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<document id="222D299E2E45D36E67D0CE7F8D4D8109" ID-DOI="10.35885/ruthenica.2023.33(1).3" ID-ISSN="2307-7336" ID-Zenodo-Dep="11041197" ID-ZooBank="63522E75-C4DF-4DFB-992B-C1ACBC803399" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713810501483" checkinUser="felipe" docAuthor="Sang, Do Duc &amp; Son, Nguyen Thanh" docDate="2023" docId="B11287D15654FF977FC5060996D7FA24" docLanguage="en" docName="Ruthenica.33.1.19-26.pdf" docOrigin="Ruthenica, Russian Malacological Journal 33 (1)" docSource="http://dx.doi.org/10.35885/ruthenica.2023.33(1).3" docStyle="DocumentStyle:3A4A24405A53639AD762166AE048D148.1:Ruthenica.2022-.journal_article.english.index" docStyleId="3A4A24405A53639AD762166AE048D148" docStyleName="Ruthenica.2022-.journal_article.english.index" docStyleVersion="1" docTitle="Chuatienpupa Sang &amp; Son 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="21" masterDocId="4D2BFFA95655FF957C610005952FFFBE" masterDocTitle="A new genus and new species of Pupinidae (Gastropoda: Caenogastropoda) from Northern Vietnam" masterLastPageNumber="26" masterPageNumber="19" pageNumber="20" updateTime="1713810820037" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EE97D6065356E34A0835CA9D37B77DAA">A new genus and new species of Pupinidae (Gastropoda: Caenogastropoda) from Northern Vietnam</mods:title>
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<mods:namePart id="F84CA18EDD7BB47FB580AB5957960E9A">Sang, Do Duc</mods:namePart>
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<mods:namePart id="805AD8DB9DF51B21857440EA7A3F0034">Son, Nguyen Thanh</mods:namePart>
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<mods:title id="0ED7CB1467D856F9E4AFB65C7D46D4A0">Ruthenica, Russian Malacological Journal</mods:title>
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<mods:date id="41B2EA307CB9347997E3EC6FAA5AF0BA">2023</mods:date>
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<subSubSection id="71A1654C5654FF947FC5060991F2F997" box="[932,1245,1548,1577]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="390436C75654FF947FC5060991F2F997" blockId="1.[932,1245,1548,1577]" box="[932,1245,1548,1577]" pageId="1" pageNumber="20">
<heading id="624C81AB5654FF947FC5060991F2F997" box="[932,1245,1548,1577]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="20" reason="6">
<taxonomicName id="FEBB4D445654FF947FC50609914EF997" ID-CoL="B3DDM" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[932,1121,1548,1577]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55654FF947FC50609914EF997" bold="true" box="[932,1121,1548,1577]" italics="true" pageId="1" pageNumber="20">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5654FF947808060891F2F997" box="[1129,1245,1549,1577]" pageId="1" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="71A1654C5654FF947F26063B9001F904" pageId="1" pageNumber="20" type="materials_examined">
<paragraph id="390436C75654FF947F26063B967EF9C7" blockId="1.[801,1376,1597,2048]" pageId="1" pageNumber="20">
<typeStatus id="E60088655654FF947F26063B9650F9E6" box="[839,895,1598,1624]" pageId="1" pageNumber="20">Type</typeStatus>
species
<taxonomicName id="FEBB4D445654FF947F9906389018F9E6" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[1016,1335,1597,1624]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="20" phylum="Mollusca" rank="species" species="megacanalis" status="sp. nov.">
<emphasis id="0BCFEAD55654FF947F9906389018F9E6" bold="true" box="[1016,1335,1597,1624]" italics="true" pageId="1" pageNumber="20">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5654FF947921063B967EF9C7" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="390436C75654FF947F26067A9001F904" blockId="1.[801,1376,1597,2048]" pageId="1" pageNumber="20">
ZooBank registration:
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urn:lsid:zoobank.org:act:
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<subSubSection id="71A1654C5654FF977F2606C596A0FB89" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="diagnosis">
<paragraph id="390436C75654FF947F2606C591EBF964" blockId="1.[801,1376,1597,2048]" box="[839,1220,1728,1754]" pageId="1" pageNumber="20">
<emphasis id="0BCFEAD55654FF947F2606C596EEF964" bold="true" box="[839,961,1728,1754]" pageId="1" pageNumber="20">Diagnosis.</emphasis>
As in the
<typeStatus id="E60088655654FF94785506C5914BF964" box="[1076,1124,1728,1754]" pageId="1" pageNumber="20">type</typeStatus>
species.
</paragraph>
<paragraph id="390436C75654FF977F2606E5944EFB66" blockId="1.[801,1376,1597,2048]" lastBlockId="2.[174,749,1020,2047]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<emphasis id="0BCFEAD55654FF947F2606E591DAF945" bold="true" box="[839,1269,1760,1787]" pageId="1" pageNumber="20">
Differential diagnosis.
<taxonomicName id="FEBB4D445654FF94783106E591DAF945" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[1104,1269,1760,1787]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55654FF94783106E591DAF945" bold="true" box="[1104,1269,1760,1787]" italics="true" pageId="1" pageNumber="20">Chuatienpupa</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="10FC57AE5654FF94789A06E4904FF945" box="[1275,1376,1761,1787]" pageId="1" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
is most similar to the genus
<taxonomicName id="FEBB4D445654FF9478300707906AF8A5" authority="Vignard, 1829" authorityName="Vignard" authorityYear="1829" box="[1105,1349,1793,1819]" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55654FF9478300707918DF8A5" box="[1105,1186,1794,1819]" italics="true" pageId="1" pageNumber="20">Pupina</emphasis>
<bibRefCitation id="5D2A4B365654FF9478C70704906AF8A5" author="Vignard M." box="[1190,1349,1793,1819]" pageId="1" pageNumber="20" pagination="439 - 440" refId="ref5053" refString="Vignard M. 1829. Description du Maillotin (Pupina), nouveau genre de coquilles. Annales des Sciences Naturelles, 18: 439 - 440." type="journal article" year="1829">
Vignard,
<quantity id="FE439B225654FF94796C0704904FF8A5" box="[1293,1376,1793,1819]" metricMagnitude="1" metricUnit="m" metricValue="4.6456599999999995" pageId="1" pageNumber="20" unit="in" value="1829.0">1829 in</quantity>
</bibRefCitation>
</taxonomicName>
general shell and aperture shape, with the presence of canals (anterior and posterior canals). However, this closely related genus lacks the basal plica, and upper palatal plica (some
<taxonomicName id="FEBB4D445654FF947825078191C2F823" box="[1092,1261,1923,1949]" pageId="1" pageNumber="20">
<emphasis id="0BCFEAD55654FF947825078191BAF823" box="[1092,1173,1924,1949]" italics="true" pageId="1" pageNumber="20">Pupina</emphasis>
species
</taxonomicName>
possesses a homologous structure to the lower palatal plica of the new genus), which divides the posterior canal into two cavities in the new genus. In addition, the anterior canal of
<taxonomicName id="FEBB4D445657FF977D1903F894E6FBA8" authorityName="Vignard" authorityYear="1829" box="[376,457,1021,1046]" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977D1903F894E6FBA8" box="[376,457,1021,1046]" italics="true" pageId="2" pageNumber="21">Pupina</emphasis>
</taxonomicName>
is narrower and situated on the columellar side. Finally,
<taxonomicName id="FEBB4D445657FF977E750418974AFB88" authorityName="Vignard" authorityYear="1829" box="[532,613,1053,1078]" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977E750418974AFB88" box="[532,613,1053,1078]" italics="true" pageId="2" pageNumber="21">Pupina</emphasis>
</taxonomicName>
possesses a circular, thin operculum, outer surface flat and con- sists of many spirals, while
<taxonomicName id="FEBB4D445657FF977DBF045997ACFBC9" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[478,643,1116,1143]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF977DBF045997ACFBC9" bold="true" box="[478,643,1116,1143]" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF977EE9045897C2FBC9" box="[648,749,1117,1143]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
has a solid, closed bowl-shaped operculum, with flat inner, and completely smooth, dome-shaped outer surface (
<figureCitation id="A1802A425657FF977D6F04BB9460FB66" box="[270,335,1214,1240]" captionStart="FIG" captionStartId="1.[169,211,938,959]" captionText="FIG. 1. A. General shell morphology of Chuatienpupa gen. nov. and its terminology. B. Details of the anterior canal (bottom view). C. Details of the posterior canal." pageId="2" pageNumber="21">Fig. 1</figureCitation>
).
</paragraph>
<caption id="6DC4664F5657FF97783E00B8906BFE86" pageId="2" pageNumber="21" startId="2.[1119,1164,189,210]" targetType="figure">
<paragraph id="390436C75657FF97783E00B8906BFE86" blockId="2.[1119,1384,189,312]" pageId="2" pageNumber="21">
FIG. 2. Map of Vietnam showing the type local- ity (black dot) of
<taxonomicName id="FEBB4D445657FF97794800EA9047FEA1" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="species" species="megacanalis" status="gen. nov. et sp. nov.">
<emphasis id="0BCFEAD55657FF97794800EA9047FEA1" bold="true" italics="true" pageId="2" pageNumber="21">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF9778E50126906BFE86" box="[1156,1348,291,312]" pageId="2" pageNumber="21" rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="6DC4664F5657FF97783E014C91C8FE63" ID-DOI="http://doi.org/10.5281/zenodo.11041204" ID-Zenodo-Dep="11041204" httpUri="https://zenodo.org/record/11041204/files/figure.png" pageId="2" pageNumber="21" startId="2.[1119,1170,329,350]" targetBox="[184,910,200,916]" targetPageId="2" targetType="figure">
<paragraph id="390436C75657FF97783E014C91C8FE63" blockId="2.[1119,1384,329,477]" pageId="2" pageNumber="21">
РИС. 2. Карта Вьетнама с типовым местона- хождением (черный кружок)
<taxonomicName id="FEBB4D445657FF9778BA0190902DFE7A" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="species" species="megacanalis" status="gen. nov. et sp. nov.">
<emphasis id="0BCFEAD55657FF9778BA0190902DFE7A" bold="true" italics="true" pageId="2" pageNumber="21">Chuatienpupa megacanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF97796D01AA91C8FE63" pageId="2" pageNumber="21" rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="390436C75657FF977CB504DB946AF9E5" blockId="2.[174,749,1020,2047]" pageId="2" pageNumber="21">
<taxonomicName id="FEBB4D445657FF977CB504DB94E5FB47" authority="Gray, 1850" authorityName="Gray" authorityYear="1850" box="[212,458,1246,1273]" class="Gastropoda" family="Pupinidae" genus="Pupinella" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" sensu="lato">
<emphasis id="0BCFEAD55657FF977CB504DB946EFB46" box="[212,321,1246,1272]" italics="true" pageId="2" pageNumber="21">Pupinella</emphasis>
<bibRefCitation id="5D2A4B365657FF977D2A04DB94E5FB47" author="Gray J. E." box="[331,458,1246,1273]" pageId="2" pageNumber="21" refId="ref4106" refString="Gray J. E. 1850. Nomenclature of molluscous animals and shells in the collection of the British Museum. Part I. Cyclophoridae. British Museum, London, 69 pp." type="book" year="1850">Gray, 1850</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF977DB504DB94D9FB46" box="[468,502,1246,1272]" pageId="2" pageNumber="21" sensu="lato">s.l.</taxonomicNameLabel>
, is somewhat similar to the new genus due to the pupoid shell shape, and the presence of the anterior canal, but the posterior canal of that genus is absent. Species of the subge- nus
<taxonomicName id="FEBB4D445657FF977CBF055A9786FAC7" authority="H. Adams, 1866" authorityName="H. Adams" authorityYear="1866" box="[222,681,1375,1402]" class="Gastropoda" family="Pupinidae" genus="Pupinella" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="subGenus" subGenus="Pupinopsis">
<emphasis id="0BCFEAD55657FF977CBF055A9464FAC7" box="[222,331,1375,1401]" italics="true" pageId="2" pageNumber="21">Pupinella</emphasis>
(
<emphasis id="0BCFEAD55657FF977D3F056594F5FAC7" box="[350,474,1376,1401]" italics="true" pageId="2" pageNumber="21">Pupinopsis</emphasis>
) H. Adams, 1866
</taxonomicName>
(
<typeStatus id="E60088655657FF977EDA055A97C2FAC7" box="[699,749,1375,1401]" pageId="2" pageNumber="21">type</typeStatus>
species:
<taxonomicName id="FEBB4D445657FF977D6F057A9788FA24" authority="H. Adams, 1866" authorityName="H. Adams" authorityYear="1866" box="[270,679,1407,1434]" class="Gastropoda" family="Pupinidae" genus="Pupinella" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="species" species="swinhoei">
<emphasis id="0BCFEAD55657FF977D6F057A94C8FA27" box="[270,487,1407,1433]" italics="true" pageId="2" pageNumber="21">Pupinella swinhoei</emphasis>
H. Adams, 1866
</taxonomicName>
) have both canals (anterior and posterior), but differs from the new genus by having a narrower anterior canal, which is situated on the columellar side and lacking both palatal plica and basal plica, a circular, thin operculum, with outer surface flat and consisting of 4 to 5 spirals.
</paragraph>
<paragraph id="390436C75657FF977CB5066796A0FB89" blockId="2.[174,749,1020,2047]" lastBlockId="2.[809,1384,1019,1434]" pageId="2" pageNumber="21">
<taxonomicName id="FEBB4D445657FF977CB506679493F9C5" authority="Thach, 2017" authorityName="Thach" authorityYear="2017" box="[212,444,1633,1659]" class="Gastropoda" family="Pupinidae" genus="Barnaia" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977CB506679401F9C5" box="[212,302,1634,1659]" italics="true" pageId="2" pageNumber="21">Barnaia</emphasis>
<bibRefCitation id="5D2A4B365657FF977D5306649493F9C5" author="Thach N. N." box="[306,444,1633,1659]" pageId="2" pageNumber="21" refId="ref4926" refString="Thach N. N. 2017. New shells of Southeast Asia. Sea shells &amp; Land snails. 48 HrBooks Company, 128 pp." type="book" year="2017">Thach, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FEBB4D445657FF977DA4066497EFF9C2" authority="Gould, 1856" authorityName="Gould" authorityYear="1856" box="[453,704,1633,1660]" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977DA40664971BF9C5" box="[453,564,1633,1659]" italics="true" pageId="2" pageNumber="21">Pollicaria</emphasis>
<bibRefCitation id="5D2A4B365657FF977E59066497EFF9C2" author="Gould A. A." box="[568,704,1633,1660]" pageId="2" pageNumber="21" pagination="11 - 16" refId="ref4009" refString="Gould A. A. 1856. Descriptions of new species of shells. Proceedings of the Boston Society of Natural History, 6: 11 - 16." type="journal article" year="1856">Gould, 1856</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FEBB4D445657FF977CCF068794FBF922" authority="Pfeiffer, 1856" authorityName="Pfeiffer" authorityYear="1856" box="[174,468,1665,1692]" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977CCF0687941BF922" box="[174,308,1666,1692]" italics="true" pageId="2" pageNumber="21">Rhaphaulus</emphasis>
Pfeiffer, 1856
</taxonomicName>
are similar to
<taxonomicName id="FEBB4D445657FF977E19068495C8F902" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF977E19068495C8F902" bold="true" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF977C8D06A7947FF902" box="[236,336,1698,1724]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
in the pupoid shell shape, but differ in the absence of canals, as well as the lack of parietal plica, and a flat, circular operculum (
<figureCitation id="A1802A425657FF977E3906E7978DF943" box="[600,674,1762,1789]" captionStart="FIG" captionStartId="1.[169,211,938,959]" captionText="FIG. 1. A. General shell morphology of Chuatienpupa gen. nov. and its terminology. B. Details of the anterior canal (bottom view). C. Details of the posterior canal." pageId="2" pageNumber="21">Figs 1</figureCitation>
,
<figureCitation id="A1802A425657FF977ED306E697CFF943" box="[690,736,1763,1789]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="3.[168,210,1924,1945]" captionStartId-1="5.[166,208,1892,1913]" captionTargetBox-0="[168,1376,189,1900]" captionTargetBox-1="[166,1374,189,1857]" captionTargetId-0="figure-11@3.[168,1378,189,1900]" captionTargetId-1="figure-11@5.[166,1376,189,1857]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIG. 3. Chuatienpupa megacanalis sp. nov.AE. Holotype VNMN-IZ 000.002.331.FH. Paratypes ZVNU.MOL 047.Photos: Nguyen T.S. and Do D.S." captionText-1="FIG. 4.A. Pupina keraudrenii Vignard, 1829 (NMR 178592). B. Pupina anceyi Bavay et Dautzenberg, 1899 (syntype MNHN- IM-2000-35833). C. Pupinella illustris (Mabille, 1887) (lectotype of Pupina tonkiniana Bavay &amp; Dautzenberg, 1899, MNHN-IM-2000-35838). D. Pupinella mansuyi (Dautzenberg et Fischer, 1908) (syntype RBINS MT970). E. Barnaia longituba (Páll-Gergely et Gargominy, 2017) (holotype MNHN 2012-27162). F. Rhaphaulus tonkinensis Páll-Gergely, Hunyadi et Maassen, 2014 (holotype HNHM 98757, from Páll-Gergely et al., 2014)." figureDoi-0="http://doi.org/10.5281/zenodo.11041208" figureDoi-1="http://doi.org/10.5281/zenodo.11041212" httpUri-0="https://zenodo.org/record/11041208/files/figure.png" httpUri-1="https://zenodo.org/record/11041212/files/figure.png" pageId="2" pageNumber="21">34</figureCitation>
). The shell of
<taxonomicName id="FEBB4D445657FF977D5407069790F8A0" authority="Mollendorff, 1885" authorityName="Mollendorff" authorityYear="1885" box="[309,703,1795,1822]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977D54070694C1F8A3" box="[309,494,1795,1821]" italics="true" pageId="2" pageNumber="21">Pseudopomatias</emphasis>
<bibRefCitation id="5D2A4B365657FF977D9507069790F8A0" author="Mollendorff O. F. von" box="[500,703,1795,1822]" pageId="2" pageNumber="21" pagination="161 - 170" refId="ref4454" refString="Mollendorff O. F. von. 1885. Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 17 (11 - 12): 161 - 170." type="journal article" year="1885">Möllendorff, 1885</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FEBB4D445657FF977CCF07219728F880" authority="Pall-Gergely, 2015" authorityName="Pall-Gergely" authorityYear="2015" box="[174,519,1827,1854]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977CCF07219407F883" box="[174,296,1828,1853]" italics="true" pageId="2" pageNumber="21">Vargapupa</emphasis>
<bibRefCitation id="5D2A4B365657FF977D5007269728F880" author="Pall-Gergely B. &amp; Feher Z. &amp; Hunyadi A. &amp; Asami T." box="[305,519,1827,1854]" pageId="2" pageNumber="21" pagination="1 - 49" refId="ref4593" refString="Pall-Gergely B., Feher Z., Hunyadi A., Asami T. 2015. Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa, 3937 (1): 1 - 49." type="journal article" year="2015">Páll-Gergely, 2015</bibRefCitation>
</taxonomicName>
differs from that of
<taxonomicName id="FEBB4D445657FF977CCF0746947CF8E0" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[174,339,1859,1886]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF977CCF0746947CF8E0" bold="true" box="[174,339,1859,1886]" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF977D3807419490F8E0" box="[345,447,1860,1886]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
by the spindle shaped with rather regularly ribbed on shell surface, the peristome consists of two circles (most of species), but lacks anterior and posterior canals, the operculum densely coiled, thin and membranous. Both
<taxonomicName id="FEBB4D445657FF977E3807C1941AF841" authority="Gould, 1862" authorityName="Gould" authorityYear="1862" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977E3807C197C2F860" box="[601,749,1988,2014]" italics="true" pageId="2" pageNumber="21">Coptocheilus</emphasis>
<bibRefCitation id="5D2A4B365657FF977CCF07E0941AF841" author="Gould A. A." box="[174,309,2021,2047]" pageId="2" pageNumber="21" pagination="280 - 284" refId="ref4038" refString="Gould A. A. 1862. Descriptions of new genera and spe- cies of shells. Proceedings of the Boston Society of Natural History, 8: 280 - 284." type="journal article" year="1862">Gould, 1862</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FEBB4D445657FF977D0707E09765F841" authority="Gray, 1847" authorityName="Gray" authorityYear="1847" box="[358,586,2021,2047]" class="Gastropoda" family="Pupinidae" genus="Tortulosa" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="0BCFEAD55657FF977D0707E094E1F841" box="[358,462,2021,2047]" italics="true" pageId="2" pageNumber="21">Tortulosa</emphasis>
<bibRefCitation id="5D2A4B365657FF977DB207E09765F841" author="Gray J. E." box="[467,586,2021,2047]" pageId="2" pageNumber="21" pagination="129 - 219" refId="ref4070" refString="Gray J. E. 1847. A list of the genera of recent Mollusca, their synonyms and types. Proceedings of the Zoo- logical Society of London, 15: 129 - 219." type="journal article" year="1847">Gray, 1847</bibRefCitation>
</taxonomicName>
lack the canals on the peristome, which is present in
<taxonomicName id="FEBB4D445657FF9778A303FE9048FBA8" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[1218,1383,1019,1046]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF9778A303FE9048FBA8" bold="true" box="[1218,1383,1019,1046]" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF977F48041896A0FB89" box="[809,911,1053,1079]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5657FF977F2E043896D0FB47" pageId="2" pageNumber="21" type="etymology">
<paragraph id="390436C75657FF977F2E043896D0FB47" blockId="2.[809,1384,1019,1434]" pageId="2" pageNumber="21">
<emphasis id="0BCFEAD55657FF977F2E043896FFFBE9" bold="true" box="[847,976,1085,1111]" pageId="2" pageNumber="21">Etymology</emphasis>
. The name
<taxonomicName id="FEBB4D445657FF977834043991D5FBE9" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[1109,1274,1084,1111]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF977834043991D5FBE9" bold="true" box="[1109,1274,1084,1111]" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FC57AE5657FF97796004389047FBE9" box="[1281,1384,1085,1111]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
is the combination of the
<typeStatus id="E60088655657FF977839045891A7FBC9" box="[1112,1160,1117,1143]" pageId="2" pageNumber="21">type</typeStatus>
locality, where the
<typeStatus id="E60088655657FF977F48047B9676FB26" box="[809,857,1150,1176]" pageId="2" pageNumber="21">type</typeStatus>
species was found (Chua Tien pagoda, Lac Thuy district,
<collectingRegion id="FB7FF8255657FF977FB5049B9196FB09" box="[980,1209,1181,1208]" country="Vietnam" name="Hoa Binh" pageId="2" pageNumber="21">Hoa Binh province</collectingRegion>
,
<collectingCountry id="41AC76575657FF9778A804989005FB06" box="[1225,1322,1181,1208]" name="Vietnam" pageId="2" pageNumber="21">Vietnam</collectingCountry>
) and
<emphasis id="0BCFEAD55657FF977F4804BA964BFB66" box="[809,868,1215,1240]" italics="true" pageId="2" pageNumber="21">Pupa</emphasis>
(generic name that used to be used for pupoid shaped
<taxonomicName id="FEBB4D445657FF977F1F04DB96DCFB47" authorityName="Pfeiffer" authorityYear="1853" box="[894,1011,1246,1273]" class="Gastropoda" family="Pupinidae" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="71A1654C5657FF977F2E04FA96D7FA24" pageId="2" pageNumber="21" type="discussion">
<paragraph id="390436C75657FF977F2E04FA96D7FA24" blockId="2.[809,1384,1019,1434]" pageId="2" pageNumber="21">
<emphasis id="0BCFEAD55657FF977F2E04FA9140FAA7" bold="true" box="[847,1135,1278,1305]" pageId="2" pageNumber="21">
Remarks.
<taxonomicName id="FEBB4D445657FF977FAB04FB9140FAA7" authority="Sang &amp; Son, 2023" authorityName="Sang &amp; Son" authorityYear="2023" box="[970,1135,1278,1305]" class="Gastropoda" family="Pupinidae" genus="Chuatienpupa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="0BCFEAD55657FF977FAB04FB9140FAA7" bold="true" box="[970,1135,1278,1305]" italics="true" pageId="2" pageNumber="21">Chuatienpupa</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="10FC57AE5657FF97781604FA91F1FAA7" box="[1143,1246,1279,1305]" pageId="2" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
is classified in the family
<taxonomicName id="FEBB4D445657FF977FDA051A9104FA87" authorityName="Pfeiffer" authorityYear="1853" box="[955,1067,1311,1337]" class="Gastropoda" family="Pupinidae" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="21" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
due to the smooth sculpture, pupoid shaped and covered by a glaze shell, the presence of two canals at the parieto-angular corner and the basal area.
</paragraph>
</subSubSection>
</treatment>
</document>