114B119DA57B1199C3DFCDF07DD" pageId="0" pageNumber="1" type="description">
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<paragraph id="0898A20AFAC94A8F7B61EC516D887DD7" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="C0E495AA57240F637EA857849FB9470B" pageId="0" pageNumber="1">
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<emphasis id="4AA79663CBDED4D814D4B28B96B90A8D" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="FA27901869A324D334C6150EA2786F56" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 3): HL: 1.14-1.24 (1.2); HW: 1.16-1.23 (1.2); SL: 0.66-0.69 (0.68); EL: 0.13-0.15 (0.14); WL: 1.07-1.1 (1.09); PSL: 0.19-0.21 (0.2); MTL: 0.68-0.73 (0.7); PNW: 0.52-0.53 (0.52); PTW: 0.14-0.17 (0.16); PPW: 0.28-0.34 (0.31); CI: 98.2-101.0 (99.7); SI: 55.5-57.1 (56.4); PSLI: 16.2-16.7 (16.5); PPI: 44.8-61.3 (51.3); PNI: 42.7-45.3 (43.7); MTI: 57.5-59.1 (58.4).
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</paragraph>
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<paragraph id="3F7A868A3B2371FD88E01E3BCCB33B52" pageId="0" pageNumber="1">
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<emphasis id="52FEDECAB660001FD017E73AC9DBD9DB" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation id="55337F3E1382FD47A0AC026D2207E34E" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth invisible. Sides of the head with dense, short, suberect pilosity; whole head with relatively dense, short, decumbent to erect pilosity. Anterior and medial parts of frons with moderately dense and thick, longitudinal, and sometimes interrupted rugae in posteromedial part, interspaces with sparse and distinct rugulae; posterolateral sides with longitudinal to irregular and thick rugae with distinctly rugulate interspaces. Occipital lobes with thick, sparse, irregular rugae, interspaces smooth or with fine, indistinct rugulae. Gena with sparse, thick, and longitudinal rugae and smooth interspaces. Area posterolateral from eyes with dense longitudinal to irregular rugofoveolae. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; pilosity decumbent to suberect (Fig.
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<figureCitation id="B8654DFB792079A9C86DAD01565A6285" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity (Fig.
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<figureCitation id="C20C96AE772F98FF02086392AB3487CE" captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64J</figureCitation>
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).
|
||
<emphasis id="36480306DF15BCB16D4A99C49E844C3D" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, thin, with acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="2AB3D09D56B30E1ECE8592CFE02528EC" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D</figureCitation>
|
||
). Surface shiny with fine rugofoveolae, dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches. Pilosity sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="A843596953A4EDD71F28E557F7433B6A" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
|
||
).
|
||
<emphasis id="CA1F314331129DA6AC08C8142C82C0D9" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with fine and sparse rugofoveolae; peduncle; node smooth to indistinctly rugulose, low, triangular, with rounded apex, in rear view node dorsoventrally convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="25FC03F22C1FF5FDA104E0281B9BDC55" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
|
||
).
|
||
<emphasis id="F244468E6BBD8D33C7BA9754FA4AAFCF" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny, with fine and sparse rugulae; in dorsal view postpetiole oval, lateral margins medially with two indistinct dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="D3D2FA2A0E701B782AA146FC62238B9D" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
|
||
).
|
||
<emphasis id="CE1DD39B38CB8767F14C0860C21204E3" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity moderately dense, long, and erect (Fig.
|
||
<figureCitation id="4BB2AAAF01773800602AE853997E76A6" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
|
||
).
|
||
<emphasis id="6DA97995D11688CEF8AAA49E982BA345" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brownish orange; legs dark yellow, gaster bright brown (Fig.
|
||
<figureCitation id="26BDE18813E7BC49AB63C070243BDF24" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F52FEBC0F34B9358BEA765235E273FE5" pageId="0" pageNumber="1">
|
||
<emphasis id="E8DCEC29A269EFAFA2F4F18DE6DDAFD1" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="A0AF8D10D51CF863162452BB0B13E0F1" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 3): HL: 0.58-0.61 (0.59); HW: 0.48-0.49 (0.49); SL: 0.6-0.63 (0.61); EL: 0.11-0.12 (0.11); WL: 0.73-0.74 (0.73); PSL: 0.07-0.09 (0.08); MTL: 0.5-0.52 (0.51); PNW: 0.3-0.34 (0.32); PTW: 0.09-0.1 (0.09); PPW: 0.13-0.16 (0.15); CI: 117.1-127.1 (121.6); SI: 122.0-129.4 (126.2); PSLI: 11.3-14.2 (12.8); PPI: 60.4-65.7 (63.0); PNI: 62.7-70.4 (66.4); MTI: 103.3-106.8 (104.7).
|
||
</paragraph>
|
||
<paragraph id="BFD373F3983B1DB54F4F0FA61E3427E6" pageId="0" pageNumber="1">
|
||
<emphasis id="840F9444AEE32557081C137D907DB993" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly convex (Fig.
|
||
<figureCitation id="D23622538CA04876D041BFEA20D630CB" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52A</figureCitation>
|
||
). Pilosity relatively sparse, long, decumbent to subdecumbent. Head foveolate with additional indistinct longitudinal rugulae on medial frons, area posterolateral from eyes smooth; antennal sockets with few thick, curved outward rugae and foveolae interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="7B73172967DE6F5EC61A7BB46AA5A0AA" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52A, C</figureCitation>
|
||
).
|
||
<emphasis id="0E96F92531858AB6D1F76E042AABC2C7" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and long, arched; promesonotal groove present but indistinct; metanotal groove present and distinct; propodeal spines indistinct, lobe-like (Fig.
|
||
<figureCitation id="D5C30F8E63C578EAD6861A0886F28D92" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C</figureCitation>
|
||
). Sculpture smooth and shiny, only promesonotal dorsum with arched rugae and indistinct foveolae, mesonotum and lateral sides of propodeum with indistinct and sparse foveolae. Pilosity sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="4029E7CACC81FB6639A26743708716DE" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis id="40B3551B0DAC52E86C2568340CD1AEFD" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Peduncle short and thin with ventral face relatively straight (Fig.
|
||
<figureCitation id="D39E42B40CF1A59A66EBACA9F3F728B7" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis id="6B72DD92EB2E8DA4CAA6FC530FC02346" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="5A7FA9C924DA4E5C2476313CE43F83A3" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis id="CD5BCDF2EB967B5F74B89A0F22F08B3F" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown; legs and lateral sides of pronotum yellowish brown (Fig.
|
||
<figureCitation id="F1D65964CD68C1CAC464BA8970D22A8F" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="854602F61A81DE815DE6A47932004AF4" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="F8DD73877C08187571A60AA4A656D2CF" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="291484B25EF9019DA1C41324ECA1B324" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FBA42C6C65D9F252A473A6A0129BADB5" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="E2A7B297C6EF2385AAE652AB72075588" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="C45D26A10C4F69EE5006A1741C023BB2" pageId="0" pageNumber="1">The species was collected at 1350 m in elevation, in montane rainforest. Nests were located in rotten logs and root mat.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8856EFF538CF08B7E27C4363B7DE362" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="EAA0EDA8532DF430BD14E338C41BAF84" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="96923DEC5C254D056417433A76D33B81" pageId="0" pageNumber="1">
|
||
<taxonomicName id="3641098DB4395E63465F75E3477A72F8" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
|
||
<emphasis id="0366013245F686123DDDB5A5421B77E6" italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from Parc National de Marojejy in Antsiranana, is most similar to
|
||
<taxonomicName id="4A6DE13154D6A77C554416D74613C4A5" lsidName="P. itremo" pageId="0" pageNumber="1" rank="species" species="itremo">
|
||
<emphasis id="C4D92ECC08E083D1990AD0F76D0F5323" italics="true" pageId="0" pageNumber="1">P. itremo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from
|
||
<normalizedToken id="AC6C8DDBC29DF85902C69994EDE71C10" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="9FE5F9673A2CE40FAAB29EE551C2043A" originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
|
||
in Fianarantsoa and
|
||
<normalizedToken id="5DA5822F06468CE9D672AA58404301DD" originalValue="Forêt">Foret</normalizedToken>
|
||
d Ambohitantely in Antananarivo. Major workers of
|
||
<taxonomicName id="3CAFA9763E02C4DEE1D84674E9D9553A" lsidName="P. sava" pageId="0" pageNumber="1" rank="species" species="sava">
|
||
<emphasis id="51F14C7E171A4F289C65F336DE7F23EA" italics="true" pageId="0" pageNumber="1">P. sava</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated based on brighter body colouration, presence of dense, short, suberect pilosity sides of the head, medial frons with sparse and distinctly rugulate interspaces, finely rugofoveolae mesosoma with dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches; minor workers can be separated based on foveolate head with additional indistinct longitudinal rugulae on medial frons, smooth area posterolateral from eyes, predominantly smooth mesosoma with promesonotal dorsum with arched rugae and indistinct foveolae, and indistinct and sparse foveolae on mesonotum and lateral sides of propodeum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |