443 lines
54 KiB
XML
443 lines
54 KiB
XML
<document id="18A383D6A8B7901046E7F91D6829EE40" ID-CLB-Dataset="8482" ID-DOI="10.11646/zootaxa.4858.1.9" ID-GBIF-Dataset="02f3688b-c8fb-446b-a79f-ba187f6b2b42" ID-ISSN="1175-5326" ID-Zenodo-Dep="4411659" ID-ZooBank="FF7B5935-823E-4187-A3FE-D3CEF7BF3E9F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609609385138" checkinUser="plazi" docAuthor="Stauffer, Jay R., Bills, Roger, Skelton, Paul H. & Weyl, Olaf L. F." docDate="2020" docId="A86987FEFFFE203AFF5819D0FC57F843" docLanguage="en" docName="zootaxa.4858.1.9.pdf" docOrigin="Zootaxa 4858 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serranochromis robustus" docType="treatment" docVersion="8" lastPageNumber="128" masterDocId="5450FF86FFFF2038FFCF1B5FFFB0FF85" masterDocTitle="Re-elevation to species level and redescription of Serranochromis jallae and Serranochromis robustus (Teleostei: Cichlidae)" masterLastPageNumber="134" masterPageNumber="126" pageNumber="127" updateTime="1698851479360" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="94703AE2F3E517CD4ECDCD1C83C17464">Re-elevation to species level and redescription of Serranochromis jallae and Serranochromis robustus (Teleostei: Cichlidae)</mods:title>
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<mods:name id="5ED83A3A187060FB5C133E2A64632722" type="personal">
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<mods:namePart id="B036075937888405B3ADCF7FAACD220A">Stauffer, Jay R.</mods:namePart>
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<mods:affiliation id="51FA24187336F5F1358B62FB0D783A83">Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802, USA. Honorary Research Associate, South African Institute for Aquatic Biodiversity, Makhanda, 6140, RSA.</mods:affiliation>
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<mods:roleTerm id="36B630E5B55A537862E556E76D077E51">Author</mods:roleTerm>
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<mods:namePart id="8DB26C40862570960F07CCA265E3F7AD">Bills, Roger</mods:namePart>
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<mods:nameIdentifier id="1317BAE3314FD2B57AF4F45C1B2FFF63" type="ORCID">0000-0001-6034-4196</mods:nameIdentifier>
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<mods:affiliation id="D2A438844F704F6BCBA0193A86561E11">South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA. & R. Bills @ saiab. ac. za; https: // orcid. org / 0000 - 0001 - 6034 - 4196</mods:affiliation>
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<mods:nameIdentifier id="84F05BE799235EF9D983E6A87C176E3B" type="email">ills@saiab.ac.za</mods:nameIdentifier>
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<mods:namePart id="85C7A0C2897D923B6B0FDBB2DC9F9C66">Skelton, Paul H.</mods:namePart>
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<mods:nameIdentifier id="221677A36531C84866CEC05917451605" type="ORCID">0000-0001-9587-2802</mods:nameIdentifier>
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<mods:affiliation id="E6460FF849BFF40A818BC0359CB4ECA3">South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA. & p. skelton @ saiab. ac. za; https: // orcid. org / 0000 - 0001 - 9587 - 2802</mods:affiliation>
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<mods:nameIdentifier id="B2A14FF8F8077F2C2364040CEB620248" type="email">p.skelton@saiab.ac.za</mods:nameIdentifier>
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<mods:namePart id="D3829F009F3AE6BB6445A023B47ED23E">Weyl, Olaf L. F.</mods:namePart>
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<mods:nameIdentifier id="0A7291198AD5597A3F7A49A0CBDBF152" type="ORCID">0000-0002-8935-3296</mods:nameIdentifier>
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<mods:affiliation id="DF9A57C5C05DFCC4617E7F3302A44ABD">South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA. & o. weyl @ saiab. ac. za; https: // orcid. org / 0000 - 0002 - 8935 - 3296</mods:affiliation>
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<mods:nameIdentifier id="6C92253C689E4BB6179B2E43FB536F0D" type="email">o.weyl@saiab.ac.za</mods:nameIdentifier>
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<mods:title id="B69D14E4913EB6E9CBA54DE39BD847FA">Zootaxa</mods:title>
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<mods:date id="54D458C5EBEA86A19D59FCB742E78813">2020</mods:date>
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<mods:number id="F79DB0665A741E013C68636E150ED02A">2020-09-29</mods:number>
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<mods:number id="23703BA94BF2EC0D54FD95F3223E0C91">4858</mods:number>
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<mods:identifier id="DA69C8CB96A19E548849F115A7F72274" type="DOI">10.11646/zootaxa.4858.1.9</mods:identifier>
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<treatment id="A86987FEFFFE203AFF5819D0FC57F843" ID-DOI="http://doi.org/10.5281/zenodo.4411627" ID-GBIF-Taxon="177209774" ID-Zenodo-Dep="4411627" LSID="urn:lsid:plazi:treatment:A86987FEFFFE203AFF5819D0FC57F843" httpUri="http://treatment.plazi.org/id/A86987FEFFFE203AFF5819D0FC57F843" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127">
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<subSubSection id="68DA6563FFFE2039FF5819D0FD22FD2C" box="[151,658,655,682]" pageId="1" pageNumber="127" type="nomenclature">
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<paragraph id="207F36E8FFFE2039FF5819D0FD22FD2C" blockId="1.[151,658,655,682]" box="[151,658,655,682]" pageId="1" pageNumber="127">
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<heading id="7B378184FFFE2039FF5819D0FD22FD2C" bold="true" box="[151,658,655,682]" fontSize="11" level="1" pageId="1" pageNumber="127" reason="1">
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<taxonomicName id="E7C04D6BFFFE2039FF5819D0FD22FD2C" ID-CoL="4WWVN" authority="(Gunther 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[151,658,655,682]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="12B4EAFAFFFE2039FF5819D0FD22FD2C" bold="true" box="[151,658,655,682]" pageId="1" pageNumber="127">
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<emphasis id="12B4EAFAFFFE2039FF5819D0FE7AFD2F" bold="true" box="[151,458,655,682]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
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(
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<bibRefCitation id="44514B19FFFE2039FE1519D0FD3AFD2C" author="Gunther, A." box="[474,650,655,681]" pageId="1" pageNumber="127" pagination="303 - 314" refId="ref4528" refString="Gunther, A. (1864) Report on a collection of reptiles and fishes made by Dr. Kirk in the Zambesi and Nyassa regions. Proceedings of the Zoological Society of London, 1864 (Pt 2), 303 - 314, pls. 26 - 27." type="journal article" year="1864">Günther 1864</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="68DA6563FFFE2039FF58198BFDD1FC8F" pageId="1" pageNumber="127" type="reference_group">
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<paragraph id="207F36E8FFFE2039FF58198BFDBEFD6E" blockId="1.[151,1232,723,871]" box="[151,526,723,748]" pageId="1" pageNumber="127">
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<taxonomicName id="E7C04D6BFFFE2039FF58198BFDBEFD6E" ID-CoL="3KMJF" authority="Gunther 1864" authorityName="Gunther" authorityYear="1864" box="[151,526,723,748]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="12B4EAFAFFFE2039FF58198BFECBFD69" box="[151,379,724,748]" italics="true" pageId="1" pageNumber="127">Hemichromis robustus</emphasis>
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<bibRefCitation id="44514B19FFFE2039FE4D198BFDBEFD6E" author="Gunther, A." box="[386,526,723,748]" pageId="1" pageNumber="127" pagination="303 - 314" refId="ref4528" refString="Gunther, A. (1864) Report on a collection of reptiles and fishes made by Dr. Kirk in the Zambesi and Nyassa regions. Proceedings of the Zoological Society of London, 1864 (Pt 2), 303 - 314, pls. 26 - 27." type="journal article" year="1864">Günther 1864</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="207F36E8FFFE2039FF5819ADFDD1FC8F" blockId="1.[151,1232,723,871]" box="[151,609,754,778]" pageId="1" pageNumber="127">
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<taxonomicName id="E7C04D6BFFFE2039FF5819ADFEEEFC8F" ID-CoL="4DFV7" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[151,350,754,778]" class="Actinopterygii" family="Cichlidae" genus="Paratilapia" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robusta">
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<emphasis id="12B4EAFAFFFE2039FF5819ADFEEEFC8F" box="[151,350,754,778]" italics="true" pageId="1" pageNumber="127">Paratilapia robusta</emphasis>
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</taxonomicName>
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(in part)
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<bibRefCitation id="44514B19FFFE2039FE7119ADFDD1FC8F" author="Boulenger, G. A." box="[446,609,754,778]" pageId="1" pageNumber="127" pagination="98 - 143" refId="ref4339" refString="Boulenger, G. A. (1899) A revision of the African and Syrian fishes of the family Cichlidae, Part II, Proceedings of the Zoological Society of London, 1899, 98 - 143." type="journal article" year="1899">Boulenger 1899</bibRefCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="68DA6563FFFE2039FF58184EFDCAFCAC" box="[151,634,785,809]" pageId="1" pageNumber="127" type="description">
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||
<paragraph id="207F36E8FFFE2039FF58184EFDCAFCAC" blockId="1.[151,1232,723,871]" box="[151,634,785,809]" pageId="1" pageNumber="127">
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<taxonomicName id="E7C04D6BFFFE2039FF58184EFEC9FCAC" authorityName=": Boulenger" authorityYear="1911" box="[151,377,785,809]" class="Actinopterygii" family="Cichlidae" genus="Paratilapia" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="thumbergi">
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<emphasis id="12B4EAFAFFFE2039FF58184EFEC9FCAC" box="[151,377,785,809]" italics="true" pageId="1" pageNumber="127">Paratilapia thumbergi</emphasis>
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</taxonomicName>
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(in part)
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<bibRefCitation id="44514B19FFFE2039FE16184EFDCAFCAC" author="Boulenger, G. A." box="[473,634,785,809]" pageId="1" pageNumber="127" pagination="399 - 418" refId="ref4378" refString="Boulenger, G. A. (1911) On a collection of fishes from the Lake Ngami basin, Bechuanaland Transactions of the Zoological Society of London, 18, 399 - 418. https: // doi. org / 10.1111 / j. 1469 - 7998.1907. tb 07828. x" type="journal article" year="1911">Boulenger 1911</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="68DA6563FFFE2039FF58186FFB60FCE3" pageId="1" pageNumber="127" type="reference_group">
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||
<paragraph id="207F36E8FFFE2039FF58186FFD96FCC2" blockId="1.[151,1232,723,871]" box="[151,550,815,840]" pageId="1" pageNumber="127">
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||
<taxonomicName id="E7C04D6BFFFE2039FF58186FFD96FCC2" ID-CoL="4WWVN" authority="Jackson 1961" authorityName="Jackson" authorityYear="1961" box="[151,550,815,840]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="12B4EAFAFFFE2039FF58186FFE26FCCD" box="[151,406,816,840]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
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<bibRefCitation id="44514B19FFFE2039FE52186FFD96FCC2" author="Jackson, P. B. N." box="[413,550,815,840]" pageId="1" pageNumber="127" pagination="535 - 561" refId="ref4649" refString="Jackson, P. B. N. (1961) Check list of the fishes of Nyassaland. Occasional Papers of the Natural Museum of Southern Rhodesia, Natural Sciences, 3, 535 - 561." type="journal article" year="1961">Jackson 1961</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="207F36E8FFFE2039FF581811FB60FCE3" blockId="1.[151,1232,723,871]" box="[151,1232,846,871]" pageId="1" pageNumber="127">
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Misidentified as
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<taxonomicName id="E7C04D6BFFFE2039FE8E1811FD64FCE3" ID-CoL="4WWVX" authority="Regan 1922" authorityName="Regan" authorityYear="1922" box="[321,724,846,871]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="thumbergi">
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<emphasis id="12B4EAFAFFFE2039FE8E1811FDE1FCE3" box="[321,593,846,870]" italics="true" pageId="1" pageNumber="127">Serranochromis thumbergi</emphasis>
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||
<bibRefCitation id="44514B19FFFE2039FD971810FD64FCE3" author="Regan, C. T." box="[600,724,846,871]" pageId="1" pageNumber="127" pagination="675 - 727" refId="ref5035" refString="Regan, C. T. (1922) The cichlid fishes of Lake Nyasa. Proceedings of the Zoological Society of London, 1921, 675 - 727. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 03287. x" type="journal article" year="1922">Regan 1922</bibRefCitation>
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||
</taxonomicName>
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||
,
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||
<bibRefCitation id="44514B19FFFE2039FD111810FCCAFCE3" author="Trewavas, E." box="[734,890,846,870]" pageId="1" pageNumber="127" pagination="65 - 118" refId="ref5319" refString="Trewavas, E. (1935) A synopsis of the cichlid fishes of Lake Nyasa. Annals and Magazine of Natural History, Series 10, 16 (91), 65 - 118. https: // doi. org / 10.1080 / 00222933508655026" type="journal article" year="1935">Trewavas 1935</bibRefCitation>
|
||
,
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||
<bibRefCitation id="44514B19FFFE2039FC4A1810FBFCFCE3" author="Bertram, C. K. & Borley, H. J. & Trewavas, E." box="[901,1100,846,870]" pageId="1" pageNumber="127" refId="ref4088" refString="Bertram, C. K., Borley, H. J. & Trewavas, E. (1942) Report on the fish and fisheries of Lake Nyasa 1945 - 1947. Crown Agents, London, 181 pp." type="book" year="1942">
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||
Bertram
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<emphasis id="12B4EAFAFFFE2039FC111810FBA1FCE3" box="[990,1041,846,870]" italics="true" pageId="1" pageNumber="127">et al.</emphasis>
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||
1942
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</bibRefCitation>
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||
,
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||
<bibRefCitation id="44514B19FFFE2039FB991810FB7CFCE3" author="Lowe, R. H." box="[1110,1228,846,870]" pageId="1" pageNumber="127" refId="ref4884" refString="Lowe, R. H. (1952) Report on the Tilapia and other Fish and Fisheries of Lake Nyasa 1945 - 47. HMSO, Colonial Office Fishery Publications 1 (2). HMSO, Colonial Office Fishery London, 126 pp." type="book" year="1952">Lowe 1952</bibRefCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="68DA6563FFFE2039FF5818CBFBA1FBBB" pageId="1" pageNumber="127" type="description">
|
||
<paragraph id="207F36E8FFFE2039FF5818CBFDABFB9F" blockId="1.[151,1437,916,2023]" pageId="1" pageNumber="127">
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Although we have not examined specimens of
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<taxonomicName id="E7C04D6BFFFE2039FD6E18CBFB8EFC2B" authority="Bordin" authorityName="Bordin" box="[673,1086,916,942]" class="Actinopterygii" family="Cichlidae" genus="Pelmatochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="tanganyicae">
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<emphasis id="12B4EAFAFFFE2039FD6E18CBFC57FC2B" box="[673,999,916,942]" italics="true" pageId="1" pageNumber="127">Pelmatochromis tanganyicae</emphasis>
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Bordin
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</taxonomicName>
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, given the current geographical ranges of what is now recognized as
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<taxonomicName id="E7C04D6BFFFE2039FDF318E6FD08FC57" authorityName="Jackson" authorityYear="1961" box="[572,696,952,979]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="12B4EAFAFFFE2039FDF318E6FD08FC57" box="[572,696,952,979]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="E7C04D6BFFFE2039FD3E18E6FCFFFC57" authorityName="Boulenger" authorityYear="1896" box="[753,847,952,979]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
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<emphasis id="12B4EAFAFFFE2039FD3E18E6FCFFFC57" box="[753,847,952,979]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
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</taxonomicName>
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it seems improbable that
|
||
<taxonomicName id="E7C04D6BFFFE2039FBBA18E6FAAAFC57" box="[1141,1306,953,978]" class="Actinopterygii" family="Cichlidae" genus="Pelmatochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="tanganyicae">
|
||
<emphasis id="12B4EAFAFFFE2039FBBA18E6FAAAFC57" box="[1141,1306,953,978]" italics="true" pageId="1" pageNumber="127">P. tanganyicae</emphasis>
|
||
</taxonomicName>
|
||
is either of these species. We thus consider it an error, and do not include this species in the synonymy, although further examination of this species is warranted.
|
||
</paragraph>
|
||
<paragraph id="207F36E8FFFE2039FF081F7AFBA1FBBB" blockId="1.[151,1437,916,2023]" box="[199,1041,1060,1087]" pageId="1" pageNumber="127">Tsungwa is the common name in use in the local languages and by anglers.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68DA6563FFFE2039FF081F17FCA3FB2E" pageId="1" pageNumber="127" type="materials_examined">
|
||
<paragraph id="207F36E8FFFE2039FF081F17FC1CFBE6" blockId="1.[151,1437,916,2023]" box="[199,940,1096,1123]" pageId="1" pageNumber="127">
|
||
<emphasis id="12B4EAFAFFFE2039FF081F17FD94FBE6" bold="true" box="[199,548,1096,1123]" pageId="1" pageNumber="127">
|
||
Material examined.
|
||
<typeStatus id="FF7B884AFFFE2039FE781F16FD94FBE6" box="[439,548,1097,1123]" pageId="1" pageNumber="127" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
of
|
||
<taxonomicName id="E7C04D6BFFFE2039FD851F16FD75FBE7" authority="BMNH" authorityName="BMNH" box="[586,709,1096,1123]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FD851F16FD75FBE7" box="[586,709,1096,1123]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
BMNH 1864.1.9.56
|
||
</paragraph>
|
||
<paragraph id="207F36E8FFFE2039FF581F33FCA3FB2E" blockId="1.[151,1437,916,2023]" pageId="1" pageNumber="127">
|
||
<emphasis id="12B4EAFAFFFE2039FF581F33FEFBFB03" bold="true" box="[151,331,1132,1158]" pageId="1" pageNumber="127">Other material</emphasis>
|
||
:
|
||
<materialsCitation id="90A83CB5FFFE2039FE941F32FC1DFB03" ID-GBIF-Occurrence="3029720301" accessionNumber="PSU 12920" box="[347,941,1132,1159]" collectionCode="PSU" location="Chembe Beach" pageId="1" pageNumber="127" specimenCode="2, 188.2" specimenCount="1">
|
||
<collectionCode id="46D1AE2DFFFE2039FE941F32FE3EFB02" box="[347,398,1133,1159]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">PSU</collectionCode>
|
||
12920,
|
||
<specimenCode id="70669E93FFFE2039FE241F32FE4BFB02" box="[491,507,1133,1159]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">2</specimenCode>
|
||
,
|
||
<specimenCode id="70669E93FFFE2039FDC81F33FDF5FB02" box="[519,581,1132,1159]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">188.2</specimenCode>
|
||
–
|
||
<quantity id="E7389B0DFFFE2039FD9B1F33FD77FB02" box="[596,711,1132,1159]" metricMagnitude="-1" metricUnit="m" metricValue="2.005" pageId="1" pageNumber="127" unit="mm" value="200.5">200.5 mm</quantity>
|
||
SL,
|
||
<location id="251F6033FFFE2039FD311F33FC1DFB03" LSID="urn:lsid:plazi:treatment:A86987FEFFFE203AFF5819D0FC57F843:251F6033FFFE2039FD311F33FC1DFB03" box="[766,941,1132,1158]" name="Chembe Beach" pageId="1" pageNumber="127">Chembe Beach</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="90A83CB5FFFE2039FC771F32FA27FB02" ID-GBIF-Occurrence="3029720302" accessionNumber="PSU 12921" box="[952,1431,1132,1159]" collectionCode="PSU" location="Nkhata Bay" pageId="1" pageNumber="127" specimenCount="1" stateProvince="Nkhata Bay">
|
||
<collectionCode id="46D1AE2DFFFE2039FC771F32FC5BFB02" box="[952,1003,1133,1159]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">PSU</collectionCode>
|
||
12921,
|
||
<quantity id="E7389B0DFFFE2039FB871F32FB68FB02" box="[1096,1240,1133,1159]" metricMagnitude="0" metricUnit="m" metricValue="1.1787" pageId="1" pageNumber="127" unit="mm" value="1178.7">1, 178.7 mm</quantity>
|
||
SL,
|
||
<collectingRegion id="E204F80AFFFE2039FAC01F33FA27FB02" box="[1295,1431,1132,1159]" country="Malawi" name="Nkhata Bay" pageId="1" pageNumber="127">Nkhata Bay</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="90A83CB5FFFE2039FF581FCEFCBFFB2E" ID-GBIF-Occurrence="3029720305" accessionNumber="PSU 12922" box="[151,783,1168,1195]" collectionCode="PSU" location="Mwalamba Rocks" pageId="1" pageNumber="127" specimenCode="1, 189.6" specimenCount="1" stateProvince="Chirombo Bay">
|
||
<collectionCode id="46D1AE2DFFFE2039FF581FCEFF7AFB2E" box="[151,202,1169,1195]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">PSU</collectionCode>
|
||
12922,
|
||
<specimenCode id="70669E93FFFE2039FEE91FCEFE84FB2E" box="[294,308,1169,1195]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">1</specimenCode>
|
||
,
|
||
<specimenCode id="70669E93FFFE2039FE8D1FCFFE34FB2F" box="[322,388,1168,1194]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="127" type="Museum">189.6</specimenCode>
|
||
,
|
||
<location id="251F6033FFFE2039FE401FCFFDEDFB2F" LSID="urn:lsid:plazi:treatment:A86987FEFFFE203AFF5819D0FC57F843:251F6033FFFE2039FE401FCFFDEDFB2F" box="[399,605,1168,1194]" name="Mwalamba Rocks" pageId="1" pageNumber="127" stateProvince="Chirombo Bay">Mwalamba Rocks</location>
|
||
,
|
||
<collectingRegion id="E204F80AFFFE2039FDA71FCFFCBFFB2E" box="[616,783,1168,1195]" inferredBySuffix="true" pageId="1" pageNumber="127">Chirombo Bay</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68DA6563FFFE203AFF081FEBFA97FEC4" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127" type="diagnosis">
|
||
<paragraph id="207F36E8FFFE2039FF081FEBFC48FA4E" blockId="1.[151,1437,916,2023]" pageId="1" pageNumber="127">
|
||
<emphasis id="12B4EAFAFFFE2039FF081FEBFEF2FB4B" bold="true" box="[199,322,1204,1230]" pageId="1" pageNumber="127">Diagnosis.</emphasis>
|
||
The
|
||
<typeStatus id="FF7B884AFFFE2039FEB51FEBFE6CFB4B" box="[378,476,1204,1230]" pageId="1" pageNumber="127" type="holotype">holotype</typeStatus>
|
||
is half of a skin (examined by JRS) collected from Lake
|
||
<collectingCountry id="58D77678FFFE2039FB951FEBFB00FB4B" box="[1114,1200,1204,1230]" name="Malawi" pageId="1" pageNumber="127">Malaŵi</collectingCountry>
|
||
by Kirk who accompanied Livingstone’s expedition (
|
||
<bibRefCitation id="44514B19FFFE2039FDDC1F86FD76FB76" author="Trewavas, E." box="[531,710,1240,1267]" pageId="1" pageNumber="127" pagination="1 - 58" refId="ref5365" refString="Trewavas, E. (1964) A revision of the genus Serranochromis Regan (Pisces, Cichlidae). Musee Royal de l'Afrique Centrale, Tervuren, Annales, Series in- 8 º, Sciences Zoologique, 125, 1 - 58." type="journal article" year="1964">Trewavas 1964</bibRefCitation>
|
||
). The presence of four or five scale rows between the posterior margin of the orbit and the ascending arm of the preoperculum, the presence of widely set unicuspid teeth on the jaws, widely separated gill rakers, and anal fins with egg ocelli places this species in
|
||
<taxonomicName id="E7C04D6BFFFE2039FBBD1E7FFA95FABF" authorityName="Regan" authorityYear="1920" box="[1138,1317,1312,1338]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis id="12B4EAFAFFFE2039FBBD1E7FFA95FABF" box="[1138,1317,1312,1338]" italics="true" pageId="1" pageNumber="127">Serranochromis</emphasis>
|
||
</taxonomicName>
|
||
. Breeding males of
|
||
<taxonomicName id="E7C04D6BFFFE2039FF321E1AFEC8FADB" box="[253,376,1348,1375]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FF321E1AFEC8FADB" box="[253,376,1348,1375]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
possess ocelli that are restricted to the posterior 4–5 membranes of the anal fin, which delimits them from all other
|
||
<taxonomicName id="E7C04D6BFFFE2039FEB81E37FD9AFA07" authorityName="Regan" authorityYear="1920" box="[375,554,1384,1410]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis id="12B4EAFAFFFE2039FEB81E37FD9AFA07" box="[375,554,1384,1410]" italics="true" pageId="1" pageNumber="127">Serranochromis</emphasis>
|
||
</taxonomicName>
|
||
spp., which have ocelli throughout the anal fin in breeding males, with the exception of
|
||
<taxonomicName id="E7C04D6BFFFE2039FEDF1ED2FEDDFA23" authorityName="Boulenger" authorityYear="1896" box="[272,365,1420,1447]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FEDF1ED2FEDDFA23" box="[272,365,1420,1447]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
. As stated above,
|
||
<taxonomicName id="E7C04D6BFFFE2039FDF31ED2FD08FA23" box="[572,696,1420,1447]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FDF31ED2FD08FA23" box="[572,696,1420,1447]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
was shown to be delimited from all other species in the genus by several authors including
|
||
<bibRefCitation id="44514B19FFFE2039FE771EEFFDD5FA4E" author="Jackson, P. B. N." box="[440,613,1456,1483]" pageId="1" pageNumber="127" pagination="535 - 561" refId="ref4649" refString="Jackson, P. B. N. (1961) Check list of the fishes of Nyassaland. Occasional Papers of the Natural Museum of Southern Rhodesia, Natural Sciences, 3, 535 - 561." type="journal article" year="1961">Jackson (1961)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="44514B19FFFE2039FDBF1EEFFD49FA4E" author="Jubb, R. A." box="[624,761,1456,1483]" pageId="1" pageNumber="127" refId="ref4766" refString="Jubb, R. A. (1961) An illustrated guide to the freshwater fishes of the Zambezi river, Lake Kariba, Pungwe, Sabi, Lundi, and Limpopo rivers. Stuart Manning, Bulawayo, 170 pp." type="book" year="1961">Jubb (1961)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="44514B19FFFE2039FCFC1EEEFC44FA4E" author="Trewavas, E." box="[819,1012,1456,1483]" pageId="1" pageNumber="127" pagination="1 - 58" refId="ref5365" refString="Trewavas, E. (1964) A revision of the genus Serranochromis Regan (Pisces, Cichlidae). Musee Royal de l'Afrique Centrale, Tervuren, Annales, Series in- 8 º, Sciences Zoologique, 125, 1 - 58." type="journal article" year="1964">Trewavas (1964)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="207F36E8FFFE203AFF081E8BFA97FEC4" blockId="1.[151,1437,916,2023]" lastBlockId="2.[151,1436,151,321]" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127">
|
||
<taxonomicName id="E7C04D6BFFFE2039FF081E8BFE51FA6B" box="[199,481,1492,1518]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FF081E8BFE51FA6B" box="[199,481,1492,1518]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
generally has a longer lower jaw (50.7–59.6% HL) than
|
||
<taxonomicName id="E7C04D6BFFFE2039FBA61E8AFB76FA6B" authorityName="Boulenger" authorityYear="1896" box="[1129,1222,1492,1519]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FBA61E8AFB76FA6B" box="[1129,1222,1492,1519]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
(49.2–52.7% HL); three specimens of
|
||
<taxonomicName id="E7C04D6BFFFE2039FEBB1EA6FE62F997" authorityName="Boulenger" authorityYear="1896" box="[372,466,1528,1555]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FEBB1EA6FE62F997" box="[372,466,1528,1555]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
had a lower jaw length greater than 52% of the head length and two specimens of
|
||
<taxonomicName id="E7C04D6BFFFE2039FA481EA6FF46F9B3" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FA481EA6FF46F9B3" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
possessed a lower jaw length of less than 52% of the head length.
|
||
<taxonomicName id="E7C04D6BFFFE2039FC131D43FB45F9B3" box="[988,1269,1564,1590]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FC131D43FB45F9B3" box="[988,1269,1564,1590]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
is not as deepbodied as
|
||
<taxonomicName id="E7C04D6BFFFE2039FEC41D1EFED8F9DF" authorityName="Boulenger" authorityYear="1896" box="[267,360,1600,1627]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FEC41D1EFED8F9DF" box="[267,360,1600,1627]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
as evidenced by the distance between the posterior insertion of the dorsal fin and the posterior insertion of the anal fin (13.4–15.1% SL in
|
||
<taxonomicName id="E7C04D6BFFFE2039FD451D3AFCB7F9FB" box="[650,775,1636,1663]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FD451D3AFCB7F9FB" box="[650,775,1636,1663]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
vs. 14.9–18.4% SL in
|
||
<taxonomicName id="E7C04D6BFFFE2039FBC11D3AFBDCF9FB" authorityName="Boulenger" authorityYear="1896" box="[1038,1132,1636,1663]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FBC11D3AFBDCF9FB" box="[1038,1132,1636,1663]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
). Additionally,
|
||
<taxonomicName id="E7C04D6BFFFE2039FAD01D3AFA2CF9FB" box="[1311,1436,1636,1663]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FAD01D3AFA2CF9FB" box="[1311,1436,1636,1663]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
has a narrower least caudal peduncle depth (10.9–12.8% SL) than
|
||
<taxonomicName id="E7C04D6BFFFE2039FC451DD6FC58F927" authorityName="Boulenger" authorityYear="1896" box="[906,1000,1672,1699]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FC451DD6FC58F927" box="[906,1000,1672,1699]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
(11.3–14.2% SL); the least caudal peduncle depth of all
|
||
<taxonomicName id="E7C04D6BFFFE2039FEA41DF2FE55F943" box="[363,485,1708,1735]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FEA41DF2FE55F943" box="[363,485,1708,1735]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
was less than 12.8% SL while, except for the smallest specimen of
|
||
<taxonomicName id="E7C04D6BFFFE2039FB1F1DF2FA9BF943" authorityName="Boulenger" authorityYear="1896" box="[1232,1323,1708,1735]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FB1F1DF2FA9BF943" box="[1232,1323,1708,1735]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="E7389B0DFFFE2039FAF71DF2FA2DF942" box="[1336,1437,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="8.809999999999999" pageId="1" pageNumber="127" unit="mm" value="88.1">88.1 mm</quantity>
|
||
SL), the least caudal peduncle depth was greater than 13.2% SL.
|
||
<taxonomicName id="E7C04D6BFFFE2039FCBE1D8FFB3AF96F" box="[881,1162,1744,1770]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FCBE1D8FFB3AF96F" box="[881,1162,1744,1770]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
has a smaller horizontal eye diameter (HED)(17.7–27.4% HL) than
|
||
<taxonomicName id="E7C04D6BFFFE2039FD4B1DAAFD51F88B" authorityName="Boulenger" authorityYear="1896" box="[644,737,1780,1807]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FD4B1DAAFD51F88B" box="[644,737,1780,1807]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
(18.6–25.5% HL). Although the HED is subject to allometric growth, there is no overlap in the minimum polygons when HED is plotted against head length (
|
||
<figureCitation id="B8FB2A6DFFFE2039FB731C47FB4DF8B6" box="[1212,1277,1816,1843]" captionStart="FIGURE 1" captionStartId="2.[151,250,1318,1342]" captionTargetBox="[179,1415,355,1290]" captionTargetId="figure-112@2.[166,1421,342,1292]" captionTargetPageId="2" captionText="FIGURE 1. Horizontal eye diameter as percent head length plotted against head length of Serranochromis robustus (+) and Serranochromis jallae (x)." figureDoi="http://doi.org/10.5281/zenodo.4411661" httpUri="https://zenodo.org/record/4411661/files/figure.png" pageId="1" pageNumber="127">Fig. 1</figureCitation>
|
||
), and only the two smallest specimens of
|
||
<taxonomicName id="E7C04D6BFFFE2039FE081C62FDF2F8D3" box="[455,578,1852,1879]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FE081C62FDF2F8D3" box="[455,578,1852,1879]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
have a HED greater than 19% HL. In general,
|
||
<taxonomicName id="E7C04D6BFFFE2039FB981C62FB63F8D3" box="[1111,1235,1852,1879]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FB981C62FB63F8D3" box="[1111,1235,1852,1879]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
has more teeth in the outer row of the left lower jaw (14–23) than
|
||
<taxonomicName id="E7C04D6BFFFE2039FD671C3EFCB3F8FF" authorityName="Boulenger" authorityYear="1896" box="[680,771,1888,1915]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FD671C3EFCB3F8FF" box="[680,771,1888,1915]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
(13–14); two specimens of
|
||
<taxonomicName id="E7C04D6BFFFE2039FBF91C3EFB00F8FF" box="[1078,1200,1888,1915]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FBF91C3EFB00F8FF" box="[1078,1200,1888,1915]" italics="true" pageId="1" pageNumber="127">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
had 14 teeth with the remainder having 15 or more. Adults in breeding color of
|
||
<taxonomicName id="E7C04D6BFFFE2039FCD01CDBFB88F81B" box="[799,1080,1924,1950]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFE2039FCD01CDBFB88F81B" box="[799,1080,1924,1950]" italics="true" pageId="1" pageNumber="127">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B8FB2A6DFFFE2039FB871CDBFB3BF81A" box="[1096,1163,1924,1951]" captionStart="FIGURE 2" captionStartId="4.[151,250,805,829]" captionTargetBox="[164,1424,181,781]" captionTargetId="figure-19@4.[164,1424,181,781]" captionTargetPageId="4" captionText="FIGURE 2. Adult male of Serranochromis robustus. Photo by Denis Tweddle." figureDoi="http://doi.org/10.5281/zenodo.4411663" httpUri="https://zenodo.org/record/4411663/files/figure.png" pageId="1" pageNumber="127">Fig. 2</figureCitation>
|
||
) are blue/green laterally with a narrow yellow marginal band on the dorsal fin and usually a small yellowish tip to the upper caudal-fin lobe. Adults in breeding color of
|
||
<taxonomicName id="E7C04D6BFFFE2039FE041C92FD96F863" authorityName="Boulenger" authorityYear="1896" box="[459,550,1996,2023]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFE2039FE041C92FD96F863" box="[459,550,1996,2023]" italics="true" pageId="1" pageNumber="127">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
are yellow/green laterally with a bright orange marginal band on the dorsal and caudal fins in fish from the
|
||
<collectingRegion id="E204F80AFFFD203AFE091BC8FD89FF34" box="[454,569,151,177]" country="Namibia" name="Kavango West" pageId="2" pageNumber="128">Okavango</collectingRegion>
|
||
River system, but creamy yellow bands in fish from the Upper
|
||
<collectingRegion id="E204F80AFFFD203AFB391BC8FA2CFF34" box="[1270,1436,151,177]" country="Namibia" name="Zambezi" pageId="2" pageNumber="128">Zambezi River</collectingRegion>
|
||
system (
|
||
<figureCitation id="B8FB2A6DFFFD203AFF3B1BE4FE87FF50" box="[244,311,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1626,1650]" captionTargetBox="[151,1436,367,1603]" captionTargetId="figure-114@5.[151,1436,345,1603]" captionTargetPageId="5" captionText="FIGURE 3. Adult males of Serranochromis jallae in breeding color (Okavango top; Upper Zambezi bottom). The anal fin in the top photo somewhat obscured by a shadow. Photos by Denis Tweddle." figureDoi="http://doi.org/10.5281/zenodo.4411665" httpUri="https://zenodo.org/record/4411665/files/figure.png" pageId="2" pageNumber="128">Fig. 3</figureCitation>
|
||
). The throat and belly of
|
||
<taxonomicName id="E7C04D6BFFFD203AFD9F1BE4FD1CFF50" authorityName="Boulenger" authorityYear="1896" box="[592,684,187,213]" class="Actinopterygii" family="Cichlidae" genus="Hemichromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFD203AFD9F1BE4FD1CFF50" box="[592,684,187,213]" italics="true" pageId="2" pageNumber="128">S. jallae</emphasis>
|
||
</taxonomicName>
|
||
becomes more intensely yellow-orange in breeding dress hence the local name ‘yellow-belly’. Rarely, specimens from the
|
||
<collectingRegion id="E204F80AFFFD203AFD341B80FCDEFF7C" box="[763,878,223,249]" country="Namibia" name="Kavango West" pageId="2" pageNumber="128">Okavango</collectingRegion>
|
||
River in
|
||
<collectingCountry id="58D77678FFFD203AFC191B80FB86FF7C" box="[982,1078,223,249]" name="Namibia" pageId="2" pageNumber="128">Namibia</collectingCountry>
|
||
also have yellow rather than orange marginal bands (D. Tweddle, pers. comm.).
|
||
<bibRefCitation id="44514B19FFFD203AFD771A5CFCC5FE9B" author="Trewavas, E." box="[696,885,259,286]" pageId="2" pageNumber="128" pagination="1 - 58" refId="ref5365" refString="Trewavas, E. (1964) A revision of the genus Serranochromis Regan (Pisces, Cichlidae). Musee Royal de l'Afrique Centrale, Tervuren, Annales, Series in- 8 º, Sciences Zoologique, 125, 1 - 58." type="journal article" year="1964">Trewavas (1964)</bibRefCitation>
|
||
noted that breeding males of
|
||
<taxonomicName id="E7C04D6BFFFD203AFB731A5CFA2CFE98" authority="robustus" box="[1212,1436,259,285]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="subSpecies" species="robustus" subSpecies="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFB731A5CFA2CFE98" box="[1212,1436,259,285]" italics="true" pageId="2" pageNumber="128">S. robustus robustus</emphasis>
|
||
</taxonomicName>
|
||
had no yellow on throat and belly while those of
|
||
<taxonomicName id="E7C04D6BFFFD203AFD731A78FCCFFEC4" authorityName=": Trewavas" authorityYear="1964" box="[700,895,295,321]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="subSpecies" species="robustus" subSpecies="jallae">
|
||
<emphasis id="12B4EAFAFFFD203AFD731A78FCCFFEC4" box="[700,895,295,321]" italics="true" pageId="2" pageNumber="128">S. robustus jallae</emphasis>
|
||
</taxonomicName>
|
||
possessed yellow on throat and belly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="74BF6660FFFD203AFF581E79FE14FAE7" ID-DOI="http://doi.org/10.5281/zenodo.4411661" ID-Zenodo-Dep="4411661" httpUri="https://zenodo.org/record/4411661/files/figure.png" pageId="2" pageNumber="128" startId="2.[151,250,1318,1342]" targetBox="[179,1415,355,1290]" targetPageId="2">
|
||
<paragraph id="207F36E8FFFD203AFF581E79FE14FAE7" blockId="2.[151,1437,1318,1378]" pageId="2" pageNumber="128">
|
||
<emphasis id="12B4EAFAFFFD203AFF581E79FEA5FABB" bold="true" box="[151,277,1318,1342]" pageId="2" pageNumber="128">FIGURE 1.</emphasis>
|
||
Horizontal eye diameter as percent head length plotted against head length of
|
||
<taxonomicName id="E7C04D6BFFFD203AFB881E79FAF8FABB" box="[1095,1352,1318,1342]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFB881E79FAF8FABB" box="[1095,1352,1318,1342]" italics="true" pageId="2" pageNumber="128">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
(+) and
|
||
<taxonomicName id="E7C04D6BFFFD203AFF581E15FECAFAE7" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[151,378,1354,1378]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="jallae">
|
||
<emphasis id="12B4EAFAFFFD203AFF581E15FECAFAE7" box="[151,378,1354,1378]" italics="true" pageId="2" pageNumber="128">Serranochromis jallae</emphasis>
|
||
</taxonomicName>
|
||
(x).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="68DA6563FFFD203AFF081ECFFAB7F907" pageId="2" pageNumber="128" type="description">
|
||
<paragraph id="207F36E8FFFD203AFF081ECFFAB7F907" blockId="2.[151,1437,1424,1991]" pageId="2" pageNumber="128">
|
||
<emphasis id="12B4EAFAFFFD203AFF081ECFFEE9FA2F" bold="true" box="[199,345,1424,1450]" pageId="2" pageNumber="128">Description.</emphasis>
|
||
Large robust
|
||
<taxonomicName id="E7C04D6BFFFD203AFE371ECFFD1BFA2F" authorityName="Regan" authorityYear="1920" box="[504,683,1424,1450]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="genus">
|
||
<emphasis id="12B4EAFAFFFD203AFE371ECFFD1BFA2F" box="[504,683,1424,1450]" italics="true" pageId="2" pageNumber="128">Serranochromis</emphasis>
|
||
</taxonomicName>
|
||
with general body shape as in
|
||
<figureCitation id="B8FB2A6DFFFD203AFBC11ECFFBE3FA2F" box="[1038,1107,1424,1450]" captionStart="FIGURE 2" captionStartId="4.[151,250,805,829]" captionTargetBox="[164,1424,181,781]" captionTargetId="figure-19@4.[164,1424,181,781]" captionTargetPageId="4" captionText="FIGURE 2. Adult male of Serranochromis robustus. Photo by Denis Tweddle." figureDoi="http://doi.org/10.5281/zenodo.4411663" httpUri="https://zenodo.org/record/4411663/files/figure.png" pageId="2" pageNumber="128">Fig. 2</figureCitation>
|
||
. Principal morphometric and meristic data in
|
||
<tableCitation id="6D420353FFFD203AFE821EEBFE11FA4B" box="[333,417,1460,1486]" captionStart="TABLE 1" captionStartId="3.[151,239,152,176]" captionText="TABLE 1. Morphometric and meristic values of Serranochromis robustus (n = 8) and Serranochromis jallae (n = 9)." pageId="2" pageNumber="128">Table 1</tableCitation>
|
||
and documented above in diagnosis. Head elongate (36.2–38.0% SL). Cheek depth 26.1– 33.5% HL. Snout elongate (35.2–39.8% HL) and posterior end of lower jaw anterior to anterior margin of orbit. Two pored scales posterior to hypural plate, and 9–11 rows of scales on cheek. Outer arch of epibranchial with 4–5 gill rakers; outer arch of ceratobranchial with 10–12. Teeth in 2 series on upper and lower jaws. One or two scales between pectoral fin and posterior margin of opercle. Caudal fin emarginate and lacking marginal band in breeding adult males. Pectoral fin directly under origin of dorsal fin. Pelvic fin posterior to origin of dorsal fin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68DA6563FFFD203AFF081DD3FC57F843" pageId="2" pageNumber="128" type="distribution">
|
||
<paragraph id="207F36E8FFFD203AFF081DD3FC57F843" blockId="2.[151,1437,1424,1991]" pageId="2" pageNumber="128">
|
||
<emphasis id="12B4EAFAFFFD203AFF081DD3FE41F923" bold="true" box="[199,497,1676,1702]" pageId="2" pageNumber="128">Distribution and habitat.</emphasis>
|
||
The native range of
|
||
<taxonomicName id="E7C04D6BFFFD203AFD151DD2FCE5F923" box="[730,853,1676,1703]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFD151DD2FCE5F923" box="[730,853,1676,1703]" italics="true" pageId="2" pageNumber="128">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
is Lake
|
||
<collectingCountry id="58D77678FFFD203AFC7A1DD3FBBBF923" box="[949,1035,1676,1702]" name="Malawi" pageId="2" pageNumber="128">Malaŵi</collectingCountry>
|
||
and the outflowing Shire River and Lake Malombe — where it occupies mostly complex habitats in water shallower than
|
||
<specimenCount id="36C6FD61FFFD203AFBAF1DEFFB11F94E" box="[1120,1185,1712,1739]" pageId="2" pageNumber="128" type="generic">
|
||
<quantity id="E7389B0DFFFD203AFBAF1DEFFB2DF94E" box="[1120,1181,1712,1739]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="128" unit="m" value="10.0">10 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
In southern Lake
|
||
<collectingCountry id="58D77678FFFD203AFAA11DEEFF7AF96B" name="Malawi" pageId="2" pageNumber="128">Malawi</collectingCountry>
|
||
,
|
||
<taxonomicName id="E7C04D6BFFFD203AFF1B1D8AFEFEF96B" box="[212,334,1748,1775]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFF1B1D8AFEFEF96B" box="[212,334,1748,1775]" italics="true" pageId="2" pageNumber="128">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
are generally associated with the rock/sand interface or, associated with submerged aquatic vegetation and among reed beds in shallow water and inlets. The
|
||
<taxonomicName id="E7C04D6BFFFD203AFCF91DA7FC59F897" authorityName="Regan" authorityYear="1920" box="[822,1001,1784,1810]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="genus">
|
||
<emphasis id="12B4EAFAFFFD203AFCF91DA7FC59F897" box="[822,1001,1784,1810]" italics="true" pageId="2" pageNumber="128">Serranochromis</emphasis>
|
||
</taxonomicName>
|
||
species in the Luangwa River may be
|
||
<taxonomicName id="E7C04D6BFFFD203AFF581C42FEA2F8B3" box="[151,274,1820,1847]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFF581C42FEA2F8B3" box="[151,274,1820,1847]" italics="true" pageId="2" pageNumber="128">S. robustus</emphasis>
|
||
</taxonomicName>
|
||
based on photographs taken by anglers but specimens are needed to confirm this (Denis Tweddle, pers. comm.). Fish larger than
|
||
<quantity id="E7389B0DFFFD203AFE791C1FFE58F8DF" box="[438,488,1856,1882]" metricMagnitude="0" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="128" unit="kg" value="2.0">2 kg</quantity>
|
||
are rare.
|
||
<taxonomicName id="E7C04D6BFFFD203AFD9B1C1FFCDFF8DF" box="[596,879,1856,1882]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="12B4EAFAFFFD203AFD9B1C1FFCDFF8DF" box="[596,879,1856,1882]" italics="true" pageId="2" pageNumber="128">Serranochromis robustus</emphasis>
|
||
</taxonomicName>
|
||
was introduced from Lake
|
||
<collectingCountry id="58D77678FFFD203AFB661C1FFB4FF8DF" box="[1193,1279,1856,1882]" name="Malawi" pageId="2" pageNumber="128">Malawi</collectingCountry>
|
||
into the Sand River Dam in
|
||
<collectingCountry id="58D77678FFFD203AFEFA1C3BFE1AF8FB" box="[309,426,1892,1918]" name="Swaziland" pageId="2" pageNumber="128">Swaziland</collectingCountry>
|
||
where it established and spread into the Komati River system in
|
||
<collectingCountry id="58D77678FFFD203AFBB21C3BFABBF8FB" box="[1149,1291,1892,1918]" name="South Africa" pageId="2" pageNumber="128">South Africa</collectingCountry>
|
||
(
|
||
<bibRefCitation id="44514B19FFFD203AFAD61C3BFE96F827" author="De Moor, I. J. & Bruton, M. N." pageId="2" pageNumber="128" pagination="78 - 133" refId="ref4432" refString="De Moor, I. J. & Bruton, M. N. (1988) Atlas of alien and translocated indigenous aquatic animals of southern Africa. South African National Scientific Programmes Report, 144, 78 - 133" type="journal article" year="1988">De Moor & Bruton 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="44514B19FFFD203AFEFF1CD7FE5CF827" author="Bills, R. & Boycott, R. & Fakudze, M. & Khumalo, N. & Msibi, J. & Scott, L. & Terry, S. & Tweddle, D." box="[304,492,1928,1955]" pageId="2" pageNumber="128" pagination="1 - 166" refId="ref4130" refString="Bills, R., Boycott, R., Fakudze, M., Khumalo, N., Msibi, J., Scott, L., Terry, S. & Tweddle, D. (2004) Fish and fisheries survey of Swaziland (2002 - 2003), Final Report-July 2004. SAIAB Investigational Report, 70, 1 - 166 pp." type="book chapter" year="2004">
|
||
Bills
|
||
<emphasis id="12B4EAFAFFFD203AFEA41CD6FE12F827" box="[363,418,1928,1954]" italics="true" pageId="2" pageNumber="128">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
). Verbal reports (2019 pers. com., Erica Tovela,
|
||
<collectingRegion id="E204F80AFFFD203AFC311CD6FBE5F826" box="[1022,1109,1929,1955]" country="Mozambique" name="Maputo" pageId="2" pageNumber="128">Maputo</collectingRegion>
|
||
Museum) from
|
||
<collectingCountry id="58D77678FFFD203AFAC71CD7FA2CF827" box="[1288,1436,1928,1954]" name="Mozambique" pageId="2" pageNumber="128">Mozambique</collectingCountry>
|
||
indicate the species has recently spread onto the
|
||
<collectingCountry id="58D77678FFFD203AFD771CF3FCFCF843" box="[696,844,1964,1990]" name="Mozambique" pageId="2" pageNumber="128">Mozambique</collectingCountry>
|
||
coastal plain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |