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<document ID-CLB-Dataset="207920" ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.e84187" ID-GBIF-Dataset="bac14654-3ac7-46b5-a981-8873c1b9e291" ID-Pensoft-Pub="2747-8106-96-179" ID-Pensoft-UUID="FE0861D71BB454999EE6637C0D9B2B0C" ID-ZooBank="35B619086E6548B09A177281C2253391" ModsDocID="2747-8106-96-179" checkinTime="1670952513976" checkinUser="pensoft" docAuthor="Gruendel, Joachim, Keupp, Helmut, Lang, Fritz &amp; Nuetzel, Alexander" docDate="2022" docId="A4826EFB1DA4529285523B16C04C865D" docLanguage="en" docName="Zitteliana 96: 179-221" docOrigin="Zitteliana 96" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.e84187" docTitle="Ptygmatis nodosa" docType="treatment" docVersion="5" id="FE0861D71BB454999EE6637C0D9B2B0C" lastPageNumber="179" masterDocId="FE0861D71BB454999EE6637C0D9B2B0C" masterDocTitle="Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)" masterLastPageNumber="221" masterPageNumber="179" pageNumber="179" updateTime="1732645988798" updateUser="ExternalLinkService">
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<mods:title id="7C4F526D2FB21A4204850B690AFDF230">Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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<mods:namePart id="CAF2349D69759081DE1A2B127462579C">Gruendel, Joachim</mods:namePart>
<mods:affiliation id="041F3BF65E544B3DBE1042884CEC23D3">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="171BD953AFBDC17C0FFAE72158ED9B76">Keupp, Helmut</mods:namePart>
<mods:affiliation id="88FAD0137543AAE19BD4175DEB265E0A">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="A8EA8803084189DF5F33F1AB9251A2E1">Lang, Fritz</mods:namePart>
<mods:affiliation id="009DCD3FEB3D7AFA59597E49B2C963EB">Drosselweg 16, 96114 Hirschaid, Germany</mods:affiliation>
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<mods:namePart id="B65453C8C5AF4E04EA492F1E543DB263">Nuetzel, Alexander</mods:namePart>
<mods:nameIdentifier id="9289B60001BB5A676312CC238F8761ED" type="ORCID">https://orcid.org/0000-0002-8852-7688</mods:nameIdentifier>
<mods:affiliation id="9DD8200D427F53B01715277E7D8D38D1">SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<mods:title id="A64F0D9565CCE76AA851BB6D4F572790">Zitteliana</mods:title>
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<mods:date id="8C977B40F34C44D1C5EA43CF7E437525">2022</mods:date>
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<mods:number id="0C44A0199C5CA1A507F29E4CD12DBD16">96</mods:number>
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<mods:start id="08268C0B6BDDE87E89C566B0F235AD6A">179</mods:start>
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<treatment id="A4826EFB1DA4529285523B16C04C865D" ID-GBIF-Taxon="206232346" LSID="urn:lsid:plazi:treatment:A4826EFB1DA4529285523B16C04C865D" httpUri="http://treatment.plazi.org/id/A4826EFB1DA4529285523B16C04C865D" lastPageNumber="179" pageId="0" pageNumber="179" scope_class="Gastropoda" scope_family="Ptygmatididae" scope_subClass="Heterobranchia" scope_subFamily="Ptygmatidinae" scope_superFamily="Nerineoidea">
<subSubSection id="2B58B754DA06F0A492BFE54579F50D09" pageId="0" pageNumber="179" type="nomenclature">
<paragraph id="9B3179F18A4725FE29CDDE42E68AB474" pageId="0" pageNumber="179">
<taxonomicName id="5B29AB0C56DCCB9F62BDFA97F4971C30" ID-CoL="CF2HR" LSID="A4826EFB-1DA4-5292-8552-3B16C04C865D" authority="(Voltz, 1836)" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">Ptygmatis nodosa (Voltz, 1836)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7AAC085AB2E5CCF8ECD831046B1F4A34" pageId="0" pageNumber="179" type="description">
<paragraph id="9DE8F237C40EF929DA0EFFCC05A5AA6C" pageId="0" pageNumber="179">
<figureCitation id="A5C3C0EC208125FB5E6E1738C17A7994" captionStart="Plate 11" captionStartId="F11" captionText="Plate 11. Ptygmatis nodosa (Voltz, 1836). (1 - 3) SNSB-BSPG 2021 XV 40, Saal (collection Lang); (1) lateral view, height 22 mm; (2) last whorl with aperture, width 6.6 mm; (3) base, width 6.6 mm. (4 - 5) SNSB-BSPG 2021 XV 41; Saal (collection Lang); (4) lateral view, height 9 mm; (5) base, width 3.5 mm. (6) SNSB-BSPG 2021 XV 42, Saal (collection Lang), aperture, width of whorl 3.7 mm. (7 - 8) SNSB-BSPG 2021 XV 43, Saal (collection Lang); (7) lateral view, height 11.5 mm; (8) last two whorls in lateral view, width 4.2 mm. (9) SNSB-BSPG 2021 XV 44, Saal (collection Lang), base, width 6 mm. (10 - 11) SNSB-BSPG 2021 XV 45, Saal (collection Lang); (10) lateral view, height 16 mm; (11) last whorls in lateral view, width 5.2 mm. (12 - 13) SNSB-BSPG 2021 XV 46, Saal (collection Keupp); (12) lateral view, height 9 mm; (13) apex in lateral view, height 2.9 mm. (14) SNSB-BSPG 2021 XV 47, Saal (collection Lang), lateral view, height 10 mm. (15) SNSB-BSPG 2021 XV 48, Saal (collection Lang), last whorls with aperture, width 7.8 mm." figureDoi="10.3897/zitteliana.96.e84187.plate11" httpUri="https://binary.pensoft.net/fig/780389" pageId="0" pageNumber="179">Plate 11: figs 1-15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2BCC14EC539BAFF5CC2949A3BD98CEE" pageId="0" pageNumber="179" type="reference_group">
<paragraph id="FF2F7B6A3C33068BF0852BF7D7A8D96B" pageId="0" pageNumber="179">
<taxonomicName id="74A6BE3B3FF0DA0D1EDEE2CAEA94F9E4" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">*1836 - Nerinea nodosa</taxonomicName>
(Voltz) - Voltz: 542.
</paragraph>
<paragraph id="AA6EC2735CA7A2C9B075421491356C76" pageId="0" pageNumber="179">
<taxonomicName id="7AEB0F2F9EB1D1C88965A940B0152409" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">1836 - Nerinea nodosa</taxonomicName>
Voltz - Bronn: 561, pl. 16, fig. 9.
</paragraph>
<paragraph id="776C4DA079BB952A749DC4C0DF01D08D" pageId="0" pageNumber="179">
<taxonomicName id="B5655352B7D9412DFE4217A288E4B95D" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">1851 - Nerinea nodosa</taxonomicName>
Voltz -
<normalizedToken id="E2795D4172F5514DD557D5E4E758BADF" originalValue="d’Orbigny">d'Orbigny</normalizedToken>
: 95, pl. 254, figs 3-5.
</paragraph>
<paragraph id="2502249C89FEF829F12D6F74B366BC63" pageId="0" pageNumber="179">
<taxonomicName id="836941C386C318584FB24F5D5942343D" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea calypso" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="calypso">*1852 - Nerinea calypso</taxonomicName>
<normalizedToken id="6A539018A41F1DECA4D95F67BED3C880" originalValue="d’Orbigny">d'Orbigny</normalizedToken>
-
<normalizedToken id="4F2C73008194034E1C6EB20EAA97AC60" originalValue="d’Orbigny">d'Orbigny</normalizedToken>
: 136, pl. 274, figs 4-6.
</paragraph>
<paragraph id="0D836D4F944CFF15D6BE37ED4CBBB437" pageId="0" pageNumber="179">
<taxonomicName id="569E9E1230EC55AB7B0C5BDF82EB78B7" authorityName="Voltz" authorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea elegans" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="elegans">*1852 - Nerinea elegans</taxonomicName>
Thurm. -
<normalizedToken id="BFA50ABC05B2348E581AFBD984890CCB" originalValue="d’Orbigny">d'Orbigny</normalizedToken>
: 146, pl. 278, figs 4-6.
</paragraph>
<paragraph id="8A39BB5A18B871C22586130728D04C42" pageId="0" pageNumber="179">
<taxonomicName id="E3AF3A20B6A69CF16C84F953469AAEB1" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea plassenensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="plassenensis">?1870 - Nerinea plassenensis</taxonomicName>
Pet. - Gemmellaro: 25, pl. 4, figs 14, 15.
</paragraph>
<paragraph id="1664BDF21D0D748D53E4E7A71067D126" pageId="0" pageNumber="179">
<taxonomicName id="827C9FD3F8E9AEA5CBD7B8003DAF4FFA" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">1889 - Nerinea nodosa</taxonomicName>
Voltz - Loriol in Loriol and Koby: 32, pl. 4, figs 5-11.
</paragraph>
<paragraph id="262365CFED066E19E0CC6B0ADBE2B239" pageId="0" pageNumber="179">
<taxonomicName id="1C6A399038B9AFE538AC88D9A57B9956" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">1997 - Ptygmatis nodosa</taxonomicName>
(Bronn ex Voltz, 1836) - Fischer and Weber: 37, pl. 11, figs 4-6.
</paragraph>
<paragraph id="08DA5F31D3CE083D41C64817DBD4B9E8" pageId="0" pageNumber="179">
<taxonomicName id="92B09AB5D1C3A7E85CAEEE2D670D0607" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">1997 - Ptygmatis nodosa</taxonomicName>
(Bronn ex Voltz, 1836) - Fischer and Weber: 54.
</paragraph>
<paragraph id="5A06A625515BDDF1BB8E418AA128B88C" pageId="0" pageNumber="179">
<taxonomicName id="D9C97B91973673E48097E1787F8FDD69" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Nerinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinella elegans" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="elegans">?1997 - Nerinella elegans</taxonomicName>
(Bronn ex Thurmann, 1836) - Fischer and Weber: 58, pl. 8, fig. 5.
</paragraph>
<paragraph id="769E0D9993AA644C9B95F5133A0BC669" pageId="0" pageNumber="179">
<taxonomicName id="22691691E6FAB6D5E0C0FDC0AE812298" authorityName="Sharpe" authorityYear="1850" class="Gastropoda" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" pageId="0" pageNumber="179" rank="superFamily" superFamily="Nerineoidea">2017 - Nerineoidea</taxonomicName>
Nr. 8 -
<normalizedToken id="7CA45D16A696BD96C6E3A563C497380C" originalValue="Gründel">Gruendel</normalizedToken>
: 33, pl.15A.
</paragraph>
</subSubSection>
<subSubSection id="2B5BD843DA74D7C644588A9E05186F3E" pageId="0" pageNumber="179" type="materials_examined">
<paragraph id="2857CBE6490C00D077DF83CAD9DDA9BA" pageId="0" pageNumber="179">Material.</paragraph>
<paragraph id="4B98D69E3CC3F68BD4DAA414186F6644" pageId="0" pageNumber="179">
142 fragments and juvenile specimens from Saal:
<specimenCount id="0B76276A693D9A70EE3D2B512A2B1210" type="generic">138 specimens</specimenCount>
collection Lang, of which eight are illustrated (SNSB-BSPG 2021 XV 40-45, 47, 48),
<specimenCount id="F521903811AC52B929653A3D066D2A0B" type="generic">four specimens</specimenCount>
collection Keupp (SNSB-BSPG 2021 XV 46, 49-51).
</paragraph>
</subSubSection>
<subSubSection id="ED9C4848C1445B51756E3BF68A284B71" pageId="0" pageNumber="179" type="description">
<paragraph id="1F013601A59D5E74FEBB3A689D707FFE" pageId="0" pageNumber="179">Description.</paragraph>
<paragraph id="42C3F3006D19FDC7769C0BE7867A72AC" pageId="0" pageNumber="179">The present material consists of fragments of larger specimens and juvenile specimens. The largest specimen is 32 mm high. The apical whorls are not preserved. The shell is slender with a somewhat variable apical angle. The ornament consists of a nodular adapical bulge forming a narrow ramp. The suture is situated on the adapical portion of the bulge. The nodular spiral cord (in some specimens only very weakly developed) is mostly close to the abapical suture or - more rarely - about half way between adapical bulge and abapical suture. The number of nodes per whorl is strongly variable (only in part due to preservation). In few specimens, an additional weakly nodular spiral cord is present between the nodular spiral cord and the adapical bulge. The base is weakly convex with a pronounced spiral cord at the almost rectangular transition to the whorl face. This bordering spiral cord is sometimes nodular. The base is covered with spiral cords. The aperture has a rhomboid outline and a distinctly oblique siphonal canal. The plait pattern is only visible in few specimens. It consists of one or two columellar plaits, one strong parietal plait, while a palatal is plait very rarely visible - perhaps due to the preservation.</paragraph>
</subSubSection>
<subSubSection id="3B8181D435E50B9A920BAA864BB83DEC" pageId="0" pageNumber="179" type="remarks">
<paragraph id="84D8B0E7CBA92BE1FDFA9E05DDE4131C" pageId="0" pageNumber="179">Remarks.</paragraph>
<paragraph id="CD504F66E9A5DF75D398C94ABCB10179" pageId="0" pageNumber="179">
In the present material, some specimens have one and others have two columellar plaits. The references listed in the chresonymy and synonymy list above probably refer to material from older strata (Oxfordian, with the exception of
<bibRefCitation id="087DA9F8E4F9E09EC067889AF28D1D00" author="Gruendel, J" journalOrPublisher="Der Steinkern" pageId="0" pageNumber="179" pagination="20 - 47" refId="B30" refString="Gruendel, J, 2017. Die Gastropoden-Fauna von Saal bei Kelheim. Der Steinkern 30: 20 - 47" title="Die Gastropoden-Fauna von Saal bei Kelheim." volume="30" year="2017">
<normalizedToken id="F8CFBC0CEFFB69D49924B640E8023AF5" originalValue="Gründel">Gruendel</normalizedToken>
2017
</bibRefCitation>
). These references note the presence of four plaits (two columellar plaits, one parietal and one palatal plait).
</paragraph>
</subSubSection>
<subSubSection id="7C3F48DC951888606997D27362FC81EB" pageId="0" pageNumber="179" type="relationships">
<paragraph id="FF4CBB5555A8B0323978C2F78071F188" pageId="0" pageNumber="179">Relationships.</paragraph>
<paragraph id="7A7316886A3B881A150F38347FBCCD88" pageId="0" pageNumber="179">
See Remarks for
<taxonomicName id="019F87A253F9224DCCEEC47332AB2F0D" authorityName="Cox" authorityYear="1947" class="Gastropoda" family="Eunerineidae" genus="Eunerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eunerinea" pageId="0" pageNumber="179" phylum="Mollusca" rank="genus">
<emphasis id="C132B02BE1A925636F16EBBF086A070B" italics="true" pageId="0" pageNumber="179">Eunerinea</emphasis>
</taxonomicName>
sp. 1 for relationships with
<taxonomicName id="A15424D9EA4215B9A36CED4C4B1D677D" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="C3A69B63B3F23C190FD5BCA50209905F" italics="true" pageId="0" pageNumber="179">Ptygmatis nodosa</emphasis>
</taxonomicName>
.
<taxonomicName id="43053ECE97FA6E4D62A05B1A180197EE" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="CCE3393A7B0606BD297A2234AE06DAE3" italics="true" pageId="0" pageNumber="179">Ptygmatis nodosa</emphasis>
</taxonomicName>
Voltz sensu
<bibRefCitation id="8B8A1CF7DF080D19EBB5150EC26BC5F5" author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
has higher whorls, its whorl face is more concave, its basal spiral rib is not as pronounced and widened, and it has three spiral cords on the base.
<taxonomicName id="89D263E111C1C7526344ECE5F1024422" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea danubiensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="danubiensis">
<emphasis id="DC007CFCD717A54C00ED719CBD621076" italics="true" pageId="0" pageNumber="179">Nerinea danubiensis</emphasis>
</taxonomicName>
Zittel sensu
<bibRefCitation id="B085D0222CEC51CE9918BDAAE60D9437" author="Schlosser, M" journalOrPublisher="Palaeontographica" pageId="0" pageNumber="179" pagination="41 - 110" refId="B88" refString="Schlosser, M, 1882. Die Fauna des Kelheimer Diceras-Kalkes. I. Vertebrata, Crustacea, Cephalopoda und Gastropoda. Palaeontographica 28: 41 - 110" title="Die Fauna des Kelheimer Diceras-Kalkes. I. Vertebrata, Crustacea, Cephalopoda und Gastropoda." volume="28" year="1882">Schlosser (1882)</bibRefCitation>
has more rapidly increasing whorls in width, its second nodular spiral cord is situated directly above the suture and forms the basal spiral cord at the transition from whorl face to base instead of having a non-nodular bulge/cord at this position. Moreover, this spiral cord is stronger than the subsutural row of knobs.
<taxonomicName id="58410F2775C9AD8FE6F23D4B37C55AE5" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea plassenensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="plassenensis">
<emphasis id="C1E16057C8B1F4C68BE3F14AE28F1A06" italics="true" pageId="0" pageNumber="179">Nerinea plassenensis</emphasis>
</taxonomicName>
Peters, 1855 has a more convex base and stronger subsutural knobs and therefore a more concave whorl face. In
<taxonomicName id="EF9727F7BC1BABF4F6AA2867F7D26146" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="D81CC2BF598B425967AD1CED41068560" italics="true" pageId="0" pageNumber="179">Nerinea nodosa</emphasis>
</taxonomicName>
Voltz sensu
<bibRefCitation id="5477D8080A305C126C7547BE888661AA" author="Thurmann, J" journalOrPublisher="Memoires de la Societe du museum d'histoire naturelle de Strasbourg" pageId="0" pageNumber="179" refId="B100" refString="Thurmann, J, Etallon, A, 1861-1864. Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18-20: 1-500." title="Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18 - 20: 1 - 500." year="1861 - 1864">
Thurmann and
<normalizedToken id="499BD9D1D8E123B1B3DF55976E28C033" originalValue="Étallon">Etallon</normalizedToken>
(1861-1864)
</bibRefCitation>
, the spiral cord at the transition from whorl face to base is distinctly knobby.
<taxonomicName id="423C2CF0AA5F366E7A324E19A4EB8F83" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="CAA3EB051FE597CFF5138A15B99EDD40" italics="true" pageId="0" pageNumber="179">Ptygmatis nodosa</emphasis>
</taxonomicName>
Voltz sensu
<bibRefCitation id="B1146524AC6A998B73E80F4DC5582E17" author="Maire, V" journalOrPublisher="Bulletin de la Societe grayloise d'emulation Lettress, Sciences, Beaux-arts" pageId="0" pageNumber="179" pagination="93 - 168" refId="B60" refString="Maire, V, 1913. Les Gastropodes du Jurassique superieur graylois (1re partie). Bulletin de la Societe grayloise d'emulation Lettress, Sciences, Beaux-arts 16: 93 - 168" title="Les Gastropodes du Jurassique superieur graylois (1 re partie)." volume="16" year="1913">Maire (1913</bibRefCitation>
,
<bibRefCitation id="3E4546EDE143D177468DDC55037AB11B" author="Maire, V" journalOrPublisher="Bulletin de la Societe grayloise d'emulation Lettress, Sciences, Beaux-arts" pageId="0" pageNumber="179" refId="B61" refString="Maire, V, 1927. Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85-173." title="Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85 - 173." year="1927">1927</bibRefCitation>
) has fewer but larger subsutural knobs, the edge from whorl face to base is at least partly knobby (Maire emphasized the great variability of this species), and it has some spiral cords on the base.
<taxonomicName id="CAE2A1A82F9DCBF67E3D7EE9113E4258" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea paronae" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="paronae">
<emphasis id="CE191BA2B1745E6463005A0B15EC6947" italics="true" pageId="0" pageNumber="179">Nerinea paronae</emphasis>
</taxonomicName>
Stefano, 1884 has more and smaller subsutural knobs, more weak, knobby spiral cord on the whorl face, knobs on the edge demarcating whorl face and base and it has spiral cords on the base.
</paragraph>
<caption id="2BD9FB314014DA24D22E9C6B1868488B" doi="10.3897/zitteliana.96.e84187.plate11" httpUri="https://binary.pensoft.net/fig/780389" pageId="0" pageNumber="179" start="Plate 11" startId="F11">
<paragraph id="3A0071F6CDE938A7A8C518E7B9AC2535" pageId="0" pageNumber="179">
<emphasis id="99CBFD9DFF394B5B977EE9C761831783" bold="true" pageId="0" pageNumber="179">Plate 11.</emphasis>
<taxonomicName id="D4E980E173CAA57E5E9CC2D6D8448559" baseAuthorityName="Voltz" baseAuthorityYear="1836" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis nodosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="4F2E7763F98AC3FD4EC05AFE6DC1979C" italics="true" pageId="0" pageNumber="179">Ptygmatis nodosa</emphasis>
</taxonomicName>
(Voltz, 1836).
<emphasis id="5A5DB51DCEFF40D56AC61924F9F7E6CB" bold="true" pageId="0" pageNumber="179">(1-3)</emphasis>
SNSB-BSPG 2021 XV 40, Saal (collection Lang);
<emphasis id="DE46976BA31C46FE642DBC924D074E34" bold="true" pageId="0" pageNumber="179">(1)</emphasis>
lateral view, height 22 mm;
<emphasis id="2C7D3EE9CD09255B7B4B273C338AD4C7" bold="true" pageId="0" pageNumber="179">(2)</emphasis>
last whorl with aperture, width 6.6 mm;
<emphasis id="7FB7E1BFB08A0CFC13D09281CCD12BEC" bold="true" pageId="0" pageNumber="179">(3)</emphasis>
base, width 6.6 mm.
<emphasis id="3D18DBEE2EBBD8666B44F99562C6C15F" bold="true" pageId="0" pageNumber="179">(4-5)</emphasis>
SNSB-BSPG 2021 XV 41; Saal (collection Lang);
<emphasis id="CF5CC778D8C3BFB8EFFE0D4CA09DD7E8" bold="true" pageId="0" pageNumber="179">(4)</emphasis>
lateral view, height 9 mm;
<emphasis id="431D5762EA003937F1EF5A5FF1E432BF" bold="true" pageId="0" pageNumber="179">(5)</emphasis>
base, width 3.5 mm.
<emphasis id="93748662C5E246EB25091F300FE1AD88" bold="true" pageId="0" pageNumber="179">(6)</emphasis>
SNSB-BSPG 2021 XV 42, Saal (collection Lang), aperture, width of whorl 3.7 mm.
<emphasis id="914E246942E3164EDC562C58E445C2DA" bold="true" pageId="0" pageNumber="179">(7-8)</emphasis>
SNSB-BSPG 2021 XV 43, Saal (collection Lang);
<emphasis id="732D506A0176C331FBF89166F19E73C4" bold="true" pageId="0" pageNumber="179">(7)</emphasis>
lateral view, height 11.5 mm;
<emphasis id="6637F5F46ABB43565FAEC5B463143AC2" bold="true" pageId="0" pageNumber="179">(8)</emphasis>
last two whorls in lateral view, width 4.2 mm.
<emphasis id="ECBCF891B85FFC5EDA9F7337BE51B50B" bold="true" pageId="0" pageNumber="179">(9)</emphasis>
SNSB-BSPG 2021 XV 44, Saal (collection Lang), base, width 6 mm.
<emphasis id="71A3510493F94A1A046E079EF3B87543" bold="true" pageId="0" pageNumber="179">(10-11)</emphasis>
SNSB-BSPG 2021 XV 45, Saal (collection Lang);
<emphasis id="3B6402C271C0474FBEF3B638322D02DD" bold="true" pageId="0" pageNumber="179">(10)</emphasis>
lateral view, height 16 mm;
<emphasis id="D47A7A58C77D7F7DBF12A366752261C8" bold="true" pageId="0" pageNumber="179">(11)</emphasis>
last whorls in lateral view, width 5.2 mm.
<emphasis id="1275978CA602CC466A8588A13CB29E31" bold="true" pageId="0" pageNumber="179">(12-13)</emphasis>
SNSB-BSPG 2021 XV 46, Saal (collection Keupp);
<emphasis id="201B6074089085750B9C3E4B40E7C079" bold="true" pageId="0" pageNumber="179">(12)</emphasis>
lateral view, height 9 mm;
<emphasis id="523D1CC462F556CAB3403ED23A903D9B" bold="true" pageId="0" pageNumber="179">(13)</emphasis>
apex in lateral view, height 2.9 mm.
<emphasis id="1A286318945B830B20C464F00D2E70AB" bold="true" pageId="0" pageNumber="179">(14)</emphasis>
SNSB-BSPG 2021 XV 47, Saal (collection Lang), lateral view, height 10 mm.
<emphasis id="1BEB740469F41802AE4DDCF3B6E5EA34" bold="true" pageId="0" pageNumber="179">(15)</emphasis>
SNSB-BSPG 2021 XV 48, Saal (collection Lang), last whorls with aperture, width 7.8 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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