870 lines
153 KiB
XML
870 lines
153 KiB
XML
<document id="1F521C1381E8CEF9279DF08D2BEFF547" ID-DOI="10.5281/zenodo.189206" ID-GBIF-Dataset="60cbb643-8df2-44b2-b799-b8b1fe7b61a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="189206" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100215462" checkinUser="plazi" docAuthor="Hervé, Christophe, Roberts, Michael J. & Murphy, John A." docDate="2009" docId="A552823FBF4AFFB8FF7E96FB1A24E857" docLanguage="en" docName="zt02171p028.pdf" docOrigin="Zootaxa 2171" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drassodex Murphy 2007" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="596BFA47BF48FFBFFFE990421B59EC7C" masterDocTitle="A taxonomic revision of the genus Drassodex Murphy, 2007 (Araneae: Gnaphosidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="3" updateTime="1698591093210" updateUser="plazi">
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<mods:titleInfo id="3F5A7C507DCCAC5DDDA8767BFA1A0BD4">
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<mods:title id="3DFF1669A3C063F56CA928296E14EB42">A taxonomic revision of the genus Drassodex Murphy, 2007 (Araneae: Gnaphosidae)</mods:title>
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<mods:namePart id="B004D354F4473D9F08C3914B851860C3">Hervé, Christophe</mods:namePart>
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<mods:namePart id="17BD16ECE7605F785DB6C99D4ED680EE">Roberts, Michael J.</mods:namePart>
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<mods:namePart id="29F22A13C3D613A2B7B15F191C8F0C6D">Murphy, John A.</mods:namePart>
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<mods:title id="26E6B378659FC537DCF7B76C993DE2D7">Zootaxa</mods:title>
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<mods:part id="35397B1E26F877368A940B8229C420D0">
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<mods:date id="BEDC1625211F9D45EE6EECB2350034EA">2009</mods:date>
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<mods:number id="FAC7C0C0AD5A13448AE8494076A712C0">2171</mods:number>
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<treatment id="A552823FBF4AFFB8FF7E96FB1A24E857" ID-DOI="http://doi.org/10.5281/zenodo.5663944" ID-GBIF-Taxon="119621558" ID-Zenodo-Dep="5663944" LSID="urn:lsid:plazi:treatment:A552823FBF4AFFB8FF7E96FB1A24E857" httpUri="http://treatment.plazi.org/id/A552823FBF4AFFB8FF7E96FB1A24E857" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
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<subSubSection id="65E160A2BF4AFFBDFF7E96FB1A95EAAE" box="[151,460,1720,1747]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="2D443329BF4AFFBDFF7E96FB1A95EAAE" blockId="2.[151,460,1720,1747]" box="[151,460,1720,1747]" pageId="2" pageNumber="3">
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<heading id="760C8445BF4AFFBDFF7E96FB1A95EAAE" bold="true" box="[151,460,1720,1747]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="EAFB48AABF4AFFBDFF7E96FB1A95EAAE" ID-CoL="486Q" ID-ENA="2338157" authority="Murphy, 2007" authorityName="Murphy" authorityYear="2007" box="[151,460,1720,1747]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E96FB1A95EAAE" bold="true" box="[151,460,1720,1747]" pageId="2" pageNumber="3">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E96FB1A4FEAAF" bold="true" box="[151,278,1721,1747]" italics="true" pageId="2" pageNumber="3">Drassodex</emphasis>
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<bibRefCitation id="496A4ED8BF4AFFBDFEF496FA1A95EAAE" author="Murphy" box="[285,460,1720,1746]" pageId="2" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy, 2007</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="65E160A2BF4AFFBDFF7E974018C8EB65" box="[151,913,1794,1818]" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="2D443329BF4AFFBDFF7E974018C8EB65" blockId="2.[151,1224,1794,1881]" box="[151,913,1794,1818]" pageId="2" pageNumber="3">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E97401A74EB66" bold="true" box="[151,301,1794,1818]" pageId="2" pageNumber="3">
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<typeStatus id="F2408D8BBF4AFFBDFF7E97401B96EB66" box="[151,207,1794,1818]" pageId="2" pageNumber="3">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="EAFB48AABF4AFFBDFEDA974019D0EB65" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[307,649,1794,1817]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hypocrita">
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<emphasis id="1F8FEF3BBF4AFFBDFEDA97401AAEEB65" box="[307,503,1794,1817]" italics="true" pageId="2" pageNumber="3">Drassus hypocrita</emphasis>
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<bibRefCitation id="496A4ED8BF4AFFBDFE17974019D0EB65" author="Simon" box="[510,649,1794,1817]" pageId="2" pageNumber="27" refString="Simon, E. (1878) Les arachnides de France. Roret, Paris, 4, 1 - 334." type="journal article" year="1878">Simon, 1878</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="65E160A2BF4AFFBDFF7E976019C1EB25" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="2D443329BF4AFFBDFF7E976019C1EB25" blockId="2.[151,1224,1794,1881]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="0DEB1407BF4AFFBDFF7E976019C1EB25" pageId="2" pageNumber="3">
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<taxonomicName id="EAFB48AABF4AFFBDFF7E97601A95EB45" ID-CoL="486P" authority="Simon, 1893: 359" authorityName="Simon" authorityPageNumber="359" authorityYear="1893" box="[151,460,1826,1849]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E97601A5EEB45" box="[151,263,1826,1849]" italics="true" pageId="2" pageNumber="3">Drassodes</emphasis>
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<treatmentCitation id="AC5A1538BF4AFFBDFEE797601A95EB45" author="Simon" box="[270,460,1826,1849]" page="359" pageId="2" pageNumber="3" year="1893">
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<bibRefCitation id="496A4ED8BF4AFFBDFEE797601AC3EB45" author="Simon" box="[270,410,1826,1849]" pageId="2" pageNumber="27" refString="Simon, E. (1893) Histoire naturelle des araignees. Roret, Paris, 1, 257 - 488." type="journal article" year="1893">Simon, 1893</bibRefCitation>
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: 359
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</treatmentCitation>
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</taxonomicName>
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(in part,
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<taxonomicName id="EAFB48AABF4AFFBDFDC7976019D4EB45" box="[558,653,1826,1849]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hispanus">
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<emphasis id="1F8FEF3BBF4AFFBDFDC7976019D4EB45" box="[558,653,1826,1849]" italics="true" pageId="2" pageNumber="3">hispanus</emphasis>
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</taxonomicName>
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-group);
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<treatmentCitation id="AC5A1538BF4AFFBDFD02976018F0EB45" author="Simon" box="[747,937,1826,1849]" page="120" pageId="2" pageNumber="3" year="1914">
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<bibRefCitation id="496A4ED8BF4AFFBDFD029760182EEB45" author="Simon" box="[747,887,1826,1849]" pageId="2" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon, 1914</bibRefCitation>
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: 120
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</treatmentCitation>
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(in part,
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<taxonomicName id="EAFB48AABF4AFFBDFBE297601F28EB45" box="[1035,1137,1826,1849]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hypocrita">
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<emphasis id="1F8FEF3BBF4AFFBDFBE297601F28EB45" box="[1035,1137,1826,1849]" italics="true" pageId="2" pageNumber="3">hypocrita</emphasis>
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</taxonomicName>
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-group).
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<taxonomicName id="EAFB48AABF4AFFBDFF7E97001AF3EB25" ID-CoL="486Q" authority="Murphy, 2007" authorityName="Murphy" authorityYear="2007" box="[151,426,1858,1881]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E97001A50EB25" box="[151,265,1858,1881]" italics="true" pageId="2" pageNumber="3">Drassodex</emphasis>
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<bibRefCitation id="496A4ED8BF4AFFBDFEE697001AF3EB25" author="Murphy" box="[271,426,1858,1881]" pageId="2" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy, 2007</bibRefCitation>
|
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</taxonomicName>
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||
: x, 11;
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||
<bibRefCitation id="496A4ED8BF4AFFBDFE1E970019CCEB25" author="Platnick" box="[503,661,1858,1881]" pageId="2" pageNumber="27" refString="Platnick, N. I. (2008) The world spider catalog, version 9.0 American Museum of Natural History, online at http: // research. amnh. org / entomology / spiders / catalog / index. html (Accessed 15 September 2008)" type="journal article" year="2008">Platnick, 2008</bibRefCitation>
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.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="65E160A2BF4AFFBDFF7E97CA1E72EBB6" pageId="2" pageNumber="3" type="discussion">
|
||
<paragraph id="2D443329BF4AFFBDFF7E97CA1E72EBB6" blockId="2.[151,1436,1928,1994]" pageId="2" pageNumber="3">
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<emphasis id="1F8FEF3BBF4AFFBDFF7E97CA1A5FEBDE" bold="true" box="[151,262,1928,1954]" pageId="2" pageNumber="3">Remark.</emphasis>
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The illustration of the male bulb of “
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<taxonomicName id="EAFB48AABF4AFFBDFD2E97CB181DEBDE" box="[711,836,1929,1954]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFD2E97CB181DEBDE" box="[711,836,1929,1954]" italics="true" pageId="2" pageNumber="3">Drassodes</emphasis>
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</taxonomicName>
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gr.
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<taxonomicName id="EAFB48AABF4AFFBDFC9A97CB18BDEBDE" box="[883,996,1929,1954]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hypocrita">
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<emphasis id="1F8FEF3BBF4AFFBDFC9A97CB18BDEBDE" box="[883,996,1929,1954]" italics="true" pageId="2" pageNumber="3">hypocrita</emphasis>
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</taxonomicName>
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” by
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<bibRefCitation id="496A4ED8BF4AFFBDFBC997CB1E1AEBDF" author="Barrientos" box="[1056,1347,1929,1955]" pageId="2" pageNumber="27" refString="Barrientos, J. A. (1986) Algunas aranas de los entornos de la Laguna de Sarinena (Huesca). Coleccion de Estudios Altoaragoneses, 6, 101 - 106." type="journal article" year="1986">Barrientos (1986: fig. 2)</bibRefCitation>
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clearly shows that the specimen doesn’t belong to the genus
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<taxonomicName id="EAFB48AABF4AFFBDFCEC97F218DBEBB5" box="[773,898,1968,1993]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFCEC97F218DBEBB5" box="[773,898,1968,1993]" italics="true" pageId="2" pageNumber="3">Drassodex</emphasis>
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</taxonomicName>
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but to the genus
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<taxonomicName id="EAFB48AABF4AFFBDFBA297F21E72EBB6" authority="Latreille." authorityName="Latreille." box="[1099,1323,1968,1994]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1F8FEF3BBF4AFFBDFBA297F21FE1EBB5" box="[1099,1208,1968,1993]" italics="true" pageId="2" pageNumber="3">Clubiona</emphasis>
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Latreille.
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="65E160A2BF4BFFBCFF2F90D51860EE48" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="2D443329BF4BFFBCFF2F90D51F98EDBC" blockId="3.[151,1437,151,1995]" pageId="3" pageNumber="4">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFF2F90D51A1FECCD" bold="true" box="[198,326,151,177]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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Members of the genus
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<taxonomicName id="EAFB48AABF4BFFBCFD9090DA19AFECCD" box="[633,758,152,177]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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||
<emphasis id="1F8FEF3BBF4BFFBCFD9090DA19AFECCD" box="[633,758,152,177]" italics="true" pageId="3" pageNumber="4">Drassodex</emphasis>
|
||
</taxonomicName>
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can be easily distinguished from all other Palaearctic gnaphosids by the combination of the following characters (those in parentheses are characters observed in
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<emphasis id="1F8FEF3BBF4BFFBCFF7E90A71AF7EC82" box="[151,430,229,254]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="EAFB48AABF4BFFBCFF7E90A71A02EC82" box="[151,347,229,254]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sensu">
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Drassodes
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<taxonomicNameLabel id="04BC5240BF4BFFBCFEF390A71A02EC82" box="[282,347,229,254]" pageId="3" pageNumber="4" sensu="stricto">sensu</taxonomicNameLabel>
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</taxonomicName>
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stricto
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</emphasis>
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): metatarsal preening combs on legs III and IV absent, replaced by preening brushes, dorsal male scutum absent, plain-coloured abdomen, dorsal spines on tibiae III and IV absent (present), trochanters slightly notched (deeply notched), retromarginal teeth as large as distal promarginal teeth (small or minute retromarginal teeth), and presence of some singular and well defined genitalic characters, as bifid RTA, highly coiled or twisted embolus, very deep epigynal atrium, bordered with slightly sclerotized lateral and hood-like anterior margins, and strongly corkscrew-shaped internal membranous coils.
|
||
</paragraph>
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||
<paragraph id="2D443329BF4BFFBCFF2F918F1860EE48" blockId="3.[151,1437,151,1995]" pageId="3" pageNumber="4">
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||
<emphasis id="1F8FEF3BBF4BFFBCFF2F918F1AC2ED9B" bold="true" box="[198,411,461,487]" pageId="3" pageNumber="4">Included species.</emphasis>
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<taxonomicName id="EAFB48AABF4BFFBCFE4A918F19A0ED9B" authority="L. Koch, 1866" authorityName="L. Koch" authorityYear="1866" box="[419,761,461,487]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hispanus">
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<emphasis id="1F8FEF3BBF4BFFBCFE4A918F1969ED9A" box="[419,560,461,486]" italics="true" pageId="3" pageNumber="4">D. hispanus</emphasis>
|
||
(
|
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<bibRefCitation id="496A4ED8BF4BFFBCFDA8918F19A9ED9B" author="Koch" box="[577,752,461,487]" pageId="3" pageNumber="27" refString="Koch, L. (1866) Die Arachniden-Familie der Drassiden. Hefte 1 - 6. Nurnberg, pp. 1 - 304." type="book chapter" year="1866">L. Koch, 1866</bibRefCitation>
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||
)
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</taxonomicName>
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,
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<taxonomicName id="EAFB48AABF4BFFBCFCEF918F1F45ED9B" authority="Pavesi, 1873" authorityName="Pavesi" authorityYear="1873" box="[774,1052,461,487]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heeri">
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||
<emphasis id="1F8FEF3BBF4BFFBCFCEF918F1831ED9A" box="[774,872,461,486]" italics="true" pageId="3" pageNumber="4">D. heeri</emphasis>
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(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFC9E918F1F4AED9B" author="Pavesi" box="[887,1043,461,487]" pageId="3" pageNumber="27" refString="Pavesi, P. (1873) Catalogo sistematico dei ragni del cantone ticino con la loro distribuzione orizontale e verticale e cenni sulla araneologica elvetica. Annali del Museo civico di storia naturale di Genova, 4, 5 - 215." type="journal article" year="1873">Pavesi, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFBC0918F1E2AED9B" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[1065,1395,461,487]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFBC0918F1FE7ED9A" box="[1065,1214,461,486]" italics="true" pageId="3" pageNumber="4">D. hypocrita</emphasis>
|
||
(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFB27918F1E33ED9B" author="Simon" box="[1230,1386,461,487]" pageId="3" pageNumber="27" refString="Simon, E. (1878) Les arachnides de France. Roret, Paris, 4, 1 - 334." type="journal article" year="1878">Simon, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFA96918F1AE9EE72" authority="Simon, 1914" authorityName="Simon" authorityYear="1914" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cervinus">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFA96918F1BA2EE71" italics="true" pageId="3" pageNumber="4">D. cervinus</emphasis>
|
||
(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFEE391B61AFFEE72" author="Simon" box="[266,422,500,526]" pageId="3" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFE5591B619A2EE72" authority="Simon, 1914" authorityName="Simon" authorityYear="1914" box="[444,763,500,526]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFE5591B6191FEE71" box="[444,582,500,525]" italics="true" pageId="3" pageNumber="4">D. fritillifer</emphasis>
|
||
(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFDBF91B619ABEE72" author="Simon" box="[598,754,500,526]" pageId="3" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFCEE91B61F64EE72" authority="Simon, 1914" authorityName="Simon" authorityYear="1914" box="[775,1085,500,526]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="validior">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFCEE91B618D0EE71" box="[775,905,500,525]" italics="true" pageId="3" pageNumber="4">D. validior</emphasis>
|
||
(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFC7091B61F6DEE72" author="Simon" box="[921,1076,500,526]" pageId="3" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFBA391B61ECEEE72" authority="Schenkel, 1936" authorityName="Schenkel" authorityYear="1936" box="[1098,1431,500,526]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lesserti">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFBA391B61F9DEE71" box="[1098,1220,500,525]" italics="true" pageId="3" pageNumber="4">D. lesserti</emphasis>
|
||
(
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFB3A91B61ED6EE72" author="Schenkel" box="[1235,1423,500,526]" pageId="3" pageNumber="27" refString="Schenkel, E. (1936) Kleine Beitrage zur Spinnenkunde. II. Teil. Revue suisse de zoolologie, 43, 307 - 333." type="journal article" year="1936">Schenkel, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFF7E92581A50EE4F" box="[151,265,538,563]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFF7E92581A50EE4F" box="[151,265,538,563]" italics="true" pageId="3" pageNumber="4">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4BFFBCFEF992581A37EE48" bold="true" box="[272,366,538,564]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="04BC5240BF4BFFBCFEF992581A37EE48" box="[272,366,538,564]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="EAFB48AABF4BFFBCFE9C92581AB7EE4F" box="[373,494,538,563]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFE9C92581AB7EE4F" box="[373,494,538,563]" italics="true" pageId="3" pageNumber="4">D. drescoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4BFFBCFE1C9258190BEE48" bold="true" box="[501,594,538,564]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="04BC5240BF4BFFBCFE1C9258190BEE48" box="[501,594,538,564]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4BFFBCFD8B9258198DEE4F" box="[610,724,538,563]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="granja" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFD8B9258198DEE4F" box="[610,724,538,563]" italics="true" pageId="3" pageNumber="4">D. granja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4BFFBCFD3292581860EE48" bold="true" box="[731,825,538,564]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="04BC5240BF4BFFBCFD3292581860EE48" box="[731,825,538,564]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E160A2BF4BFFBBFF2F92021F69E82E" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="2D443329BF4BFFBBFF2F920219E7ECA4" blockId="3.[151,1437,151,1995]" lastBlockId="4.[151,1437,152,1106]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<emphasis id="1F8FEF3BBF4BFFBCFF2F92021A06EE26" bold="true" box="[198,351,576,602]" pageId="3" pageNumber="4">Description.</emphasis>
|
||
Total length: 8.09–17.57. Carapace (
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFCC992031FC2EE27" author="Murphy" box="[800,1179,577,603]" pageId="3" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007" yearSuffix="p">Murphy 2007: figs p. 232, 234</bibRefCitation>
|
||
) oval in dorsal view, widest between coxae II and III, with ocular area narrowed, light brown posteriorly, darker anteriorly, without darkened lateral edge; cephalic area not elevated; thoracic groove long, well developed, longitudinal. From front, anterior eye row slightly recurved, almost straight, posterior row recurved; from above, anterior row slightly recurved, posterior row straight; AME circular, dark; PME irregularly circular, light; lateral eyes oval, light; eyes almost subequal; AME separated by their diameter or more, by slightly less to slightly more than their diameter from ALE; PME separated by slightly more their diameter, and by at least twice their diameter from PLE; lateral eyes of each side separated by their diameter or more; MOQ slightly longer than wide, roughly rectangular, often shallowly narrowed anteriorly; clypeal height equal or greater than AME diameter. Chelicerae (
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFECA93DF19D7EFCB" author="Murphy" box="[291,654,925,951]" pageId="3" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007" yearSuffix="p">Murphy 2007: figs p. 232, 234</bibRefCitation>
|
||
) convex anteriorly near the base, with small lateral condyle, partly hidden under the carapace; three discrete promarginal teeth, apical and medium large, subequal or apical very slightly smaller, basal mostly weaker, sometimes minute, situated just near medium tooth; two discrete and large retromarginal teeth, separated much as the apical and medium promarginal teeth; in males, chelicerae often more elongated, laterally slightly more convex in the middle, fang longer, teeth smaller, especially apical of each margin; chilum triangular, partly divided. Endites slightly depressed in females, more in males, straight, parallel, with long internal distal scopula and weak external distal serrula; labium roughly square or slightly longer than wide; sternum shield-shaped, with dark setae covering entire surface, weakly rebordered, with short extensions to and between coxae; trochanters only shallowly notched. Leg formula
|
||
<quantity id="EA039ECCBF4BFFBCFB0C94901E67E890" box="[1253,1342,1234,1260]" metricMagnitude="2" metricUnit="m" metricValue="1.047242" pageId="3" pageNumber="4" unit="in" value="4123.0">4123 in</quantity>
|
||
female,
|
||
<quantity id="EA039ECCBF4BFFBCFF7E94BB1BB6E96F" box="[151,239,1273,1299]" metricMagnitude="1" metricUnit="m" metricValue="3.61442" pageId="3" pageNumber="4" unit="in" value="1423.0">1423 in</quantity>
|
||
male. Typical leg spination pattern (only surfaces bearing spines listed): femora: I, II d1-0-0; I p0-0-1; II p0-1-1; III, IV d1-1-0; III p0-1-1, r0-0(1)-1; IV r0-0-1; tibiae: I
|
||
<date id="594515E9BF4BFFBCFC429562189FE946" box="[939,966,1312,1338]" pageId="3" pageNumber="4">v1</date>
|
||
p-2-0; II v0-1p-0; III, IV p1-1, r1-1; III
|
||
<date id="594515E9BF4BFFBCFF7E95041BEDE91C" box="[151,180,1350,1376]" pageId="3" pageNumber="4">v1</date>
|
||
p-1p-2; IV
|
||
<date id="594515E9BF4BFFBCFEAC95041AD8E91C" box="[325,385,1350,1376]" pageId="3" pageNumber="4">v2-2</date>
|
||
-2; metatarsi: I, II
|
||
<date id="594515E9BF4BFFBCFD81950419D2E91C" box="[616,651,1350,1376]" pageId="3" pageNumber="4">v2</date>
|
||
-0-0; III, IV p1-2-2; III
|
||
<date id="594515E9BF4BFFBCFC5F95041881E91C" box="[950,984,1350,1376]" pageId="3" pageNumber="4">v2</date>
|
||
-0-2, r0(1)-2-2; IV
|
||
<date id="594515E9BF4BFFBCFB2195041FBCE91C" box="[1224,1253,1350,1376]" pageId="3" pageNumber="4">v1</date>
|
||
r-1p-2, r1-2-2. Metatarsal preening comb lacking, but presence of a distal preening brush under metatarsi of both posterior legs. Metatarsi I and II and all the tarsi thickly scopulate, tibiae I and II weakly scopulate, often only the prolateral side, between the median spine(s) and the tip. Tarsi with two weakly pectinate claws and with claw tufts. Tarsi and metatarsi of the first pair of male legs ventrally with a series of long sensitive setae, retrolaterally directed. Palpal femur with 1 dorsomedial spine and 1 dorsodistal spine, often a second spine in prolateral position. Abdomen (
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFDED966C182FEA34" author="Murphy" box="[516,886,1582,1608]" pageId="3" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007" yearSuffix="p">Murphy 2007: figs p. 232, 234</bibRefCitation>
|
||
) usually pale to dark grey, sometimes with an inconspicuous darkened cardiac mark and dots, and with conspicuous anterior tuft of hairs; venter same colour or lighter, with two closely spaced longitudinal rows of dark spots, beginning behind the epigastric fold, and two short wider spaced rows, beginning in front of the anterior lateral spinnerets; males without dorsal scutum. Six spinnerets (
|
||
<figureCitation id="B5C02FACBF4BFFBCFDE1968B1909EA9F" box="[520,592,1737,1763]" captionStart="FIGURE 1" captionStartId="5.[151,255,1141,1165]" captionTargetBox="[230,1354,205,1110]" captionTargetId="figure@5.[219,1367,194,1117]" captionTargetPageId="5" captionText="FIGURE 1. Drassodex simoni sp. nov. Spinnerets and spigots of adult female (left) and male (right), showing position of the anterior lateral (ALS), posterior median (PMS), and posterior lateral (PLS) spinnerets, along with the distribution of spigots presumed to serve the major ampullate (MA), piriform (P), minor ampullate (mA), cylindrical (C) and aciniform (A) glands. The position of the annular crescent (AC) is indicated by an arrow (after Murphy 2007: fig. p. 235)." httpUri="https://zenodo.org/record/189207/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="496A4ED8BF4BFFBCFD89968B1851EA9F" author="Murphy" box="[608,776,1737,1763]" pageId="3" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
|
||
: figs p. 232–235), ALS large, cylindrical, separated by slightly less than their diameter, apically with two major ampullate gland spigots and with eleven to sixteen piriform gland spigots, situated around the edge and middle of the spinneret tip; a pair of small sclerotized crescent, from which emerges a fringe of closely spaced and thin setae, is present on the ALS, on the prolateral and retrolateral sides of the tip of those spinnerets; PMS are the smallest, apically with minor ampullate gland spigots (two in female, one in male) and numerous aciniform gland spigots, and subapically, on the posterior side, with five cylindrical gland spigots, exclusively in female; PLS are smaller and narrow than the ALS, apically with one minor ampullate gland spigot, two cylindrical gland spigots, exclusively in female, and numerous aciniform gland spigots.
|
||
</paragraph>
|
||
<paragraph id="2D443329BF4CFFBBFF2F90A719B1EE27" blockId="4.[151,1437,152,1106]" pageId="4" pageNumber="5">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF2F90A71A67EC82" box="[198,318,229,254]" italics="true" pageId="4" pageNumber="5">Male palp</emphasis>
|
||
. RTA bifid (
|
||
<figureCitation id="B5C02FACBF4CFFBBFE3A90A71967EC83" box="[467,574,229,255]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 5, 9</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFDA290A71933EC83" box="[587,618,229,255]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">13</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD9E90A7199DEC83" box="[631,708,229,255]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">23, 27</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD3990A719B6EC83" box="[720,751,229,255]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">31</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD1590A71810EC83" box="[764,841,229,255]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">44, 48</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFCBC90A7182FEC83" box="[853,886,229,255]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">52</figureCitation>
|
||
), inferior prong lamellar, rounded, often blunt, sometimes absent (
|
||
<taxonomicName id="EAFB48AABF4CFFBBFE94914E1950ED59" box="[381,521,268,293]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFE94914E1950ED59" box="[381,521,268,293]" italics="true" pageId="4" pageNumber="5">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
only,
|
||
<figureCitation id="B5C02FACBF4CFFBBFDBC914E19F7ED5A" box="[597,686,268,294]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">Fig. 37</figureCitation>
|
||
); superior prong more sclerotized and sharp, regularly curved, cat-claw shaped (
|
||
<figureCitation id="B5C02FACBF4CFFBBFE8891701AE2ED30" box="[353,443,306,332]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">Figs 13</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFE2E9170194BED30" box="[455,530,306,332]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">23, 27</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFDF791701967ED30" box="[542,574,306,332]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">37</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFDA0917019CDED30" box="[585,660,306,332]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">44, 48</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD49917019E6ED30" box="[672,703,306,332]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">52</figureCitation>
|
||
) or sometimes almost straight (
|
||
<figureCitation id="B5C02FACBF4CFFBBFBC591701FCDED30" box="[1068,1172,306,332]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 5, 9</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFB4991701F99ED30" box="[1184,1216,306,332]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">31</figureCitation>
|
||
), often longer than inferior prong. In ventral view, the embolus, arising mesally, is long and corkscrew-shaped, and is associated for almost its entire length with a haematodocha, together forming a huge coiled structure (
|
||
<figureCitation id="B5C02FACBF4CFFBBFB3891C21E65EDE6" box="[1233,1340,384,410]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 4, 8</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFAA391C21E33EDE6" box="[1354,1386,384,410]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">12</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFA9E91C21BEEEDBC" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">22, 26</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFF2B91E41BBBEDBC" box="[194,226,422,448]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">30</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFF0791E41A60EDBC" box="[238,313,422,448]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">43, 47</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFEAD91E41A3DEDBC" box="[324,356,422,448]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">51</figureCitation>
|
||
), which spirals upwards either along one axis (
|
||
<figureCitation id="B5C02FACBF4CFFBBFC6D91E41886EDBC" box="[900,991,422,448]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFC0291E41F6FEDBC" box="[1003,1078,422,448]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">22, 26</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFBA891E41F39EDBC" box="[1089,1120,422,448]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">30</figureCitation>
|
||
) or two axes (
|
||
<figureCitation id="B5C02FACBF4CFFBBFAEA91E41E33EDBC" box="[1283,1386,422,448]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 4, 8</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFA9E91E41BEEED9B" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">43, 47</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFF2D918F1BBDED9B" box="[196,228,461,487]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">51</figureCitation>
|
||
); sometimes the embolar coil is very compact and twisted (
|
||
<taxonomicName id="EAFB48AABF4CFFBBFC4D918F1F74ED9A" box="[932,1069,461,486]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFC4D918F1F74ED9A" box="[932,1069,461,486]" italics="true" pageId="4" pageNumber="5">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
only,
|
||
<figureCitation id="B5C02FACBF4CFFBBFB9F918F1F97ED9B" box="[1142,1230,461,487]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">Fig. 36</figureCitation>
|
||
); terminal part of the coil is either simple or provided with an enveloping scoop-shaped embolar apophysis, projecting forward (
|
||
<figureCitation id="B5C02FACBF4CFFBBFF7692581A55EE48" box="[159,268,538,564]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 4, 8</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFEF392581A62EE48" box="[282,315,538,564]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">47</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFEA192581A33EE48" box="[328,362,538,564]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">51</figureCitation>
|
||
). Median apophysis arising near the embolar coil, stout and hook-shaped, either pointing inwards or prolaterally (
|
||
<taxonomicName id="EAFB48AABF4CFFBBFE5A92031962EE26" box="[435,571,577,602]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFE5A92031962EE26" box="[435,571,577,602]" italics="true" pageId="4" pageNumber="5">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
only,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD6D92031983EE27" box="[644,730,577,603]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2D443329BF4CFFBBFF2F922A1F69E82E" blockId="4.[151,1437,152,1106]" pageId="4" pageNumber="5">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF2F922A1A7FEEFD" box="[198,294,616,641]" italics="true" pageId="4" pageNumber="5">Epigyne</emphasis>
|
||
. Epigynal atrium often very deep, higher than wide to strongly wider than high (0.28–0.71 wide, 0.23–0.64 high, ratio 0.80–1.91) (see
|
||
<tableCitation id="60790692BF4CFFBBFD8A92CC19E4EED4" box="[611,701,654,680]" captionStart="TABLE 1" captionStartId="4.[151,239,1156,1180]" captionTargetBox="[165,1342,1213,1783]" captionTargetPageId="4" captionText="TABLE 1. Width, height (in mm) and ratios width / height of the atrium of the species of Drassodex." httpUri="http://table.plazi.org/id/798463A1BF4CFFBBFF7E94C61F99E8E0" pageId="4" pageNumber="5" tableUuid="798463A1BF4CFFBBFF7E94C61F99E8E0">Table 1</tableCitation>
|
||
for details), variously shaped; atrium bordered with slightly sclerotized and often very curved lateral margins, which are notched near the epigastric furrow, forming sockets (
|
||
<figureCitation id="B5C02FACBF4CFFBBFEEA929E1A0DEE8A" box="[259,340,732,758]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFE8A929E1ADCEE8A" box="[355,389,732,758]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">10</figureCitation>
|
||
); lateral margins converging anteriorly to a variously shaped hood-like arch, simply convex or straight (
|
||
<figureCitation id="B5C02FACBF4CFFBBFE9593401A81EF60" box="[380,472,770,796]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFE0D93401969EF60" box="[484,560,770,796]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">20, 24</figureCitation>
|
||
) or often provided with a tongue-shaped (
|
||
<figureCitation id="B5C02FACBF4CFFBBFBC993401FD3EF60" box="[1056,1162,770,796]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 2, 6</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFB7193401E49EF60" box="[1176,1296,770,796]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">10, 14, 18</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFAF793401E67EF60" box="[1310,1342,770,796]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">28</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFAA393401BEEEF3F" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">32, 34, 39</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFF2A936B1BB8EF3F" box="[195,225,809,835]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">41</figureCitation>
|
||
) or tubercle-shaped projection (
|
||
<figureCitation id="B5C02FACBF4CFFBBFDBE936B19EBEF3F" box="[599,690,809,835]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">Figs 45</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD54936B186FEF3F" box="[701,822,809,835]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">49, 53, 55</figureCitation>
|
||
); lateral margins connected posteriorly by a thin and slightly chitinized ridge; copulatory openings situated anteriorly, separated by a thin to large septum, often partially hidden by the anterior arch or tongue when viewed from above, but easily visible from a posterior view (
|
||
<figureCitation id="B5C02FACBF4CFFBBFF3693DF1A7EEFCB" box="[223,295,925,951]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
); membranous coils wide and very high, corkscrew-shaped much as the embolar coils of the male palps, associated with long and thin coiled ducts, which are connected posteriorly to flattened lanceolate structures and distally to very small bursae (
|
||
<figureCitation id="B5C02FACBF4CFFBBFD5F93A8185CE878" box="[694,773,1002,1028]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFCFD93A8186DE878" box="[788,820,1002,1028]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">11</figureCitation>
|
||
); major part of the coiled ducts situated inside the membranous coils (axial, spiral-column shaped) (
|
||
<figureCitation id="B5C02FACBF4CFFBBFD099453189BE857" box="[736,962,1041,1067]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="4" pageNumber="5">Figs 11, 15, 17, 19</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFC2694531F45E857" box="[975,1052,1041,1067]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="4" pageNumber="5">21, 25</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFBC094531F10E857" box="[1065,1097,1041,1067]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="4" pageNumber="5">29</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFBBF94531F2CE857" box="[1110,1141,1041,1067]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">33</figureCitation>
|
||
) or joined with the outer walls of these coils (external, peripheral) (
|
||
<figureCitation id="B5C02FACBF4CFFBBFD6D947A19B4E82E" box="[644,749,1080,1106]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="4" pageNumber="5">Figs 3, 7</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFD13947A181FE82E" box="[762,838,1080,1106]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="4" pageNumber="5">35, 40</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFCBB947A18C7E82E" box="[850,926,1080,1106]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="4" pageNumber="5">42, 46</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4CFFBBFC43947A1F7AE82E" box="[938,1059,1080,1106]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="4" pageNumber="5">50, 54, 56</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="798463A1BF4CFFBBFF7E94C61F99E8E0" ID-Table-UUID="798463A1BF4CFFBBFF7E94C61F99E8E0" box="[151,1216,1156,1180]" httpUri="http://table.plazi.org/id/798463A1BF4CFFBBFF7E94C61F99E8E0" pageId="4" pageNumber="5" targetBox="[165,1342,1213,1783]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="2D443329BF4CFFBBFF7E94C61F99E8E0" blockId="4.[151,1216,1156,1180]" box="[151,1216,1156,1180]" pageId="4" pageNumber="5">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF7E94C61A51E8E0" bold="true" box="[151,264,1156,1180]" pageId="4" pageNumber="5">TABLE 1.</emphasis>
|
||
Width, height (in mm) and ratios width/height of the atrium of the species of
|
||
<taxonomicName id="EAFB48AABF4CFFBBFBAE94C71FE0E8E0" box="[1095,1209,1157,1180]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFBAE94C71FE0E8E0" box="[1095,1209,1157,1180]" italics="true" pageId="4" pageNumber="5">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D443329BF4CFFBBFF4C94961E67EA8B" pageId="4" pageNumber="5">
|
||
<table id="5FFBC189BF4C0040FF4C94FF1E67EA8B" box="[165,1342,1213,1783]" gridcols="4" gridrows="12" pageId="4" pageNumber="5">
|
||
<tr id="93CB316BBF4C0040FF4C94FF1E67E97E" box="[165,1342,1213,1282]" gridrow="0" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C94FF1A2FE97E" box="[165,374,1213,1282]" gridcol="0" gridrow="0" pageId="4" pageNumber="5">Species</th>
|
||
<th id="D01A5817BF4C0040FE1C94FF199AE97E" box="[501,707,1213,1282]" gridcol="1" gridrow="0" pageId="4" pageNumber="5">Atrium width min max</th>
|
||
<th id="D01A5817BF4C0040FCDA94FF1F58E97E" box="[819,1025,1213,1282]" gridcol="2" gridrow="0" pageId="4" pageNumber="5">Atrium height min max</th>
|
||
<th id="D01A5817BF4C0040FB9994FF1E67E97E" box="[1136,1342,1213,1282]" gridcol="3" gridrow="0" pageId="4" pageNumber="5">Ratio width/height min max</th>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C955B1E67E94C" box="[165,1342,1305,1328]" gridrow="1" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C955B1A2FE94C" box="[165,374,1305,1328]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C955B1A7CE94C" box="[165,293,1305,1328]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hispanus">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C955B1A7CE94C" box="[165,293,1305,1328]" italics="true" pageId="4" pageNumber="5">D. hispanus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C955B199AE94C" box="[501,707,1305,1328]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">0.65 0.70</td>
|
||
<td id="D01A5817BF4C0040FCDA955B1F58E94C" box="[819,1025,1305,1328]" gridcol="2" gridrow="1" pageId="4" pageNumber="5">0.51 0.55</td>
|
||
<td id="D01A5817BF4C0040FB99955B1E67E94C" box="[1136,1342,1305,1328]" gridcol="3" gridrow="1" pageId="4" pageNumber="5">1.23 1.31</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C95041E67E921" box="[165,1342,1350,1373]" gridrow="2" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C95041A2FE921" box="[165,374,1350,1373]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C95041A7BE921" box="[165,290,1350,1373]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C95041A7BE921" box="[165,290,1350,1373]" italics="true" pageId="4" pageNumber="5">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C9504199AE921" box="[501,707,1350,1373]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">0.68 0.71</td>
|
||
<td id="D01A5817BF4C0040FCDA95041F58E921" box="[819,1025,1350,1373]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">0.60 0.64</td>
|
||
<td id="D01A5817BF4C0040FB9995041E67E921" box="[1136,1342,1350,1373]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">1.06 1.18</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C95361E67E9F7" box="[165,1342,1396,1419]" gridrow="3" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C95361A2FE9F7" box="[165,374,1396,1419]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C95361BA4E9F7" box="[165,253,1396,1419]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="heeri">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C95361BA4E9F7" box="[165,253,1396,1419]" italics="true" pageId="4" pageNumber="5">D. heeri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C9536199AE9F7" box="[501,707,1396,1419]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">0.37 0.49</td>
|
||
<td id="D01A5817BF4C0040FCDA95361F58E9F7" box="[819,1025,1396,1419]" gridcol="2" gridrow="3" pageId="4" pageNumber="5">0.32 0.36</td>
|
||
<td id="D01A5817BF4C0040FB9995361E67E9F7" box="[1136,1342,1396,1419]" gridcol="3" gridrow="3" pageId="4" pageNumber="5">1.09 1.38</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C95E31E67E9C4" box="[165,1342,1441,1464]" gridrow="4" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C95E31A2FE9C4" box="[165,374,1441,1464]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C95E31BBDE9C4" box="[165,228,1441,1464]" italics="true" pageId="4" pageNumber="5">D. sp.</emphasis>
|
||
(
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF1B95E31A35E9C4" box="[242,364,1441,1464]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="drescoi">
|
||
aff.
|
||
<emphasis id="1F8FEF3BBF4CFFBBFEF795E31A35E9C4" box="[286,364,1441,1464]" italics="true" pageId="4" pageNumber="5">drescoi</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C95E3199AE9C4" box="[501,707,1441,1464]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">0.57 0.67</td>
|
||
<td id="D01A5817BF4C0040FCDA95E31F58E9C4" box="[819,1025,1441,1464]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">0.49 0.58</td>
|
||
<td id="D01A5817BF4C0040FB9995E31E67E9C4" box="[1136,1342,1441,1464]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">1.09 1.17</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C958D1E67E99A" box="[165,1342,1487,1510]" gridrow="5" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C958D1A2FE99A" box="[165,374,1487,1510]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C958D1A4DE99A" box="[165,276,1487,1510]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="drescoi">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C958D1A4DE99A" box="[165,276,1487,1510]" italics="true" pageId="4" pageNumber="5">D. drescoi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C958D199AE99A" box="[501,707,1487,1510]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">0.50 0.66</td>
|
||
<td id="D01A5817BF4C0040FCDA958D1F58E99A" box="[819,1025,1487,1510]" gridcol="2" gridrow="5" pageId="4" pageNumber="5">0.45 0.58</td>
|
||
<td id="D01A5817BF4C0040FB99958D1E67E99A" box="[1136,1342,1487,1510]" gridcol="3" gridrow="5" pageId="4" pageNumber="5">0.86 1.34</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C95BE1E67EA6F" box="[165,1342,1532,1555]" gridrow="6" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C95BE1A2FEA6F" box="[165,374,1532,1555]" gridcol="0" gridrow="6" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C95BE1A54EA6F" box="[165,269,1532,1555]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C95BE1A54EA6F" box="[165,269,1532,1555]" italics="true" pageId="4" pageNumber="5">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C95BE199AEA6F" box="[501,707,1532,1555]" gridcol="1" gridrow="6" pageId="4" pageNumber="5">0.28 0.44</td>
|
||
<td id="D01A5817BF4C0040FCDA95BE1F58EA6F" box="[819,1025,1532,1555]" gridcol="2" gridrow="6" pageId="4" pageNumber="5">0.33 0.46</td>
|
||
<td id="D01A5817BF4C0040FB9995BE1E67EA6F" box="[1136,1342,1532,1555]" gridcol="3" gridrow="6" pageId="4" pageNumber="5">0.80 1.14</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C96681E67EA3D" box="[165,1342,1578,1601]" gridrow="7" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C96681A2FEA3D" box="[165,374,1578,1601]" gridcol="0" gridrow="7" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C96681A75EA3D" box="[165,300,1578,1601]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C96681A75EA3D" box="[165,300,1578,1601]" italics="true" pageId="4" pageNumber="5">D. hypocrita</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C9668199AEA3D" box="[501,707,1578,1601]" gridcol="1" gridrow="7" pageId="4" pageNumber="5">0.37 0.52</td>
|
||
<td id="D01A5817BF4C0040FCDA96681F58EA3D" box="[819,1025,1578,1601]" gridcol="2" gridrow="7" pageId="4" pageNumber="5">0.23 0.32</td>
|
||
<td id="D01A5817BF4C0040FB9996681E67EA3D" box="[1136,1342,1578,1601]" gridcol="3" gridrow="7" pageId="4" pageNumber="5">1.37 1.91</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C96151E67EA12" box="[165,1342,1623,1646]" gridrow="8" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C96151A2FEA12" box="[165,374,1623,1646]" gridcol="0" gridrow="8" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C96151A4AEA12" box="[165,275,1623,1646]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lesserti">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C96151A4AEA12" box="[165,275,1623,1646]" italics="true" pageId="4" pageNumber="5">D. lesserti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C9615199AEA12" box="[501,707,1623,1646]" gridcol="1" gridrow="8" pageId="4" pageNumber="5">0.52 0.68</td>
|
||
<td id="D01A5817BF4C0040FCDA96151F58EA12" box="[819,1025,1623,1646]" gridcol="2" gridrow="8" pageId="4" pageNumber="5">0.31 0.41</td>
|
||
<td id="D01A5817BF4C0040FB9996151E67EA12" box="[1136,1342,1623,1646]" gridcol="3" gridrow="8" pageId="4" pageNumber="5">1.45 1.77</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C96C61E67EAE0" box="[165,1342,1668,1692]" gridrow="9" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C96C61A2FEAE0" box="[165,374,1668,1692]" gridcol="0" gridrow="9" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C96C71A43EAE0" box="[165,282,1669,1692]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="validior">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C96C71A43EAE0" box="[165,282,1669,1692]" italics="true" pageId="4" pageNumber="5">D. validior</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C96C6199AEAE0" box="[501,707,1668,1692]" gridcol="1" gridrow="9" pageId="4" pageNumber="5">0.43 0.55</td>
|
||
<td id="D01A5817BF4C0040FCDA96C61F58EAE0" box="[819,1025,1668,1692]" gridcol="2" gridrow="9" pageId="4" pageNumber="5">0.34 0.50</td>
|
||
<td id="D01A5817BF4C0040FB9996C61E67EAE0" box="[1136,1342,1668,1692]" gridcol="3" gridrow="9" pageId="4" pageNumber="5">1.05 1.34</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C96F01E67EAB5" box="[165,1342,1714,1737]" gridrow="10" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C96F01A2FEAB5" box="[165,374,1714,1737]" gridcol="0" gridrow="10" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C96F01A54EAB5" box="[165,269,1714,1737]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granja">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C96F01A54EAB5" box="[165,269,1714,1737]" italics="true" pageId="4" pageNumber="5">D. granja</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C96F0199AEAB5" box="[501,707,1714,1737]" gridcol="1" gridrow="10" pageId="4" pageNumber="5">0.41 0.53</td>
|
||
<td id="D01A5817BF4C0040FCDA96F01F58EAB5" box="[819,1025,1714,1737]" gridcol="2" gridrow="10" pageId="4" pageNumber="5">0.26 0.33</td>
|
||
<td id="D01A5817BF4C0040FB9996F01E67EAB5" box="[1136,1342,1714,1737]" gridcol="3" gridrow="10" pageId="4" pageNumber="5">1.24 1.80</td>
|
||
</tr>
|
||
<tr id="93CB316BBF4C0040FF4C96A21E67EA8B" box="[165,1342,1760,1783]" gridrow="11" pageId="4" pageNumber="5">
|
||
<th id="D01A5817BF4C0040FF4C96A21A2FEA8B" box="[165,374,1760,1783]" gridcol="0" gridrow="11" pageId="4" pageNumber="5">
|
||
<taxonomicName id="EAFB48AABF4CFFBBFF4C96A21A79EA8B" box="[165,288,1760,1783]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cervinus">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF4C96A21A79EA8B" box="[165,288,1760,1783]" italics="true" pageId="4" pageNumber="5">D. cervinus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D01A5817BF4C0040FE1C96A2199AEA8B" box="[501,707,1760,1783]" gridcol="1" gridrow="11" pageId="4" pageNumber="5">0.40 0.59</td>
|
||
<td id="D01A5817BF4C0040FCDA96A21F58EA8B" box="[819,1025,1760,1783]" gridcol="2" gridrow="11" pageId="4" pageNumber="5">0.30 0.41</td>
|
||
<td id="D01A5817BF4C0040FB9996A21E67EA8B" box="[1136,1342,1760,1783]" gridcol="3" gridrow="11" pageId="4" pageNumber="5">1.14 1.53</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="65E160A2BF4CFFB9FF2F977C1E26EB02" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="2D443329BF4CFFBBFF2F977C1E2CEB8F" blockId="4.[151,1437,1854,2035]" pageId="4" pageNumber="5">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFF2F977C1884EB24" bold="true" box="[198,989,1854,1880]" pageId="4" pageNumber="5">Remarks about the structures and the functions of the genitalia.</emphasis>
|
||
The recent overview of the gnaphosid genera by
|
||
<bibRefCitation id="496A4ED8BF4CFFBBFEFF97271A89EB03" author="Murphy" box="[278,464,1893,1919]" pageId="4" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy (2007)</bibRefCitation>
|
||
shows the high diversity of genitalic structures occurring in the family. Among these genera, the genitalia of the genus
|
||
<taxonomicName id="EAFB48AABF4CFFBBFD8997C91984EBD8" box="[608,733,1931,1956]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4CFFBBFD8997C91984EBD8" box="[608,733,1931,1956]" italics="true" pageId="4" pageNumber="5">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
appear singular, atypical, the males having huge corkscrewshaped embolar structure which enter the equally corkscrew-shaped internal structure of the females; the illustrations of this paper clearly show, for each species, that the male coils closely match the female coils.
|
||
</paragraph>
|
||
<caption id="798463A1BF4DFFBAFF7E94371B96E96B" httpUri="https://zenodo.org/record/189207/files/figure.png" pageId="5" pageNumber="6" targetBox="[230,1354,205,1110]" targetPageId="5">
|
||
<paragraph id="2D443329BF4DFFBAFF7E94371B96E96B" blockId="5.[151,1436,1141,1303]" pageId="5" pageNumber="6">
|
||
<emphasis id="1F8FEF3BBF4DFFBAFF7E94371A43E8F1" bold="true" box="[151,282,1141,1165]" pageId="5" pageNumber="6">FIGURE 1.</emphasis>
|
||
<taxonomicName id="EAFB48AABF4DFFBAFEC894341ABBE8F1" box="[289,482,1142,1165]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4DFFBAFEC894341ABBE8F1" box="[289,482,1142,1165]" italics="true" pageId="5" pageNumber="6">Drassodex simoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4DFFBAFE0394371919E8F1" bold="true" box="[490,576,1141,1165]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="04BC5240BF4DFFBAFE0394371919E8F1" box="[490,576,1141,1165]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Spinnerets and spigots of adult female (left) and male (right), showing position of the anterior lateral (ALS), posterior median (PMS), and posterior lateral (PLS) spinnerets, along with the distribution of spigots presumed to serve the major ampullate (MA), piriform (P), minor ampullate (mA), cylindrical (C) and aciniform (A) glands. The position of the annular crescent (AC) is indicated by an arrow (after Murphy 2007: fig. p. 235).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D443329BF4DFFBAFF2F95081A44EB48" blockId="5.[151,1437,1354,2037]" pageId="5" pageNumber="6">
|
||
The mechanism of the introduction of the male coil inside the female had been studied by M. J. Roberts, who artificially expanded a male palp of
|
||
<taxonomicName id="EAFB48AABF4DFFBAFD979533185CE9F6" box="[638,773,1393,1418]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cervinus">
|
||
<emphasis id="1F8FEF3BBF4DFFBAFD979533185CE9F6" box="[638,773,1393,1418]" italics="true" pageId="5" pageNumber="6">D. cervinus</emphasis>
|
||
</taxonomicName>
|
||
in hot lactic acid. It appeared that three haematodochae are implicated in this mechanism: a basal haematodocha, in the cymbium, which pushes out and rotates the palpal organs; a middle one, which pushes the median apophysis out to the side and the more distal organs further; and a distal one, which fills the entire length of the coil carrying the embolus. The embolus itself lies in the “upper” edge of the coil and emerges as a free structure near the tip. The distal haematodocha is found as a distensible membrane on the inner surface of the coil. When expanded, this latter pushes out, swells and partially uncoils the embolus. The median apophysis is a hook structure lying just below the coil and must probably engage in the postero-lateral sockets of the female epigyne, allowing a firm grip. To summarize, the whole coiled male embolus is inserted through the coil of the epigyne, mechanical stability being provided by the median apophysis hooking into the postero-lateral epigyne socket. Finally, if this inference is correct, the free embolus tip should reach somewhere near the apex of the female coil, where the sperm is hence discharged.
|
||
</paragraph>
|
||
<paragraph id="2D443329BF4DFFB9FF2F97021E40EFEC" blockId="5.[151,1437,1354,2037]" lastBlockId="6.[151,1437,152,2034]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Concerning the female, two independent investigations were carried out by two of the authors to describe the structure of the vulva (i.e., the dorsal view of the epigyne). In entelegyne spiders, the vulva is typically constituted by a pair of seminal receptacles, commonly called spermathecae, associated with two pairs of ducts, one connecting the copulatory openings to the spermathecae, which permit to transfer the sperm by the introduction of the embolus, the insemination ducts, the other pair connecting the spermathecae to the oviduct, the fertilization ducts. Following, in part, the interpretation of Wiehle (1967: 13), one can think that the very small bursae attached near the distal part of the large coils of the epigyne are the spermathecae, and the highly visible, narrow, coiled ducts, running back from these spermathecae to the flattened lanceolate structures, are the fertilization ducts. The insemination ducts, connecting the apex of the large coils to these small spermathecae, had not been found with classical observation; we can thus hypothesize that they are probably minute or transparent, perhaps sometimes hidden by the dark secretions often observed in the inseminated females. Another interpretation can be proposed for the structures above-mentioned. Indeed, the small bursae can be seen as some glandular organs rather than some seminal receptacles. This hypothesis was proposed by
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FF7E918F1A1DED9B" author="Grimm" box="[151,324,461,487]" pageId="6" pageNumber="27" refString="Grimm, U. (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF), 26, 1 - 318." type="journal article" year="1985">Grimm (1985)</bibRefCitation>
|
||
. Regarding this opinion, and considering that the sperm is discharged near the apical part of the large coils of the female (indicated by the presence of the dark secretions, presumably equivalent to a “plug”,
|
||
<figureCitation id="B5C02FACBF4EFFB9FF7E92581BADEE48" box="[151,244,538,564]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="6" pageNumber="7">Figs 34</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4EFFB9FEE992581A78EE48" box="[256,289,538,564]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="6" pageNumber="7">46</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4EFFB9FEC792581A16EE48" box="[302,335,538,564]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="6" pageNumber="7">56</figureCitation>
|
||
), the spermathecae could then be located in this latter area, but not clearly identified at this time. An hypothesis was then made that the spermathecae could correspond to the slight enlargements, sometimes observed of the apical part of the narrow coiled ducts (see vulva of
|
||
<taxonomicName id="EAFB48AABF4EFFB9FBC6922A1F9AEEFD" box="[1071,1219,616,641]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FBC6922A1F9AEEFD" box="[1071,1219,616,641]" italics="true" pageId="6" pageNumber="7">D. hypocrita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="B5C02FACBF4EFFB9FB3B922A1E41EEFE" box="[1234,1304,616,642]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
|
||
), or maybe even correspond to the entire length of these ducts, implicating consequently that of the spermathecae is not globular but tubular receptacles! (Note that Grimm located the spermathecae only as the proximal part of this duct, near the epigastric furrow, see “R” in fig. 127). Complementary, the membranous large coils is hence considered as the copulatory openings (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FD9D93401848EF60" author="Grimm" box="[628,785,770,796]" pageId="6" pageNumber="27" refString="Grimm, U. (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF), 26, 1 - 318." type="journal article" year="1985">Grimm 1985</bibRefCitation>
|
||
) or the insemination ducts, the small bursae are some anterior glandular groups (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FE3B936B1936EF3F" author="Grimm" box="[466,623,809,835]" pageId="6" pageNumber="27" refString="Grimm, U. (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF), 26, 1 - 318." type="journal article" year="1985">Grimm 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FD95936B1843EF3F" author="Ledoux" box="[636,794,809,835]" pageId="6" pageNumber="27" refString="Ledoux, J. - C. (1997) Sur l'origine de la structure entelegyne: essai de morphologie comparee. Published by the author (with English extended summary), Solignac-sur-Loire, 42 pp." type="book" year="1997">Ledoux 1997</bibRefCitation>
|
||
) (a preliminary microscope examination seems reveal discrete gland ducts), the narrow coiled ducts is considered, in part or in totality, as the spermathecae, and the flattened lanceolate structures is considered as the fertilization ducts (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FC2D93341F39EFEC" author="Grimm" box="[964,1120,886,912]" pageId="6" pageNumber="27" refString="Grimm, U. (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF), 26, 1 - 318." type="journal article" year="1985">Grimm 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FB8593341E55EFEC" author="Ledoux" box="[1132,1292,886,912]" pageId="6" pageNumber="27" refString="Ledoux, J. - C. (1997) Sur l'origine de la structure entelegyne: essai de morphologie comparee. Published by the author (with English extended summary), Solignac-sur-Loire, 42 pp." type="book" year="1997">Ledoux 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2D443329BF4EFFB9FF2F93DF1A48E857" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="7">Without investigation in the fine observation of the ultrastructure of the female genitalia, it was decided to name the different parts of the vulva without connotation about their functions, by the following terms: the copulatory openings, the membranous coils, the coiled ducts, the small bursae and the flattened lanceolate structures.</paragraph>
|
||
<paragraph id="2D443329BF4EFFB9FF2F94751E26EB02" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="7">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FF2F94751942E82D" bold="true" box="[198,539,1079,1105]" pageId="6" pageNumber="7">
|
||
Relationships of
|
||
<taxonomicName id="EAFB48AABF4EFFB9FE7C9475194DE82D" box="[405,532,1079,1105]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FE7C9475194DE82D" bold="true" box="[405,532,1079,1105]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In a recent proposition for a new arrangement of the gnaphosid genera of the world,
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FF02941C1AF8E804" author="Murphy" box="[235,417,1118,1144]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy (2007)</bibRefCitation>
|
||
erected the genus
|
||
<taxonomicName id="EAFB48AABF4EFFB9FD96941D19A4E804" box="[639,765,1119,1144]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FD96941D19A4E804" box="[639,765,1119,1144]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
to receive the species of the
|
||
<taxonomicName id="EAFB48AABF4EFFB9FBB1941D1E14E804" box="[1112,1357,1119,1144]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FBB1941D1E14E804" box="[1112,1357,1119,1144]" italics="true" pageId="6" pageNumber="7">Drassodes hypocrita</emphasis>
|
||
</taxonomicName>
|
||
-group of
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FF5194C71A04E8E3" author="Simon" box="[184,349,1157,1183]" pageId="6" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon (1914)</bibRefCitation>
|
||
(
|
||
<taxonomicName id="EAFB48AABF4EFFB9FE8694C71AB4E8E2" box="[367,493,1157,1182]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vinosus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FE8694C71AB4E8E2" box="[367,493,1157,1182]" italics="true" pageId="6" pageNumber="7">D. vinosus</emphasis>
|
||
</taxonomicName>
|
||
excluded). Indeed, although the species belonging to
|
||
<taxonomicName id="EAFB48AABF4EFFB9FB9C94C71FADE8E2" box="[1141,1268,1157,1182]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FB9C94C71FADE8E2" box="[1141,1268,1157,1182]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
possess some features also observable in
|
||
<taxonomicName id="EAFB48AABF4EFFB9FE3294EE190EE8B9" box="[475,599,1196,1221]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FE3294EE190EE8B9" box="[475,599,1196,1221]" italics="true" pageId="6" pageNumber="7">Drassodes</emphasis>
|
||
</taxonomicName>
|
||
(presence of notched trochanters, similar cheliceral dentition and lack of a dorsal scutum on the male abdomen), a special feature of the ALS was employed to unite
|
||
<taxonomicName id="EAFB48AABF4EFFB9FB0D94911E38E890" box="[1252,1377,1235,1260]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FB0D94911E38E890" box="[1252,1377,1235,1260]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
with the genera
|
||
<taxonomicName id="EAFB48AABF4EFFB9FEF394BB191CE96F" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[282,581,1273,1299]" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FEF394BB1AFDE96E" box="[282,420,1273,1298]" italics="true" pageId="6" pageNumber="7">Anagraphis</emphasis>
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FE4594BB191CE96F" author="Simon" box="[428,581,1273,1299]" pageId="6" pageNumber="27" refString="Simon, E. (1893) Histoire naturelle des araignees. Roret, Paris, 1, 257 - 488." type="journal article" year="1893">Simon, 1893</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4EFFB9FD6994BB18D6E96F" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[640,911,1273,1299]" class="Arachnida" family="Gnaphosidae" genus="Talanites" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FD6994BB19B5E96E" box="[640,748,1273,1298]" italics="true" pageId="6" pageNumber="7">Talanites</emphasis>
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FD1D94BB18D6E96F" author="Simon" box="[756,911,1273,1299]" pageId="6" pageNumber="27" refString="Simon, E. (1893) Histoire naturelle des araignees. Roret, Paris, 1, 257 - 488." type="journal article" year="1893">Simon, 1893</bibRefCitation>
|
||
</taxonomicName>
|
||
, all three forming the informal
|
||
<taxonomicName id="EAFB48AABF4EFFB9FAE194BB1ECBE96E" box="[1288,1426,1273,1298]" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FAE194BB1ECBE96E" box="[1288,1426,1273,1298]" italics="true" pageId="6" pageNumber="7">Anagraphis</emphasis>
|
||
</taxonomicName>
|
||
- group (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FF0495621AC0E946" author="Murphy" box="[237,409,1312,1338]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
|
||
). The feature of the ALS concerned is the presence of a sclerotized annular crescent, bearing thin setae, on both the prolateral and retrolateral sides of the spinneret tip, situated between the sclerotized edge of the basal segment and the area of the piriform gland spigots (
|
||
<figureCitation id="B5C02FACBF4EFFB9FBA5952F1FCBE9FB" box="[1100,1170,1389,1415]" captionStart="FIGURE 1" captionStartId="5.[151,255,1141,1165]" captionTargetBox="[230,1354,205,1110]" captionTargetId="figure@5.[219,1367,194,1117]" captionTargetPageId="5" captionText="FIGURE 1. Drassodex simoni sp. nov. Spinnerets and spigots of adult female (left) and male (right), showing position of the anterior lateral (ALS), posterior median (PMS), and posterior lateral (PLS) spinnerets, along with the distribution of spigots presumed to serve the major ampullate (MA), piriform (P), minor ampullate (mA), cylindrical (C) and aciniform (A) glands. The position of the annular crescent (AC) is indicated by an arrow (after Murphy 2007: fig. p. 235)." httpUri="https://zenodo.org/record/189207/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FB4B952F1E1EE9FB" author="Murphy" box="[1186,1351,1389,1415]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
|
||
: figs p. 233, 235). This character is unique among the family
|
||
<taxonomicName id="EAFB48AABF4EFFB9FCAD95D61FE8E9D2" authority="Murphy 2007" authorityName="Murphy" authorityYear="2007" box="[836,1201,1428,1454]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">
|
||
Gnaphosidae (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FC1195D61FF1E9D2" author="Murphy" box="[1016,1192,1428,1454]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and it presumably represents a remnant of the sclerotized distal segment of the ALS, which is present in the more plesiomorphic gnaphosoid families and their outgroups (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FD7095A31810E987" author="Platnick" box="[665,841,1505,1531]" pageId="6" pageNumber="27" refString="Platnick, N. I. (1990) Spinneret morphology and the phylogeny of grounds spiders (Araneae, Gnaphosoidea). American Museum Novitates, 2978, 1 - 42." type="journal article" year="1990">Platnick 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FCBE95A31F2EE987" author="Platnick" box="[855,1143,1505,1531]" pageId="6" pageNumber="27" refString="Platnick, N. I. & Baehr, B. (2006) A revision of the australasian ground spiders of the family Prodidomidae (Araneae: Gnaphosoidea). Bulletin of the American Museum of Natural History, 298, 1 - 287." type="journal article" year="2006">Platnick & Baehr 2006</bibRefCitation>
|
||
). Two other remarkable features, also presumably plesiomorphic in gnaphosids (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FCD6964A18B2EA5E" author="Platnick" box="[831,1003,1544,1570]" pageId="6" pageNumber="27" refString="Platnick, N. I. (1990) Spinneret morphology and the phylogeny of grounds spiders (Araneae, Gnaphosoidea). American Museum Novitates, 2978, 1 - 42." type="journal article" year="1990">Platnick 1990</bibRefCitation>
|
||
), are observable in the
|
||
<taxonomicName id="EAFB48AABF4EFFB9FAEF964A1ECBEA5D" box="[1286,1426,1544,1569]" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FAEF964A1ECBEA5D" box="[1286,1426,1544,1569]" italics="true" pageId="6" pageNumber="7">Anagraphis</emphasis>
|
||
</taxonomicName>
|
||
- group: the presence of some piriform gland spigots in the middle, rather than exclusively around the edge, of the ALS tip (only observed in
|
||
<taxonomicName id="EAFB48AABF4EFFB9FDEB961719D9EA12" box="[514,640,1621,1646]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FDEB961719D9EA12" box="[514,640,1621,1646]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4EFFB9FD529617187FEA12" box="[699,806,1621,1646]" class="Arachnida" family="Gnaphosidae" genus="Talanites" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FD529617187FEA12" box="[699,806,1621,1646]" italics="true" pageId="6" pageNumber="7">Talanites</emphasis>
|
||
</taxonomicName>
|
||
) and the general shape of these spigots, which differ from the typical form seen in most gnaphosid genera (
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FCCA963E1897EAEA" author="Murphy" box="[803,974,1660,1686]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
|
||
). Consequently, the
|
||
<taxonomicName id="EAFB48AABF4EFFB9FB2B963E1E14EAE9" box="[1218,1357,1660,1685]" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FB2B963E1E14EAE9" box="[1218,1357,1660,1685]" italics="true" pageId="6" pageNumber="7">Anagraphis</emphasis>
|
||
</taxonomicName>
|
||
-group seems to be united only by plesiomorphic features, and hence is potentially a non-natural (i.e., nonmonophyletic) group. However, as pointed out by
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FD1A968B18F7EA9F" author="Murphy" box="[755,942,1737,1763]" pageId="6" pageNumber="27" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs. 2 volumes. Volume 1 (text), i - xii + 1 - 92; Volume 2 (plates), i - ii + 93 - 605." type="journal article" year="2007">Murphy (2007)</bibRefCitation>
|
||
, these genera share some similar somatic and genitalic features, as the configuration of the male palp (coiled embolus in
|
||
<taxonomicName id="EAFB48AABF4EFFB9FBBC96B21F8CEB75" box="[1109,1237,1776,1801]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FBBC96B21F8CEB75" box="[1109,1237,1776,1801]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4EFFB9FAF896B21AD2EB4C" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FAF896B21BB5EB4C" italics="true" pageId="6" pageNumber="7">Anagraphis pallens</emphasis>
|
||
<bibRefCitation id="496A4ED8BF4EFFB9FF1B97541AD2EB4C" author="Simon" box="[242,395,1814,1840]" pageId="6" pageNumber="27" refString="Simon, E. (1878) Les arachnides de France. Roret, Paris, 4, 1 - 334." type="journal article" year="1878">Simon, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
), the shape of the RTA (bifid), the eye pattern (in particular the subcircular PME) and the cheliceral dentition pattern (three discrete teeth on the promargin, two on the retromargin.). Until new considerations, the genus
|
||
<taxonomicName id="EAFB48AABF4EFFB9FE2C9726191BEB01" box="[453,578,1892,1917]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FE2C9726191BEB01" box="[453,578,1892,1917]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
is hence considered as closest to the genera
|
||
<taxonomicName id="EAFB48AABF4EFFB9FBA597261F8CEB01" box="[1100,1237,1892,1917]" class="Arachnida" family="Gnaphosidae" genus="Anagraphis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FBA597261F8CEB01" box="[1100,1237,1892,1917]" italics="true" pageId="6" pageNumber="7">Anagraphis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4EFFB9FAE797261E2FEB01" box="[1294,1398,1892,1917]" class="Arachnida" family="Gnaphosidae" genus="Talanites" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FAE797261E2FEB01" box="[1294,1398,1892,1917]" italics="true" pageId="6" pageNumber="7">Talanites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E160A2BF4EFFB8FF2F97C81FFBEED4" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="2D443329BF4EFFB8FF2F97C81FFBEED4" blockId="6.[151,1437,152,2034]" lastBlockId="7.[151,1437,152,1067]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FF2F97C81936EBD8" bold="true" box="[198,623,1930,1956]" pageId="6" pageNumber="7">Relationships between the species.</emphasis>
|
||
Examination of the genitalic characters of the members of
|
||
<taxonomicName id="EAFB48AABF4EFFB9FAF797C91EC2EBD8" box="[1310,1435,1931,1956]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FAF797C91EC2EBD8" box="[1310,1435,1931,1956]" italics="true" pageId="6" pageNumber="7">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
allowed us to associate some of them into two hypothetically monophyletic groups: one of these groups includes
|
||
<taxonomicName id="EAFB48AABF4EFFB9FEE9979A1ACDEB8D" box="[256,404,2008,2033]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FEE9979A1ACDEB8D" box="[256,404,2008,2033]" italics="true" pageId="6" pageNumber="7">D. hypocrita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4EFFB9FE4D979A1945EB8D" box="[420,540,2008,2033]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lesserti">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FE4D979A1945EB8D" box="[420,540,2008,2033]" italics="true" pageId="6" pageNumber="7">D. lesserti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4EFFB9FDC5979A19F4EB8D" box="[556,685,2008,2033]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="validior">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FDC5979A19F4EB8D" box="[556,685,2008,2033]" italics="true" pageId="6" pageNumber="7">D. validior</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4EFFB9FD0F979A1800EB8D" box="[742,857,2008,2033]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granja" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4EFFB9FD0F979A1800EB8D" box="[742,857,2008,2033]" italics="true" pageId="6" pageNumber="7">D. granja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4EFFB9FC88979518E7EB8D" bold="true" box="[865,958,2007,2033]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="04BC5240BF4EFFB9FC88979518E7EB8D" box="[865,958,2007,2033]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
These species exhibit the same shape of male embolar coil, with the distal loop very much narrowed and shifted more retrolaterally than the basal loop, spiralling upwards along a different axis (
|
||
<figureCitation id="B5C02FACBF4FFFB8FD3790FC1814ECA4" box="[734,845,190,216]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="7" pageNumber="8">Figs 4, 8</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FCB590FC18F2ECA4" box="[860,939,190,216]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="7" pageNumber="8">43, 47</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FC5090FC1883ECA4" box="[953,986,190,216]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="7" pageNumber="8">51</figureCitation>
|
||
), the presence of a terminal embolar apophysis (in the first three species only, absent in
|
||
<taxonomicName id="EAFB48AABF4FFFB8FD0490A71839EC82" box="[749,864,229,254]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="granja">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FD0490A71839EC82" box="[749,864,229,254]" italics="true" pageId="7" pageNumber="8">D. granja</emphasis>
|
||
</taxonomicName>
|
||
), and the position of the coiled duct, joined with the outer walls of the membranous coil (
|
||
<figureCitation id="B5C02FACBF4FFFB8FD91914E19BDED5A" box="[632,740,268,294]" captionStart="FIGURES 2 – 9" captionStartId="10.[151,274,1864,1888]" captionTargetBox="[186,1400,188,1838]" captionTargetId="figure@10.[173,1417,175,1846]" captionTargetPageId="10" captionText="FIGURES 2 – 9. Drassodex hypocrita (Simon, 1878) (2 – 5 Ledoux collection, after Murphy 2007: figs p. 233; 6 – 9 MNHN AR 9437). 2 – 3, 6 – 7 epigyne (2, 6 ventral view; 3, 7 dorsal view); 4 – 5, 8 – 9 left male palp (4, 8 ventral view; 5, 9 retrolateral view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; E = embolus; EA = embolar apophysis; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189208/files/figure.png" pageId="7" pageNumber="8">Figs 3, 7</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FD18914E1867ED5A" box="[753,830,268,294]" captionStart="FIGURES 41 – 48. 41 – 44" captionStartId="21.[151,269,1898,1922]" captionTargetBox="[205,1373,208,1862]" captionTargetId="figure@21.[194,1387,197,1863]" captionTargetPageId="21" captionText="FIGURES 41 – 48. 41 – 44 Drassodex granja sp. nov. (MNHN AR 9256). 41 – 42 epigyne (paratype) (41 ventral view, 42 dorsal view); 43 – 44 left male palp (holotype) (43 ventral view, 44 retrolateral view). 45 – 48 Drassodex validior (Simon, 1914) (45 – 46 MNHN AR 9248; 47 – 48 MNHN AR 9241, lectotype). 45 – 46 epigyne (45 ventral view, 46 dorsal view); 47 – 48 left male palp (47 ventral view, 48 retrolateral view)." httpUri="https://zenodo.org/record/189213/files/figure.png" pageId="7" pageNumber="8">42, 46</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FCA2914E1891ED5A" box="[843,968,268,294]" captionStart="FIGURES 49 – 57" captionStartId="24.[151,271,1930,1954]" captionTargetBox="[173,1413,206,1900]" captionTargetId="figure@24.[164,1422,194,1906]" captionTargetPageId="24" captionText="FIGURES 49 – 57. Drassodex lesserti (Schenkel, 1936) (MNHN AR 9246). 49 – 50, 53 – 56 epigyne (49, 53, 55 ventral view; 50, 54, 56 dorsal view); 51 – 52, 57 left male palp (51 ventral view, 52 retrolateral view, 57 details of the embolar coil tip, showing variation of terminal apophysis)." httpUri="https://zenodo.org/record/189214/files/figure.png" pageId="7" pageNumber="8">50, 54, 56</figureCitation>
|
||
). The second group includes at least
|
||
<taxonomicName id="EAFB48AABF4FFFB8FA96914E1B8DED37" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="heeri">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FA96914E1B8DED37" italics="true" pageId="7" pageNumber="8">D. heeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FF0D91701A39ED37" box="[228,352,306,331]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FF0D91701A39ED37" box="[228,352,306,331]" italics="true" pageId="7" pageNumber="8">D. drescoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4FFFB8FE8191701A91ED30" bold="true" box="[360,456,306,332]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="04BC5240BF4FFFB8FE8191701A91ED30" box="[360,456,306,332]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4FFFB8FDED91701920ED37" box="[516,633,306,331]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FDED91701920ED37" box="[516,633,306,331]" italics="true" pageId="7" pageNumber="8">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4FFFB8FD68917019B8ED30" bold="true" box="[641,737,306,332]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="04BC5240BF4FFFB8FD68917019B8ED30" box="[641,737,306,332]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, all these species having a regularly looped male embolar coil, the distal loop large and positioned in the same axis as the basal loop (
|
||
<figureCitation id="B5C02FACBF4FFFB8FBE7911B1F33ED0F" box="[1038,1130,345,371]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="7" pageNumber="8">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FB9E911B1F9AED0F" box="[1143,1219,345,371]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="7" pageNumber="8">22, 26</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FB39911B1FA8ED0F" box="[1232,1265,345,371]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="7" pageNumber="8">30</figureCitation>
|
||
), the tip of the embolar coil very simple, only attenuated or truncated, without terminal apophysis, and the position of the coiled duct, running freely inside the longitudinal axis of the membranous coil (
|
||
<figureCitation id="B5C02FACBF4FFFB8FBD191E41E4AEDBC" box="[1080,1299,422,448]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="7" pageNumber="8">Figs 11, 15, 17, 19</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FAC991E41E32EDBC" box="[1312,1387,422,448]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="7" pageNumber="8">21, 25</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FA9E91E41ECEEDBC" box="[1399,1431,422,448]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="7" pageNumber="8">29</figureCitation>
|
||
,
|
||
<figureCitation id="B5C02FACBF4FFFB8FF7E918F1BE3ED9B" box="[151,186,461,487]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="7" pageNumber="8">33</figureCitation>
|
||
). These three species occur only in and around the Alps. We can probably bring closer
|
||
<taxonomicName id="EAFB48AABF4FFFB8FAE5918F1EC4ED9A" box="[1292,1437,461,486]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cervinus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FAE5918F1EC4ED9A" box="[1292,1437,461,486]" italics="true" pageId="7" pageNumber="8">D. cervinus</emphasis>
|
||
</taxonomicName>
|
||
(Pyrenees and
|
||
<collectingCountry id="55EC73B9BF4FFFB8FEAE91B61AD4EE72" box="[327,397,500,526]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
|
||
) to these species, based on the great similarity in their genitalia (regular male embolar coil,
|
||
<figureCitation id="B5C02FACBF4FFFB8FF3892581A7FEE48" box="[209,294,538,564]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="7" pageNumber="8">Fig. 30</figureCitation>
|
||
, and female spiral-column coiled duct,
|
||
<figureCitation id="B5C02FACBF4FFFB8FD1B92581811EE48" box="[754,840,538,564]" captionStart="FIGURES 28 – 31" captionStartId="17.[151,273,1040,1064]" captionTargetBox="[171,1414,203,1005]" captionTargetId="figure@17.[164,1423,194,1016]" captionTargetPageId="17" captionText="FIGURES 28 – 31. Drassodex cervinus (Simon, 1914) (28 – 29 MNHN AR 9217, paralectotype; 30 – 31 MNHN AR 14600, lectotype). 28 – 29 epigyne (28 ventral view, 29 dorsal view); 30 – 31 left male palp (30 ventral view, 31 retrolateral view)." httpUri="https://zenodo.org/record/189211/files/figure.png" pageId="7" pageNumber="8">Fig. 29</figureCitation>
|
||
); this latter species is hence presumably a vicariant of the former species. It is not easy to define the affinities of the two similar species
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB4692031E18EE26" box="[1199,1345,577,602]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hispanus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB4692031E18EE26" box="[1199,1345,577,602]" italics="true" pageId="7" pageNumber="8">D. hispanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4FFFB8FA9692031BA5EEFD" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FA9692031BA5EEFD" italics="true" pageId="7" pageNumber="8">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
with the others, but the coiled duct, which is apparently joined to the outer walls of the membranous coil (
|
||
<figureCitation id="B5C02FACBF4FFFB8FF3B92CC1A02EED4" box="[210,347,654,680]" captionStart="FIGURES 32 – 40. 32 – 33" captionStartId="18.[151,273,1722,1746]" captionTargetBox="[196,1386,203,1689]" captionTargetId="figure@18.[185,1402,194,1698]" captionTargetPageId="18" captionText="FIGURES 32 – 40. 32 – 33 Drassodex cervinus (Simon, 1914) (MNHN AR 9241, paralectotype D. validior). 32 – 33 epigyne (32 ventral view, 33 dorsal view). 34 – 38 Drassodex fritillifer (Simon, 1914) (MNHN AR 1646, syntypes). 34 – 35 epigyne (34 ventral view, 35 dorsal view); 36 – 37 left male palp (36 ventral view, 37 retrolateral view); 38 left male chelicerae. 39 – 40 Drassodex hispanus (L. Koch, 1866) (MNHN AR 9247). 39 – 40 epigyne (39 ventral view, 40 dorsal view)." httpUri="https://zenodo.org/record/189212/files/figure.png" pageId="7" pageNumber="8">Figs 35, 40</figureCitation>
|
||
), seems to be much closer to the first group than that of the second one.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E160A2BF4FFFB8FF2F92F71A24E857" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="2D443329BF4FFFB8FF2F92F71A24E857" blockId="7.[151,1437,152,1067]" pageId="7" pageNumber="8">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FF2F92F71A3EEEB3" bold="true" box="[198,359,693,719]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
Considering the great number of misidentifications encountered in the MNHN collection, we can suppose that similar mistakes occurred in other European collections, and subsequently in catalogues or faunistic studies. It could be particularly the case for the species described under
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB9193401F80EF67" box="[1144,1241,770,795]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB9193401F80EF67" box="[1144,1241,770,795]" italics="true" pageId="7" pageNumber="8">Drassus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB0393401E2EEF67" box="[1258,1399,770,795]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hispanus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB0393401E2EEF67" box="[1258,1399,770,795]" italics="true" pageId="7" pageNumber="8">D. hispanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FA6E93401B8FEF3E" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="heeri">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FA6E93401B8FEF3E" italics="true" pageId="7" pageNumber="8">D heeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FF0E936B1ADBEF3E" box="[231,386,809,834]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FF0E936B1ADBEF3E" box="[231,386,809,834]" italics="true" pageId="7" pageNumber="8">D. hypocrita</emphasis>
|
||
</taxonomicName>
|
||
). Nevertheless, representatives of
|
||
<taxonomicName id="EAFB48AABF4FFFB8FCD8936B18EDEF3E" box="[817,948,809,834]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FCD8936B18EDEF3E" box="[817,948,809,834]" italics="true" pageId="7" pageNumber="8">Drassodex</emphasis>
|
||
</taxonomicName>
|
||
mainly occur in south-western Europe, especially in and around mountain systems, at medium to high elevations:
|
||
<taxonomicName id="EAFB48AABF4FFFB8FBFE93121FF8EF15" box="[1047,1185,848,873]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cervinus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FBFE93121FF8EF15" box="[1047,1185,848,873]" italics="true" pageId="7" pageNumber="8">D. cervinus</emphasis>
|
||
</taxonomicName>
|
||
is recorded from the Pyrenees and north-eastern
|
||
<collectingCountry id="55EC73B9BF4FFFB8FE369334197CEFEC" box="[479,549,886,912]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FDDB933519E4EFEC" box="[562,701,887,912]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fritillifer">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FDDB933519E4EFEC" box="[562,701,887,912]" italics="true" pageId="7" pageNumber="8">D. fritillifer</emphasis>
|
||
</taxonomicName>
|
||
from the Pyrenees and eastern
|
||
<collectingCountry id="55EC73B9BF4FFFB8FBDA93341F20EFEC" box="[1075,1145,886,912]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB6C93351FBFEFEC" box="[1157,1254,887,912]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="heeri">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB6C93351FBFEFEC" box="[1157,1254,887,912]" italics="true" pageId="7" pageNumber="8">D. heeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB1F93351E28EFEC" box="[1270,1393,887,912]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB1F93351E28EFEC" box="[1270,1393,887,912]" italics="true" pageId="7" pageNumber="8">D. drescoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4FFFB8FA9093341B92EFCB" bold="true" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="04BC5240BF4FFFB8FA9093341B92EFCB" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4FFFB8FEEE93DF1A24EFCA" box="[263,381,925,950]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FEEE93DF1A24EFCA" box="[263,381,925,950]" italics="true" pageId="7" pageNumber="8">D. simoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4FFFB8FE6D93DF1ABCEFCB" bold="true" box="[388,485,925,951]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="04BC5240BF4FFFB8FE6D93DF1ABCEFCB" box="[388,485,925,951]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the Alps,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FD4C93DF186DEFCA" box="[677,820,925,950]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hispanus">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FD4C93DF186DEFCA" box="[677,820,925,950]" italics="true" pageId="7" pageNumber="8">D. hispanus</emphasis>
|
||
</taxonomicName>
|
||
from central and eastern
|
||
<collectingCountry id="55EC73B9BF4FFFB8FB8393DF1FE8EFCB" box="[1130,1201,925,951]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FB5793DF1E6DEFCA" box="[1214,1332,925,950]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="granja" status="sp. nov.">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FB5793DF1E6DEFCA" box="[1214,1332,925,950]" italics="true" pageId="7" pageNumber="8">D. granja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8FEF3BBF4FFFB8FAD593DF1EC5EFCB" bold="true" box="[1340,1436,925,951]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="04BC5240BF4FFFB8FAD593DF1EC5EFCB" box="[1340,1436,925,951]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from central
|
||
<collectingCountry id="55EC73B9BF4FFFB8FED293861ADFEFA2" box="[315,390,964,990]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
|
||
,
|
||
<taxonomicName id="EAFB48AABF4FFFB8FE7A93861947EFA1" box="[403,542,964,989]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="validior">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FE7A93861947EFA1" box="[403,542,964,989]" italics="true" pageId="7" pageNumber="8">D. validior</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAFB48AABF4FFFB8FDB4938619B9EFA1" box="[605,736,964,989]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lesserti">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FDB4938619B9EFA1" box="[605,736,964,989]" italics="true" pageId="7" pageNumber="8">D. lesserti</emphasis>
|
||
</taxonomicName>
|
||
from western Alps and South
|
||
<collectingCountry id="55EC73B9BF4FFFB8FB8A93861FE5EFA2" box="[1123,1212,964,990]" name="France" pageId="7" pageNumber="8">France</collectingCountry>
|
||
. The most widely distributed species is apparently
|
||
<taxonomicName id="EAFB48AABF4FFFB8FDC693A81995E87F" box="[559,716,1002,1027]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="1F8FEF3BBF4FFFB8FDC693A81995E87F" box="[559,716,1002,1027]" italics="true" pageId="7" pageNumber="8">D. hypocrita</emphasis>
|
||
</taxonomicName>
|
||
, recorded from
|
||
<collectingCountry id="55EC73B9BF4FFFB8FC7C93A818B0E878" box="[917,1001,1002,1028]" name="France" pageId="7" pageNumber="8">France</collectingCountry>
|
||
to West part of
|
||
<collectingCountry id="55EC73B9BF4FFFB8FB5D93A81E53E878" box="[1204,1290,1002,1028]" name="Russia" pageId="7" pageNumber="8">Russia</collectingCountry>
|
||
, at lower to medium elevations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |