264 lines
33 KiB
XML
264 lines
33 KiB
XML
<document id="720ADC8F0C2153A1BD5751C9818C265D" ID-CLB-Dataset="54646" ID-DOI="10.11646/zootaxa.5278.1.1" ID-GBIF-Dataset="1528757a-5499-40b9-a728-c5bf0c770b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFBAFFFEFF6D6BC2CCC1FA37" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma longitenebricum Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docUuid="82407841-CBF9-4647-A04B-A12A242B7AE0" docUuidSource="ZooBank" docVersion="9" lastPageNumber="20" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="18" updateTime="1728574925831" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="A6DA0A14FF85FE9E7491D6D55CCFB7ED">Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:name id="63A40A0D48FA7F3F6C451002C8D6445B" type="personal">
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<mods:namePart id="78576780CFB1B13E64A5105E8C4E4F5B">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="5D7539C7EACF1D9B06A9F69CBD6FF937" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation id="DC4B5D939018A8407F2E01418981CC0F">Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier id="D023C249CC649135569B855AE1CB1238" type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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<mods:namePart id="8CD043E404F37CD9A8C06C3C7B8DC0C7">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="1773405DB9DF731ED377BF2CF6831B75">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:title id="4A77EA3058C4AB9831737EF60D37EC09">Zootaxa</mods:title>
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<mods:date id="73F534DE4523448B4EAB4AC85B0C3193">2023</mods:date>
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<mods:number id="210FB8540777F9A90273C904BE0758CF">2023-05-04</mods:number>
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<treatment id="A148D509FFBAFFFEFF6D6BC2CCC1FA37" ID-DOI="http://doi.org/10.5281/zenodo.7894870" ID-GBIF-Taxon="209583693" ID-Zenodo-Dep="7894870" LSID="urn:lsid:zoobank.org:act:82407841-CBF9-4647-A04B-A12A242B7AE0" httpUri="http://treatment.plazi.org/id/A148D509FFBAFFFEFF6D6BC2CCC1FA37" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="61FB3794FFBAFFFCFF6D6BC2CA3BFF78" box="[151,579,151,178]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="295E641FFFBAFFFCFF6D6BC2CA3BFF78" blockId="17.[151,1179,151,249]" box="[151,579,151,178]" pageId="17" pageNumber="18">
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<heading id="7216D373FFBAFFFCFF6D6BC2CA3BFF78" bold="true" box="[151,579,151,178]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="1B95B80DFFBAFFFCFF6D6BC2CA3BFF78" bold="true" box="[151,579,151,178]" pageId="17" pageNumber="18">
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<taxonomicName id="EEE11F9CFFBAFFFCFF6D6BC2C982FF7B" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,506,151,178]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
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<emphasis id="1B95B80DFFBAFFFCFF6D6BC2C982FF7B" bold="true" box="[151,506,151,178]" italics="true" pageId="17" pageNumber="18">Leptostigma longitenebricum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00A60576FFBAFFFCFDFB6BC2CA3BFF78" box="[513,579,151,177]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="61FB3794FFBAFFFCFF6D6BEEC919FF30" pageId="17" pageNumber="18" type="description">
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<paragraph id="295E641FFFBAFFFCFF6D6BEECCE3FF1C" blockId="17.[151,1179,151,249]" box="[151,1179,187,213]" pageId="17" pageNumber="18">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<uuid id="5D475ECAFFBAFFFCFD7A6BEECCE3FF1C" box="[640,1179,187,213]" pageId="17" pageNumber="18">82407841-CBF9-4647-A04B-A12A242B7AE0</uuid>
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</uri>
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</paragraph>
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<paragraph id="295E641FFFBAFFFCFF6D6B8AC919FF30" blockId="17.[151,1179,151,249]" box="[151,353,223,249]" pageId="17" pageNumber="18">
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<figureCitation id="B1DA789AFFBAFFFCFF6D6B8AC970FF30" box="[151,264,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="17" pageNumber="18">Figs. 3I–J</figureCitation>
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,
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<figureCitation id="B1DA789AFFBAFFFCFEEE6B8AC925FF30" box="[276,349,223,249]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="17.[151,250,1733,1757]" captionStartId-1="18.[151,250,1928,1952]" captionStartId-2="19.[151,250,1365,1389]" captionTargetBox-0="[160,1428,561,1699]" captionTargetBox-1="[151,1436,633,1894]" captionTargetBox-2="[151,1436,489,1335]" captionTargetId-0="figure-128@17.[151,1436,561,1709]" captionTargetId-1="figure-244@18.[151,1436,633,1905]" captionTargetId-2="figure-160@19.[151,1436,489,1342]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURE 16. Leptostigma longitenebricum holotype F-1565, A, photograph, B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm." captionText-1="FIGURE 17. F-1563, tentative Leptostigma longitenebricum, A, photograph and B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm." captionText-2="FIGURE 18. GSC 142842, tentative Leptostigma longitenebricum forewing: A, photograph; B, drawing. Scale bar 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7894894" figureDoi-1="http://doi.org/10.5281/zenodo.7894898" figureDoi-2="http://doi.org/10.5281/zenodo.7894902" httpUri-0="https://zenodo.org/record/7894894/files/figure.png" httpUri-1="https://zenodo.org/record/7894898/files/figure.png" httpUri-2="https://zenodo.org/record/7894902/files/figure.png" pageId="17" pageNumber="18">16–18</figureCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="61FB3794FFBAFFFCFF6D6A72C90DFE43" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="295E641FFFBAFFFCFF6D6A72C90DFE43" blockId="17.[151,1437,295,538]" pageId="17" pageNumber="18">
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<materialsCitation id="99896E42FFBAFFFCFF6D6A72C90DFE43" ID-GBIF-Occurrence="4100387319" collectionCode="TRU" collectorName="John Leahy" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
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<emphasis id="1B95B80DFFBAFFFCFF6D6A72C97CFE88" bold="true" box="[151,260,295,321]" pageId="17" pageNumber="18">
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<typeStatus id="F65ADABDFFBAFFFCFF6D6A72C97CFE88" box="[151,260,295,321]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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</emphasis>
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. F-1565 (part, counterpart) (
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<figureCitation id="B1DA789AFFBAFFFCFDAA6A72CABDFE88" box="[592,709,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="17" pageNumber="18">Figs. 3I–J</figureCitation>
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,
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<figureCitation id="B1DA789AFFBAFFFCFD296A72CA8AFE88" box="[723,754,295,321]" captionStart="FIGURE 16" captionStartId="17.[151,250,1733,1757]" captionTargetBox="[160,1428,561,1699]" captionTargetId="figure-128@17.[151,1436,561,1709]" captionTargetPageId="17" captionText="FIGURE 16. Leptostigma longitenebricum holotype F-1565, A, photograph, B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894894" httpUri="https://zenodo.org/record/7894894/files/figure.png" pageId="17" pageNumber="18">16</figureCitation>
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).
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<collectorName id="841401C9FFBAFFFCFCF26A72CB22FE88" box="[776,858,295,321]" pageId="17" pageNumber="18">Almost</collectorName>
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complete, in dorsal aspect, but legs, one antenna, body mostly poorly preserved, hind wings poorly preserved; from
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<collectorName id="841401C9FFBAFFFCFC696A1ECB8EFEAC" box="[915,1014,331,357]" pageId="17" pageNumber="18">McAbee</collectorName>
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, collector
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<collectorName id="841401C9FFBAFFFCFB946A1ECC8EFEAC" box="[1134,1270,331,357]" pageId="17" pageNumber="18">John Leahy</collectorName>
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; housed in the collections of
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<collectionCode id="4FF0FCDAFFBAFFFCFECF6A25C909FE43" box="[309,369,368,394]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="17" pageNumber="18" type="Herbarium">TRU</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="61FB3794FFBAFFFCFF3D6AC6CBD1FDD0" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="295E641FFFBAFFFCFF3D6AC6CBD1FDD0" blockId="17.[151,1437,295,538]" pageId="17" pageNumber="18">
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<emphasis id="1B95B80DFFBAFFFCFF3D6AC6C942FE64" bold="true" box="[199,314,403,429]" pageId="17" pageNumber="18">Diagnosis</emphasis>
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. Distinguished from other species of
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<taxonomicName id="EEE11F9CFFBAFFFCFD1C6AC1CB08FE64" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[742,880,404,429]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="1B95B80DFFBAFFFCFD1C6AC1CB08FE64" box="[742,880,404,429]" italics="true" pageId="17" pageNumber="18">Leptostigma</emphasis>
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</taxonomicName>
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by antenna with a combination of club elongate, darker than flagellar base, base two long, pale segments; forewing lightly, evenly infuscate throughout. Tentatively diagnostic: R+M longer than 1-M, 2r-rs enters cell 3rm at or before its middle, 2r-m enters M far behind 2m-cu, anal cells separated by 1A joining 2A+3A for at least half height of cell 2a.
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</paragraph>
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</subSubSection>
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<caption id="7D9E3497FFBAFFFCFF6D6D90C9B8F8C8" ID-DOI="http://doi.org/10.5281/zenodo.7894894" ID-Zenodo-Dep="7894894" httpUri="https://zenodo.org/record/7894894/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1733,1757]" targetBox="[160,1428,561,1699]" targetPageId="17" targetType="figure">
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<paragraph id="295E641FFFBAFFFCFF6D6D90C9B8F8C8" blockId="17.[151,1436,1733,1793]" pageId="17" pageNumber="18">
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<emphasis id="1B95B80DFFBAFFFCFF6D6D90C959F914" bold="true" box="[151,289,1733,1757]" pageId="17" pageNumber="18">FIGURE 16.</emphasis>
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<taxonomicName id="EEE11F9CFFBAFFFCFED36D93CA2DF914" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[297,597,1733,1757]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="longitenebricum">
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<emphasis id="1B95B80DFFBAFFFCFED36D93CA2DF914" box="[297,597,1733,1757]" italics="true" pageId="17" pageNumber="18">Leptostigma longitenebricum</emphasis>
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</taxonomicName>
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holotype F-1565, A, photograph, B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="61FB3794FFBAFFFFFF3D6C7ACAF1FDA8" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
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<paragraph id="295E641FFFBAFFFFFF3D6C7ACAF1FDA8" blockId="17.[151,1437,1839,2010]" lastBlockId="18.[151,1437,151,609]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<emphasis id="1B95B80DFFBAFFFCFF3D6C7AC929F880" bold="true" box="[199,337,1839,1865]" pageId="17" pageNumber="18">Description</emphasis>
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.
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<typeStatus id="F65ADABDFFBAFFFCFE9B6C7AC9B5F880" box="[353,461,1839,1865]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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. Body length
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<emphasis id="1B95B80DFFBAFFFCFD906C65CAFCF880" box="[618,644,1840,1865]" italics="true" pageId="17" pageNumber="18">ca</emphasis>
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.
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<quantity id="EE19C9FAFFBAFFFCFD6E6C7ACA83F883" box="[660,763,1839,1866]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="18" unit="mm" value="15.5">15.5 mm</quantity>
|
||
, body, legs, pterostigma dark (tarsi less dark), antenna with club darkened, funicle paler, wing membrane lightly infuscate throughout. Head roundish, eyes occupying most of head sides, oval, converging ventrally, with temples weak (scarcely visible in front view), toruli close to each other. Antenna with eight antennomeres, club elongate, narrowed apically, two basal flagellar segments cylindrical, next three widened, apical one narrowed distally; basal, apical segments longest, penultimate shortest.Pronotum with wide lateral lobes; median longitudinal mesonotal sulcus, notauli well developed, meeting well before scarcely transverse scutellum; metanotum rather narrow, with wide triangular scutellum, small, roundish cenchri. Forewing length
|
||
<typeStatus id="F65ADABDFFB9FFFFFF6D6B8AC881FF30" box="[151,249,223,249]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
<quantity id="EE19C9FAFFB9FFFFFEF86B8BC910FF30" box="[258,360,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="19" unit="mm" value="15.5">15.5 mm</quantity>
|
||
(F-1563,
|
||
<quantity id="EE19C9FAFFB9FFFFFE216B8BCA3CFF30" box="[475,580,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="18" pageNumber="19" unit="mm" value="11.5">11.5 mm</quantity>
|
||
), maximum width estimated
|
||
<quantity id="EE19C9FAFFB9FFFFFC6B6B8BCC7FFF30" box="[913,1031,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="18" pageNumber="19" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 mm</quantity>
|
||
, length/width: 3.1–3.3; pterostigma narrow, tapering, similar to width of C-R before pterostigma; R+M longer than 1-M; maximum length of cell 2r shorter than 3r; 2r-rs shorter than Rs between 2r-m, 2r-rs, joining Rs at middle of cell 3rm; 2r-rm curved, sometimes slightly sigmoidal, joins Rs at or slightly distal to 2m-cu joins M, joins M behind 2m-cu more than for half length of 2r-m; 1-M curved or (GSC 142842) almost straight; cell 1mcu with M, 1m-cu slightly divergent toward anterior margin; 3r-m near straight or (GSC 142842) distinctly bowed toward base; cell 2mcu with M similar length or (GSC 142842) shorter than 1m-cu; 2m-cu slightly curved toward base; cu-a postfurcal (
|
||
<typeStatus id="F65ADABDFFB9FFFFFBD06AE2CCF7FE18" box="[1066,1167,439,465]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
) or interstitial; anal cell separated by 1A fusion with 2A+3A (no crossvein) for length of almost half width of cell 2a in GSC 142842, for more than its width in F-1563. Mid, hind femora long, oval,
|
||
<emphasis id="1B95B80DFFB9FFFFFCC46AAACB18FDD1" box="[830,864,511,536]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
|
||
3.5 times as long as wide; tibiae thin, only slightly longer than their femora, basitarsi
|
||
<emphasis id="1B95B80DFFB9FFFFFDE06976CA43FDF5" box="[538,571,547,572]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
|
||
0.35 as long as their tibiae. First abdominal tergum if correctly identified (F-1563 only) entire, with straight hind margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFB9FFFFFF6D6CDDC9B8F80D" ID-DOI="http://doi.org/10.5281/zenodo.7894898" ID-Zenodo-Dep="7894898" httpUri="https://zenodo.org/record/7894898/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1928,1952]" targetBox="[151,1436,633,1894]" targetPageId="18" targetType="figure">
|
||
<paragraph id="295E641FFFB9FFFFFF6D6CDDC9B8F80D" blockId="18.[151,1436,1928,1988]" pageId="18" pageNumber="19">
|
||
<emphasis id="1B95B80DFFB9FFFFFF6D6CDDC967F869" bold="true" box="[151,287,1928,1952]" pageId="18" pageNumber="19">FIGURE 17.</emphasis>
|
||
F-1563, tentative
|
||
<taxonomicName id="EEE11F9CFFB9FFFFFE2F6CDCCA86F869" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[469,766,1928,1952]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="longitenebricum">
|
||
<emphasis id="1B95B80DFFB9FFFFFE2F6CDCCA86F869" box="[469,766,1928,1952]" italics="true" pageId="18" pageNumber="19">Leptostigma longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
, A, photograph and B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFB8FFFEFF3D6BC2C933FEAC" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="295E641FFFB8FFFEFF3D6BC2C933FEAC" blockId="19.[151,1437,151,465]" pageId="19" pageNumber="20">
|
||
<emphasis id="1B95B80DFFB8FFFEFF3D6BC2C94CFF78" bold="true" box="[199,308,151,177]" pageId="19" pageNumber="20">Remarks</emphasis>
|
||
. Differs from
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFE296BC2CA1EFF78" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[467,614,151,177]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFE296BC2CA1EFF78" box="[467,614,151,177]" italics="true" pageId="19" pageNumber="20">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFD966BC2CAD2FF78" bold="true" box="[620,682,151,177]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFD966BC2CAD2FF78" box="[620,682,151,177]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFD1A6BC2CBE1FF78" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[736,921,151,177]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFD1A6BC2CBE1FF78" box="[736,921,151,177]" italics="true" pageId="19" pageNumber="20">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFC5A6BC2CBA6FF78" bold="true" box="[928,990,151,177]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFC5A6BC2CBA6FF78" box="[928,990,151,177]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by flagellar base pale and, venationally, in more the distal position of 2r-rs (behind middle of cell 3rm). Differs from
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFBF66BEECCDBFF1C" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1036,1187,187,213]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFBF66BEECCDBFF1C" box="[1036,1187,187,213]" italics="true" pageId="19" pageNumber="20">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFB566BEECC94FF1C" bold="true" box="[1196,1260,187,213]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFB566BEECC94FF1C" box="[1196,1260,187,213]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its antennal club being long, also venationally by its R+M long and anal cells distinctly separated by fusion of 1A with 2A+3A. Differs from
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFECB6A56C9BAFED5" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[305,450,259,284]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="proxivena" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFECB6A56C9BAFED5" box="[305,450,259,284]" italics="true" pageId="19" pageNumber="20">L. proxivena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFE366A56CA76FED4" bold="true" box="[460,526,259,285]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFE366A56CA76FED4" box="[460,526,259,285]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFDDA6A56CAD6FED4" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[544,686,259,285]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fasciatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFDDA6A56CAD6FED4" box="[544,686,259,285]" italics="true" pageId="19" pageNumber="20">L. fasciatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFD436A56CA83FED4" bold="true" box="[697,763,259,285]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFD436A56CA83FED4" box="[697,763,259,285]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFCF76A56CBC9FED4" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[781,945,259,285]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alaemacula" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFB8FFFEFCF76A56CBC9FED4" box="[781,945,259,285]" italics="true" pageId="19" pageNumber="20">L. alaemacula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFB8FFFEFC416A56CB85FED4" bold="true" box="[955,1021,259,285]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="00A60576FFB8FFFEFC416A56CB85FED4" box="[955,1021,259,285]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with unknown antenna by its wing membrane lightly, evenly infuscate throughout (without fascia or macula) and by 2r-m entering M far behind, not close to, 2m-cu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFB8FFFEFF3D6A3BCA57FE18" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="295E641FFFB8FFFEFF3D6A3BCA57FE18" blockId="19.[151,1437,151,465]" pageId="19" pageNumber="20">
|
||
<emphasis id="1B95B80DFFB8FFFEFF3D6A3BC957FE41" bold="true" box="[199,303,366,392]" pageId="19" pageNumber="20">Material</emphasis>
|
||
.
|
||
<materialsCitation id="99896E42FFB8FFFEFEC56A3ACA57FE18" ID-GBIF-Occurrence="4100387304" collectingDate="2000-07-12" collectionCode="TRU, GSC" pageId="19" pageNumber="20" specimenCode="GSC 142842" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="F65ADABDFFB8FFFEFEC56A3AC9DFFE40" box="[319,423,367,393]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
and tentatively from venation F-1563 (
|
||
<figureCitation id="B1DA789AFFB8FFFEFC916A3ACBC7FE40" box="[875,959,367,393]" captionStart="FIGURE 17" captionStartId="18.[151,250,1928,1952]" captionTargetBox="[151,1436,633,1894]" captionTargetId="figure-244@18.[151,1436,633,1905]" captionTargetPageId="18" captionText="FIGURE 17. F-1563, tentative Leptostigma longitenebricum, A, photograph and B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894898" httpUri="https://zenodo.org/record/7894898/files/figure.png" pageId="19" pageNumber="20">Fig. 17</figureCitation>
|
||
), collector unknown, in the collections of
|
||
<collectionCode id="4FF0FCDAFFB8FFFEFF6D6AC6C8AAFE64" box="[151,210,403,429]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="19" pageNumber="20" type="Herbarium">TRU</collectionCode>
|
||
, and
|
||
<specimenCode id="7947CC64FFB8FFFEFEF66AC6C9D9FE64" box="[268,417,403,429]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC 142842</specimenCode>
|
||
, collector SBA at McAbee (
|
||
<collectionCode id="4FF0FCDAFFB8FFFEFD206AC6CB6AFE64" box="[730,786,403,429]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
|
||
locality V-016800),
|
||
<date id="5D5F42DFFFB8FFFEFC036AC6CCE9FE64" box="[1017,1169,403,429]" pageId="19" pageNumber="20" value="2000-07-12">
|
||
<collectingDate id="4D1BBB37FFB8FFFEFC036AC6CCE9FE64" box="[1017,1169,403,429]" pageId="19" pageNumber="20" value="2000-07-12">July 12, 2000</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="B1DA789AFFB8FFFEFB5A6AC6CC8AFE64" box="[1184,1266,403,429]" captionStart="FIGURE 18" captionStartId="19.[151,250,1365,1389]" captionTargetBox="[151,1436,489,1335]" captionTargetId="figure-160@19.[151,1436,489,1342]" captionTargetPageId="19" captionText="FIGURE 18. GSC 142842, tentative Leptostigma longitenebricum forewing: A, photograph; B, drawing. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894902" httpUri="https://zenodo.org/record/7894902/files/figure.png" pageId="19" pageNumber="20">Fig. 18</figureCitation>
|
||
), housed in the collections of the
|
||
<collectionCode id="4FF0FCDAFFB8FFFEFEA56AE2C9EDFE18" box="[351,405,439,465]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
|
||
(Vancouver).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFB8FFFEFF6D6E00CD16FAA4" ID-DOI="http://doi.org/10.5281/zenodo.7894902" ID-Zenodo-Dep="7894902" box="[151,1390,1365,1390]" httpUri="https://zenodo.org/record/7894902/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1365,1389]" targetBox="[151,1436,489,1335]" targetPageId="19" targetType="figure">
|
||
<paragraph id="295E641FFFB8FFFEFF6D6E00CD16FAA4" blockId="19.[151,1390,1365,1390]" box="[151,1390,1365,1390]" pageId="19" pageNumber="20">
|
||
<emphasis id="1B95B80DFFB8FFFEFF6D6E00C958FAA7" bold="true" box="[151,288,1365,1390]" pageId="19" pageNumber="20">FIGURE 18.</emphasis>
|
||
GSC 142842, tentative
|
||
<taxonomicName id="EEE11F9CFFB8FFFEFDEE6E03CB45FAA7" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[532,829,1366,1390]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longitenebricum">
|
||
<emphasis id="1B95B80DFFB8FFFEFDEE6E03CB45FAA7" box="[532,829,1366,1390]" italics="true" pageId="19" pageNumber="20">Leptostigma longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
forewing: A, photograph; B, drawing. Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFB8FFFEFF3D6EC9C9EAFA13" pageId="19" pageNumber="20" type="etymology">
|
||
<paragraph id="295E641FFFB8FFFEFF3D6EC9C9EAFA13" blockId="19.[151,1437,1436,1534]" pageId="19" pageNumber="20">
|
||
<emphasis id="1B95B80DFFB8FFFEFF3D6EC9C936FA7F" bold="true" box="[199,334,1436,1462]" pageId="19" pageNumber="20">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="1B95B80DFFB8FFFEFDF56EC9CA22FA7F" box="[527,602,1436,1462]" italics="true" pageId="19" pageNumber="20">longus</emphasis>
|
||
, meaning long, and
|
||
<emphasis id="1B95B80DFFB8FFFEFCB36EC9CBC6FA7F" box="[841,958,1436,1462]" italics="true" pageId="19" pageNumber="20">tenebricus</emphasis>
|
||
, dark, referring to the antennal club. The epithet is an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFB8FFFEFF3D6EB1CCC1FA37" box="[199,1209,1508,1534]" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="295E641FFFB8FFFEFF3D6EB1CCC1FA37" blockId="19.[151,1437,1436,1534]" box="[199,1209,1508,1534]" pageId="19" pageNumber="20">
|
||
<emphasis id="1B95B80DFFB8FFFEFF3D6EB1C9ECFA37" bold="true" box="[199,404,1508,1534]" pageId="19" pageNumber="20">Locality and age.</emphasis>
|
||
McAbee,
|
||
<collectingRegion id="EB25AAFDFFB8FFFEFDF36EB1CAB5FA37" box="[521,717,1508,1534]" country="Canada" name="British Columbia" pageId="19" pageNumber="20">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="51F6248FFFB8FFFEFD236EB1CB48FA37" box="[729,816,1508,1534]" name="Canada" pageId="19" pageNumber="20">Canada</collectingCountry>
|
||
, Hoodoo Face beds; mid-Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |