184 lines
17 KiB
XML
184 lines
17 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFB3FFF4FF6D69E7C979FE64" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eopachylosticta byrami Cockerell 1925" docType="treatment" docVersion="2" lastPageNumber="26" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="25" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:A148D509FFB3FFF4FF6D69E7C979FE64" httpUri="http://treatment.plazi.org/id/A148D509FFB3FFF4FF6D69E7C979FE64" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<subSubSection box="[151,656,690,717]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph blockId="24.[151,656,690,753]" box="[151,656,690,717]" pageId="24" pageNumber="25">
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<heading bold="true" box="[151,656,690,717]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<taxonomicName authority="(Cockerell, 1925)" authorityName="Cockerell" authorityYear="1925" baseAuthorityName="Cockerell" baseAuthorityYear="1925" box="[151,656,690,717]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="byrami">
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<emphasis bold="true" box="[151,656,690,717]" pageId="24" pageNumber="25">
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<emphasis bold="true" box="[151,436,690,717]" italics="true" pageId="24" pageNumber="25">Eopachylosticta byrami</emphasis>
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(
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<bibRefCitation author="Cockerell, T. D. A." box="[451,647,690,716]" pageId="24" pageNumber="25" pagination="1 - 15" refId="ref17014" refString="Cockerell, T. D. A. (1925) Fossil insects in the United States National Museum. Proceedings of the United States National Museum, 64, 1 - 15. [https: // www. biodiversitylibrary. org / item / 32410 # page / 7 / mode / 1 up]" type="journal article" year="1925">Cockerell, 1925</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection box="[151,238,727,753]" pageId="24" pageNumber="25" type="description">
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<paragraph blockId="24.[151,656,690,753]" box="[151,238,727,753]" pageId="24" pageNumber="25">
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<figureCitation box="[151,234,727,753]" captionStart="FIGURE 23" captionStartId="24.[151,250,1555,1579]" captionTargetBox="[151,1426,777,1532]" captionTargetId="figure-227@24.[151,1436,777,1532]" captionTargetPageId="24" captionText="FIGURE 23. Eopachylosticta byrami (Cockerell) holotype, USNM-69181, Pachylostictinae, from the Green River Formation: A, photograph; B, drawing. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894925" httpUri="https://zenodo.org/record/7894925/files/figure.png" pageId="24" pageNumber="25">Fig. 23</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7894925" ID-Zenodo-Dep="7894925" httpUri="https://zenodo.org/record/7894925/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1555,1579]" targetBox="[151,1426,777,1532]" targetPageId="24" targetType="figure">
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<paragraph blockId="24.[151,1437,1555,1615]" pageId="24" pageNumber="25">
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<emphasis bold="true" box="[151,288,1555,1579]" pageId="24" pageNumber="25">FIGURE 23.</emphasis>
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<taxonomicName authority="(Cockerell)" authorityName="Cockerell" authorityYear="1925" baseAuthorityName="Cockerell" baseAuthorityYear="1925" box="[294,657,1555,1579]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="byrami">
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<emphasis box="[294,535,1555,1579]" italics="true" pageId="24" pageNumber="25">Eopachylosticta byrami</emphasis>
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(Cockerell)
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</taxonomicName>
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holotype, USNM-69181,
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<taxonomicName authorityName="Benson" authorityYear="1938" box="[922,1091,1555,1579]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
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, from the Green River Formation: A, photograph; B, drawing. Scale bar 5 mm.
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</paragraph>
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</caption>
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<subSubSection pageId="24" pageNumber="25" type="reference_group">
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||
<paragraph blockId="24.[151,687,1657,1712]" box="[151,687,1657,1682]" pageId="24" pageNumber="25">
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<treatmentCitationGroup box="[151,687,1657,1682]" pageId="24" pageNumber="25">
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<treatmentCitation author="Cockerell, T. D. A." box="[151,536,1657,1682]" page="10" pageId="24" pageNumber="25" year="1925">
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<taxonomicName authority="Cockerell, 1925, p. 10" authorityName="Cockerell" authorityPageNumber="10" authorityYear="1925" box="[151,536,1657,1682]" class="Insecta" family="Cimbicidae" genus="Amasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="byrami">
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<emphasis box="[151,303,1658,1682]" italics="true" pageId="24" pageNumber="25">Amasis byrami</emphasis>
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<bibRefCitation author="Cockerell, T. D. A." box="[309,473,1657,1682]" pageId="24" pageNumber="25" pagination="1 - 15" refId="ref17014" refString="Cockerell, T. D. A. (1925) Fossil insects in the United States National Museum. Proceedings of the United States National Museum, 64, 1 - 15. [https: // www. biodiversitylibrary. org / item / 32410 # page / 7 / mode / 1 up]" type="journal article" year="1925">Cockerell, 1925</bibRefCitation>
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, p. 10
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</taxonomicName>
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</treatmentCitation>
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, plate 1,
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<figureCitation box="[628,687,1658,1682]" captionStart="FIGURE 4" captionStartId="6.[151,250,1327,1351]" captionTargetBox="[155,1430,525,1300]" captionTargetId="figure-85@6.[151,1436,525,1304]" captionTargetPageId="6" captionText="FIGURE 4. Allenbycimbex morrisae, holotype RBCM.EH2007.002.0001: A, photograph; B, drawing; C, drawing of right wing; D, drawing of left wing. Scale bar for A and B is 5 mm (as the fossil has been stretched by geologic shear forces, C and D are altered to the presumed unstretched configuration)." figureDoi="http://doi.org/10.5281/zenodo.7894842" httpUri="https://zenodo.org/record/7894842/files/figure.png" pageId="24" pageNumber="25">Fig. 4</figureCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="24.[151,687,1657,1712]" box="[151,612,1688,1712]" pageId="24" pageNumber="25">
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<treatmentCitationGroup box="[151,612,1688,1712]" pageId="24" pageNumber="25">
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<treatmentCitation author="Malaise, R." box="[151,608,1688,1712]" page="14" pageId="24" pageNumber="25" year="1945">
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<taxonomicName authority="Malaise, 1945, p. 14" authorityName="Malaise" authorityPageNumber="14" authorityYear="1945" box="[151,608,1688,1712]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="byrami">
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<emphasis box="[151,393,1688,1712]" italics="true" pageId="24" pageNumber="25">Eopachylosticta byrami</emphasis>
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<bibRefCitation author="Malaise, R." box="[399,545,1688,1712]" pageId="24" pageNumber="25" pagination="1 - 288" refId="ref18378" refString="Malaise, R. (1945) Tenthredinoidoa of south-eastern Asia with a general zoogeographical review. Opuscula Entomologica Suppl, 4 (288), 1 - 288. https: // doi. org / 10.1080 / 11035894509446460" type="journal article" year="1945">Malaise, 1945</bibRefCitation>
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, p. 14
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="24" pageNumber="25" type="materials_examined">
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<paragraph blockId="24.[151,1437,1758,2037]" pageId="24" pageNumber="25">
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||
<materialsCitation collectionCode="USNM" county="Green River Formation" location="River Formation" municipality="Green River" pageId="24" pageNumber="25" specimenCode="USNM 69181" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
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<emphasis bold="true" box="[151,267,1759,1785]" pageId="24" pageNumber="25">
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<typeStatus box="[151,262,1759,1785]" pageId="24" pageNumber="25">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[275,438,1758,1785]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">USNM 69181</specimenCode>
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. A complete female with its body mostly well preserved, but almost no legs evident and poorly preserved wings with venation too faint to interpret.
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<collectingCounty box="[848,1118,1794,1821]" pageId="24" pageNumber="25">
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<collectingMunicipality box="[848,988,1794,1821]" pageId="24" pageNumber="25">
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Green
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<location LSID="urn:lsid:plazi:treatment:A148D509FFB3FFF4FF6D69E7C979FE64:2C3E32C4FFB3FFF5FC646C57CBA4F8D5" box="[926,988,1794,1820]" county="Green River Formation" municipality="Green River" name="River Formation" pageId="24" pageNumber="25" stateProvince="Colorado">River</location>
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</collectingMunicipality>
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Formation
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</collectingCounty>
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, about
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<quantity box="[1205,1290,1794,1821]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="24" pageNumber="25" unit="mi" value="8.0">8 miles</quantity>
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south of
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<location LSID="urn:lsid:plazi:treatment:A148D509FFB3FFF4FF6D69E7C979FE64:2C3E32C4FFB3FFF5FA866C56C899F888" county="Green River Formation" municipality="Green River" name="De Beque" pageId="24" pageNumber="25" stateProvince="Colorado">De Beque</location>
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,
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<collectingRegion box="[237,341,1830,1856]" country="United States of America" name="Colorado" pageId="24" pageNumber="25">Colorado</collectingRegion>
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(
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<bibRefCitation author="Cockerell, T. D. A." box="[355,532,1830,1857]" pageId="24" pageNumber="25" pagination="1 - 15" refId="ref17014" refString="Cockerell, T. D. A. (1925) Fossil insects in the United States National Museum. Proceedings of the United States National Museum, 64, 1 - 15. [https: // www. biodiversitylibrary. org / item / 32410 # page / 7 / mode / 1 up]" type="journal article" year="1925">Cockerell 1925</bibRefCitation>
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), Ypresian.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,477,1866,1893]" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph blockId="24.[151,1437,1758,2037]" box="[199,477,1866,1893]" pageId="24" pageNumber="25">
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<emphasis bold="true" box="[199,322,1866,1892]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
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<paragraph blockId="24.[151,1437,1758,2037]" lastBlockId="25.[151,1437,151,429]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<emphasis bold="true" box="[199,345,1902,1928]" pageId="24" pageNumber="25">Description.</emphasis>
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Body length
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<emphasis box="[508,534,1903,1928]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
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.
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<quantity box="[552,645,1903,1929]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="24" pageNumber="25" unit="mm" value="8.2">8.2 mm</quantity>
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; head, antennae dark; thorax dark, hind femora somewhat infuscate (otherwise legs not visibly preserved, presumably pale), anterior abdomen light, posterior segments lightly infuscate. Head transverse, temples short; compound eyes not preserved; antenna five-segmented, with pedicel transverse, distinctly wider than flagellar base, flagellomere 1 thin, slightly widened apically, twice as long as wide, trapezoid, flagellomere 2 wider, together about as long as thick, unsegmented club. Pronotum not distinct, mesonotum only partially visible, with median longitudinal mesonotal sulcus and notauli well developed, meeting well before presumed position of scutellum (poorly preserved); mesopleuron with percurrent mesopleural groove; otherwise, thoracic structure obscure. Hind femora short. Forewing almost entirely invisible as preserved. Abdomen short, wide; ovipositor short, not exceeding abdominal apex, sheaths not distinct.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="materials_examined">
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<paragraph blockId="25.[151,1437,151,429]" box="[199,429,330,357]" pageId="25" pageNumber="26">
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<emphasis bold="true" box="[199,303,330,356]" pageId="25" pageNumber="26">Material</emphasis>
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.
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<typeStatus box="[317,425,331,357]" pageId="25" pageNumber="26">Holotype</typeStatus>
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.
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</paragraph>
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<paragraph blockId="25.[151,1437,151,429]" pageId="25" pageNumber="26">
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<emphasis bold="true" box="[199,399,366,393]" pageId="25" pageNumber="26">Locality and age</emphasis>
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.
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<materialsCitation county="Green River" location="De Beque" municipality="River Formation" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Colorado">
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<collectingCounty box="[414,553,367,393]" pageId="25" pageNumber="26">
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Green
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<collectingMunicipality box="[491,553,367,393]" pageId="25" pageNumber="26">River</collectingMunicipality>
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</collectingCounty>
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Formation, about eight miles south of
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<location LSID="urn:lsid:plazi:treatment:A148D509FFB3FFF4FF6D69E7C979FE64:2C3E32C4FFB2FFF4FC1F6A3ACC20FE40" box="[997,1112,367,393]" county="Green River" municipality="River Formation" name="De Beque" pageId="25" pageNumber="26" stateProvince="Colorado">De Beque</location>
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,
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<collectingRegion box="[1125,1229,367,393]" country="United States of America" name="Colorado" pageId="25" pageNumber="26">Colorado</collectingRegion>
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(
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<bibRefCitation author="Cockerell, T. D. A." box="[1245,1423,366,393]" pageId="25" pageNumber="26" pagination="1 - 15" refId="ref17014" refString="Cockerell, T. D. A. (1925) Fossil insects in the United States National Museum. Proceedings of the United States National Museum, 64, 1 - 15. [https: // www. biodiversitylibrary. org / item / 32410 # page / 7 / mode / 1 up]" type="journal article" year="1925">Cockerell 1925</bibRefCitation>
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||
), Ypresian
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||
</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |