treatments-xml/data/B1/08/8E/B1088E654E241F25C6D6FD42FF6AFE9A.xml

<document id="6A58D44DC87138A9B29F14D3F4D9B081" ID-CLB-Dataset="27947" ID-DOI="10.11646/zootaxa.4521.4.6" ID-GBIF-Dataset="609e1f9a-16f8-4821-b45b-55ea3f1b8e78" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610164" ID-ZooBank="3AD5E6D5-F35B-43DB-8D56-586E15DF9B21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553624435604" checkinUser="plazi" docAuthor="Král, David, Hillert, Oliver, Drožová, Dana &amp; Šípek, Petr" docDate="2018" docId="B1088E654E241F25C6D6FD42FF6AFE9A" docLanguage="en" docName="zootaxa.4521.4.6.pdf" docOrigin="Zootaxa 4521 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lethrus (Lethrus) thracicus Král &amp; Hillert &amp; Drožová &amp; Šípek 2018, new species" docType="treatment" docVersion="7" lastPageNumber="582" masterDocId="4D31F61D4E251F2CC641FFF2FF81FFDA" masterDocTitle="Lethrus (Lethrus) thracicus (Coleoptera: Geotrupidae), a new species from the European part of Turkey" masterLastPageNumber="583" masterPageNumber="573" pageNumber="574" updateTime="1698713797296" updateUser="ExternalLinkService">
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<mods:title id="49EF2FD97BC078C3A9AC4EB6A84D9BAB">Lethrus (Lethrus) thracicus (Coleoptera: Geotrupidae), a new species from the European part of Turkey</mods:title>
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<mods:namePart id="B9117B1CEC01A3031DFF538228127920">Král, David</mods:namePart>
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<mods:namePart id="6B80A85DC89057BE6402D30D18780CEF">Drožová, Dana</mods:namePart>
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<mods:namePart id="697E25BA8827E66C5578FCD9132C03F1">Šípek, Petr</mods:namePart>
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<taxonomicName id="FEA144F04E241F2DC6D6FD42FD25FD11" ID-CoL="63YCT" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[151,676,688,715]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Lethrus">
<emphasis id="0BD5E3614E241F2DC6D6FD42FF74FD10" bold="true" box="[151,245,688,714]" italics="true" pageId="1" pageNumber="574">Lethrus</emphasis>
(
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) 
<emphasis id="0BD5E3614E241F2DC737FD42FE67FD10" bold="true" box="[374,486,688,714]" italics="true" pageId="1" pageNumber="574">thracicus</emphasis>
Král &amp; Hillert
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, 
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(
<figureCitation id="A19A23F64E241F2DC6DEFD26FF69FD36" box="[159,232,724,749]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="1" pageNumber="574">Figs. 1</figureCitation>
a–e; 2d, h; 3d, h; 4d, h; 5d; 6; 7)
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<materialsCitation id="89C9352E4E241F2DC6D6FCEEFEA6FC82" ID-GBIF-Occurrence="2028521106" country="Turkey" elevation="121" latitude="41.674435" location="Habiller" longLatPrecision="1" longitude="26.786764" municipality="European" pageId="1" pageNumber="574" specimenCount="1" stateProvince="Edirne" typeStatus="holotype">
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<typeStatus id="E61A81D14E241F2DC6D6FCEEFF53FCEF" box="[151,210,796,821]" pageId="1" pageNumber="574">Type</typeStatus>
locality
</emphasis>
. 
<collectingMunicipality id="D97AA5094E241F2DC77DFCEFFE26FCEE" box="[316,423,797,820]" pageId="1" pageNumber="574">European</collectingMunicipality>
<collectingCountry id="41B67FE34E241F2DC7EEFCEEFD81FCEF" box="[431,512,796,821]" name="Turkey" pageId="1" pageNumber="574">Turkey</collectingCountry>
, 
<collectingRegion id="FB65F1914E241F2DC44DFCEEFDD6FCEF" box="[524,599,796,821]" country="Turkey" name="Edirne" pageId="1" pageNumber="574">Edirne</collectingRegion>
prov., 
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E of 
<collectingRegion id="FB65F1914E241F2DC56BFCEEFCF9FCEF" box="[810,888,796,821]" country="Turkey" name="Edirne" pageId="1" pageNumber="574">Edirne</collectingRegion>
, 
<location id="3C7E69A84E241F2DC5C5FCEEFC61FCEF" LSID="urn:lsid:plazi:treatment:B1088E654E241F25C6D6FD42FF6AFE9A:3C7E69A84E241F2DC5C5FCEEFC61FCEF" box="[900,992,796,821]" country="Turkey" latitude="41.674435" longLatPrecision="1" longitude="26.786764" municipality="European" name="Habiller" pageId="1" pageNumber="574" stateProvince="Edirne">Habiller</location>
env., 
<geoCoordinate id="5C9559B44E241F2DC262FCEEFB4BFCEF" box="[1059,1226,796,821]" direction="north" orientation="latitude" pageId="1" pageNumber="574" precision="1" value="41.674435">41°40'27.96&quot;N</geoCoordinate>
<geoCoordinate id="5C9559B44E241F2DC290FCEEFAF9FCEF" box="[1233,1400,796,821]" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="1" value="26.786764">26°47'12.35&quot;E</geoCoordinate>
, ca 
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<quantity id="FE5992964E241F2DC6D6FCB2FF5FFC8D" box="[151,222,832,856]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="574" unit="m" value="121.0">121 m</quantity>
a. s. l.
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</paragraph>
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<collectingCounty id="D07F47FF4E241F2DC686FC96FE83FCA7" box="[199,258,868,893]" pageId="1" pageNumber="574">Type</collectingCounty>
material
</emphasis>
. 
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<collectingCounty id="D07F47FF4E241F2DC7C9FC96FE7FFCA7" box="[392,510,868,893]" pageId="1" pageNumber="574">European</collectingCounty>
<materialsCitation id="89C9352E4E241F2DC44AFC96FC9AFCA7" ID-GBIF-Occurrence="2028521099" box="[523,795,868,893]" collectionCode="DKCP" country="Turkey" county="Prov." location="Prov." pageId="1" pageNumber="574" specimenCount="1" specimenCount-male="1" stateProvince="Edirne" typeStatus="holotype">
<collectingCountry id="41B67FE34E241F2DC44AFC96FDE0FCA7" box="[523,609,868,893]" name="Turkey" pageId="1" pageNumber="574">Turkey</collectingCountry>
, 
<collectingRegion id="FB65F1914E241F2DC435FC96FD46FCA7" box="[628,711,868,893]" country="Turkey" name="Edirne" pageId="1" pageNumber="574">Edirne</collectingRegion>
<collectingCounty id="D07F47FF4E241F2DC495FC96FC97FCA7" box="[724,790,868,893]" pageId="1" pageNumber="574">Prov.</collectingCounty>
:
</materialsCitation>
</emphasis>
<typeStatus id="E61A81D14E241F2DC568FC96FC15FCA7" box="[809,916,868,893]" pageId="1" pageNumber="574" type="holotype">Holotype</typeStatus>
, 
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(DKCP), 
<materialsCitation id="89C9352E4E241F2DC27BFC96FA9CFC7A" ID-GBIF-Occurrence="2028521086" collectingDate="2011-04-26" collectorName="P. Jansta &amp; D. Drozova" country="Turkey" county="Prov." elevation="121" latitude="41.674435" location="Habiller" longLatPrecision="1" longitude="26.786764" municipality="European Turkey" pageId="1" pageNumber="574" specimenCount="1" stateProvince="Edirne" typeStatus="holotype">
“ 
<collectingMunicipality id="D97AA5094E241F2DC205FC96FA91FCA7" box="[1092,1296,868,893]" pageId="1" pageNumber="574">European Turkey</collectingMunicipality>
, 
<collectingDate id="5D5BE05B4E241F2DC360FC96FA1BFCA7" box="[1313,1434,868,893]" pageId="1" pageNumber="574" value="2011-04-26">
<date id="4D1F19B34E241F2DC360FC96FA1BFCA7" box="[1313,1434,868,893]" pageId="1" pageNumber="574" value="2011-04-26">26.iv.2011</date>
</collectingDate>
| 
<location id="3C7E69A84E241F2DC6D6FC75FF72FC7A" LSID="urn:lsid:plazi:treatment:B1088E654E241F25C6D6FD42FF6AFE9A:3C7E69A84E241F2DC6D6FC75FF72FC7A" box="[151,243,903,928]" country="Turkey" county="Prov." latitude="41.674435" longLatPrecision="1" longitude="26.786764" municipality="European Turkey" name="Habiller" pageId="1" pageNumber="574" stateProvince="Edirne">Habiller</location>
env. | ca 
<elevation id="B28CD8404E241F2DC719FC7AFE20FC45" box="[344,417,904,928]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="574" unit="m" value="121.0">
<quantity id="FE5992964E241F2DC719FC7AFE20FC45" box="[344,417,904,928]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="574" unit="m" value="121.0">121 m</quantity>
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, 
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<geoCoordinate id="5C9559B44E241F2DC418FC75FD7CFC7A" box="[601,765,903,928]" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="1" value="26.786764">26°47'12.35&quot;E</geoCoordinate>
| 
<collectorName id="94545AA54E241F2DC551FC7AFCF0FC7A" box="[784,881,904,928]" pageId="1" pageNumber="574">P. Janšta</collectorName>
, 
<collectorName id="94545AA54E241F2DC53DFC7AFC7CFC7A" box="[892,1021,904,928]" pageId="1" pageNumber="574">D. Drožová</collectorName>
&amp; P. Šípek lgt. [printed]”
</materialsCitation>
. 
<materialsCitation id="89C9352E4E241F2DC366FC7AFE60FC1E" ID-GBIF-Occurrence="2028521088" collectionCode="DKCP" country="Turkey" county="Prov." pageId="1" pageNumber="574" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Edirne" typeStatus="paratype">
<typeStatus id="E61A81D14E241F2DC366FC7AFA19FC7A" box="[1319,1432,904,928]" pageId="1" pageNumber="574" type="paratype">Paratypes</typeStatus>
: 
<typeStatus id="E61A81D14E241F2DC6D6FC5EFF75FC1E" box="[151,244,940,964]" pageId="1" pageNumber="574" type="allotype">allotype</typeStatus>
, 
<specimenCount id="2FA7F4FA4E241F2DC741FC5EFE94FC19" box="[256,277,940,963]" pageId="1" pageNumber="574" type="female">♀</specimenCount>
and 
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(
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)
</materialsCitation>
, 
<materialsCitation id="89C9352E4E241F2DC7AFFC5EFD7BFC1E" ID-GBIF-Occurrence="2028521122" box="[494,762,940,964]" collectionCode="OHCB" country="Turkey" county="Prov." isTailEnumeration="true" pageId="1" pageNumber="574" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Edirne" typeStatus="paratype">
<specimenCount id="2FA7F4FA4E241F2DC7AFFC5EFDAFFC19" box="[494,558,940,964]" pageId="1" pageNumber="574" type="male">2 ♂♂</specimenCount>
and 
<specimenCount id="2FA7F4FA4E241F2DC427FC5EFD10FC19" box="[614,657,940,964]" pageId="1" pageNumber="574" type="female">1 ♀</specimenCount>
(
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)
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<materialsCitation id="89C9352E4E241F2DC543FC5EFC5BFC1E" ID-GBIF-Occurrence="2028521092" box="[770,986,940,964]" collectionCode="CUPC" country="Turkey" county="Prov." pageId="1" pageNumber="574" specimenCount="2" specimenCount-male="2" stateProvince="Edirne" typeStatus="paratype">
and 
<specimenCount id="2FA7F4FA4E241F2DC572FC5EFCF2FC19" box="[819,883,940,964]" pageId="1" pageNumber="574" type="male">2 ♂♂</specimenCount>
(
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)
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, 
<materialsCitation id="89C9352E4E241F2DC5A7FC5EFA19FC1E" ID-GBIF-Occurrence="2028521084" box="[998,1432,940,964]" collectionCode="DKCP" country="Turkey" county="Prov." location="Prov." pageId="1" pageNumber="574" specimenCount="3" specimenCount-male="3" stateProvince="Edirne" typeStatus="paratype">
same data as holotype; 
<specimenCount id="2FA7F4FA4E241F2DC2B1FC5EFAB1FC19" box="[1264,1328,940,964]" pageId="1" pageNumber="574" type="male">3 ♂♂</specimenCount>
(
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)
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, 
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“TR [= 
<collectingMunicipality id="D97AA5094E241F2DC6BCFC3DFEC9FC32" box="[253,328,975,1000]" pageId="1" pageNumber="574">Turcia</collectingMunicipality>
]—ca 15 
<location id="3C7E69A84E241F2DC7FDFC22FE67FC3D" LSID="urn:lsid:plazi:treatment:B1088E654E241F25C6D6FD42FF6AFE9A:3C7E69A84E241F2DC7FDFC22FE67FC3D" box="[444,486,976,999]" country="Turkey" county="Prov." latitude="41.666668" longLatPrecision="1155" longitude="26.666666" municipality="Turcia" name="Km" pageId="1" pageNumber="574" stateProvince="Edirne">Km</location>
| e.[east of] 
<collectingRegion id="FB65F1914E241F2DC4CEFC3DFD5BFC32" box="[655,730,975,1000]" country="Turkey" name="Edirne" pageId="1" pageNumber="574">Edirne</collectingRegion>
[ca 
<geoCoordinate id="5C9559B44E241F2DC558FC22FCF7FC32" box="[793,886,976,1000]" direction="north" orientation="latitude" pageId="1" pageNumber="574" precision="925" value="41.666668">41°40'N</geoCoordinate>
<geoCoordinate id="5C9559B44E241F2DC5C5FC3DFC60FC32" box="[900,993,975,1000]" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="925" value="26.666666">26°40'E</geoCoordinate>
] | 27.3.[19]88 [Walter] Heinz [legit] [handwritten, black ink]”
</materialsCitation>
.
</paragraph>
<paragraph id="391E3F734E241F2DC686FBE5FE44FB8F" blockId="1.[151,1438,796,2009]" pageId="1" pageNumber="574">
<emphasis id="0BD5E3614E241F2DC686FBE5FC64FBEA" bold="true" box="[199,997,1047,1072]" pageId="1" pageNumber="574">
<collectingCounty id="D07F47FF4E241F2DC686FBE5FEC6FBEA" box="[199,327,1047,1072]" pageId="1" pageNumber="574">Additional</collectingCounty>
material examined. 
<collectingCounty id="D07F47FF4E241F2DC402FBE5FD38FBEA" box="[579,697,1047,1072]" pageId="1" pageNumber="574">European</collectingCounty>
<materialsCitation id="89C9352E4E241F2DC483FBE5FC64FBEA" ID-GBIF-Occurrence="2028521132" box="[706,997,1047,1072]" collectingDate="1987-04" collectionCode="NMPC" country="Turkey" county="Prov." latitude="40.983334" location="Prov." longLatPrecision="1159" longitude="27.5" pageId="1" pageNumber="574" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tekirdag">
<collectingCountry id="41B67FE34E241F2DC483FBE5FC99FBEA" box="[706,792,1047,1072]" name="Turkey" pageId="1" pageNumber="574">Turkey</collectingCountry>
, 
<collectingRegion id="FB65F1914E241F2DC569FBE5FC15FBEA" box="[808,916,1047,1072]" country="Turkey" name="Tekirdag" pageId="1" pageNumber="574">Tekirdağ</collectingRegion>
<collectingCounty id="D07F47FF4E241F2DC5DCFBE5FC61FBEA" box="[925,992,1047,1072]" pageId="1" pageNumber="574">Prov.</collectingCounty>
:
</materialsCitation>
</emphasis>
<specimenCount id="2FA7F4FA4E241F2DC5AFFBEAFB9CFBF5" box="[1006,1053,1048,1072]" pageId="1" pageNumber="574" type="male">1 ♂</specimenCount>
, 
<specimenCount id="2FA7F4FA4E241F2DC26DFBEAFBD8FBF5" box="[1068,1113,1048,1072]" pageId="1" pageNumber="574" type="female">1 ♀</specimenCount>
(NMPC), 
<collectingRegion id="FB65F1914E241F2DC294FBE5FAB9FBEA" box="[1237,1336,1047,1072]" country="Turkey" name="Tekirdag" pageId="1" pageNumber="574">Tekirdağ</collectingRegion>
env. [ca 
<geoCoordinate id="5C9559B44E241F2DC6D6FBCEFF75FB8F" box="[151,244,1084,1109]" direction="north" orientation="latitude" pageId="1" pageNumber="574" precision="925" value="40.983334">40°59'N</geoCoordinate>
<geoCoordinate id="5C9559B44E241F2DC6BBFBCEFED8FB8E" box="[250,345,1084,1108]" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="925" value="27.5">27°30'E</geoCoordinate>
], 
<collectingDate id="5D5BE05B4E241F2DC72AFBCEFE40FB8F" box="[363,449,1084,1109]" pageId="1" pageNumber="574" value="1987-04">
<date id="4D1F19B34E241F2DC72AFBCEFE40FB8F" box="[363,449,1084,1109]" pageId="1" pageNumber="574" value="1987-04">iv.1987</date>
</collectingDate>
.
</paragraph>
</subSubSection>
<subSubSection id="71BB6CF84E241F2AC686FBADFED5F876" lastPageId="6" lastPageNumber="579" pageId="1" pageNumber="574" type="description">
<paragraph id="391E3F734E241F2DC686FBADFA0FFB1A" blockId="1.[151,1438,796,2009]" pageId="1" pageNumber="574">
<emphasis id="0BD5E3614E241F2DC686FBADFE5FFBA2" bold="true" box="[199,478,1119,1144]" pageId="1" pageNumber="574">
Description of 
<typeStatus id="E61A81D14E241F2DC739FBADFE5FFBA2" box="[376,478,1119,1144]" pageId="1" pageNumber="574" type="holotype">holotype</typeStatus>
</emphasis>
(
<specimenCount id="2FA7F4FA4E241F2DC7B1FB92FD86FBA2" box="[496,519,1120,1144]" pageId="1" pageNumber="574" type="male">♂</specimenCount>
). Maximally developed (hyperthelic) male with well developed ventral mandible processes (
<figureCitation id="A19A23F64E241F2DC750FB76FE20FB46" box="[273,417,1156,1181]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="1" pageNumber="574">Figs. 1a, c, e</figureCitation>
). Oblong, strongly convex; dorsal surface black, except moderately shiny pronotum almost alutaceous; ventral surface black with fine blue tinge, moderately shiny, claws black-brown; macrosetation black.
</paragraph>
<paragraph id="391E3F734E241F2DC686FB3EFF68F997" blockId="1.[151,1438,796,2009]" pageId="1" pageNumber="574">
<emphasis id="0BD5E3614E241F2DC686FB3EFE85FB3E" box="[199,260,1228,1252]" italics="true" pageId="1" pageNumber="574">Head</emphasis>
(
<figureCitation id="A19A23F64E241F2DC752FB3EFE27FB3E" box="[275,422,1228,1253]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="1" pageNumber="574">Figs. 1a, c, e</figureCitation>
). Labrum bilobed, asymmetrical, right lobe remarkably more developed; surface rugosely and coarsely, shallowly and sparsely punctate, each puncture bearing short recumbent macroseta; anterior margin with dense row of long macrosetae. Clypeus transverse, trapezoidal with anterior angles rounded. Frontal impressions vague, frontal tubercles indistinct. Frontoclypeal suture present only laterally; keels separating eye canthus from frons only slightly developed but distinct, slightly divergent posteriad. Eye canthus exceeding eyes, projecting anterolaterad, almost rectangular, lateral margins divergent posteriad, anterolateral angle rounded, oblique keel above eyes absent. Pleurostomal process evenly arcuate, hardly exceeding ventrolateral mandible outline. Punctation of frons double, consisting of coarse, transversally rugose, regularly and densely distributed punctures, intermixed with fine, irregularly distributed ones; coarse punctures separated by approximately less than their diameter, punctation becoming distinctly sparser posteriad and on occiput; clypeus and eye canthus distinctly rugose.
</paragraph>
<paragraph id="391E3F734E241F2DC686F9AAFC0FF94F" blockId="1.[151,1438,796,2009]" pageId="1" pageNumber="574">
<emphasis id="0BD5E3614E241F2DC686F9AAFEBCF9AA" box="[199,317,1624,1648]" italics="true" pageId="1" pageNumber="574">Mandibles</emphasis>
symmetrical, external outline almost semicircular, pointed subapically in dorsal aspect (
<figureCitation id="A19A23F64E241F2DC35FF9A5FA12F9AA" box="[1310,1427,1623,1648]" captionStart="FIGURES 2" captionStartId="3.[151,263,1909,1931]" captionTargetBox="[214,1374,222,1893]" captionTargetId="figure@3.[214,1377,193,1893]" captionTargetPageId="3" captionText="FIGURES 2a–h. Male head in dorsal view. a, e—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b, f—L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c, g—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d, h—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." httpUri="https://zenodo.org/record/2610168/files/figure.png" pageId="1" pageNumber="574">Figs. 2a, e</figureCitation>
) with maximum width approximately at middle of mandibles length.
</paragraph>
<paragraph id="391E3F734E241F2DC686F952FEA1F803" blockId="1.[151,1438,796,2009]" pageId="1" pageNumber="574">
<emphasis id="0BD5E3614E241F2DC686F952FD84F962" box="[199,517,1696,1720]" italics="true" pageId="1" pageNumber="574">Ventral mandible processes</emphasis>
(
<figureCitation id="A19A23F64E241F2DC45BF96DFD21F962" box="[538,672,1695,1720]" captionStart="FIGURES 2" captionStartId="3.[151,263,1909,1931]" captionTargetBox="[214,1374,222,1893]" captionTargetId="figure@3.[214,1377,193,1893]" captionTargetPageId="3" captionText="FIGURES 2a–h. Male head in dorsal view. a, e—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b, f—L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c, g—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d, h—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." httpUri="https://zenodo.org/record/2610168/files/figure.png" pageId="1" pageNumber="574">Figs. 2d, h</figureCitation>
; 
<figureCitation id="A19A23F64E241F2DC4F0F96DFD4EF962" box="[689,719,1695,1720]" captionStart="FIGURES 3" captionStartId="4.[151,263,1916,1938]" captionTargetBox="[205,1377,224,1895]" captionTargetId="figure@4.[197,1389,193,1895]" captionTargetPageId="4" captionText="FIGURES 3a–h. Male head in frontal view. a, e—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b, f—L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c, g—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d, h—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." httpUri="https://zenodo.org/record/2610170/files/figure.png" pageId="1" pageNumber="574">3d</figureCitation>
, 
<figureCitation id="A19A23F64E241F2DC4A0F96DFD70F962" box="[737,753,1695,1720]" captionStart="FIGURES 5" captionStartId="6.[151,263,1097,1119]" captionText="FIGURES 5a–d. Aedeagi in dorsal and lateral aspect. a—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b— L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." pageId="1" pageNumber="574">5</figureCitation>
; 
<figureCitation id="A19A23F64E241F2DC545F96DFCC3F962" box="[772,834,1695,1720]" captionStart="FIGURES 4" captionStartId="5.[151,263,1916,1938]" captionTargetBox="[218,1374,244,1889]" captionTargetId="figure@5.[211,1376,193,1895]" captionTargetPageId="5" captionText="FIGURES 4a–h. Male head in left lateral view. a, e—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b, f—L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c, g—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d, h—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." httpUri="https://zenodo.org/record/2610172/files/figure.png" pageId="1" pageNumber="574">4d, h</figureCitation>
) weakly asymmetrical, right process slightly more developed than left one and with different angle in lateral aspect. Both processes distinctly longer than length of mandible; base thickened, not exceeding lateral mandibular outline in dorsal aspect, with slightly concave external outline in basal half in frontal aspect; longitudinal keel on base laterally present, straight and approximately parallel to lateral mandibular outline in dorsal aspect, distinctly broader as maximum width of mandibles outline basally; in lateral aspect weakly arcuate, approximately subparallel to lateral mandibular outline, slightly divergent gradually basad approximately from middle of its length. Inferiobasal tooth absent; both processes bent inward aproximately in middle of mandibles length in frontal view; anterior subapical tooth absent; apical emargination absent; apical tooth round.
</paragraph>
<caption id="6DDE6FFB4E271F2EC6D6F86EFC4CF835" httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="2" pageNumber="575" startId="2.[151,263,1948,1970]" targetBox="[221,1370,217,1869]" targetPageId="2">
<paragraph id="391E3F734E271F2EC6D6F86EFC4CF835" blockId="2.[151,1436,1948,2031]" pageId="2" pageNumber="575">
<emphasis id="0BD5E3614E271F2EC6D6F86EFEC8F868" bold="true" box="[151,329,1948,1970]" pageId="2" pageNumber="575">FIGURES 1a–e.</emphasis>
<taxonomicName id="FEA144F04E271F2EC715F86EFC98F868" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[340,793,1948,1970]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Lethrus">
<emphasis id="0BD5E3614E271F2EC715F86EFE20F86B" box="[340,417,1948,1969]" italics="true" pageId="2" pageNumber="575">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E271F2EC7F5F86EFD80F86B" box="[436,513,1948,1969]" italics="true" pageId="2" pageNumber="575">Lethrus</emphasis>
) 
<emphasis id="0BD5E3614E271F2EC454F86EFDF2F86B" box="[533,627,1948,1969]" italics="true" pageId="2" pageNumber="575">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E271F2EC566F86FFC24F868" box="[807,933,1948,1970]" pageId="2" pageNumber="575" rank="species">new species</taxonomicNameLabel>
(European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale.
</paragraph>
</caption>
<caption id="6DDE6FFB4E261F2FC6D6F887FDB3F83D" httpUri="https://zenodo.org/record/2610168/files/figure.png" pageId="3" pageNumber="576" startId="3.[151,263,1909,1931]" targetBox="[214,1374,222,1893]" targetPageId="3">
<paragraph id="391E3F734E261F2FC6D6F887FDB3F83D" blockId="3.[151,1436,1909,2023]" pageId="3" pageNumber="576">
<emphasis id="0BD5E3614E261F2FC6D6F887FEC9F851" bold="true" box="[151,328,1909,1931]" pageId="3" pageNumber="576">FIGURES 2a–h.</emphasis>
Male head in dorsal view. a, e— 
<taxonomicName id="FEA144F04E261F2FC4D2F887FCDFF851" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[659,862,1909,1931]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E261F2FC4D2F887FD61F850" box="[659,736,1909,1930]" italics="true" pageId="3" pageNumber="576">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E261F2FC4AEF884FC85F851" box="[751,772,1910,1931]" italics="true" pageId="3" pageNumber="576">L.</emphasis>
) 
<emphasis id="0BD5E3614E261F2FC553F884FCDFF851" box="[786,862,1910,1931]" italics="true" pageId="3" pageNumber="576">apterus</emphasis>
</taxonomicName>
(Slovakia, Kamenica nad Hronom, DKCP), b, f— 
<taxonomicName id="FEA144F04E261F2FC31DF884FF43F873" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E261F2FC31DF884FAF0F851" box="[1372,1393,1910,1931]" italics="true" pageId="3" pageNumber="576">L.</emphasis>
(
<emphasis id="0BD5E3614E261F2FC33EF884FA15F851" box="[1407,1428,1910,1931]" italics="true" pageId="3" pageNumber="576">L.</emphasis>
) 
<emphasis id="0BD5E3614E261F2FC6D6F866FF43F873" box="[151,194,1940,1961]" italics="true" pageId="3" pageNumber="576">ares</emphasis>
</taxonomicName>
(Greece, Evros dist., Polía, holotype, NMPC); c, g— 
<taxonomicName id="FEA144F04E261F2FC4A0F866FC10F873" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[737,913,1940,1962]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E261F2FC4A0F866FD77F873" box="[737,758,1940,1961]" italics="true" pageId="3" pageNumber="576">L.</emphasis>
(
<emphasis id="0BD5E3614E261F2FC545F866FC98F873" box="[772,793,1940,1961]" italics="true" pageId="3" pageNumber="576">L.</emphasis>
) 
<emphasis id="0BD5E3614E261F2FC566F866FC10F873" box="[807,913,1940,1961]" italics="true" pageId="3" pageNumber="576">schneideri</emphasis>
</taxonomicName>
(Greece: Thrace distr., Karydia, holotype, NMPC); d, h— 
<emphasis id="0BD5E3614E261F2FC69BF841FF69F812" box="[218,232,1971,1992]" italics="true" pageId="3" pageNumber="576">L</emphasis>
. 
<taxonomicName id="FEA144F04E261F2FC6B8F840FDAAF812" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[249,555,1970,1992]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E261F2FC740F841FE8EF812" box="[257,271,1971,1992]" italics="true" pageId="3" pageNumber="576">L</emphasis>
.) 
<emphasis id="0BD5E3614E261F2FC769F841FE07F812" box="[296,390,1971,1992]" italics="true" pageId="3" pageNumber="576">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E261F2FC478F841FD37F812" box="[569,694,1970,1992]" pageId="3" pageNumber="576" rank="species">new species</taxonomicNameLabel>
(European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale.
</paragraph>
</caption>
<caption id="6DDE6FFB4E211F28C6D6F88EFDB3F834" httpUri="https://zenodo.org/record/2610170/files/figure.png" pageId="4" pageNumber="577" startId="4.[151,263,1916,1938]" targetBox="[205,1377,224,1895]" targetPageId="4">
<paragraph id="391E3F734E211F28C6D6F88EFDB3F834" blockId="4.[151,1436,1916,2030]" pageId="4" pageNumber="577">
<emphasis id="0BD5E3614E211F28C6D6F88EFEC6F848" bold="true" box="[151,327,1916,1938]" pageId="4" pageNumber="577">FIGURES 3a–h.</emphasis>
Male head in frontal view. a, e— 
<taxonomicName id="FEA144F04E211F28C4D7F88EFCE1F848" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[662,864,1916,1938]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E211F28C4D7F88EFD62F84B" box="[662,739,1916,1937]" italics="true" pageId="4" pageNumber="577">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E211F28C4B3F88FFC86F848" box="[754,775,1917,1938]" italics="true" pageId="4" pageNumber="577">L.</emphasis>
) 
<emphasis id="0BD5E3614E211F28C555F88FFCE1F848" box="[788,864,1917,1938]" italics="true" pageId="4" pageNumber="577">apterus</emphasis>
</taxonomicName>
(Slovakia, Kamenica nad Hronom, DKCP), b, f— 
<taxonomicName id="FEA144F04E211F28C31DF88FFF43F86B" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E211F28C31DF88FFAF3F848" box="[1372,1394,1917,1938]" italics="true" pageId="4" pageNumber="577">L.</emphasis>
(
<emphasis id="0BD5E3614E211F28C33EF88FFA15F848" box="[1407,1428,1917,1938]" italics="true" pageId="4" pageNumber="577">L.</emphasis>
) 
<emphasis id="0BD5E3614E211F28C6D6F86EFF43F86B" box="[151,194,1948,1969]" italics="true" pageId="4" pageNumber="577">ares</emphasis>
</taxonomicName>
(Greece, Evros dist., Polía, holotype, NMPC); c, g— 
<taxonomicName id="FEA144F04E211F28C4A0F86EFC10F86A" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[737,913,1947,1969]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E211F28C4A0F86EFD77F86B" box="[737,758,1948,1969]" italics="true" pageId="4" pageNumber="577">L.</emphasis>
(
<emphasis id="0BD5E3614E211F28C545F86EFC98F86B" box="[772,793,1948,1969]" italics="true" pageId="4" pageNumber="577">L.</emphasis>
) 
<emphasis id="0BD5E3614E211F28C566F869FC10F86A" box="[807,913,1947,1968]" italics="true" pageId="4" pageNumber="577">schneideri</emphasis>
</taxonomicName>
(Greece: Thrace distr., Karydia, holotype, NMPC); d, h— 
<emphasis id="0BD5E3614E211F28C69BF848FF69F815" box="[218,232,1978,1999]" italics="true" pageId="4" pageNumber="577">L</emphasis>
. 
<taxonomicName id="FEA144F04E211F28C6B8F84BFDAAF815" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[249,555,1977,1999]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E211F28C740F848FE8EF815" box="[257,271,1978,1999]" italics="true" pageId="4" pageNumber="577">L</emphasis>
.) 
<emphasis id="0BD5E3614E211F28C769F848FE07F815" box="[296,390,1978,1999]" italics="true" pageId="4" pageNumber="577">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E211F28C478F848FD37F815" box="[569,694,1977,1999]" pageId="4" pageNumber="577" rank="species">new species</taxonomicNameLabel>
(European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale.
</paragraph>
</caption>
<caption id="6DDE6FFB4E201F29C6D6F88EFD2FF835" httpUri="https://zenodo.org/record/2610172/files/figure.png" pageId="5" pageNumber="578" startId="5.[151,263,1916,1938]" targetBox="[218,1374,244,1889]" targetPageId="5">
<paragraph id="391E3F734E201F29C6D6F88EFD2FF835" blockId="5.[151,1437,1916,2031]" pageId="5" pageNumber="578">
<emphasis id="0BD5E3614E201F29C6D6F88EFEC9F848" bold="true" box="[151,328,1916,1938]" pageId="5" pageNumber="578">FIGURES 4a–h.</emphasis>
Male head in left lateral view. a, e— 
<taxonomicName id="FEA144F04E201F29C4FCF88FFC09F848" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[701,904,1916,1938]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E201F29C4FCF88FFC8BF848" box="[701,778,1917,1938]" italics="true" pageId="5" pageNumber="578">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E201F29C558F88FFCAFF848" box="[793,814,1917,1938]" italics="true" pageId="5" pageNumber="578">L.</emphasis>
) 
<emphasis id="0BD5E3614E201F29C57DF88FFC09F848" box="[828,904,1917,1938]" italics="true" pageId="5" pageNumber="578">apterus</emphasis>
</taxonomicName>
(Slovakia, Kamenica nad Hronom, DKCP), b, f— 
<taxonomicName id="FEA144F04E201F29C3C6F88FFF70F86B" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E201F29C3C6F88FFA1CF848" box="[1415,1437,1917,1938]" italics="true" pageId="5" pageNumber="578">L.</emphasis>
(
<emphasis id="0BD5E3614E201F29C6DEF86EFF35F86B" box="[159,180,1948,1969]" italics="true" pageId="5" pageNumber="578">L.</emphasis>
) 
<emphasis id="0BD5E3614E201F29C687F86EFF70F86B" box="[198,241,1948,1969]" italics="true" pageId="5" pageNumber="578">ares</emphasis>
</taxonomicName>
(Greece, Evros dist., Polía, holotype, NMPC); c, g— 
<taxonomicName id="FEA144F04E201F29C568F86EFC5EF86A" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[809,991,1947,1969]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E201F29C568F86EFCBFF86B" box="[809,830,1948,1969]" italics="true" pageId="5" pageNumber="578">L.</emphasis>
(
<emphasis id="0BD5E3614E201F29C50EF86EFCE5F86B" box="[847,868,1948,1969]" italics="true" pageId="5" pageNumber="578">L.</emphasis>
) 
<emphasis id="0BD5E3614E201F29C534F869FC5EF86A" box="[885,991,1947,1968]" italics="true" pageId="5" pageNumber="578">schneideri</emphasis>
</taxonomicName>
(Greece: Thrace distr., Karydia, holotype, NMPC); d, h— 
<emphasis id="0BD5E3614E201F29C77EF849FECCF80A" box="[319,333,1979,2000]" italics="true" pageId="5" pageNumber="578">L</emphasis>
. 
<taxonomicName id="FEA144F04E201F29C721F848FD1BF80A" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[352,666,1978,2000]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E201F29C729F849FEF7F80A" box="[360,374,1979,2000]" italics="true" pageId="5" pageNumber="578">L</emphasis>
.) 
<emphasis id="0BD5E3614E201F29C7D3F848FE71F815" box="[402,496,1978,1999]" italics="true" pageId="5" pageNumber="578">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E201F29C4EBF849FCA8F80A" box="[682,809,1978,2000]" pageId="5" pageNumber="578" rank="species">new species</taxonomicNameLabel>
(European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale.
</paragraph>
</caption>
<caption id="6DDE6FFB4E231F2AC6D6FBBBFDB3FB60" pageId="6" pageNumber="579">
<paragraph id="391E3F734E231F2AC6D6FBBBFDB3FB60" blockId="6.[151,1436,1097,1210]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC6D6FBBBFEC6FB85" bold="true" box="[151,327,1097,1119]" pageId="6" pageNumber="579">FIGURES 5a–d.</emphasis>
Aedeagi in dorsal and lateral aspect. a— 
<taxonomicName id="FEA144F04E231F2AC4A7FBBBFC30FB85" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[742,945,1097,1119]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E231F2AC4A7FBBBFCB2FB84" box="[742,819,1097,1118]" italics="true" pageId="6" pageNumber="579">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E231F2AC503FBB8FCD6FB85" box="[834,855,1098,1119]" italics="true" pageId="6" pageNumber="579">L.</emphasis>
) 
<emphasis id="0BD5E3614E231F2AC524FBB8FC30FB85" box="[869,945,1098,1119]" italics="true" pageId="6" pageNumber="579">apterus</emphasis>
</taxonomicName>
(Slovakia, Kamenica nad Hronom, DKCP), b— 
<taxonomicName id="FEA144F04E231F2AC6D6FB9AFE8EFBA7" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[151,271,1127,1149]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E231F2AC6D6FB9AFF2DFBA7" box="[151,172,1128,1149]" italics="true" pageId="6" pageNumber="579">L.</emphasis>
(
<emphasis id="0BD5E3614E231F2AC6FCFB9AFF53FBA7" box="[189,210,1128,1149]" italics="true" pageId="6" pageNumber="579">L.</emphasis>
) 
<emphasis id="0BD5E3614E231F2AC6A5FB9AFE8EFBA7" box="[228,271,1128,1149]" italics="true" pageId="6" pageNumber="579">ares</emphasis>
</taxonomicName>
(Greece, Evros dist., Polía, holotype, NMPC); c— 
<taxonomicName id="FEA144F04E231F2AC568FB9AFC5EFBA7" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[809,991,1127,1149]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E231F2AC568FB9AFCBFFBA7" box="[809,830,1128,1149]" italics="true" pageId="6" pageNumber="579">L.</emphasis>
(
<emphasis id="0BD5E3614E231F2AC511FB9AFCE4FBA7" box="[848,869,1128,1149]" italics="true" pageId="6" pageNumber="579">L.</emphasis>
) 
<emphasis id="0BD5E3614E231F2AC537FB9AFC5EFBA7" box="[886,991,1128,1149]" italics="true" pageId="6" pageNumber="579">schneideri</emphasis>
</taxonomicName>
(Greece: Thrace distr., Karydia, holotype, NMPC); d— 
<emphasis id="0BD5E3614E231F2AC75BFB75FEA9FB46" box="[282,296,1159,1180]" italics="true" pageId="6" pageNumber="579">L</emphasis>
. 
<taxonomicName id="FEA144F04E231F2AC775FB74FDD7FB46" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[308,598,1158,1180]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E231F2AC77CFB75FECAFB46" box="[317,331,1159,1180]" italics="true" pageId="6" pageNumber="579">L</emphasis>
.) 
<emphasis id="0BD5E3614E231F2AC721FB74FE3FFB41" box="[352,446,1158,1179]" italics="true" pageId="6" pageNumber="579">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E231F2AC41EFB75FD58FB46" box="[607,729,1158,1180]" pageId="6" pageNumber="579" rank="species">new species</taxonomicNameLabel>
(European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale.
</paragraph>
</caption>
<paragraph id="391E3F734E231F2AC686FB1AFCC5FA6E" blockId="6.[151,1436,1255,2036]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686FB1AFEB7FB25" box="[199,310,1256,1279]" italics="true" pageId="6" pageNumber="579">Pronotum</emphasis>
(
<figureCitation id="A19A23F64E231F2AC704FB15FE52FADA" box="[325,467,1255,1280]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="6" pageNumber="579">Figs. 1a, c, e</figureCitation>
) transverse, distinctly broader than base of elytra, broadest just behind middle; margin entirely bordered, slightly crenulate in anterior parts. Anterior corners weakly but distinctly projecting anterolaterad, with angulate outline; lateral margin approximately weakly emarginate anteriorly, than straight to rounded posterior angle; basal margin straight. Punctation of dorsal surface simple, consisting of deep, sparsely and irregularly distributed punctures; punctures separated by approximately two to four their diameters discally, surface near lateral margins considerably shagreened and alutaceous.
</paragraph>
<paragraph id="391E3F734E231F2AC686FA4DFC8CFA02" blockId="6.[151,1436,1255,2036]" box="[199,781,1471,1496]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686FA4DFEEAFA0D" box="[199,363,1471,1495]" italics="true" pageId="6" pageNumber="579">Scutellar plate</emphasis>
widely triangular, finely shagreened.
</paragraph>
<paragraph id="391E3F734E231F2AC686FA16FD5DF99E" blockId="6.[151,1436,1255,2036]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686FA16FE8CFA26" box="[199,269,1508,1532]" italics="true" pageId="6" pageNumber="579">Elytra</emphasis>
almost semicircular, apices not prominent, each apex forming independent arc. Epipleuron strongly narrowed apicad, epipleural keel not reaching elytral apex. Whole surface alutaceous, finely transversally rugose; striae not indicated, entirely vanishing in rugosities.
</paragraph>
<paragraph id="391E3F734E231F2AC686F9A2FD06F956" blockId="6.[151,1436,1255,2036]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F9A2FF7DF9BD" box="[199,252,1616,1639]" italics="true" pageId="6" pageNumber="579">Legs</emphasis>
. Profemur not armed, protibia with row of eight gradually proximad diminishing external denticles, and with row of tubercles on ventromedial edge.
</paragraph>
<paragraph id="391E3F734E231F2AC686F965FA92F96A" blockId="6.[151,1436,1255,2036]" box="[199,1299,1687,1712]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F965FEF2F975" box="[199,371,1687,1711]" italics="true" pageId="6" pageNumber="579">Male genitalia.</emphasis>
Aedeagus with parameres shallowly sinuate distally in lateral aspect (
<figureCitation id="A19A23F64E231F2AC2C5F965FB61F96A" box="[1156,1248,1687,1712]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="6" pageNumber="579">Figs. 1b</figureCitation>
, 
<figureCitation id="A19A23F64E231F2AC2ACF965FA8BF96A" box="[1261,1290,1687,1712]" captionStart="FIGURES 5" captionStartId="6.[151,263,1097,1119]" captionText="FIGURES 5a–d. Aedeagi in dorsal and lateral aspect. a—Lethrus (L.) apterus (Slovakia, Kamenica nad Hronom, DKCP), b— L. (L.) ares (Greece, Evros dist., Polía, holotype, NMPC); c—L. (L.) schneideri (Greece: Thrace distr., Karydia, holotype, NMPC); d—L. (L.) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype). Differential characters shown by arrow. Not to scale." pageId="6" pageNumber="579">5d</figureCitation>
)
</paragraph>
<paragraph id="391E3F734E231F2AC686F949FAC1F90E" blockId="6.[151,1436,1255,2036]" box="[199,1344,1723,1748]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F949FEFCF90E" bold="true" box="[199,381,1723,1748]" pageId="6" pageNumber="579">Measurements.</emphasis>
Total body length 
<quantity id="FE5992964E231F2AC411F94EFD4AF909" box="[592,715,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="6" pageNumber="579" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">18–24 mm</quantity>
, holotype— 
<quantity id="FE5992964E231F2AC516F949FC27F909" box="[855,934,1723,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="579" unit="mm" value="23.0">23 mm</quantity>
, allotype— 
<quantity id="FE5992964E231F2AC269F949FBF7F909" box="[1064,1142,1723,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="579" unit="mm" value="24.0">24 mm</quantity>
(
<figureCitation id="A19A23F64E231F2AC2C4F949FAB3F90E" box="[1157,1330,1723,1748]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="6" pageNumber="579">Figs. 1a, c, d, e</figureCitation>
).
</paragraph>
<paragraph id="391E3F734E231F2AC686F92DFC01F8C6" blockId="6.[151,1436,1255,2036]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F92DFE31F922" bold="true" box="[199,432,1759,1784]" pageId="6" pageNumber="579">Variability in males</emphasis>
. Mandible processes in medium developed and underdeveloped (hypothelic) males short, more or less straight with simply rounded to almost acute apically.
</paragraph>
<paragraph id="391E3F734E231F2AC686F8D5FED5F876" blockId="6.[151,1436,1255,2036]" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F8D5FE3AF89A" bold="true" box="[199,443,1831,1856]" pageId="6" pageNumber="579">Sexual dimorphism.</emphasis>
Female (
<figureCitation id="A19A23F64E231F2AC46AF8D5FDFFF89A" box="[555,638,1831,1856]" captionStart="FIGURES 1" captionStartId="2.[151,263,1948,1970]" captionTargetBox="[221,1370,217,1869]" captionTargetId="figure@2.[203,1390,195,1924]" captionTargetPageId="2" captionText="FIGURES 1a–e. Lethrus (Lethrus) thracicus Král &amp; Hillert, new species (European Turkey, Edirne prov., Habiller env., holotype (male), excepting allotype (female)—d). a—habitus, left lateral aspect; b—aedeagus, dorsal and left lateral view; c, d—habitus dorsal view; e—habitus, left oblique antero-lateral view. Not to scale." httpUri="https://zenodo.org/record/2610166/files/figure.png" pageId="6" pageNumber="579">Fig. 1d</figureCitation>
) differ from males as follows: external outline of mandibles almost straight, in apical quarter rounded in dorsal aspect; ventral mandibular process absent; pronotum anterior corners more angulate, pronotum lateral margins broadly rounded, protibia broader, row of tubercles on ventromedial edge less pronounced.
</paragraph>
</subSubSection>
<subSubSection id="71BB6CF84E231F2BC686F845FAACFD17" lastPageId="7" lastPageNumber="580" pageId="6" pageNumber="579" type="diagnosis">
<paragraph id="391E3F734E231F2BC686F845FAACFD17" blockId="6.[151,1436,1255,2036]" lastBlockId="7.[151,1437,151,933]" lastPageId="7" lastPageNumber="580" pageId="6" pageNumber="579">
<emphasis id="0BD5E3614E231F2AC686F845FE44F80A" bold="true" box="[199,453,1975,2000]" pageId="6" pageNumber="579">Differential diagnosis</emphasis>
. Among the species distributed in the Balkan Peninsula, the new species is most similar and probably closely related to 
<taxonomicName id="FEA144F04E231F2AC7B9F82EFDCCF82E" authorityName="Scopoli" authorityYear="1777" box="[504,589,2012,2036]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="0BD5E3614E231F2AC7B9F82EFDCCF82E" box="[504,589,2012,2036]" italics="true" pageId="6" pageNumber="579">Lethrus</emphasis>
</taxonomicName>
<taxonomicName id="FEA144F04E231F2AC414F82EFBA4F82E" authority="Kral, Rejsek &amp; Schneider, 2001" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[597,1061,2011,2036]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E231F2AC41FF82EFDECF829" box="[606,621,2012,2035]" italics="true" pageId="6" pageNumber="579">L</emphasis>
.) 
<emphasis id="0BD5E3614E231F2AC4C4F82EFD35F829" box="[645,692,2012,2035]" italics="true" pageId="6" pageNumber="579">ares</emphasis>
<bibRefCitation id="5D3042824E231F2AC4FDF829FBA4F82E" author="Kral, D. &amp; Rejsek, J. &amp; Schneider, J." box="[700,1061,2011,2036]" pageId="6" pageNumber="579" pagination="253 - 260" refId="ref2826" refString="Kral, D., Rejsek, J. &amp; Schneider, J. (2001) Lethrus (Lethrus) ares sp. n. (Coleoptera: Geotrupidae) from Greece. Klapalekiana, 37, 253 - 260." type="journal article" year="2001">Král, Rejsek &amp; Schneider, 2001</bibRefCitation>
</taxonomicName>
, 
<emphasis id="0BD5E3614E231F2AC271F82EFBBEF829" box="[1072,1087,2012,2035]" italics="true" pageId="6" pageNumber="579">L</emphasis>
. 
<taxonomicName id="FEA144F04E231F2AC20CF82EFA1DF82E" authority="(Laxmann, 1770)" authorityName="Laxmann" authorityYear="1770" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[1101,1436,2012,2036]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E231F2AC216F82EFBE7F829" box="[1111,1126,2012,2035]" italics="true" pageId="6" pageNumber="579">L</emphasis>
.) 
<emphasis id="0BD5E3614E231F2AC23FF82EFB53F829" box="[1150,1234,2012,2035]" italics="true" pageId="6" pageNumber="579">apterus</emphasis>
(Laxmann, 1770)
</taxonomicName>
and 
<emphasis id="0BD5E3614E221F2BC688FF6BFF59FF6A" box="[201,216,153,176]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. 
<taxonomicName id="FEA144F04E221F2BC6ABFF6AFD0FFF6A" authority="Kral &amp; Hillert, 2013" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[234,654,151,176]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC6B2FF6BFE83FF6A" box="[243,258,153,176]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC75CFF6AFE13FF6A" box="[285,402,152,176]" italics="true" pageId="7" pageNumber="580">schneideri</emphasis>
Král &amp; Hillert, 2013
</taxonomicName>
. Distinguishing features in males are as follow: absence of anterior subbasal tooth of ventral mandibular processes (
<emphasis id="0BD5E3614E221F2BC485FF4FFD52FF0E" box="[708,723,189,212]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC4AFFF4FFD7CFF0E" box="[750,765,189,212]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC556FF4EFCFDFF0E" box="[791,892,188,212]" italics="true" pageId="7" pageNumber="580">schaumii</emphasis>
Reitter, 1890 and 
<emphasis id="0BD5E3614E221F2BC217FF4FFBE4FF0E" box="[1110,1125,189,212]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC23EFF4FFB0FFF0E" box="[1151,1166,189,212]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC2E8FF4EFB7FFF0E" box="[1193,1278,188,212]" italics="true" pageId="7" pageNumber="580">elephas</emphasis>
Reitter, 1890 have distinct anterior subbasal tooth); absence of anterior subapical tooth of ventral mandibular processes (
<emphasis id="0BD5E3614E221F2BC315FF13FAE2FF22" box="[1364,1379,225,248]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC33DFF13FA0AFF22" box="[1404,1419,225,248]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC6D6FEF6FEB0FEC6" box="[151,305,260,284]" italics="true" pageId="7" pageNumber="580">halkidikiensis</emphasis>
Hillert &amp; Král, 2013, 
<emphasis id="0BD5E3614E221F2BC470FEF7FDC1FEC6" box="[561,576,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC41BFEF7FDE8FEC6" box="[602,617,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC4C3FEF7FD42FEC6" box="[642,707,261,284]" italics="true" pageId="7" pageNumber="580">perun</emphasis>
Hillert &amp; Král, 2013, 
<emphasis id="0BD5E3614E221F2BC585FEF7FC52FEC6" box="[964,979,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC5ADFEF7FC7AFEC6" box="[1004,1019,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC255FEF6FBFEFEC6" box="[1044,1151,260,284]" italics="true" pageId="7" pageNumber="580">raymondi</emphasis>
Reitter, 1890 and 
<emphasis id="0BD5E3614E221F2BC315FEF7FAE2FEC6" box="[1364,1379,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC33DFEF7FA0AFEC6" box="[1404,1419,261,284]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC6D6FEDAFEA8FEE5" box="[151,297,296,319]" italics="true" pageId="7" pageNumber="580">strymonensis</emphasis>
Hillert &amp; Král, 2013 have distinct anterior subapical tooth); presence of approximately symmetrical ventral mandibular processes and regularly rounded or obtuse-angular anterior pronotal angles (
<emphasis id="0BD5E3614E221F2BC345FEBFFA92FEBE" box="[1284,1299,333,356]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC371FEBFFABEFEBE" box="[1328,1343,333,356]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC31DFEBEFA1DFEBE" box="[1372,1436,332,356]" italics="true" pageId="7" pageNumber="580">fallax</emphasis>
Nikolajev, 1975 and 
<emphasis id="0BD5E3614E221F2BC7C7FE83FE14FE52" box="[390,405,369,392]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. 
<taxonomicName id="FEA144F04E221F2BC7E4FE82FD31FE52" authority="Pittino, 2011" authorityName="Pittino" authorityYear="2011" box="[421,688,367,392]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="liviae" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC7EEFE83FE3FFE52" box="[431,446,369,392]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC799FE82FD97FE52" box="[472,534,368,392]" italics="true" pageId="7" pageNumber="580">liviae</emphasis>
<bibRefCitation id="5D3042824E221F2BC45EFE9DFD31FE52" author="Pittino, R." box="[543,688,367,392]" pageId="7" pageNumber="580" pagination="381 - 394" refId="ref2992" refString="Pittino, R. (2011) Lethrus (Lethrus) liviae n. sp. from European Turkey (Coleoptera Geotrupidae). Giornale Italiano di Entomologia, 12, 381 - 394." type="journal article" year="2011">Pittino, 2011</bibRefCitation>
</taxonomicName>
have remarkably asymmetrical ventral mandibular processes and strongly produced acute-angular anterior pronotal angle). For characters to separate 
<emphasis id="0BD5E3614E221F2BC209FE67FBD6FE76" box="[1096,1111,405,428]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. 
<taxonomicName id="FEA144F04E221F2BC227FE66FB6AFE76" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[1126,1259,404,428]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC22EFE67FBFFFE76" box="[1135,1150,405,428]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC2D6FE67FB6AFE76" box="[1175,1259,405,428]" italics="true" pageId="7" pageNumber="580">apterus</emphasis>
</taxonomicName>
, 
<taxonomicName id="FEA144F04E221F2BC2B8FE67FAF6FE76" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[1273,1399,404,428]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E221F2BC2B8FE67FA89FE76" box="[1273,1288,405,428]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC361FE67FAAEFE76" box="[1312,1327,405,428]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC309FE67FAF6FE76" box="[1352,1399,405,428]" italics="true" pageId="7" pageNumber="580">ares</emphasis>
</taxonomicName>
, 
<emphasis id="0BD5E3614E221F2BC3C4FE67FA15FE76" box="[1413,1428,405,428]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC6E1FE4BFF2EFE0A" box="[160,175,441,464]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) s 
<emphasis id="0BD5E3614E221F2BC692FE4AFEBCFE0A" box="[211,317,440,464]" italics="true" pageId="7" pageNumber="580">chneideri</emphasis>
and 
<emphasis id="0BD5E3614E221F2BC734FE4BFE05FE0A" box="[373,388,441,464]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. 
<taxonomicName id="FEA144F04E221F2BC7D5FE4AFD5BFE0A" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[404,730,439,464]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC7DCFE4BFE2DFE0A" box="[413,428,441,464]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC787FE4AFDAFFE0A" box="[454,558,440,464]" italics="true" pageId="7" pageNumber="580">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E221F2BC4A7FE4AFCF0FE0A" box="[742,881,439,464]" pageId="7" pageNumber="580" rank="species">new species</taxonomicNameLabel>
, see the character matrix (
<tableCitation id="74230AC84E221F2BC2DAFE45FB71FE0A" box="[1179,1264,439,464]" captionStart="TABLE 1" captionStartId="8.[147,224,550,570]" captionText="TABLE 1. Character matrix for separation of males of Lethrus (L.) apterus, L. (L.) ares, L. (L.) schneideri and L. (L.) thracicus Král &amp; Hillert, new species species character Lethrus (L.) apterus Lethrus (L.) ares Lethrus (L.) schneideri Lethrus (L.) thracicus shape of mandibles in lateral outline angulate at middle lateral outline semicircular (Figs. 2b, f) lateral outline semicircular (Figs. 2c, g) lateral outline angulate at middle (Figs. dorsal view (Figs. 2a, e) 2d, h) lateral longitudinal keel on straight and approximately parallel to straight and distinctly subparallel to straight and distinctly subparallel to straight and distinctly subparallel to base of ventral mandible lateral mandibular outline, distinctly lateral mandibular outline, approximately lateral mandibular outline, approximately lateral mandibular outline, approximately process in dorsal aspect broader as maximum width of so broad as maximum width of so broad as maximum width of so broad as maximum width of mandibles outline basally (Figs. 2a, e) mandibles outline basally (Figs.2 b, f) mandibles outline basally (Figs. 2 c, g) mandibles outline basally (Figs. 2d, h)" pageId="7" pageNumber="580">Table 1</tableCitation>
). Additionally, these four species exhibit probably a strictly allopatric distribution. 
<taxonomicName id="FEA144F04E221F2BC5EFFE2EFADFFE2F" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[942,1374,476,501]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="582" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="0BD5E3614E221F2BC5EFFE2EFB85FE2E" box="[942,1028,476,500]" italics="true" pageId="7" pageNumber="580">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E221F2BC259FE2FFBA6FE2E" box="[1048,1063,477,500]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC203FE2EFB2BFE2E" box="[1090,1194,476,500]" italics="true" pageId="7" pageNumber="580">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E221F2BC32CFE2FFF69FDC2" pageId="7" pageNumber="580" rank="species">new species</taxonomicNameLabel>
is restricted to the European part of 
<collectingCountry id="41B67FE34E221F2BC4E8FE0DFD78FDC2" box="[681,761,511,536]" name="Turkey" pageId="7" pageNumber="580">Turkey</collectingCountry>
(it means the lowland east of the Maritza river); 
<emphasis id="0BD5E3614E221F2BC311FDF3FADEFDC2" box="[1360,1375,513,536]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC33DFDF3FA0AFDC2" box="[1404,1419,513,536]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) s 
<emphasis id="0BD5E3614E221F2BC6E3FDD6FE8DFDE6" box="[162,268,548,572]" italics="true" pageId="7" pageNumber="580">chneideri</emphasis>
is an endemic species of the southernmost slopes of the Rhodope Mountains approximately between the towns of Xánthi and Komotiní (
<collectingCountry id="41B67FE34E221F2BC450FDBAFDE6FDBA" box="[529,615,584,608]" name="Greece" pageId="7" pageNumber="580">Greece</collectingCountry>
); 
<taxonomicName id="FEA144F04E221F2BC43FFDBBFC82FDBA" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[638,771,584,608]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E221F2BC43FFDBBFD0CFDBA" box="[638,653,585,608]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC4E8FDBBFD39FDBA" box="[681,696,585,608]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC495FDBBFC82FDBA" box="[724,771,585,608]" italics="true" pageId="7" pageNumber="580">ares</emphasis>
</taxonomicName>
is known so far only from four spots all situated in the Eridropótamos river basin (
<collectingCountry id="41B67FE34E221F2BC795FD9EFDA4FD5E" box="[468,549,620,644]" name="Greece" pageId="7" pageNumber="580">Greece</collectingCountry>
and 
<collectingCountry id="41B67FE34E221F2BC422FD9EFD44FD5F" box="[611,709,620,645]" name="Bulgaria" pageId="7" pageNumber="580">Bulgaria</collectingCountry>
) and 
<emphasis id="0BD5E3614E221F2BC54BFD9FFC98FD5E" box="[778,793,621,644]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. 
<taxonomicName id="FEA144F04E221F2BC56AFD9EFC32FD5E" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[811,947,620,644]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC575FD9FFCC2FD5E" box="[820,835,621,644]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC51EFD9FFC32FD5E" box="[863,947,621,644]" italics="true" pageId="7" pageNumber="580">apterus</emphasis>
</taxonomicName>
is a widely distributed Pannonian species known from 
<collectingRegion id="FB65F1914E221F2BC768FD7DFE2AFD72" box="[297,427,655,680]" country="Austria" name="Burgenland" pageId="7" pageNumber="580">Burgenland</collectingRegion>
(
<collectingCountry id="41B67FE34E221F2BC7FAFD7DFD92FD72" box="[443,531,655,680]" name="Austria" pageId="7" pageNumber="580">Austria</collectingCountry>
), 
<collectingRegion id="FB65F1914E221F2BC466FD7DFD07FD72" box="[551,646,655,680]" country="Czech Republic" name="Jihomoravsky" pageId="7" pageNumber="580">Moravia</collectingRegion>
(
<collectingCountry id="41B67FE34E221F2BC4D4FD7DFCCDFD72" box="[661,844,655,680]" name="Czech Republic" pageId="7" pageNumber="580">Czech Republic</collectingCountry>
) and 
<collectingCountry id="41B67FE34E221F2BC5CAFD7DFC52FD72" box="[907,979,655,680]" name="Serbia" pageId="7" pageNumber="580">Serbia</collectingCountry>
in the west to the Don river basin in the east (see e.g., Král 
<emphasis id="0BD5E3614E221F2BC72DFD47FE1FFD16" box="[364,414,692,716]" italics="true" pageId="7" pageNumber="580">et al</emphasis>
. 2013, 
<bibRefCitation id="5D3042824E221F2BC7B3FD46FD23FD16" author="Nikolajev, G. V." box="[498,674,692,717]" pageId="7" pageNumber="580" refId="ref2868" refString="Nikolajev, G. V. (2003) Zhuki-kravchiki (Scarabaeidae, Geotrupinae, Lethrini): biologiya, sistematika, rasprostraneniye, opredelitel' [Coleoptera, Scarabaeidae, Geotrupinae, Lethrini: biology, taxonomy, distribution, key]. Kazak universiteti, Almaty, 254 pp. [in Russian]" type="book" year="2003">Nikolajev 2003</bibRefCitation>
, 
<bibRefCitation id="5D3042824E221F2BC4ECFD47FC72FD17" author="Bagaturov, M. F. &amp; Nikolajev, G. V." box="[685,1011,692,717]" pageId="7" pageNumber="580" pagination="303 - 314" refId="ref2534" refString="Bagaturov, M. F. &amp; Nikolajev, G. V. (2015) Obzor rasprostraneniya zhukov-kravchikov roda Lethrus, 1777 (Coleoptera: Geotrupidae). Overview of distribution of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae). Kavkazskiy Entomologicheskiy Byulleten, 11, 303 - 314. [in Russian, English abstract]" type="journal article" year="2015">Bagaturov &amp; Nikolajev 2015</bibRefCitation>
and 
<bibRefCitation id="5D3042824E221F2BC26BFD46FAA0FD17" author="Chehlarov, E. &amp; Gueorguiev, B. &amp; Hristovski, S. &amp; Fancello, L. &amp; Cvetkovska-Gorgievska, A. &amp; Prelic, D." box="[1066,1313,692,717]" pageId="7" pageNumber="580" pagination="331 - 338" refId="ref2596" refString="Chehlarov, E., Gueorguiev, B., Hristovski, S., Fancello, L., Cvetkovska-Gorgievska, A. &amp; Prelic, D. (2016) New Country Records and Rare and Interesting Species of Coleoptera from the Balkan Peninsula. Acta Zoologica Bulgarica, 68, 331 - 338." type="journal article" year="2016">
Chehlarov 
<emphasis id="0BD5E3614E221F2BC2E4FD47FB5CFD16" box="[1189,1245,692,716]" italics="true" pageId="7" pageNumber="580">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="71BB6CF84E221F2BC686FD25FBF6FCA5" pageId="7" pageNumber="580" type="etymology">
<paragraph id="391E3F734E221F2BC686FD25FE0FFCCF" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="580">
<emphasis id="0BD5E3614E221F2BC686FD25FEC7FD2A" bold="true" box="[199,326,727,752]" pageId="7" pageNumber="580">Etymology</emphasis>
. Toponymic; the specific name 
<taxonomicName id="FEA144F04E221F2BC480FD2AFCA8FD2A" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[705,809,728,752]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="thracicus">
<emphasis id="0BD5E3614E221F2BC480FD2AFCA8FD2A" box="[705,809,728,752]" italics="true" pageId="7" pageNumber="580">thracicus</emphasis>
</taxonomicName>
reflects the Thracian lowlands region where the new species was collected.
</paragraph>
<paragraph id="391E3F734E221F2BC686FCEDFBF6FCA5" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="580">
<emphasis id="0BD5E3614E221F2BC686FCEDFE12FCE2" bold="true" box="[199,403,799,824]" pageId="7" pageNumber="580">Collecting events</emphasis>
. All specimens originating from the 
<typeStatus id="E61A81D14E221F2BC577FCD2FCE7FCE2" box="[822,870,800,824]" pageId="7" pageNumber="580">type</typeStatus>
locality were collected from an uncultivated field margin and / or pasture with a disturbed surface mostly in places with very sparse herbaceous layer (
<figureCitation id="A19A23F64E221F2BC2BFFCB6FAC2FC86" box="[1278,1347,836,861]" captionStart="FIGURE 7" captionStartId="9.[151,249,1268,1290]" captionTargetBox="[151,1436,593,1246]" captionTargetId="figure@9.[151,1436,375,1246]" captionTargetPageId="9" captionText="FIGURE 7. Type locality of Lethrus (L.) thracicus Král &amp; Hillert, new species. European Turkey, Edirne prov., 18 km E of Edirne, Habiller env., 26.iv.2011 (photo by Petr Šípek)." httpUri="https://zenodo.org/record/2610176/files/figure.png" pageId="7" pageNumber="580">Fig. 7</figureCitation>
). It was a sunny day and some specimens were observed to be active on the surface at 11–12 am.
</paragraph>
</subSubSection>
<subSubSection id="71BB6CF84E221F2BC686FC7EFD0AFC7F" box="[199,651,908,933]" pageId="7" pageNumber="580" type="distribution">
<paragraph id="391E3F734E221F2BC686FC7EFD0AFC7F" blockId="7.[151,1437,151,933]" box="[199,651,908,933]" pageId="7" pageNumber="580">
<emphasis id="0BD5E3614E221F2BC686FC7EFED8FC7F" bold="true" box="[199,345,908,933]" pageId="7" pageNumber="580">Distribution</emphasis>
. European 
<collectingCountry id="41B67FE34E221F2BC798FC7EFDA8FC7F" box="[473,553,908,933]" name="Turkey" pageId="7" pageNumber="580">Turkey</collectingCountry>
(
<figureCitation id="A19A23F64E221F2BC478FC7EFDFCFC7F" box="[569,637,908,933]" captionStart="FIGURE 6" captionStartId="7.[151,249,1917,1939]" captionTargetBox="[151,1436,985,1896]" captionTargetId="figure@7.[151,1436,985,1896]" captionTargetPageId="7" captionText="FIGURE 6. Sketch map of the Thracian lowlands (parts of Bulgaria, Greece and European Turkey) with marked distribution of Lethrus (L.) ares—red marks, L. (L.) schneideri—yellow marks, L. (L.) thracicus Král &amp; Hillert, new species —blue marks; triangles represent the type localities. Compiled from Král et al. (2001, 2013) and Chehlarov et al. (2016)." httpUri="https://zenodo.org/record/2610174/files/figure.png" pageId="7" pageNumber="580">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6DDE6FFB4E221F2BC6D6F88FFB4DF80B" httpUri="https://zenodo.org/record/2610174/files/figure.png" pageId="7" pageNumber="580" startId="7.[151,249,1917,1939]" targetBox="[151,1436,985,1896]" targetPageId="7">
<paragraph id="391E3F734E221F2BC6D6F88FFB4DF80B" blockId="7.[151,1436,1917,2001]" pageId="7" pageNumber="580">
<emphasis id="0BD5E3614E221F2BC6D6F88FFE8DF849" bold="true" box="[151,268,1917,1939]" pageId="7" pageNumber="580">FIGURE 6</emphasis>
. Sketch map of the Thracian lowlands (parts of Bulgaria, Greece and European Turkey) with marked distribution of 
<taxonomicName id="FEA144F04E221F2BC6D6F86EFF65F86B" authorityName="Scopoli" authorityYear="1777" box="[151,228,1948,1969]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="0BD5E3614E221F2BC6D6F86EFF65F86B" box="[151,228,1948,1969]" italics="true" pageId="7" pageNumber="580">Lethrus</emphasis>
</taxonomicName>
(
<emphasis id="0BD5E3614E221F2BC6B5F86FFE83F868" box="[244,258,1949,1970]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC758F86FFEDBF868" box="[281,346,1949,1970]" italics="true" pageId="7" pageNumber="580">ares—</emphasis>
red marks, 
<taxonomicName id="FEA144F04E221F2BC78EF86FFD03F86B" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[463,642,1948,1970]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E221F2BC78EF86FFE5CF868" box="[463,477,1949,1970]" italics="true" pageId="7" pageNumber="580">L</emphasis>
. (
<emphasis id="0BD5E3614E221F2BC7B5F86FFD83F868" box="[500,514,1949,1970]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC458F86EFD03F86B" box="[537,642,1948,1969]" italics="true" pageId="7" pageNumber="580">schneideri</emphasis>
</taxonomicName>
—yellow marks, L. 
<taxonomicName id="FEA144F04E221F2BC513F86EFBFAF868" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[850,1147,1948,1970]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E221F2BC51AF86FFCE8F868" box="[859,873,1949,1970]" italics="true" pageId="7" pageNumber="580">L</emphasis>
.) 
<emphasis id="0BD5E3614E221F2BC53EF86EFC5CF86B" box="[895,989,1948,1969]" italics="true" pageId="7" pageNumber="580">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E221F2BC2C6F86FFA83F868" box="[1159,1282,1948,1970]" pageId="7" pageNumber="580" rank="species">new species</taxonomicNameLabel>
—blue marks; triangles represent the type localities. Compiled from Král 
<emphasis id="0BD5E3614E221F2BC4ADF84EFC99F80A" box="[748,792,1979,2001]" italics="true" pageId="7" pageNumber="580">et al</emphasis>
. (2001, 2013) and Chehlarov 
<emphasis id="0BD5E3614E221F2BC209F84EFBF4F80A" box="[1096,1141,1979,2001]" italics="true" pageId="7" pageNumber="580">et al</emphasis>
. (2016).
</paragraph>
</caption>
<caption id="6DDE6FFB4E2D1F24C6D2FDD4F8D1FCD0" pageId="8" pageNumber="581">
<paragraph id="391E3F734E2D1F24C6D2FDD4F8D1FCD0" blockId="8.[147,1900,549,778]" pageId="8" pageNumber="581">
<emphasis id="0BD5E3614E2D1F24C6D2FDD4FF77FDE0" bold="true" box="[147,246,550,570]" pageId="8" pageNumber="581">TABLE 1.</emphasis>
Character matrix for separation of males of 
<taxonomicName id="FEA144F04E2D1F24C4C7FDD7FCBDFDE0" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[646,828,549,570]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E2D1F24C4C7FDD7FD4AFDE0" box="[646,715,549,570]" italics="true" pageId="8" pageNumber="581">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E2D1F24C498FDD7FD67FDE0" box="[729,742,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
.) 
<emphasis id="0BD5E3614E2D1F24C4B9FDD7FCBDFDE0" box="[760,828,549,570]" italics="true" pageId="8" pageNumber="581">apterus</emphasis>
</taxonomicName>
, 
<taxonomicName id="FEA144F04E2D1F24C506FDD7FC2CFDE0" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[839,941,549,570]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="ares" subGenus="Lethrus">
<emphasis id="0BD5E3614E2D1F24C506FDD7FCD5FDE0" box="[839,852,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
. (
<emphasis id="0BD5E3614E2D1F24C527FDD7FCF2FDE0" box="[870,883,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
.) 
<emphasis id="0BD5E3614E2D1F24C5C7FDD7FC2CFDE0" box="[902,941,549,570]" italics="true" pageId="8" pageNumber="581">ares</emphasis>
</taxonomicName>
, 
<taxonomicName id="FEA144F04E2D1F24C5F9FDD7FBD4FDE0" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[952,1109,549,570]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="schneideri" subGenus="Lethrus">
<emphasis id="0BD5E3614E2D1F24C5F9FDD7FC44FDE0" box="[952,965,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
. (
<emphasis id="0BD5E3614E2D1F24C596FDD7FC65FDE0" box="[983,996,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
.) 
<emphasis id="0BD5E3614E2D1F24C5B6FDD7FBD4FDE0" box="[1015,1109,549,570]" italics="true" pageId="8" pageNumber="581">schneideri</emphasis>
</taxonomicName>
and 
<emphasis id="0BD5E3614E2D1F24C2C1FDD7FB15FDE0" box="[1152,1172,549,570]" italics="true" pageId="8" pageNumber="581">L.</emphasis>
<taxonomicName id="FEA144F04E2D1F24C2D9FDD4FA1BFDE0" authority="Kral &amp; Hillert" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[1176,1434,549,570]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="581" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov." subGenus="Lethrus">
(
<emphasis id="0BD5E3614E2D1F24C2E1FDD7FB2DFDE0" box="[1184,1196,549,570]" italics="true" pageId="8" pageNumber="581">L</emphasis>
.) 
<emphasis id="0BD5E3614E2D1F24C2FEFDD7FA92FDE0" box="[1215,1299,549,570]" italics="true" pageId="8" pageNumber="581">thracicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="10E65E1A4E2D1F24C3E5FDD4F991FDE0" box="[1444,1552,550,570]" pageId="8" pageNumber="581" rank="species">new species</taxonomicNameLabel>
species character 
<taxonomicName id="FEA144F04E2D1F24C7D3FDBFFDC9FDB8" baseAuthorityName="Laxmann" baseAuthorityYear="1770" box="[402,584,589,610]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="apterus" subGenus="Lethrus">
<emphasis id="0BD5E3614E2D1F24C7D3FDBFFE56FDB8" box="[402,471,589,610]" italics="true" pageId="8" pageNumber="581">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E2D1F24C7A4FDBFFE70FDB8" box="[485,497,589,610]" italics="true" pageId="8" pageNumber="581">L</emphasis>
.) 
<emphasis id="0BD5E3614E2D1F24C445FDBFFDC9FDB8" box="[516,584,589,610]" italics="true" pageId="8" pageNumber="581">apterus</emphasis>
</taxonomicName>
<taxonomicName id="FEA144F04E2D1F24C4B9FDBFFC11FDB8" authorityName="Kral, Rejsek &amp; Schneider" authorityYear="2001" box="[760,912,589,610]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="0BD5E3614E2D1F24C4B9FDBFFCBCFDB8" box="[760,829,589,610]" italics="true" pageId="8" pageNumber="581">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E2D1F24C50BFDBFFCDCFDB8" box="[842,861,589,610]" italics="true" pageId="8" pageNumber="581">L.</emphasis>
) 
<emphasis id="0BD5E3614E2D1F24C528FDBFFC11FDB8" box="[873,912,589,610]" italics="true" pageId="8" pageNumber="581">ares</emphasis>
</taxonomicName>
<taxonomicName id="FEA144F04E2D1F24C236FDBFFAC7FDB8" authorityName="Kral &amp; Hillert" authorityYear="2013" box="[1143,1350,589,610]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="0BD5E3614E2D1F24C236FDBFFB3DFDB8" box="[1143,1212,589,610]" italics="true" pageId="8" pageNumber="581">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E2D1F24C288FDBFFB5DFDB8" box="[1225,1244,589,610]" italics="true" pageId="8" pageNumber="581">L.</emphasis>
) 
<emphasis id="0BD5E3614E2D1F24C2A9FDBFFAC7FDB8" box="[1256,1350,589,610]" italics="true" pageId="8" pageNumber="581">schneideri</emphasis>
</taxonomicName>
<taxonomicName id="FEA144F04E2D1F24C3B7FDBFF93DFDB8" authorityName="Král &amp; Hillert &amp; Drožová &amp; Šípek" authorityYear="2018" box="[1526,1724,589,610]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="0BD5E3614E2D1F24C3B7FDBFF9BAFDB8" box="[1526,1595,589,610]" italics="true" pageId="8" pageNumber="581">Lethrus</emphasis>
(
<emphasis id="0BD5E3614E2D1F24C009FDBFF9DAFDB8" box="[1608,1627,589,610]" italics="true" pageId="8" pageNumber="581">L.</emphasis>
) 
<emphasis id="0BD5E3614E2D1F24C029FDBFF93DFDB8" box="[1640,1724,589,610]" italics="true" pageId="8" pageNumber="581">thracicus</emphasis>
</taxonomicName>
shape of mandibles in lateral outline angulate at middle lateral outline semicircular (Figs. 2b, f) lateral outline semicircular (Figs. 2c, g) lateral outline angulate at middle (Figs. dorsal view (Figs. 2a, e) 2d, h) lateral longitudinal keel on straight and approximately parallel to straight and distinctly subparallel to straight and distinctly subparallel to straight and distinctly subparallel to base of ventral mandible lateral mandibular outline, distinctly lateral mandibular outline, approximately lateral mandibular outline, approximately lateral mandibular outline, approximately process in dorsal aspect broader as maximum width of so broad as maximum width of so broad as maximum width of so broad as maximum width of mandibles outline basally (Figs. 2a, e) mandibles outline basally (Figs.2 b, f) mandibles outline basally (Figs. 2 c, g) mandibles outline basally (Figs. 2d, h)
</paragraph>
</caption>
<paragraph id="391E3F734E2D1F24C6D2FCE3F9E4FBCA" pageId="8" pageNumber="581">
<table id="4BA1CDD34E2DE0D3C6D2FCE3F8ECFBCA" box="[147,1901,785,1040]" gridcols="5" gridrows="10" pageId="8" pageNumber="581">
<tr id="87913D314E2DE0D3C6D2FCE3F8ECFCFF" box="[147,1901,785,805]" gridrow="0" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FCE3FE07FCFF" box="[147,390,785,805]" gridcol="0" gridrow="0" pageId="8" pageNumber="581">mandibular processes</th>
<th id="C440544D4E2DE0D3C7D3FCE3FD67FCFF" box="[402,742,785,805]" gridcol="1" gridrow="0" pageId="8" pageNumber="581">weakly asymmetrical, right process</th>
<th id="C440544D4E2DE0D3C4B9FCE3F8ECFCFF" box="[760,1901,785,805]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="8" pageNumber="581">both processes almost symmetrical (Figs. both processes almost symmetrical (Figs. weakly asymmetrical, right process</th>
</tr>
<tr id="87913D314E2DE0D3C6D2FCD9F8ECFCE5" box="[147,1901,811,831]" gridrow="1" pageId="8" pageNumber="581" rowspan-0="1">
<td id="C440544D4E2DE0D3C7D3FCD9FD67FCE5" box="[402,742,811,831]" gridcol="1" gridrow="1" pageId="8" pageNumber="581">slightly more developed than left one</td>
<td id="C440544D4E2DE0D3C4B9FCD9FBE9FCE5" box="[760,1128,811,831]" gridcol="2" gridrow="1" pageId="8" pageNumber="581">3b, f; 4b, f)</td>
<td id="C440544D4E2DE0D3C236FCD9FA67FCE5" box="[1143,1510,811,831]" gridcol="3" gridrow="1" pageId="8" pageNumber="581">3c, g; 4c, g)</td>
<td id="C440544D4E2DE0D3C3B7FCD9F8ECFCE5" box="[1526,1901,811,831]" gridcol="4" gridrow="1" pageId="8" pageNumber="581">slightly more developed than left one and</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FCB7F8ECFC83" box="[147,1901,837,857]" gridrow="2" pageId="8" pageNumber="581" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="C440544D4E2DE0D3C7D3FCB7FD67FC83" box="[402,742,837,857]" gridcol="1" gridrow="2" pageId="8" pageNumber="581">and with different angle in lateral and</td>
<td id="C440544D4E2DE0D3C3B7FCB7F8ECFC83" box="[1526,1901,837,857]" gridcol="4" gridrow="2" pageId="8" pageNumber="581">with different angle in lateral aspect</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FCADF8ECFCA9" box="[147,1901,863,883]" gridrow="3" pageId="8" pageNumber="581" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="C440544D4E2DE0D3C7D3FCADFD67FCA9" box="[402,742,863,883]" gridcol="1" gridrow="3" pageId="8" pageNumber="581">frontal aspect</td>
<td id="C440544D4E2DE0D3C3B7FCADF8ECFCA9" box="[1526,1901,863,883]" gridcol="4" gridrow="3" pageId="8" pageNumber="581">(Figs. 3d, h; 4d, h)</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FC8BF8ECFC57" box="[147,1901,889,909]" gridrow="4" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FC8BF8ECFC57" box="[147,1901,889,909]" colspan="5" colspanRight="4" gridcol="0" gridrow="4" pageId="8" pageNumber="581">(Figs. 3a, e; 4a, e)</th>
</tr>
<tr id="87913D314E2DE0D3C6D2FC66F8ECFC72" box="[147,1901,916,936]" gridrow="5" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FC66FE07FC72" box="[147,390,916,936]" gridcol="0" gridrow="5" pageId="8" pageNumber="581">lateral longitudinal keel on</th>
<td id="C440544D4E2DE0D3C7D3FC66FD67FC72" box="[402,742,916,936]" gridcol="1" gridrow="5" pageId="8" pageNumber="581">weakly arcuate, approximately</td>
<td id="C440544D4E2DE0D3C4B9FC66FBE9FC72" box="[760,1128,916,936]" gridcol="2" gridrow="5" pageId="8" pageNumber="581">almost straight, distinctly subparallel to</td>
<td id="C440544D4E2DE0D3C236FC66FA67FC72" box="[1143,1510,916,936]" gridcol="3" gridrow="5" pageId="8" pageNumber="581">weakly arcuate, approximately</td>
<td id="C440544D4E2DE0D3C3B7FC66F8ECFC72" box="[1526,1901,916,936]" gridcol="4" gridrow="5" pageId="8" pageNumber="581">almost straight, approximately parallel to</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FC5CF8ECFC18" box="[147,1901,942,962]" gridrow="6" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FC5CFE07FC18" box="[147,390,942,962]" gridcol="0" gridrow="6" pageId="8" pageNumber="581">base of ventral mandible</th>
<td id="C440544D4E2DE0D3C7D3FC5CFD67FC18" box="[402,742,942,962]" gridcol="1" gridrow="6" pageId="8" pageNumber="581">parallel to lateral mandibular outline,</td>
<td id="C440544D4E2DE0D3C4B9FC5CFBE9FC18" box="[760,1128,942,962]" gridcol="2" gridrow="6" pageId="8" pageNumber="581">lateral mandibular outline, distinctly</td>
<td id="C440544D4E2DE0D3C236FC5CFA67FC18" box="[1143,1510,942,962]" gridcol="3" gridrow="6" pageId="8" pageNumber="581">subparallel to lateral mandibular outline,</td>
<td id="C440544D4E2DE0D3C3B7FC5CF8ECFC18" box="[1526,1901,942,962]" gridcol="4" gridrow="6" pageId="8" pageNumber="581">lateral mandibular outline, slightly</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FC3AF8ECFC06" box="[147,1901,968,988]" gridrow="7" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FC3AFE07FC06" box="[147,390,968,988]" gridcol="0" gridrow="7" pageId="8" pageNumber="581">process in lateral aspect</th>
<td id="C440544D4E2DE0D3C7D3FC3AFD67FC06" box="[402,742,968,988]" gridcol="1" gridrow="7" pageId="8" pageNumber="581">divergent gradually basad</td>
<td id="C440544D4E2DE0D3C4B9FC3AFBE9FC06" box="[760,1128,968,988]" gridcol="2" gridrow="7" pageId="8" pageNumber="581">divergent gradually basad approximately</td>
<td id="C440544D4E2DE0D3C236FC3AFA67FC06" box="[1143,1510,968,988]" gridcol="3" gridrow="7" pageId="8" pageNumber="581">slightly divergent gradually basad</td>
<td id="C440544D4E2DE0D3C3B7FC3AF8ECFC06" box="[1526,1901,968,988]" gridcol="4" gridrow="7" pageId="8" pageNumber="581">divergent gradually basad approximately</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FC10F8ECFC2C" box="[147,1901,994,1014]" gridrow="8" pageId="8" pageNumber="581" rowspan-0="1">
<td id="C440544D4E2DE0D3C7D3FC10FD67FC2C" box="[402,742,994,1014]" gridcol="1" gridrow="8" pageId="8" pageNumber="581">approximately from middle of its</td>
<td id="C440544D4E2DE0D3C4B9FC10FBE9FC2C" box="[760,1128,994,1014]" gridcol="2" gridrow="8" pageId="8" pageNumber="581">from middle of its length (Figs. 4b, f)</td>
<td id="C440544D4E2DE0D3C236FC10FA67FC2C" box="[1143,1510,994,1014]" gridcol="3" gridrow="8" pageId="8" pageNumber="581">approximately from middle of its length</td>
<td id="C440544D4E2DE0D3C3B7FC10F8ECFC2C" box="[1526,1901,994,1014]" gridcol="4" gridrow="8" pageId="8" pageNumber="581">from middle of its length</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FC0EF8ECFBCA" box="[147,1901,1020,1040]" gridrow="9" pageId="8" pageNumber="581" rowspan-0="1" rowspan-2="1">
<td id="C440544D4E2DE0D3C7D3FC0EFD67FBCA" box="[402,742,1020,1040]" gridcol="1" gridrow="9" pageId="8" pageNumber="581">length (Figs. 4a, e)</td>
<td id="C440544D4E2DE0D3C236FC0EFA67FBCA" box="[1143,1510,1020,1040]" gridcol="3" gridrow="9" pageId="8" pageNumber="581">(Figs. 4c, g)</td>
<td id="C440544D4E2DE0D3C3B7FC0EF8ECFBCA" box="[1526,1901,1020,1040]" gridcol="4" gridrow="9" pageId="8" pageNumber="581">(Figs. 4d, h)</td>
</tr>
</table>
</paragraph>
<paragraph id="391E3F734E2D1F24C6D2FBE5F9ABFACC" blockId="8.[147,1902,1047,1302]" pageId="8" pageNumber="581">
<tableNote id="C4473EFD4E2D1F24C6D2FBE5F9ABFACC" pageId="8" pageNumber="581" targetBox="[147,1894,1309,1643]" targetPageId="8">shape of ventral mandible external outline concave basally, external outline strongly concave basally, external outline concave basally, external outline concave basally, processes in frontal aspect inferiobasal tooth present, rounded; inferiobasal tooth present, rounded; inferiobasal tooth absent; subapical tooth inferiobasal tooth shallowly, rounded; subapical tooth distinct (more subapical tooth distinct, apical absent, apical emargination absent subapical tooth distinct, apical developed on right appendix), apical emargination present, shallow (Fig. 3b, (Fig. 3c, g) emargination present, shallow (Fig. 3d, emargination present, remarkably f) h) deep on right, shallow on left one (Fig. 3a, e) shape of left ventral anterior subapical tooth present, anterior subapical tooth present, anterior subapical tooth absent, apical anterior subapical tooth present, rounded; mandible process in lateral rounded; apical tooth not projected broadened distad, angulate; apical tooth tooth not projected apically (Figs. 4c, g) apical tooth not projected apically (Figs. aspect apically (Figs. 4a, e) projected apically (Figs. 4b, f) 4d, h)</tableNote>
</paragraph>
<paragraph id="391E3F734E2D1F24C6D2FAEFFD21F9B1" pageId="8" pageNumber="581">
<table id="4BA1CDD34E2DE0D3C6D2FAEFF8E7F9B1" box="[147,1894,1309,1643]" gridcols="5" gridrows="13" pageId="8" pageNumber="581">
<tr id="87913D314E2DE0D3C6D2FAEFF8E7FAEB" box="[147,1894,1309,1329]" gridrow="0" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FAEFFEFAFAEB" box="[147,379,1309,1329]" gridcol="0" gridrow="0" pageId="8" pageNumber="581">shape of anterior pronotal</th>
<th id="C440544D4E2DE0D3C7D3FAEFFD62FAEB" box="[402,739,1309,1329]" gridcol="1" gridrow="0" pageId="8" pageNumber="581">projected anterolaterad, angulate (see</th>
<th id="C440544D4E2DE0D3C4B9FAEFFBCFFAEB" box="[760,1102,1309,1329]" gridcol="2" gridrow="0" pageId="8" pageNumber="581">not projected anterolaterad, broadly</th>
<th id="C440544D4E2DE0D3C236FAEFFA5EFAEB" box="[1143,1503,1309,1329]" gridcol="3" gridrow="0" pageId="8" pageNumber="581">projected anterolaterad, angulate (see</th>
<th id="C440544D4E2DE0D3C3B7FAEFF8E7FAEB" box="[1526,1894,1309,1329]" gridcol="4" gridrow="0" pageId="8" pageNumber="581">projected anterolaterad, angulate (Figs</th>
</tr>
<tr id="87913D314E2DE0D3C6D2FAC4F8E7FA91" box="[147,1894,1334,1355]" gridrow="1" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FAC4FEFAFA91" box="[147,379,1334,1355]" gridcol="0" gridrow="1" pageId="8" pageNumber="581">angle</th>
<td id="C440544D4E2DE0D3C7D3FAC4FD62FA91" box="[402,739,1334,1355]" gridcol="1" gridrow="1" pageId="8" pageNumber="581">
Král 
<emphasis id="0BD5E3614E2D1F24C781FAC4FE69FA91" box="[448,488,1334,1355]" italics="true" pageId="8" pageNumber="581">et al</emphasis>
. 2013: fig. 2D)
</td>
<td id="C440544D4E2DE0D3C4B9FAC4FBCFFA91" box="[760,1102,1334,1355]" gridcol="2" gridrow="1" pageId="8" pageNumber="581">
round (see Král 
<emphasis id="0BD5E3614E2D1F24C5CBFAC4FC30FA91" box="[906,945,1334,1355]" italics="true" pageId="8" pageNumber="581">et al</emphasis>
. 2013: fig. 2E)
</td>
<td id="C440544D4E2DE0D3C236FAC4FA5EFA91" box="[1143,1503,1334,1355]" gridcol="3" gridrow="1" pageId="8" pageNumber="581">
Král 
<emphasis id="0BD5E3614E2D1F24C2E5FAC4FB4DFA91" box="[1188,1228,1334,1355]" italics="true" pageId="8" pageNumber="581">et al</emphasis>
. 2013: fig. 2F)
</td>
<td id="C440544D4E2DE0D3C3B7FAC4F8E7FA91" box="[1526,1894,1334,1355]" gridcol="4" gridrow="1" pageId="8" pageNumber="581">1a, c, e)</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FAA0F8E7FABC" box="[147,1894,1362,1382]" gridrow="2" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FAA0FEFAFABC" box="[147,379,1362,1382]" gridcol="0" gridrow="2" pageId="8" pageNumber="581">shape of parameres in</th>
<td id="C440544D4E2DE0D3C7D3FAA0FD62FABC" box="[402,739,1362,1382]" gridcol="1" gridrow="2" pageId="8" pageNumber="581">shallowly sinuate distally (Fig. 5a)</td>
<td id="C440544D4E2DE0D3C4B9FAA0FBCFFABC" box="[760,1102,1362,1382]" gridcol="2" gridrow="2" pageId="8" pageNumber="581">distinctly sinuate distally</td>
<td id="C440544D4E2DE0D3C236FAA0FA5EFABC" box="[1143,1503,1362,1382]" gridcol="3" gridrow="2" pageId="8" pageNumber="581">shallowly sinuate distally</td>
<td id="C440544D4E2DE0D3C3B7FAA0F8E7FABC" box="[1526,1894,1362,1382]" gridcol="4" gridrow="2" pageId="8" pageNumber="581">shallowly sinuate distally</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA9EF8E7FA5A" box="[147,1894,1388,1408]" gridrow="3" pageId="8" pageNumber="581" rowspan-1="1">
<th id="C440544D4E2DE0D3C6D2FA9EFEFAFA5A" box="[147,379,1388,1408]" gridcol="0" gridrow="3" pageId="8" pageNumber="581">lateral aspect</th>
<td id="C440544D4E2DE0D3C4B9FA9EFBCFFA5A" box="[760,1102,1388,1408]" gridcol="2" gridrow="3" pageId="8" pageNumber="581">(Fig. 5b)</td>
<td id="C440544D4E2DE0D3C236FA9EFA5EFA5A" box="[1143,1503,1388,1408]" gridcol="3" gridrow="3" pageId="8" pageNumber="581">(Fig. 5c)</td>
<td id="C440544D4E2DE0D3C3B7FA9EF8E7FA5A" box="[1526,1894,1388,1408]" gridcol="4" gridrow="3" pageId="8" pageNumber="581">(Fig. 5d)</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA75F8E7FA41" box="[147,1894,1415,1435]" gridrow="4" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FA75FEFAFA41" box="[147,379,1415,1435]" gridcol="0" gridrow="4" pageId="8" pageNumber="581">distribution</th>
<td id="C440544D4E2DE0D3C7D3FA75FD62FA41" box="[402,739,1415,1435]" gridcol="1" gridrow="4" pageId="8" pageNumber="581">widely distributed from Austria</td>
<td id="C440544D4E2DE0D3C4B9FA75FBCFFA41" box="[760,1102,1415,1435]" gridcol="2" gridrow="4" pageId="8" pageNumber="581">restricted only to south-easternmost</td>
<td id="C440544D4E2DE0D3C236FA75FA5EFA41" box="[1143,1503,1415,1435]" gridcol="3" gridrow="4" pageId="8" pageNumber="581">restricted only to southernmost foothills</td>
<td id="C440544D4E2DE0D3C3B7FA75F8E7FA41" box="[1526,1894,1415,1435]" gridcol="4" gridrow="4" pageId="8" pageNumber="581">restricted to the European part of Turkey</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA53F8E7FA6F" box="[147,1894,1441,1461]" gridrow="5" pageId="8" pageNumber="581" rowspan-0="1">
<td id="C440544D4E2DE0D3C7D3FA53FD62FA6F" box="[402,739,1441,1461]" gridcol="1" gridrow="5" pageId="8" pageNumber="581">(Burgenland), Czech Republic</td>
<td id="C440544D4E2DE0D3C4B9FA53FBCFFA6F" box="[760,1102,1441,1461]" gridcol="2" gridrow="5" pageId="8" pageNumber="581">foothills of the Rhodope Mts. (river</td>
<td id="C440544D4E2DE0D3C236FA53FA5EFA6F" box="[1143,1503,1441,1461]" gridcol="3" gridrow="5" pageId="8" pageNumber="581">of the Rhodope Mts. (Komotiní and</td>
<td id="C440544D4E2DE0D3C3B7FA53F8E7FA6F" box="[1526,1894,1441,1461]" gridcol="4" gridrow="5" pageId="8" pageNumber="581">(lowland east of the Maritza river) (Fig.</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA49F8E7FA15" box="[147,1894,1467,1487]" gridrow="6" pageId="8" pageNumber="581" rowspan-0="1">
<td id="C440544D4E2DE0D3C7D3FA49FD62FA15" box="[402,739,1467,1487]" gridcol="1" gridrow="6" pageId="8" pageNumber="581">(Moravia) and Serbia to</td>
<td id="C440544D4E2DE0D3C4B9FA49FBCFFA15" box="[760,1102,1467,1487]" gridcol="2" gridrow="6" pageId="8" pageNumber="581">Eridropótamos basin region in Greece</td>
<td id="C440544D4E2DE0D3C236FA49FA5EFA15" box="[1143,1503,1467,1487]" gridcol="3" gridrow="6" pageId="8" pageNumber="581">Xánthi regions in Greece) (Fig. 6)</td>
<td id="C440544D4E2DE0D3C3B7FA49F8E7FA15" box="[1526,1894,1467,1487]" gridcol="4" gridrow="6" pageId="8" pageNumber="581">6)</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA27F8E7FA33" box="[147,1894,1493,1513]" gridrow="7" pageId="8" pageNumber="581" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<td id="C440544D4E2DE0D3C7D3FA27FD62FA33" box="[402,739,1493,1513]" gridcol="1" gridrow="7" pageId="8" pageNumber="581">approximately Kursk, Rostov,</td>
<td id="C440544D4E2DE0D3C4B9FA27FBCFFA33" box="[760,1102,1493,1513]" gridcol="2" gridrow="7" pageId="8" pageNumber="581">and Bulgaria) (Fig. 6)</td>
</tr>
<tr id="87913D314E2DE0D3C6D2FA1DF8E7F9D9" box="[147,1894,1519,1539]" gridrow="8" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2FA1DF8E7F9D9" box="[147,1894,1519,1539]" colspan="5" colspanRight="4" gridcol="0" gridrow="8" pageId="8" pageNumber="581">Volgograd and Voronezh regions in</th>
</tr>
<tr id="87913D314E2DE0D3C6D2F9FBF8E7F9C7" box="[147,1894,1545,1565]" gridrow="9" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2F9FBF8E7F9C7" box="[147,1894,1545,1565]" colspan="5" colspanRight="4" gridcol="0" gridrow="9" pageId="8" pageNumber="581">Russia, southernmost to the northern</th>
</tr>
<tr id="87913D314E2DE0D3C6D2F9D1F8E7F9ED" box="[147,1894,1571,1591]" gridrow="10" pageId="8" pageNumber="581" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="C440544D4E2DE0D3C7D3F9D1FD62F9ED" box="[402,739,1571,1591]" gridcol="1" gridrow="10" pageId="8" pageNumber="581">foothills of the Stara planina Mts. in</td>
</tr>
<tr id="87913D314E2DE0D3C6D2F9CEF8E7F98B" box="[147,1894,1596,1617]" gridrow="11" pageId="8" pageNumber="581" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="C440544D4E2DE0D3C7D3F9CEFD62F98B" box="[402,739,1596,1617]" gridcol="1" gridrow="11" pageId="8" pageNumber="581">
Bulgaria (see Král 
<emphasis id="0BD5E3614E2D1F24C47CF9CEFDE4F98B" box="[573,613,1596,1617]" italics="true" pageId="8" pageNumber="581">et al</emphasis>
. 2013: fig. 6;
</td>
</tr>
<tr id="87913D314E2DE0D3C6D2F9A5F8E7F9B1" box="[147,1894,1623,1643]" gridrow="12" pageId="8" pageNumber="581">
<th id="C440544D4E2DE0D3C6D2F9A5F8E7F9B1" box="[147,1894,1623,1643]" colspan="5" colspanRight="4" gridcol="0" gridrow="12" pageId="8" pageNumber="581">Bagaturov &amp; Nikolajev 2015)</th>
</tr>
</table>
</paragraph>
<subSubSection id="71BB6CF84E2C1F25C686FF65FF6AFE9A" pageId="9" pageNumber="582" type="discussion">
<paragraph id="391E3F734E2C1F25C686FF65FF6AFE9A" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="582">
<emphasis id="0BD5E3614E2C1F25C686FF65FEB5FF6A" bold="true" box="[199,308,151,176]" pageId="9" pageNumber="582">Remarks</emphasis>
. 
<bibRefCitation id="5D3042824E2C1F25C700FF65FDB6FF6A" author="Fairmaire, L." box="[321,567,151,176]" pageId="9" pageNumber="582" pagination="249 - 280" refId="ref2653" refString="Fairmaire, L. (1866) Notice sur les Coleopteres recoltes par M. J. Lederer sur le Bosz-Dagh (Asie Mineure). Annales de la Societe Entomologique de France, Series 4, 6, 249 - 280." type="journal article" year="1866">Fairmaire (1866: 257)</bibRefCitation>
dealt in his work with the fauna of beetles of 
<collectingCountry id="41B67FE34E2C1F25C270FF65FB00FF6A" box="[1073,1153,151,176]" name="Turkey" pageId="9" pageNumber="582">Turkey</collectingCountry>
in a broader sense, 
<emphasis id="0BD5E3614E2C1F25C31DFF6AFAF6FF75" box="[1372,1399,152,175]" italics="true" pageId="9" pageNumber="582">i.e</emphasis>
. of the territory from which the new species above is described. He introduced also the name 
<taxonomicName id="FEA144F04E2C1F25C2CDFF4EFADFFF0E" authorityName="Fairmaire" authorityYear="1866" box="[1164,1374,188,212]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="582" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="0BD5E3614E2C1F25C2CDFF4EFADFFF0E" box="[1164,1374,188,212]" italics="true" pageId="9" pageNumber="582">Lethrus subaeneus</emphasis>
</taxonomicName>
from “Asia Mineure”. However, this name was published without any mention of description or reference to it. Thus, according to 
<bibRefCitation id="5D3042824E2C1F25C76BFEF6FDE2FEC6" author="ICZN" box="[298,611,260,285]" pageId="9" pageNumber="582" refId="ref2696" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN (1999; cf. Article 12)</bibRefCitation>
, we consider the name 
<taxonomicName id="FEA144F04E2C1F25C52AFEF6FB76FEC7" authority="Fairmaire, 1866" authorityName="Fairmaire" authorityYear="1866" box="[875,1271,260,285]" class="Insecta" family="Geotrupidae" genus="Lethrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="582" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="0BD5E3614E2C1F25C52AFEF6FBBCFEC6" box="[875,1085,260,284]" italics="true" pageId="9" pageNumber="582">Lethrus subaeneus</emphasis>
<bibRefCitation id="5D3042824E2C1F25C205FEF6FB76FEC7" author="Fairmaire, L." box="[1092,1271,260,285]" pageId="9" pageNumber="582" pagination="249 - 280" refId="ref2653" refString="Fairmaire, L. (1866) Notice sur les Coleopteres recoltes par M. J. Lederer sur le Bosz-Dagh (Asie Mineure). Annales de la Societe Entomologique de France, Series 4, 6, 249 - 280." type="journal article" year="1866">Fairmaire, 1866</bibRefCitation>
</taxonomicName>
to be a 
<emphasis id="0BD5E3614E2C1F25C310FEF7FF6AFE9A" italics="true" pageId="9" pageNumber="582">nomen nudum.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>