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<document id="BD394555819018BF1F355E8B81396996" ID-DOI="10.35885/ruthenica.2023.33(1).3" ID-ISSN="2307-7336" ID-Zenodo-Dep="11041197" ID-ZooBank="63522E75-C4DF-4DFB-992B-C1ACBC803399" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1713810501483" checkinUser="felipe" docAuthor="Sang, Do Duc &amp; Son, Nguyen Thanh" docDate="2023" docId="B11287D15657FF917F3F05B491B9FD1C" docLanguage="en" docName="Ruthenica.33.1.19-26.pdf" docOrigin="Ruthenica, Russian Malacological Journal 33 (1)" docSource="http://dx.doi.org/10.35885/ruthenica.2023.33(1).3" docStyle="DocumentStyle:3A4A24405A53639AD762166AE048D148.1:Ruthenica.2022-.journal_article.english.index" docStyleId="3A4A24405A53639AD762166AE048D148" docStyleName="Ruthenica.2022-.journal_article.english.index" docStyleVersion="1" docTitle="Chuatienpupa megacanalis Sang &amp; Son 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="4D2BFFA95655FF957C610005952FFFBE" masterDocTitle="A new genus and new species of Pupinidae (Gastropoda: Caenogastropoda) from Northern Vietnam" masterLastPageNumber="26" masterPageNumber="19" pageNumber="21" updateTime="1715166209063" updateUser="tatiana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A199B2E1860DF4F4869A9CD7FB8FB94E">A new genus and new species of Pupinidae (Gastropoda: Caenogastropoda) from Northern Vietnam</mods:title>
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<mods:namePart id="87F9824F3FD501094957B34068994212">Sang, Do Duc</mods:namePart>
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<mods:namePart id="7DF35EC643D8DEFB46A0EE942E2A7EE1">Son, Nguyen Thanh</mods:namePart>
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<mods:title id="A18C96F499BBE272FCAEF1356E63CE73">Ruthenica, Russian Malacological Journal</mods:title>
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<mods:date id="A29F915E8909D36F22B1DB5ABD05774F">2023</mods:date>
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<treatment id="B11287D15657FF917F3F05B491B9FD1C" LSID="urn:lsid:plazi:treatment:B11287D15657FF917F3F05B491B9FD1C" httpUri="http://treatment.plazi.org/id/B11287D15657FF917F3F05B491B9FD1C" lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21">
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<paragraph id="390436C75657FF977F3F05B4901DFA70" blockId="2.[862,1330,1457,1516]" box="[862,1330,1457,1486]" pageId="2" pageNumber="21">
<heading id="624C81AB5657FF977F3F05B4901DFA70" box="[862,1330,1457,1486]" centered="true" fontSize="12" level="2" pageId="2" pageNumber="21" reason="6">
<taxonomicName id="FEBB4D445657FF977F3F05B491E7FA70" ID-CoL="B3F2T" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[862,1224,1457,1486]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="species" species="megacanalis" status="sp. nov.">
<emphasis id="0BCFEAD55657FF977F3F05B491E7FA70" bold="true" box="[862,1224,1457,1486]" italics="true" pageId="2" pageNumber="21">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF9778B105B7901DFA70" box="[1232,1330,1458,1486]" pageId="2" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="390436C75657FF977FD705D491F5FA55" blockId="2.[862,1330,1457,1516]" box="[950,1242,1489,1516]" pageId="2" pageNumber="21">
(
<figureCitation id="A1802A425657FF977FDF05D4912AFA55" box="[958,1029,1489,1515]" captionStart="FIG" captionStartId="1.[169,211,938,959]" captionTargetId="figure-512@1.[168,735,189,908]" captionTargetPageId="1" captionText="FIG. 1. A. General shell morphology of Chuatienpupa gen. nov. and its terminology. B. Details of the anterior canal (bottom view). C. Details of the posterior canal. РИС. 1. A. ОбЩаЯ морфологиЯ раковины Chuatienpupa gen.nov.и термины,испольЗованные при ее описании. B. Детали переднего канала (вид сниЗу). C. Детали Заднего канала." pageId="2" pageNumber="21">Figs 1</figureCitation>
,
<figureCitation id="A1802A425657FF97787205D4910CFA55" box="[1043,1059,1489,1515]" captionStart="FIG" captionStartId="2.[1119,1164,189,210]" captionTargetId="figure-490@2.[184,910,200,924]" captionTargetPageId="2" captionText="FIG. 2. Map of Vietnam showing the type locality (black dot) of Chuatienpupa megacanalis gen. nov. et sp. nov. РИС. 2. Карта Вьетнама с типовым местона- хождением (черный кружок) Chuatienpupa megacanalis gen. nov. et sp. nov." pageId="2" pageNumber="21">2</figureCitation>
,
<figureCitation id="A1802A425657FF97784E05D49153FA55" box="[1071,1148,1489,1515]" captionStart="FIG" captionStartId="3.[168,210,1924,1945]" captionTargetBox="[168,1376,189,1900]" captionTargetId="figure-11@3.[168,1378,189,1900]" captionTargetPageId="3" captionText="FIG. 3. Chuatienpupa megacanalis sp. nov.AE. Holotype VNMN-IZ 000.002.331.FH. Paratypes ZVNU.MOL 047.Photos: Nguyen T.S. and Do D.S. РИС. 3. Chuatienpupa megacanalis sp. nov. AE. Голотип VNMN-IZ 000.002.331. FH. Паратипы ZVNU.MOL 047. Фото: Нгуен Т.С. и До Д.С." figureDoi="http://doi.org/10.5281/zenodo.11041208" httpUri="https://zenodo.org/record/11041208/files/figure.png" pageId="2" pageNumber="21">3 AH</figureCitation>
,
<figureCitation id="A1802A425657FF9778E605D491FDFA55" box="[1159,1234,1489,1516]" captionStart="FIG" captionStartId="6.[175,217,1061,1082]" captionTargetId="figure-173@6.[174,1384,189,1041]" captionTargetPageId="6" captionText="FIG. 5. Chuatienpupa megacanalis sp. nov., AC. Living specimen (paratype ZVNU.MOL 047). D. Juvenile shells. Photos: Nguyen T.S. and Do D.S. РИС. 5. Chuatienpupa megacanalis sp. nov., AC. ЖиваЯ особь (паратип ZVNU.MOL 047). D. Ювенильные раковины. Photos: Nguyen T.S. and Do D.S." pageId="2" pageNumber="21">5 AD</figureCitation>
)
</paragraph>
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ZooBank registration:
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urn:lsid:zoobank.org:act:
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<subSubSection id="71A1654C5657FF977F2E064A9048F8B1" pageId="2" pageNumber="21" type="materials_examined">
<paragraph id="390436C75657FF977F2E064A9048F8B1" blockId="2.[809,1384,1614,1807]" pageId="2" pageNumber="21">
<emphasis id="0BCFEAD55657FF977F2E064A96CCF9DA" bold="true" box="[847,995,1614,1636]" pageId="2" pageNumber="21">Type material.</emphasis>
<materialsCitation id="89D33C9A5657FF977F88064B9673F94F" collectingDate="2022-02-04" collectionCode="VNMN-IZ" collectorName="D. S. Do" country="Vietnam" county="Lac Thuy district" latitude="20.554167" location="Chua Tien" longLatPrecision="12" longitude="105.74167" municipality="Phu Lao" pageId="2" pageNumber="21" specimenCount="1" stateProvince="Hoa Binh" typeStatus="holotype">
<typeStatus id="E60088655657FF977F88064B916DF9DA" box="[1001,1090,1614,1636]" pageId="2" pageNumber="21" type="holotype">Holotype</typeStatus>
<collectionCode id="5FAAAE025657FF977826064A9180F9DA" box="[1095,1199,1615,1636]" pageId="2" pageNumber="21">VNMN-IZ</collectionCode>
000.002.331 (shell height
<quantity id="FE439B225657FF977F09066E969EF93E" box="[872,945,1643,1664]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="21" unit="mm" value="9.1">9.1 mm</quantity>
, shell width
<quantity id="FE439B225657FF977842066E9143F93E" box="[1059,1132,1643,1665]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="21" unit="mm" value="4.6">4.6 mm</quantity>
,
<figureCitation id="A1802A425657FF977815066E91F7F93F" box="[1140,1240,1643,1665]" captionStart="FIG" captionStartId="3.[168,210,1924,1945]" captionTargetBox="[168,1376,189,1900]" captionTargetId="figure-11@3.[168,1378,189,1900]" captionTargetPageId="3" captionText="FIG. 3. Chuatienpupa megacanalis sp. nov.AE. Holotype VNMN-IZ 000.002.331.FH. Paratypes ZVNU.MOL 047.Photos: Nguyen T.S. and Do D.S. РИС. 3. Chuatienpupa megacanalis sp. nov. AE. Голотип VNMN-IZ 000.002.331. FH. Паратипы ZVNU.MOL 047. Фото: Нгуен Т.С. и До Д.С." figureDoi="http://doi.org/10.5281/zenodo.11041208" httpUri="https://zenodo.org/record/11041208/files/figure.png" pageId="2" pageNumber="21">Fig. 3AE</figureCitation>
),
<collectingCountry id="41AC76575657FF977887066E9017F93F" box="[1254,1336,1643,1665]" name="Vietnam" pageId="2" pageNumber="21">Vietnam</collectingCountry>
,
<collectingRegion id="FB7FF8255657FF97795E066E969CF923" country="Vietnam" name="Hoa Binh" pageId="2" pageNumber="21">Hoa Binh province</collectingRegion>
,
<collectingCounty id="D0654E4B5657FF977FDA068D914EF923" box="[955,1121,1671,1693]" pageId="2" pageNumber="21">Lac Thuy district</collectingCounty>
,
<collectingMunicipality id="D960ACBD5657FF97780806829197F923" box="[1129,1208,1671,1693]" pageId="2" pageNumber="21">Phu Lao</collectingMunicipality>
commune, forests over limestone and limestone hills around the
<location id="3C64601C5657FF97788106A6906CF907" LSID="urn:lsid:plazi:treatment:B11287D15657FF917F3F05B491B9FD1C:3C64601C5657FF97788106A6906CF907" box="[1248,1347,1699,1721]" country="Vietnam" county="Lac Thuy district" latitude="20.554167" longLatPrecision="12" longitude="105.74167" municipality="Phu Lao" name="Chua Tien" pageId="2" pageNumber="21" stateProvince="Hoa Binh">Chua Tien</location>
pagoda (
<geoCoordinate id="5C8F50005657FF977F0206C596F8F968" box="[867,983,1728,1750]" degrees="20" direction="north" minutes="33.25" orientation="latitude" pageId="2" pageNumber="21" precision="9" value="20.554167">20°33.25N</geoCoordinate>
,
<geoCoordinate id="5C8F50005657FF977F8106C59172F968" box="[992,1117,1728,1750]" degrees="105" direction="east" minutes="44.50" orientation="longitude" pageId="2" pageNumber="21" precision="9" value="105.74167">105°44.50E</geoCoordinate>
), coll.
<collectorName id="944E53115657FF9778FD06C591C3F96B" box="[1180,1260,1728,1749]" pageId="2" pageNumber="21">D.S. Do</collectorName>
,
<date id="4D0510075657FF97789506C59673F94F" pageId="2" pageNumber="21" value="2022-02-04">
<collectingDate id="5D41E9EF5657FF97789506C59673F94F" pageId="2" pageNumber="21" value="2022-02-04">04 February 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="89D33C9A5657FF977F0406D8904CF8B1" collectingDate="2022-02-04" collectionCode="VNMN-IZ, ZVNU" collectorName="D. S. Do" country="Vietnam" county="Lac Thuy district" latitude="20.554167" location="Chua Tien" longLatPrecision="12" longitude="105.74167" municipality="Phu Lao" pageId="2" pageNumber="21" specimenCode="MOL 047" specimenCount="1" stateProvince="Hoa Binh" typeStatus="paratype">
<typeStatus id="E60088655657FF977F0406D896E9F94C" box="[869,966,1757,1778]" pageId="2" pageNumber="21" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="5FAAAE025657FF977FAF06D89119F94C" box="[974,1078,1757,1778]" pageId="2" pageNumber="21">VNMN-IZ</collectionCode>
000.002.332 (2 shells),
<collectionCode id="5FAAAE025657FF97797E06D8904AF94C" box="[1311,1381,1757,1778]" pageId="2" pageNumber="21">ZVNU</collectionCode>
.
<specimenCode id="691D9EBC5657FF977F4806FC96A8F8B0" box="[809,903,1785,1806]" collectionCode="ZVNU" pageId="2" pageNumber="21">MOL 047</specimenCode>
(3 shells), DDS (8 shells), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5657FF917F2E07269767FEA9" lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21" type="diagnosis">
<paragraph id="390436C75657FF917F2E07269767FEA9" blockId="2.[809,1384,1827,2048]" lastBlockId="4.[174,749,188,2047]" lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21">
<emphasis id="0BCFEAD55657FF977F2E072696E6F883" bold="true" box="[847,969,1827,1853]" pageId="2" pageNumber="21">Diagnosis.</emphasis>
Shell pupoid shaped, rather solid and glossy. Peristome thickened, not expanded, interrupted by anterior and posterior canals. Posterior canal separated by palatal lip by lower palatal plica, and an additional plica (upper palatal plica) divides posterior canal into two. Columellar area strongly thickened and expanded, with lower part curving into hook-like plica. Anterior canal very wide, twisted funnel-shaped, and opened on basal side. Operculum outside bowl-shaped, and flat inside.
</paragraph>
</subSubSection>
<caption id="6DC4664F5656FF967CC9078194CBF84D" ID-DOI="http://doi.org/10.5281/zenodo.11041208" ID-Zenodo-Dep="11041208" httpUri="https://zenodo.org/record/11041208/files/figure.png" pageId="3" pageNumber="22" startId="3.[168,210,1924,1945]" targetBox="[168,1376,189,1900]" targetPageId="3" targetType="figure">
<paragraph id="390436C75656FF967CC9078194EDF80D" blockId="3.[168,1378,1924,1971]" pageId="3" pageNumber="22">
FIG. 3.
<taxonomicName id="FEBB4D445656FF967C8E078194D2F824" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[239,509,1924,1946]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="22" phylum="Mollusca" rank="species" species="megacanalis" status="sp. nov.">
<emphasis id="0BCFEAD55656FF967C8E078194D2F824" bold="true" box="[239,509,1924,1946]" italics="true" pageId="3" pageNumber="22">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5656FF967E6007809765F824" box="[513,586,1925,1946]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0BCFEAD55656FF967E2C078197AFF827" bold="true" box="[589,640,1924,1945]" pageId="3" pageNumber="22">AE.</emphasis>
Holotype VNMN-IZ 000.002.331.
<emphasis id="0BCFEAD55656FF967FB10781912CF827" bold="true" box="[976,1027,1924,1945]" pageId="3" pageNumber="22">FH.</emphasis>
Paratypes ZVNU.MOL 047. Photos: Nguyen T.S. and Do D.S.
</paragraph>
<paragraph id="390436C75656FF967CC907C194CBF84D" blockId="3.[168,1378,1988,2035]" pageId="3" pageNumber="22">
РИС. 3.
<taxonomicName id="FEBB4D445656FF967D6107C1973DF864" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[256,530,1988,2010]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="22" phylum="Mollusca" rank="species" species="megacanalis" status="sp. nov.">
<emphasis id="0BCFEAD55656FF967D6107C1973DF864" bold="true" box="[256,530,1988,2010]" italics="true" pageId="3" pageNumber="22">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5656FF967E7B07C09747F864" box="[538,616,1989,2010]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0BCFEAD55656FF967E0E07C1978DF867" bold="true" box="[623,674,1988,2009]" pageId="3" pageNumber="22">AE.</emphasis>
Голотип VNMN-IZ 000.002.331.
<emphasis id="0BCFEAD55656FF967F9F07C1911EF867" bold="true" box="[1022,1073,1988,2009]" pageId="3" pageNumber="22">FH.</emphasis>
Паратипы ZVNU.MOL 047. Фото: Нгуен Т.С. и До Д.С.
</paragraph>
</caption>
<subSubSection id="71A1654C5651FF917CB5011895DEF800" pageId="4" pageNumber="23" type="description">
<paragraph id="390436C75651FF917CB50118974FFAC2" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917CB50118944AFE89" bold="true" box="[212,357,285,311]" pageId="4" pageNumber="23">Description.</emphasis>
Shell horny or yellowish, smooth, glossy and slightly transparent, covered by a glaze, pupoid shape with slightly pointed apex. Peristome and its surrounding area clearly distinguished from the other areas by its characteristic ivory colour or transparency. The increase of the whorls is rather regular, slow at the beginning, but suddenly increasing in the last whorls, but not bulging. Shell rather solid, composed of 55.5 whorls. Protoconch quite smooth and approximately of 1.5 whorls, although the boundary between the protoconch and the teleoconch is not easily discernible. Suture superficial, margined. The upper whorls moderately bulging, last two whorls less bulging; last whorl occupies more than 2/3 of the total shell height. Aperture circular, slightly oblique and basal part protruding. Peristome complex, thickened, not expanded, and interrupted by two canals: one at parieto-angular corner (posterior canal), and one at basal area (anterior canal). Parietal callus strong, margin slightly elevated and with large parietal tooth. Palatal area quite thickened, the upper part less expanded than on the lower part; the upper part quite concave (observed from lateral view), with two large triangular-like plicae, which are directed toward the parietal plica. Basal area extends obliquely, largely occupied by anterior canal, a strong scythe-shaped plica (basal plica), which seems to bisect the anterior canal (more obvious from basal view). Columella strongly thickened and expanded, lower part curving into hook-like plica (columellar plica) that covers anterior canal. Posterior canal continues vertically upward and forming wide groove divided into two cavities. Anterior canal twisted funnel-shaped, appearing as very wide groove that opens on basal side. Umbilicus closed.
</paragraph>
<paragraph id="390436C75651FF917CB5058794CAF983" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
Operculum quite solid, closed bowl-shaped, but hollow inside. Outer surface smooth, glossy, strongly convex, and covered by a glaze. Inner surface flat, slightly smooth, multispiral, with a low nipple on its central. Operculum uniform yellow-orange on outer surface (
<figureCitation id="A1802A425651FF917D6F0627947BF983" box="[270,340,1570,1597]" captionStart="FIG" captionStartId="1.[169,211,938,959]" captionTargetId="figure-512@1.[168,735,189,908]" captionTargetPageId="1" captionText="FIG. 1. A. General shell morphology of Chuatienpupa gen. nov. and its terminology. B. Details of the anterior canal (bottom view). C. Details of the posterior canal. РИС. 1. A. ОбЩаЯ морфологиЯ раковины Chuatienpupa gen.nov.и термины,испольЗованные при ее описании. B. Детали переднего канала (вид сниЗу). C. Детали Заднего канала." pageId="4" pageNumber="23">Figs 1</figureCitation>
,
<figureCitation id="A1802A425651FF917D03062694AFF983" box="[354,384,1571,1597]" captionStart="FIG" captionStartId="3.[168,210,1924,1945]" captionTargetBox="[168,1376,189,1900]" captionTargetId="figure-11@3.[168,1378,189,1900]" captionTargetPageId="3" captionText="FIG. 3. Chuatienpupa megacanalis sp. nov.AE. Holotype VNMN-IZ 000.002.331.FH. Paratypes ZVNU.MOL 047.Photos: Nguyen T.S. and Do D.S. РИС. 3. Chuatienpupa megacanalis sp. nov. AE. Голотип VNMN-IZ 000.002.331. FH. Паратипы ZVNU.MOL 047. Фото: Нгуен Т.С. и До Д.С." figureDoi="http://doi.org/10.5281/zenodo.11041208" httpUri="https://zenodo.org/record/11041208/files/figure.png" pageId="4" pageNumber="23">3F</figureCitation>
,
<figureCitation id="A1802A425651FF917DED062794F7F983" box="[396,472,1570,1597]" captionStart="FIG" captionStartId="6.[175,217,1061,1082]" captionTargetId="figure-173@6.[174,1384,189,1041]" captionTargetPageId="6" captionText="FIG. 5. Chuatienpupa megacanalis sp. nov., AC. Living specimen (paratype ZVNU.MOL 047). D. Juvenile shells. Photos: Nguyen T.S. and Do D.S. РИС. 5. Chuatienpupa megacanalis sp. nov., AC. ЖиваЯ особь (паратип ZVNU.MOL 047). D. Ювенильные раковины. Photos: Nguyen T.S. and Do D.S." pageId="4" pageNumber="23">5 AC</figureCitation>
).
</paragraph>
<paragraph id="390436C75651FF917CB506469467F903" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
Living specimens of
<taxonomicName id="FEBB4D445651FF917DD8064697C3F9E3" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[441,748,1602,1629]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="megacanalis" status="sp. nov.">
<emphasis id="0BCFEAD55651FF917DD8064697C3F9E3" box="[441,748,1602,1629]" italics="true" pageId="4" pageNumber="23">
Chuatienpupa
<emphasis id="0BCFEAD55651FF917E3D064797C3F9E3" bold="true" box="[604,748,1602,1629]" italics="true" pageId="4" pageNumber="23">megacanalis</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5651FF917CCF0666942AF9C3" box="[174,261,1635,1661]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
has a light grey to light brown body with a pinkish-brown to pale grey head and black tentacles (
<figureCitation id="A1802A425651FF917CD706A69414F903" box="[182,315,1699,1726]" captionStart="FIG" captionStartId="6.[175,217,1061,1082]" captionTargetId="figure-173@6.[174,1384,189,1041]" captionTargetPageId="6" captionText="FIG. 5. Chuatienpupa megacanalis sp. nov., AC. Living specimen (paratype ZVNU.MOL 047). D. Juvenile shells. Photos: Nguyen T.S. and Do D.S. РИС. 5. Chuatienpupa megacanalis sp. nov., AC. ЖиваЯ особь (паратип ZVNU.MOL 047). D. Ювенильные раковины. Photos: Nguyen T.S. and Do D.S." pageId="4" pageNumber="23">Figs 5 AC</figureCitation>
).
</paragraph>
<paragraph id="390436C75651FF917CB506C69493F881" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917CB506C69476F963" bold="true" box="[212,345,1731,1757]" pageId="4" pageNumber="23">Variability.</emphasis>
This species varies slightly in shell dimensions (smallest adult
<typeStatus id="E60088655651FF917DB506E1971CF940" box="[468,563,1764,1790]" pageId="4" pageNumber="23" type="paratype">paratype</typeStatus>
shell is
<quantity id="FE439B225651FF917EE606E197F6F940" box="[647,729,1764,1790]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="23" unit="mm" value="8.4">8.4 mm</quantity>
x
<quantity id="FE439B225651FF917CCF0701942DF8A0" box="[174,258,1796,1822]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="23" unit="mm" value="4.3">4.3 mm</quantity>
) and in shell color (the most of the specimens are horny or yellowish).
</paragraph>
<paragraph id="390436C75651FF917CB5074195DEF800" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917CB5074194ACF8E0" bold="true" box="[212,387,1860,1886]" pageId="4" pageNumber="23">Measurements</emphasis>
(in mm). Shell height 8.49.3, shell width 4.35.0, aperture height 2.62.8, aperture width 2.12.4, and operculum diameter 1.92.0 (n = 14).
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5651FF917CB507C19457F841" pageId="4" pageNumber="23" type="distribution">
<paragraph id="390436C75651FF917CB507C19457F841" blockId="4.[174,749,188,2047]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917CB507C19442F860" bold="true" box="[212,365,1988,2014]" pageId="4" pageNumber="23">Distribution.</emphasis>
This species is known from the
<typeStatus id="E60088655651FF917CCF07E095F1F841" box="[174,222,2021,2047]" pageId="4" pageNumber="23">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5651FF917F2E00B99686FE86" pageId="4" pageNumber="23" type="etymology">
<paragraph id="390436C75651FF917F2E00B99686FE86" blockId="4.[809,1384,187,674]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917F2E00B996FAFF68" bold="true" box="[847,981,188,214]" pageId="4" pageNumber="23">Etymology.</emphasis>
The species name is derived from its characteristic anterior canal, from the Latin words
<emphasis id="0BCFEAD55651FF917F5300FB9640FEA9" box="[818,879,254,279]" italics="true" pageId="4" pageNumber="23">mega</emphasis>
, meaning large, great and
<emphasis id="0BCFEAD55651FF9178DB00F89024FEA9" box="[1210,1291,253,279]" italics="true" pageId="4" pageNumber="23">canalis</emphasis>
, meaning canal.
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5651FF917F2E013A9136FDA0" pageId="4" pageNumber="23" type="biology_ecology">
<paragraph id="390436C75651FF917F2E013A9136FDA0" blockId="4.[809,1384,187,674]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917F2E013A969CFEE7" bold="true" box="[847,947,319,345]" pageId="4" pageNumber="23">Ecology.</emphasis>
This species collected from leaf litter at the foot of limestone rocks with dense disturbed vegetation. Three other pupinid species,
<taxonomicName id="FEBB4D445651FF9178A701849145FE05" authority="Bavay &amp; Dautzenberg, 1899" authorityName="Bavay &amp; Dautzenberg" authorityYear="1899" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="23" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="0BCFEAD55651FF9178A701849048FE24" box="[1222,1383,385,410]" italics="true" pageId="4" pageNumber="23">Pupina anceyi</emphasis>
<bibRefCitation id="5D2A4B365651FF917F4801A79145FE05" author="Bavay A. &amp; Dautzenberg P." box="[809,1130,417,444]" pageId="4" pageNumber="23" pagination="28 - 55" refId="ref3655" refString="Bavay A., Dautzenberg P. 1899. Description de co- quilles nouvelles de l' Indo-Chine. Journal de Conchyliologie, 47 (1): 28 - 55." type="journal article" year="1899">Bavay &amp; Dautzenberg, 1899</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FEBB4D445651FF91781401A796E9FE62" authority="Mabille, 1887" authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="23" phylum="Mollusca" rank="species" species="exclamationis">
<emphasis id="0BCFEAD55651FF91781401A79047FE05" box="[1141,1384,417,443]" italics="true" pageId="4" pageNumber="23">Pupina exclamationis</emphasis>
<bibRefCitation id="5D2A4B365651FF917F4801C796E9FE62" author="Mabille J." box="[809,966,450,476]" pageId="4" pageNumber="23" pagination="73 - 164" refId="ref4429" refString="Mabille J. 1887. Sur quelques mollusques du Tonkin. Bulletins de la Societe Malacologique de France, 4: 73 - 164." type="journal article" year="1887">Mabille, 1887</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FEBB4D445651FF917F9D01C69690FE43" authority="Dautzenberg &amp; Fischer, 1906" authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="23" phylum="Mollusca" rank="species" species="verneaui">
<emphasis id="0BCFEAD55651FF917F9D01C6919AFE62" box="[1020,1205,451,476]" italics="true" pageId="4" pageNumber="23">Pupina verneaui</emphasis>
Dautzenberg &amp; Fischer, 1906
</taxonomicName>
, were also recorded in the same habitat with the new species.
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5651FF917F2E022091B9FD1C" pageId="4" pageNumber="23" type="discussion">
<paragraph id="390436C75651FF917F2E022091B9FD1C" blockId="4.[809,1384,187,674]" pageId="4" pageNumber="23">
<emphasis id="0BCFEAD55651FF917F2E022096ECFD81" bold="true" box="[847,963,549,575]" pageId="4" pageNumber="23">Remarks.</emphasis>
In juvenile shells, as not yet covered by the thick glaze, the sculpture is absent or weak, the palatal plica has not appeared, and the basal area also has not expanded (
<figureCitation id="A1802A425651FF91784E028291A7FD1C" box="[1071,1160,647,674]" captionStart="FIG" captionStartId="6.[175,217,1061,1082]" captionTargetId="figure-173@6.[174,1384,189,1041]" captionTargetPageId="6" captionText="FIG. 5. Chuatienpupa megacanalis sp. nov., AC. Living specimen (paratype ZVNU.MOL 047). D. Juvenile shells. Photos: Nguyen T.S. and Do D.S. РИС. 5. Chuatienpupa megacanalis sp. nov., AC. ЖиваЯ особь (паратип ZVNU.MOL 047). D. Ювенильные раковины. Photos: Nguyen T.S. and Do D.S." pageId="4" pageNumber="23">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>