144 lines
15 KiB
XML
144 lines
15 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFB4FFF2FF6D6977CB6CF9EE" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocimbex clavatus Rohwer 1908" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="32" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Archibald, S. Bruce</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
|
||
<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
|
||
<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-05-04</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5278</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>38</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
|
||
<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894927" ID-Zenodo-Dep="7894927" LSID="urn:lsid:plazi:treatment:A148D509FFB4FFF2FF6D6977CB6CF9EE" httpUri="http://treatment.plazi.org/id/A148D509FFB4FFF2FF6D6977CB6CF9EE" lastPageNumber="32" pageId="31" pageNumber="32">
|
||
<subSubSection box="[151,610,546,573]" pageId="31" pageNumber="32" type="nomenclature">
|
||
<paragraph blockId="31.[151,610,546,609]" box="[151,610,546,573]" pageId="31" pageNumber="32">
|
||
<heading bold="true" box="[151,610,546,573]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
|
||
<taxonomicName authority="Rohwer, 1908" authorityName="Rohwer" authorityYear="1908" box="[151,610,546,573]" class="Insecta" family="Cimbicidae" genus="Pseudocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="clavatus">
|
||
<emphasis bold="true" box="[151,610,546,573]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[151,432,546,573]" italics="true" pageId="31" pageNumber="32">Pseudocimbex clavatus</emphasis>
|
||
<bibRefCitation author="Rohwer, S. A." box="[440,610,546,573]" pageId="31" pageNumber="32" pagination="521 - 530" refId="ref19150" refString="Rohwer, S. A. (1908) On the Tenthredinoidea of the Florissant Shales. Bulletin of the American Museum of Natural History, 24, 521 - 530." type="journal article" year="1908">Rohwer, 1908</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,238,583,609]" pageId="31" pageNumber="32" type="description">
|
||
<paragraph blockId="31.[151,610,546,609]" box="[151,238,583,609]" pageId="31" pageNumber="32">
|
||
<figureCitation box="[151,234,583,609]" captionStart="FIGURE 28" captionStartId="30.[151,250,1468,1492]" captionTargetBox="[151,1430,777,1439]" captionTargetId="figure-263@30.[151,1436,777,1445]" captionTargetPageId="30" captionText="FIGURE 28. Pseudocimbex clavatus Rohwer holotype, UCM-18614, Tenthredinoidea incertae sedis, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing; E, drawing of left hind wing. Scale bars A–B, 5 mm; C–D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7894937" httpUri="https://zenodo.org/record/7894937/files/figure.png" pageId="31" pageNumber="32">Fig. 28</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,727,651,675]" pageId="31" pageNumber="32" type="reference_group">
|
||
<paragraph blockId="31.[151,727,651,675]" box="[151,727,651,675]" pageId="31" pageNumber="32">
|
||
<treatmentCitationGroup box="[151,727,651,675]" pageId="31" pageNumber="32">
|
||
<treatmentCitation author="Rohwer, S. A." box="[151,615,651,675]" page="527" pageId="31" pageNumber="32" year="1908">
|
||
<taxonomicName authority="Rohwer, 1908, p. 527" authorityName="Rohwer" authorityPageNumber="527" authorityYear="1908" box="[151,615,651,675]" class="Insecta" family="Cimbicidae" genus="Pseudocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="clavatus">
|
||
<emphasis box="[151,388,651,675]" italics="true" pageId="31" pageNumber="32">Pseudocimbex clavatus</emphasis>
|
||
<bibRefCitation author="Rohwer, S. A." box="[395,539,651,675]" pageId="31" pageNumber="32" pagination="521 - 530" refId="ref19150" refString="Rohwer, S. A. (1908) On the Tenthredinoidea of the Florissant Shales. Bulletin of the American Museum of Natural History, 24, 521 - 530." type="journal article" year="1908">Rohwer, 1908</bibRefCitation>
|
||
, p. 527
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
,
|
||
<figureCitation box="[626,723,651,675]" captionStart="FIGURE 1" captionStartId="1.[151,250,1979,2003]" captionTargetBox="[159,1427,1145,1945]" captionTargetId="figure-357@1.[151,1436,1137,1956]" captionTargetPageId="1" captionText="FIGURE 1. Forewing and hind wing of Corynis crassicornis (Rossi) (Corynidinae) with veins (roman font) and cells (italic font) labelled. Drawn from Vilhelmsen (2019) figure 15B. Scale = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894832" httpUri="https://zenodo.org/record/7894832/files/figure.png" pageId="31" pageNumber="32">Fig. 1a, e</figureCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,1385,721,748]" pageId="31" pageNumber="32" type="materials_examined">
|
||
<paragraph blockId="31.[151,1437,721,1575]" box="[151,1385,721,748]" pageId="31" pageNumber="32">
|
||
<materialsCitation box="[151,1385,721,748]" collectionCode="UCM" pageId="31" pageNumber="32" specimenCode="UCM 18614" specimenCount="1" typeStatus="holotype">
|
||
<emphasis bold="true" box="[151,267,721,747]" pageId="31" pageNumber="32">
|
||
<typeStatus box="[151,262,721,747]" pageId="31" pageNumber="32">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode box="[274,418,721,747]" collectionCode="UCM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="31" pageNumber="32" type="University or college">UCM 18614</specimenCode>
|
||
,
|
||
<bibRefCitation author="Rohwer, S. A." box="[429,603,721,747]" pageId="31" pageNumber="32" pagination="521 - 530" refId="ref19150" refString="Rohwer, S. A. (1908) On the Tenthredinoidea of the Florissant Shales. Bulletin of the American Museum of Natural History, 24, 521 - 530." type="journal article" year="1908">Rohwer (1908)</bibRefCitation>
|
||
. An almost complete specimen missing most of the legs, one antenna.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,477,757,784]" pageId="31" pageNumber="32" type="diagnosis">
|
||
<paragraph blockId="31.[151,1437,721,1575]" box="[199,477,757,784]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[199,322,757,783]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
|
||
As for genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="32" type="description">
|
||
<paragraph blockId="31.[151,1437,721,1575]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[199,345,793,819]" pageId="31" pageNumber="32">Description.</emphasis>
|
||
<typeStatus box="[350,457,793,819]" pageId="31" pageNumber="32">Holotype</typeStatus>
|
||
, female. Body: length
|
||
<quantity box="[706,805,793,820]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="31" pageNumber="32" unit="mm" value="17.6">17.6 mm</quantity>
|
||
as preserved; head, thorax darker than abdomen. Surface sculpture not discernible. Structure of head, thorax obscure. Antenna very short (flagellum
|
||
<quantity box="[1180,1266,829,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="mm" value="1.8">1.8 mm</quantity>
|
||
long, less than half head width), capitate, apparently 11-segmented, scape and pedicel very short, with funicle of four segments gradually shortening distally from twice as long as wide up to strongly transverse and almost round club of possibly five segments, four very short and apical one longer. Hind tibia
|
||
<quantity box="[853,938,937,964]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="31" pageNumber="32" unit="mm" value="4.6">4.6 mm</quantity>
|
||
long, other leg segments incompletely or not preserved. Forewing length
|
||
<quantity box="[463,565,973,1000]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="31" pageNumber="32" unit="mm" value="12.9">12.9 mm</quantity>
|
||
, maximum width
|
||
<quantity box="[765,853,973,1000]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="mm" value="5.5">5.5 mm</quantity>
|
||
, length/width 2.4; membrane lightly infuscate, more so toward base; pterostigma dark, narrow, short, wedge-shaped; C, R basal to pterostigma thin, widely separated, slightly more so distally, with crossvein-like Sc between; pterostigma small, wedge-shaped; 1-M at or very close to Rs+M on R; 1r-rs short, 2r-rs lost; Rs+M very short; 2-Rs (separating cells 1r and 2rm) possibly detected (as dotted line in
|
||
<figureCitation box="[229,313,1117,1143]" captionStart="FIGURE 28" captionStartId="30.[151,250,1468,1492]" captionTargetBox="[151,1430,777,1439]" captionTargetId="figure-263@30.[151,1436,777,1445]" captionTargetPageId="30" captionText="FIGURE 28. Pseudocimbex clavatus Rohwer holotype, UCM-18614, Tenthredinoidea incertae sedis, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing; E, drawing of left hind wing. Scale bars A–B, 5 mm; C–D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7894937" httpUri="https://zenodo.org/record/7894937/files/figure.png" pageId="31" pageNumber="32">Fig. 28</figureCitation>
|
||
); Rs distal of 1r smoothly curved, meeting R
|
||
<subScript attach="left" box="[817,825,1136,1150]" fontSize="6" pageId="31" pageNumber="32">1</subScript>
|
||
at wing margin; 2r-rs absent; 2r-m, 3r-m present, well spaced, straight, near 0.4 and 0.7 length of cell 3r; M notably angled at 1m-cu; 1m-cu short, just more than half length of 1-M, diverging with 1-M toward anterior margin, 1m-cu and 2m-cu join cells 2rm and 3rm, respectively; 1m-cu converging to 1-M forward; 2m-cu slightly curved toward base; cu-a interstitial; anal cells incompletely preserved: cell 1a apparently open posteriorly, distally (2+3A not reaching a1-a2 crossveins), cell 2a closed basally, not preserved distally (distal portion of 2+3A obscured by fore margin of hind wing). Hind wing partially known, with unusual venation; uniformly infuscate as forewing; 1r-m about its length distant from Rs base; with Rs, M+Cu, M, Cu, cu-a and 1A detected, M+Cu forking far basally of RS base, with fork narrow, M and Cu both almost aligned with M+Cu; cu-a slightly reclival, long, placed slightly basal of Rs base; 2r-m, m-cu not evident; 1A evident, with at least short free end preserved. Abdomen moderately narrow, with few details discernible; ovipositor short (length
|
||
<quantity box="[151,255,1477,1504]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="31" pageNumber="32" unit="mm" value="0.65">0.65 mm</quantity>
|
||
), barely exceeding abdomen apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="32" type="materials_examined">
|
||
<paragraph blockId="31.[151,1437,721,1575]" box="[199,429,1513,1540]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[199,303,1513,1539]" pageId="31" pageNumber="32">Material</emphasis>
|
||
.
|
||
<typeStatus box="[317,425,1513,1539]" pageId="31" pageNumber="32">Holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="31.[151,1437,721,1575]" box="[199,788,1549,1575]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[199,404,1549,1575]" pageId="31" pageNumber="32">Locality and age.</emphasis>
|
||
Florissant Formation, Priabonian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |