187 lines
16 KiB
XML
187 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.168.1919" ID-GBIF-Dataset="9aa99d5a-4db4-407d-9d91-f2839063aeb0" ID-PMC="PMC3293443" ID-Pensoft-Pub="1313-2970-168-45" ID-PubMed="22423190" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-168-45" ModsDocOrigin="ZooKeys 168" ModsDocTitle="Two new and rare mountain door-snails (Gastropoda, Pulmonata, Clausiliidae) from high mountain areas in Macedonia" checkinTime="1451249349062" checkinUser="pensoft" docAuthor="Dedov, Ivaylo Kanev" docDate="2012" docId="A80AEA021861ACEEF605E5F7A6CEA449" docLanguage="en" docName="ZooKeys 168: 45-53" docOrigin="ZooKeys 168" docSource="http://dx.doi.org/10.3897/zookeys.168.1919" docTitle="Vestia lazarovii Dedov, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="316A23360127FFB5DA54FFE872302402" masterDocTitle="Two new and rare mountain door-snails (Gastropoda, Pulmonata, Clausiliidae) from high mountain areas in Macedonia" masterLastPageNumber="53" masterPageNumber="45" pageNumber="48" updateTime="1668153228659" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new and rare mountain door-snails (Gastropoda, Pulmonata, Clausiliidae) from high mountain areas in Macedonia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dedov, Ivaylo Kanev</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>168</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.168.1919</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.168.1919</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-168-45</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152033216" LSID="urn:lsid:zoobank.org:act:3338D8CD-0D44-4C78-8511-0198A58E3A10" httpUri="http://treatment.plazi.org/id/A80AEA021861ACEEF605E5F7A6CEA449" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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<subSubSection pageId="3" pageNumber="48" type="nomenclature">
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<paragraph pageId="3" pageNumber="48">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:3338D8CD-0D44-4C78-8511-0198A58E3A10" class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia lazarovii" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="lazarovii">Vestia lazarovii</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="48">sp. n.</taxonomicNameLabel>
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Fig. 3
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="locus typicu">
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<paragraph pageId="3" pageNumber="48">Locus typicu.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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Republic of Macedonia, Baba (= Pelister) Mountains near Kopanke hut, 41°01'59.7'N, 21°13'09.0'E, 1639 m a.s.l.,
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus peuce" order="Pinales" pageId="3" pageNumber="48" phylum="Tracheophyta" rank="species" species="peuce">Pinus peuce</taxonomicName>
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forestecotone,under logs of dead wood and fallen trunks, 03. September 2002, leg. S. Lazarov, (2 empty shells); from the same site, 16. June 2009, leg. I. K. Dedov (12 specimens, collected alive, dried).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="additional material">
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<paragraph pageId="3" pageNumber="48">Additional material.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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Republic of Macedonia, Pelister (= Baba) Mountains, Palisnopje area, 1450 m a.s.l.,
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus peuce" order="Pinales" pageId="3" pageNumber="48" phylum="Tracheophyta" rank="species" species="peuce">Pinus peuce</taxonomicName>
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forestecotone, under logs and fallen trunks, 16. June 2009, leg. T. Mitev, (2 empty shells).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="type material">
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<paragraph pageId="3" pageNumber="48">Type material.</paragraph>
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<paragraph pageId="3" pageNumber="48">holotype SMF 336343, paratypes (n = 13 specimens) SMF 336344/2 specimens; NMNHS/2 specimens; DED/MK 453/2 specimens; DED/MK 636/9 specimens; Pelister Mountains, Palisnopje area, 1450 m. a.s.l., DED/MK637/2 specimens).</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="differential diagnosis">
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<paragraph pageId="3" pageNumber="48">Differential diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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This species differs from
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia roschitzi" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="roschitzi">Vestia roschitzi</taxonomicName>
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(Brancsik, 1890) and
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia ranojevici" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="ranojevici">Vestia ranojevici</taxonomicName>
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(Pavlovic, 1912) by the wide spiral turn of its inferior lamella; from
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia elata" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="elata">Vestia elata</taxonomicName>
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(Rossmassler, 1836),
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia gulo" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="gulo">Vestia gulo</taxonomicName>
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(E. Bielz, 1859) and
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia turgida" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="turgida">Vestia turgida</taxonomicName>
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(Rossmassler, 1836) by the missing lunella.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="description">
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<paragraph pageId="3" pageNumber="48">Description of type series.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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shell relatively small, spindle shaped, yellow-brownish coloured; whorls 8.5-9.5, including 2-2.5 smooth protoconch whorls; teleoconch ribbed (R = 38-54); aperture oval pear-shaped with a whitish, weekly reflected lip; a pale palatal callus present in some specimens; basal canal and keel missing; sinulus wide, not inclined to the shell axis; superior lamella connected with spiralis or close to it; inferior lamella turning widely-spirally; lunella and basalis missing; principal and upper palatal plica usually present; principal plica very short to about 1/3 of the last whorl; upper palatal plica short or missing; clausilium plate varying from hook-shaped in its end as is typical for
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="genus">Vestia</taxonomicName>
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, or with a weak hook and thin clausilium plate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="etymology">
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<paragraph pageId="3" pageNumber="48">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="48">This species is named after the Bulgarian arachnologist Dr. Stoyan Lazarov-Panagyrsky, B. A. S., Institute of Zoology, who was the first to collect this species.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="distribution">
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<paragraph pageId="3" pageNumber="48">Distribution.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia lazarovii" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="lazarovii">Vestia lazarovii</taxonomicName>
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sp. n. is currently only known from two sites at 1450 and 1650 m a.s.l. from the Pelister (= Baba) Mountains, Republic of Macedonia.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="ecology">
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<paragraph pageId="3" pageNumber="48">Ecology.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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This species occurs in the
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus peuce" order="Pinales" pageId="3" pageNumber="48" phylum="Tracheophyta" rank="species" species="peuce">Pinus peuce</taxonomicName>
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forest ecotone, under logs of dead wood near Kopanke hut, as well as in the
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus peuce" order="Pinales" pageId="3" pageNumber="48" phylum="Tracheophyta" rank="species" species="peuce">Pinus peuce</taxonomicName>
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forest ecotone in the Palisnopje area, under logs and fallen trunks.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="comments">
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<paragraph pageId="3" pageNumber="48">Comments.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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The first species of genus
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="genus">Vestia</taxonomicName>
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to bereported from Macedonia (
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<bibRefCitation author="Urbanski, J" journalOrPublisher="Bulletin de la Societe des Amis des Sciences et Lettres de Poznan" pageId="4" pageNumber="49" pagination="113 - 147" title="Bemerkenswerte Clausiliiden (Moll., Pulm.) aus Bulgarien (Systematische, zoogeographische und oekologische Studien ueber der Mollusken der Balkan-Halbinsel VI)." volume="14" year="1960">Urbanski 1960</bibRefCitation>
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) was
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia ranojevici" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="ranojevici">Vestia ranojevici</taxonomicName>
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.
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="4" pageNumber="49" pagination="123 - 170" title="Neue Clausilien der Balkan-Halbinsel (mit taxonomischer Revision einiger Gruppen der Alopiinae und Baleinae). Zur Anatomie und Systematik der Clausilien XV." volume="104" year="1974">Nordsieck (1974)</bibRefCitation>
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reported it from the Osogovo Mountains, Kalin Kamen area, 1560 m a.s.l., Kriva Palanka district, near to the border with Bulgaria.
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia lazarovii" order="Stylommatophora" pageId="3" pageNumber="48" phylum="Mollusca" rank="species" species="lazarovii">Vestia lazarovii</taxonomicName>
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sp. n. is the second representative of the genus from the Republic of Macedonia and occurs relatively high up in the mountains (in coniferous forests and its ecotone) and is characterized by a quite strong reduction of the clausilium apparatus (reduced lunella, short principal and short or missing upper palatal plicae, missing basalis, somethimes very fine and thin clausilium plate with weakly developed hook at its end). A connection between superior and spiral lamellae is typical for the
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<pageBreakToken pageId="4" pageNumber="49" start="start">genus</pageBreakToken>
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia" order="Stylommatophora" pageId="4" pageNumber="49" phylum="Mollusca" rank="genus">Vestia</taxonomicName>
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, so the specimens with disconnected superior and spiral lamellae could be also interpreted as showing initial reduction in this part of the clausilium apparatus.
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</paragraph>
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<caption ID-Table-UUID="69F01BF9FB5F568D115B0AB79DA0D0AD" httpUri="http://table.plazi.org/id/69F01BF9FB5F568D115B0AB79DA0D0AD" pageId="4" pageNumber="49">
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<paragraph pageId="4" pageNumber="49">
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Table 2. Measurements (mm) of the
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia lazarovii" order="Stylommatophora" pageId="4" pageNumber="49" phylum="Mollusca" rank="species" species="lazarovii">Vestia lazarovii</taxonomicName>
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sp. n. and variation of the clausilium apparatus. Abbreviation: H - height of shell, D - diameter of shell, W - number of whorls, We - number of whorls of the protoconch, HP - height of peristome, DP - diameter of aperture, R - ribs on the last whorl. Holotype -
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<normalizedToken originalValue="№">No</normalizedToken>
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6.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="49">
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<table pageId="4" pageNumber="49">
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<tr pageId="4" pageNumber="49">
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">H</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">D</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">W</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">We</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">
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<sub pageId="4" pageNumber="49">H</sub>
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P
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</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">
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<sub pageId="4" pageNumber="49">D</sub>
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P
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</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">R</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">superior+spiralis</th>
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<th colspan="1" pageId="4" pageNumber="49" rowspan="1">hook shape of clausilium</th>
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</tr>
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<tr pageId="4" pageNumber="49">
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">Average</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">10</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">3.13</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">8.95</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">2.35</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">2.88</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">2.22</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">45.4</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">--</td>
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</tr>
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<tr pageId="4" pageNumber="49">
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">Variance</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.41</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.01</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.19</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.06</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.01</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">0.02</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">17.6</td>
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<td colspan="1" pageId="4" pageNumber="49" rowspan="1">--</td>
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</tr>
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</table>
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</paragraph>
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<caption pageId="4" pageNumber="49">
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<paragraph pageId="4" pageNumber="49">
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Figure 3.
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<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Vestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vestia lazarovii" order="Stylommatophora" pageId="4" pageNumber="49" phylum="Mollusca" rank="species" species="lazarovii">Vestia lazarovii</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |