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<mods:title id="C2912383FE41D1841CE622F65E1137D1">Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India</mods:title>
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<emphasis id="12966804FFBBFF81C1C7FBBFFD72A72B" bold="true" box="[473,646,1029,1053]" italics="true" pageId="15" pageNumber="1294">Gubernatoriana</emphasis>
<bibRefCitation id="4473C9E7FFBBFF81C291FBBCFCF1A72B" author="Bott R" box="[655,773,1029,1053]" pageId="15" pageNumber="1294" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott, 1970</bibRefCitation>
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<taxonomicName id="E7E2CF95FFBBFF81C08EFBFFFD7AA76B" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[144,654,1093,1117]" class="Malacostraca" family="Gecarcinucidae" genus="Paratelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="1294" phylum="Arthropoda" rank="subGenus" subGenus="Globitelphusa">
<emphasis id="12966804FFBBFF81C08EFBFFFED5A76B" box="[144,289,1093,1117]" italics="true" pageId="15" pageNumber="1294">Paratelphusa</emphasis>
(
<emphasis id="12966804FFBBFF81C125FBFFFE26A76B" box="[315,466,1093,1117]" italics="true" pageId="15" pageNumber="1294">Globitelphusa</emphasis>
)
<bibRefCitation id="4473C9E7FFBBFF81C1F3FBFFFD7AA76B" author="Alcock A" box="[493,654,1093,1117]" pageId="15" pageNumber="1294" pagination="375 - 381" refId="ref7352" refString="Alcock A. 1909. Diagnoses of new species and varieties of freshwater crabs. No. 4. Rec Indian Mus. 3: 375 - 381." type="journal article" year="1909">Alcock, 1909</bibRefCitation>
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;
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<bibRefCitation id="4473C9E7FFBBFF81C2BAFBFFFCEFA76B" author="Alcock A" box="[676,795,1093,1117]" pageId="15" pageNumber="1294" pagination="1 - 14" refId="ref7381" refString="Alcock A. 1910. Brachyura I. Fasc. II. The Indian freshwater crabs- Potamonidae. Catalogue of the Indian Decapod Crustacea in the Collection of the Indian Museum. Calcutta: 1 - 135, pls. 1 - 14." type="book chapter" year="1910">1910: 117</bibRefCitation>
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(part) (not
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<emphasis id="12966804FFBBFF81C3A1FBFFFBA4A76B" box="[959,1104,1093,1117]" italics="true" pageId="15" pageNumber="1294">Paratelphusa</emphasis>
(
<emphasis id="12966804FFBBFF81C0A6FBDFFEBBA74B" box="[184,335,1125,1149]" italics="true" pageId="15" pageNumber="1294">Globitelphusa</emphasis>
)
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).
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<emphasis id="12966804FFBBFF81C08EFB3FFEC9A7AB" box="[144,317,1157,1181]" italics="true" pageId="15" pageNumber="1294">Gubernatoriana</emphasis>
<bibRefCitation id="4473C9E7FFBBFF81C15BFB3FFE1EA7AB" author="Bott R" box="[325,490,1157,1181]" pageId="15" pageNumber="1294" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott, 1970: 44</bibRefCitation>
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.
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<emphasis id="12966804FFBBFF81C08EFB1FFEC9A78B" box="[144,317,1189,1213]" italics="true" pageId="15" pageNumber="1294">Gubernatoriana</emphasis>
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.
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<emphasis id="12966804FFBBFF81C08EFB7FFEC9A7EB" box="[144,317,1221,1245]" italics="true" pageId="15" pageNumber="1294">Gubernatoriana</emphasis>
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species
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<taxonomicName id="E7E2CF95FFBBFF81C08EFAFEFCF0A66A" ID-CoL="75RWX" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[144,772,1348,1372]" class="Malacostraca" family="Gecarcinucidae" genus="Paratelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="1294" phylum="Arthropoda" rank="species" species="gubernatoris" subGenus="Globitelphusa">
<emphasis id="12966804FFBBFF81C08EFAFEFED5A66A" box="[144,289,1348,1372]" italics="true" pageId="15" pageNumber="1294">Paratelphusa</emphasis>
(
<emphasis id="12966804FFBBFF81C12CFAFEFE3DA66A" box="[306,457,1348,1372]" italics="true" pageId="15" pageNumber="1294">Globitelphusa</emphasis>
)
<emphasis id="12966804FFBBFF81C1C4FAFEFD97A66A" box="[474,611,1348,1372]" italics="true" pageId="15" pageNumber="1294">gubernatoris</emphasis>
<bibRefCitation id="4473C9E7FFBBFF81C272FAFEFCF0A66A" author="Alcock A" box="[620,772,1348,1372]" pageId="15" pageNumber="1294" pagination="375 - 381" refId="ref7352" refString="Alcock A. 1909. Diagnoses of new species and varieties of freshwater crabs. No. 4. Rec Indian Mus. 3: 375 - 381." type="journal article" year="1909">Alcock, 1909</bibRefCitation>
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, by original designation.
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Diagnosis (revised after
<bibRefCitation id="4473C9E7FFBBFF81C1BDFA27FDE5A682" author="Bott R" box="[419,529,1436,1460]" pageId="15" pageNumber="1294" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
; based on
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species and present material)
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Carapace squarish (cl/cw 0.780.92) (cw 1.11.3 times cl), flat (ch/cl 0.6); anterolateral margin short, nearly straight; front vertically deflexed, narrow (fw/cw 0.350.42); epigastric crests slightly developed; postorbital crests indistinct; external orbital angle poorly developed; epibranchial tooth indistinct; cervical grooves indistinct; mesogastric groove long, without bifurcation posteriorly; frontal median triangle incomplete with only dorsal margin visible; epistomal median lobe broadly triangular, lacking median tooth (
<figureCitation id="B8D9A893FFBBFF81C0C5F939FE9CA5AD" box="[219,360,1667,1691]" captionStart="Figure 6" captionStartId="8.[212,279,1553,1575]" captionTargetBox="[213,1170,168,1522]" captionTargetId="figure@8.[213,1170,168,1522]" captionTargetPageId="8" captionText="Figure 6. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Dorsal view; (B) frontal view; (C) ventral view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4006926" httpUri="https://zenodo.org/record/4006926/files/figure.png" pageId="15" pageNumber="1294">Figure 6A,B</figureCitation>
;
<bibRefCitation id="4473C9E7FFBBFF81C16DF93EFE14A5AD" author="Bott R" box="[371,480,1667,1691]" pageId="15" pageNumber="1294" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 6, figs 60, 61; pl. 34, figs 33, 35). Chelipeds unequal. Ambulatory legs with fine bristles. Suture between thoracic sternites s2/s3 indistinct or visible as a small groove and between s3/s4 visible as grooves on sides only (
<figureCitation id="B8D9A893FFA4FF9EC40CFF72FB79A3D6" box="[1042,1165,200,224]" captionStart="Figure 6" captionStartId="8.[212,279,1553,1575]" captionTargetBox="[213,1170,168,1522]" captionTargetId="figure@8.[213,1170,168,1522]" captionTargetPageId="8" captionText="Figure 6. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Dorsal view; (B) frontal view; (C) ventral view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4006926" httpUri="https://zenodo.org/record/4006926/files/figure.png" pageId="16" pageNumber="1295">Figures 6C</figureCitation>
,
<figureCitation id="B8D9A893FFA4FF9EC0CAFF52FF02A236" box="[212,246,232,256]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">7A</figureCitation>
;
<bibRefCitation id="4473C9E7FFA4FF9EC116FF53FE8EA236" author="Bott R" box="[264,378,232,256]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 6, fig. 62; pl. 34, fig. 34). Male abdomen short T-shaped; fifth abdominal somite much broader than long; sixth abdominal somite slightly broader than long, subequal in length to telson; telson short; male sternoabdominal cavity deep, long, extending beyond level of cheliped bases or even beyond level of third maxilliped bases (
<figureCitation id="B8D9A893FFA4FF9EC100FEDDFE6DA249" box="[286,409,359,383]" captionStart="Figure 6" captionStartId="8.[212,279,1553,1575]" captionTargetBox="[213,1170,168,1522]" captionTargetId="figure@8.[213,1170,168,1522]" captionTargetPageId="8" captionText="Figure 6. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Dorsal view; (B) frontal view; (C) ventral view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4006926" httpUri="https://zenodo.org/record/4006926/files/figure.png" pageId="16" pageNumber="1295">Figures 6C</figureCitation>
,
<figureCitation id="B8D9A893FFA4FF9EC1B9FED2FE32A249" box="[423,454,360,383]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">7B</figureCitation>
;
<bibRefCitation id="4473C9E7FFA4FF9EC1CDFED2FDB4A249" author="Bott R" box="[467,576,359,383]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 6, fig. 62; pl. 34, fig. 34). Exopods of first, second maxillipeds with long flagellum; exopod of third maxilliped lacking flagellum, exopod longer than ischium (
<figureCitation id="B8D9A893FFA4FF9EC1DFFE1DFDC2A289" box="[449,566,423,447]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">Figure 7D</figureCitation>
). G1 short, stout with long terminal article (0.350.50 times length of subterminal segment); subterminal segment much broader than terminal article (
<figureCitation id="B8D9A893FFA4FF9EC137FE5DFE4AA2C9" box="[297,446,487,511]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">Figure 7EG</figureCitation>
; see
<bibRefCitation id="4473C9E7FFA4FF9EC1EBFE52FD96A2C9" author="Bott R" box="[501,610,487,511]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 27, fig. 27; pl. 34, fig. 36). G2 short with very short or vestigial terminal article (
<figureCitation id="B8D9A893FFA4FF9EC251FDBDFD37A129" box="[591,707,519,543]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">Figure 7H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="68F8E79DFFA4FF9EC0CAFDD9FC62A7D3" pageId="16" pageNumber="1295" type="discussion">
<paragraph id="205DB416FFA4FF9EC0CAFDD9FEC2A14D" blockId="16.[212,1172,611,1089]" box="[212,310,611,635]" pageId="16" pageNumber="1295">
<heading id="7B15037AFFA4FF9EC0CAFDD9FEC2A14D" box="[212,310,611,635]" fontSize="10" level="3" pageId="16" pageNumber="1295" reason="5">
<emphasis id="12966804FFA4FF9EC0CAFDD9FEC2A14D" box="[212,310,611,635]" italics="true" pageId="16" pageNumber="1295">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="205DB416FFA4FF9EC0CAFD31FEEDA0D7" blockId="16.[212,1172,611,1089]" pageId="16" pageNumber="1295">
<taxonomicName id="E7E2CF95FFA4FF9EC0CAFD31FE75A195" box="[212,385,651,675]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC0CAFD31FE75A195" box="[212,385,651,675]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="E7E2CF95FFA4FF9EC2D5FD31FCC4A195" authority="Pati &amp; Sharma, 2014" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[715,816,651,675]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="12966804FFA4FF9EC2D5FD31FCC4A195" box="[715,816,651,675]" italics="true" pageId="16" pageNumber="1295">Ghatiana</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="09A5D57FFFA4FF9EC35FFD31FC48A195" box="[833,956,651,675]" pageId="16" pageNumber="1295" rank="genus">new genus</taxonomicNameLabel>
, but can be differentiated based on some significant external morphological characters (see Remarks for
<taxonomicName id="E7E2CF95FFA4FF9EC11DFD71FE9CA1D5" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[259,360,715,739]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC11DFD71FE9CA1D5" box="[259,360,715,739]" italics="true" pageId="16" pageNumber="1295">Ghatiana</emphasis>
</taxonomicName>
). This genus is distinguished from
<taxonomicName id="E7E2CF95FFA4FF9EC307FD71FCAAA1D5" authorityName="Bahir and Yeo" authorityYear="2007" box="[793,862,715,739]" class="Malacostraca" family="Gecarcinucidae" genus="Snaha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC307FD71FCAAA1D5" box="[793,862,715,739]" italics="true" pageId="16" pageNumber="1295">Snaha</emphasis>
</taxonomicName>
and
<taxonomicName id="E7E2CF95FFA4FF9EC3BAFD71FC05A1D5" authorityName="Bahir and Yeo" authorityYear="2007" box="[932,1009,715,739]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC3BAFD71FC05A1D5" box="[932,1009,715,739]" italics="true" pageId="16" pageNumber="1295">Pilarta</emphasis>
</taxonomicName>
by the short terminal article of G2 (
<figureCitation id="B8D9A893FFA4FF9EC1FCFD50FDAEA034" box="[482,602,746,770]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">Figure 7H</figureCitation>
) (vs long terminal article of G2) (
<bibRefCitation id="4473C9E7FFA4FF9EC3F9FD50FEFDA014" author="Bahir MM &amp; Yeo DCJ" pageId="16" pageNumber="1295" pagination="309 - 354" refId="ref7425" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
: figs. 25H, 27G, 29E).
<taxonomicName id="E7E2CF95FFA4FF9EC203FCB0FD3EA014" box="[541,714,778,802]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC203FCB0FD3EA014" box="[541,714,778,802]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="E7E2CF95FFA4FF9EC3D0FCB0FB99A014" authorityName="Bott" authorityYear="1970" box="[974,1133,778,802]" class="Malacostraca" family="Gecarcinucidae" genus="Inglethelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC3D0FCB0FB99A014" box="[974,1133,778,802]" italics="true" pageId="16" pageNumber="1295">Inglethelphusa</emphasis>
</taxonomicName>
by the long third maxilliped exopod and short, stout G1 (vs short third maxilliped exopod and long, narrow G1) (
<figureCitation id="B8D9A893FFA4FF9EC242FCF0FD0DA054" box="[604,761,842,866]" captionStart="Figure 7" captionStartId="9.[144,211,1507,1529]" captionTargetBox="[155,1097,168,1475]" captionTargetId="figure@9.[145,1102,168,1475]" captionTargetPageId="9" captionText="Figure 7. Gubernatoriana triangulus sp. nov., holotype male (ZSI, WRC-C.1134). (A) Anterior thoracic sternites; (B) abdomen; (C) left cheliped; (D) left third maxilliped; (E) dorsal view of right G1; (F) ventral view of right G1; (G) dorsal view of terminal article of right G1; (H) right G2. Scale bars: 5 mm (A, B); 10 mm (C); 2.5 mm (D) and 1 mm (EH)." figureDoi="http://doi.org/10.5281/zenodo.4006928" httpUri="https://zenodo.org/record/4006928/files/figure.png" pageId="16" pageNumber="1295">Figure 7DF</figureCitation>
;
<bibRefCitation id="4473C9E7FFA4FF9EC311FCF1FC71A054" author="Bott R" box="[783,901,842,866]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 34, figs. 26, 28).
<taxonomicName id="E7E2CF95FFA4FF9EC0CAFCD0FD71A0B4" authority="(Roux, 1931)" baseAuthorityName="Roux" baseAuthorityYear="1931" box="[212,645,874,898]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="escheri">
<emphasis id="12966804FFA4FF9EC0CAFCD0FE2FA0B4" box="[212,475,874,898]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana escheri</emphasis>
(
<bibRefCitation id="4473C9E7FFA4FF9EC1ECFCD0FD89A0B4" author="Roux J" box="[498,637,874,898]" pageId="16" pageNumber="1295" pagination="31 - 62" refId="ref7606" refString="Roux J. 1931. Crustaces Decapodes d' eau douce de l' Inde meridionale. Rev Suisse Zool. 38: 31 - 62." type="journal article" year="1931">Roux, 1931</bibRefCitation>
)
</taxonomicName>
was recently assigned to the genus
<taxonomicName id="E7E2CF95FFA4FF9EC459FCD0FB78A0B4" authorityName="Bahir and Yeo" authorityYear="2007" box="[1095,1164,874,898]" class="Malacostraca" family="Gecarcinucidae" genus="Snaha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC459FCD0FB78A0B4" box="[1095,1164,874,898]" italics="true" pageId="16" pageNumber="1295">Snaha</emphasis>
</taxonomicName>
, while
<taxonomicName id="E7E2CF95FFA4FF9EC102FC30FD07A094" authority="(Roux, 1931)" baseAuthorityName="Roux" baseAuthorityYear="1931" box="[284,755,906,930]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="nilgiriensis">
<emphasis id="12966804FFA4FF9EC102FC30FDBFA094" box="[284,587,906,930]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana nilgiriensis</emphasis>
(
<bibRefCitation id="4473C9E7FFA4FF9EC27FFC30FD1FA094" author="Roux J" box="[609,747,906,930]" pageId="16" pageNumber="1295" pagination="31 - 62" refId="ref7606" refString="Roux J. 1931. Crustaces Decapodes d' eau douce de l' Inde meridionale. Rev Suisse Zool. 38: 31 - 62." type="journal article" year="1931">Roux, 1931</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="E7E2CF95FFA4FF9EC329FC30FEE6A0F7" authority="(Roux, 1931)" baseAuthorityName="Roux" baseAuthorityYear="1931" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="12966804FFA4FF9EC329FC30FBCCA094" box="[823,1080,906,930]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana pusilla</emphasis>
(
<bibRefCitation id="4473C9E7FFA4FF9EC453FC30FEFEA0F7" author="Roux J" pageId="16" pageNumber="1295" pagination="31 - 62" refId="ref7606" refString="Roux J. 1931. Crustaces Decapodes d' eau douce de l' Inde meridionale. Rev Suisse Zool. 38: 31 - 62." type="journal article" year="1931">Roux, 1931</bibRefCitation>
)
</taxonomicName>
were both transferred to the genus
<taxonomicName id="E7E2CF95FFA4FF9EC2D6FC11FCFEA0F4" authorityName="Bahir &amp; Yeo" authorityYear="2007" box="[712,778,939,962]" class="Malacostraca" family="Gecarcinucidae" genus="Vanni" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC2D6FC11FCFEA0F4" box="[712,778,939,962]" italics="true" pageId="16" pageNumber="1295">Vanni</emphasis>
</taxonomicName>
(
<bibRefCitation id="4473C9E7FFA4FF9EC300FC13FBE7A0F7" author="Bahir MM &amp; Yeo DCJ" box="[798,1043,937,961]" pageId="16" pageNumber="1295" pagination="309 - 354" refId="ref7425" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
;
<bibRefCitation id="4473C9E7FFA4FF9EC439FC10FEFDA0D7" author="Ng PKL &amp; Guinot D &amp; Davie PFJ" pageId="16" pageNumber="1295" pagination="1 - 286" refId="ref7512" refString="Ng PKL, Guinot D, Davie PFJ. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool. 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008</bibRefCitation>
).
</paragraph>
<paragraph id="205DB416FFA4FF9EC0E0FC50FE50A777" blockId="16.[212,1172,611,1089]" pageId="16" pageNumber="1295">
<taxonomicName id="E7E2CF95FFA4FF9EC0E0FC50FDDBA737" authority="Bott, 1970" authorityName="Bott" authorityYear="1970" box="[254,559,1001,1026]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="genus">
<emphasis id="12966804FFA4FF9EC0E0FC50FE5FA734" box="[254,427,1002,1026]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana</emphasis>
<bibRefCitation id="4473C9E7FFA4FF9EC1ABFC50FDDBA737" author="Bott R" box="[437,559,1001,1025]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott, 1970</bibRefCitation>
</taxonomicName>
, now consists of three species:
<taxonomicName id="E7E2CF95FFA4FF9EC389FC50FE39A717" authority="(Alcock 1909)" baseAuthorityName="Alcock" baseAuthorityYear="1909" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="12966804FFA4FF9EC389FC50FED4A717" italics="true" pageId="16" pageNumber="1295">Gubernatoriana gubernatoris</emphasis>
(
<bibRefCitation id="4473C9E7FFA4FF9EC12CFBB3FE37A717" author="Alcock A" box="[306,451,1033,1057]" pageId="16" pageNumber="1295" pagination="375 - 381" refId="ref7352" refString="Alcock A. 1909. Diagnoses of new species and varieties of freshwater crabs. No. 4. Rec Indian Mus. 3: 375 - 381." type="journal article" year="1909">Alcock 1909</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="E7E2CF95FFA4FF9EC1C0FBB3FC55A717" authority="(Alcock 1909)" baseAuthorityName="Alcock" baseAuthorityYear="1909" box="[478,929,1033,1057]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="12966804FFA4FF9EC1C0FBB3FD07A717" box="[478,755,1033,1057]" italics="true" pageId="16" pageNumber="1295">Gubernatoriana pilosipes</emphasis>
(
<bibRefCitation id="4473C9E7FFA4FF9EC31BFBB3FC63A717" author="Alcock A" box="[773,919,1033,1057]" pageId="16" pageNumber="1295" pagination="375 - 381" refId="ref7352" refString="Alcock A. 1909. Diagnoses of new species and varieties of freshwater crabs. No. 4. Rec Indian Mus. 3: 375 - 381." type="journal article" year="1909">Alcock 1909</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="E7E2CF95FFA4FF9EC3F9FBB3FEB5A777" authority="Pati &amp; Sharma, 2014" authorityName="Pati &amp; Sharma" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1295" phylum="Arthropoda" rank="species" species="triangulus" status="sp. nov.">
<emphasis id="12966804FFA4FF9EC3F9FBB3FEB5A777" italics="true" pageId="16" pageNumber="1295">Gubernatoriana triangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09A5D57FFFA4FF9EC154FB90FE50A777" box="[330,420,1066,1089]" pageId="16" pageNumber="1295" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="205DB416FFA4FF9EC0CAFB3FFE7DA7AB" blockId="16.[212,1172,1157,1253]" box="[212,393,1157,1181]" pageId="16" pageNumber="1295">
<heading id="7B15037AFFA4FF9EC0CAFB3FFE7DA7AB" box="[212,393,1157,1181]" fontSize="10" level="3" pageId="16" pageNumber="1295" reason="5">
<emphasis id="12966804FFA4FF9EC0CAFB3FFE7DA7AB" box="[212,393,1157,1181]" italics="true" pageId="16" pageNumber="1295">Ecological notes</emphasis>
</heading>
</paragraph>
<paragraph id="205DB416FFA4FF9EC0CAFB17FC62A7D3" blockId="16.[212,1172,1157,1253]" pageId="16" pageNumber="1295">
These crabs are found in small streams, under small stones in high-altitude areas in Western Ghats of
<collectingCountry id="58F5F486FFA4FF9EC1B8FB77FE15A7D3" box="[422,481,1229,1253]" name="India" pageId="16" pageNumber="1295">India</collectingCountry>
(Pati and Sharma, unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="68F8E79DFFA4FF9EC0CAFA93FE4CA6BE" pageId="16" pageNumber="1295" type="distribution">
<paragraph id="205DB416FFA4FF9EC0CAFA93FEACA677" blockId="16.[212,1172,1321,1416]" box="[212,344,1321,1345]" pageId="16" pageNumber="1295">
<heading id="7B15037AFFA4FF9EC0CAFA93FEACA677" box="[212,344,1321,1345]" fontSize="10" level="3" pageId="16" pageNumber="1295" reason="5">
<emphasis id="12966804FFA4FF9EC0CAFA93FEACA677" box="[212,344,1321,1345]" italics="true" pageId="16" pageNumber="1295">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="205DB416FFA4FF9EC0CAFAEAFE4CA6BE" blockId="16.[212,1172,1321,1416]" pageId="16" pageNumber="1295">
<materialsCitation id="908ABE4BFFA4FF9EC0CAFAEAFE47A6BE" ID-GBIF-Occurrence="2850101301" location="Northern Western Ghat Mountains" pageId="16" pageNumber="1295" specimenCount="1" stateProvince="Maharashtra">
<location id="253DE2CDFFA4FF9EC0CAFAEAFD91A65E" LSID="urn:lsid:plazi:treatment:A84B0500FFBBFF9EC1C7FBBFFE4CA6BE:253DE2CDFFA4FF9EC0CAFAEAFD91A65E" box="[212,613,1360,1384]" name="Northern Western Ghat Mountains" pageId="16" pageNumber="1295" stateProvince="Maharashtra">Northern Western Ghat Mountains</location>
(
<collectingRegion id="E2267AF4FFA4FF9EC26AFAEAFCFFA65E" box="[628,779,1360,1384]" country="India" name="Maharashtra" pageId="16" pageNumber="1295">Maharashtra</collectingRegion>
: Satara, Pune);
<collectingRegion id="E2267AF4FFA4FF9EC3A3FAEAFBCFA65E" box="[957,1083,1360,1384]" country="India" name="Karnataka" pageId="16" pageNumber="1295">Karnataka</collectingRegion>
: North- Canara (
<bibRefCitation id="4473C9E7FFA4FF9EC126FACAFE5CA6BE" author="Bott R" box="[312,424,1392,1416]" pageId="16" pageNumber="1295" pagination="1 - 338" refId="ref7462" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abh Senckenb Naturforsch Ges (Frankfurt). 526: 1 - 338. pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>