165 lines
17 KiB
XML
165 lines
17 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFB1FFF6FF6D6E76CB6CFE88" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichiosomites obliviosus Brues, 1908 Brues 1908" docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="27" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894892" ID-Zenodo-Dep="7894892" LSID="urn:lsid:plazi:treatment:A148D509FFB1FFF6FF6D6E76CB6CFE88" httpUri="http://treatment.plazi.org/id/A148D509FFB1FFF6FF6D6E76CB6CFE88" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<subSubSection box="[151,612,1315,1342]" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph blockId="26.[151,612,1315,1378]" box="[151,612,1315,1342]" pageId="26" pageNumber="27">
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<heading bold="true" box="[151,612,1315,1342]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<taxonomicName authority="Brues, 1908" authorityName="Brues, 1908 Brues" authorityYear="1908" box="[151,612,1315,1342]" class="Insecta" family="Cimbicidae" genus="Trichiosomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obliviosus">
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<emphasis bold="true" box="[151,612,1315,1342]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[151,458,1315,1342]" italics="true" pageId="26" pageNumber="27">Trichiosomites obliviosus</emphasis>
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<bibRefCitation author="Brues, C. T." box="[465,612,1315,1341]" pageId="26" pageNumber="27" pagination="259 - 276" refId="ref16853" refString="Brues, C. T. (1908) New Phytophagous Hymenoptera from the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College, 51, 259 - 276. https: // doi. org / 10.5962 / bhl. title. 19923" type="journal article" year="1908">Brues, 1908</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,291,1351,1378]" pageId="26" pageNumber="27" type="description">
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<paragraph blockId="26.[151,612,1315,1378]" box="[151,291,1351,1378]" pageId="26" pageNumber="27">
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<figureCitation box="[151,287,1351,1378]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStartId-0="25.[151,250,1608,1632]" captionStartId-1="26.[151,250,1173,1197]" captionTargetBox-0="[151,1431,741,1577]" captionTargetBox-1="[151,1427,181,1138]" captionTargetId-0="figure-119@25.[151,1436,741,1585]" captionTargetId-1="figure-20@26.[151,1436,181,1149]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 24. Trichiosomites obliviosus Brues holotype, MCZ-2036, Abiinae, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing. Scale bars A–B, 5 mm; C–D, 3 mm." captionText-1="FIGURE 25. Trichiosomites obliviosus Brues, UCM-4570, Abiinae, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing. Scale bars A–B, 5 mm; C–D, 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7894929" figureDoi-1="http://doi.org/10.5281/zenodo.7894931" httpUri-0="https://zenodo.org/record/7894929/files/figure.png" httpUri-1="https://zenodo.org/record/7894931/files/figure.png" pageId="26" pageNumber="27">Figs. 24–25</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,686,1420,1444]" pageId="26" pageNumber="27" type="reference_group">
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<paragraph blockId="26.[151,686,1420,1444]" box="[151,686,1420,1444]" pageId="26" pageNumber="27">
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<taxonomicName authority="Brues, 1908 p. 260" authorityName="Brues, 1908 Brues" authorityPageNumber="260" authorityYear="1908" box="[151,610,1420,1444]" class="Insecta" family="Cimbicidae" genus="Trichiosomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obliviosus">
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<emphasis box="[151,410,1420,1444]" italics="true" pageId="26" pageNumber="27">Trichiosomites obliviosus</emphasis>
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<bibRefCitation author="Brues, C. T." box="[416,610,1420,1444]" pageId="26" pageNumber="27" pagination="259 - 276" refId="ref16853" refString="Brues, C. T. (1908) New Phytophagous Hymenoptera from the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College, 51, 259 - 276. https: // doi. org / 10.5962 / bhl. title. 19923" type="journal article" year="1908">Brues, 1908 p. 260</bibRefCitation>
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</taxonomicName>
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,
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<figureCitation box="[621,681,1420,1444]" captionStart="FIGURE 1" captionStartId="1.[151,250,1979,2003]" captionTargetBox="[159,1427,1145,1945]" captionTargetId="figure-357@1.[151,1436,1137,1956]" captionTargetPageId="1" captionText="FIGURE 1. Forewing and hind wing of Corynis crassicornis (Rossi) (Corynidinae) with veins (roman font) and cells (italic font) labelled. Drawn from Vilhelmsen (2019) figure 15B. Scale = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894832" httpUri="https://zenodo.org/record/7894832/files/figure.png" pageId="26" pageNumber="27">Fig. 1</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="materials_examined">
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<paragraph blockId="26.[151,1437,1490,2020]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[151,267,1490,1516]" pageId="26" pageNumber="27">
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<typeStatus box="[151,262,1490,1516]" pageId="26" pageNumber="27">Holotype</typeStatus>
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.
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</emphasis>
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<typeStatus box="[275,379,1490,1516]" pageId="26" pageNumber="27">Holotype</typeStatus>
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<collectionCode box="[388,449,1490,1516]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="26" pageNumber="27" type="Museum">MCZ</collectionCode>
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2036,
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<bibRefCitation author="Brues, C. T." box="[529,681,1490,1516]" pageId="26" pageNumber="27" pagination="259 - 276" refId="ref16853" refString="Brues, C. T. (1908) New Phytophagous Hymenoptera from the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College, 51, 259 - 276. https: // doi. org / 10.5962 / bhl. title. 19923" type="journal article" year="1908">Brues (1908)</bibRefCitation>
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. A rather complete female with head present, but positioned under thorax (not missing as
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<bibRefCitation author="Brues, C. T." box="[406,535,1526,1552]" pageId="26" pageNumber="27" pagination="259 - 276" refId="ref16853" refString="Brues, C. T. (1908) New Phytophagous Hymenoptera from the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College, 51, 259 - 276. https: // doi. org / 10.5962 / bhl. title. 19923" type="journal article" year="1908">Brues 1908</bibRefCitation>
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presumed).
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,477,1562,1588]" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph blockId="26.[151,1437,1490,2020]" box="[199,477,1562,1588]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[199,322,1562,1588]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
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<paragraph blockId="26.[151,1437,1490,2020]" lastBlockId="27.[151,1437,151,321]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[199,345,1598,1624]" pageId="26" pageNumber="27">Description.</emphasis>
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<typeStatus box="[353,457,1598,1624]" pageId="26" pageNumber="27">Holotype</typeStatus>
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MCZ 2036 (
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<figureCitation box="[608,692,1598,1624]" captionStart="FIGURE 24" captionStartId="25.[151,250,1608,1632]" captionTargetBox="[151,1431,741,1577]" captionTargetId="figure-119@25.[151,1436,741,1585]" captionTargetPageId="25" captionText="FIGURE 24. Trichiosomites obliviosus Brues holotype, MCZ-2036, Abiinae, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing. Scale bars A–B, 5 mm; C–D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7894929" httpUri="https://zenodo.org/record/7894929/files/figure.png" pageId="26" pageNumber="27">Fig. 24</figureCitation>
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), female. Body: head length
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<quantity box="[1022,1110,1598,1625]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="0.5">0.5 mm</quantity>
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or slightly more, width
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="26" pageNumber="27" unit="mm" value="0.87">0.87 mm</quantity>
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; thorax length
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<emphasis box="[367,393,1635,1660]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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.
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<quantity box="[409,498,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="mm" value="2.5">2.5 mm</quantity>
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; abdomen length
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<emphasis box="[699,725,1635,1660]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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.
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<quantity box="[740,809,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="5.0">5 mm</quantity>
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; body colouration unknown by preservation. Forewing: length
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<emphasis box="[231,257,1671,1696]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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. 6.0 mm, estimated maximum width
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<emphasis box="[693,719,1671,1696]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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. 2.0 mm, length/width:
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<emphasis box="[1001,1027,1671,1696]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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. 3.0. UCM 4570 (
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<figureCitation box="[1244,1330,1670,1697]" captionStart="FIGURE 25" captionStartId="26.[151,250,1173,1197]" captionTargetBox="[151,1427,181,1138]" captionTargetId="figure-20@26.[151,1436,181,1149]" captionTargetPageId="26" captionText="FIGURE 25. Trichiosomites obliviosus Brues, UCM-4570, Abiinae, from the Florissant Formation:A, photograph; B, drawing; C, drawing of right forewing; D, drawing of left forewing. Scale bars A–B, 5 mm; C–D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7894931" httpUri="https://zenodo.org/record/7894931/files/figure.png" pageId="26" pageNumber="27">Fig. 25</figureCitation>
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), female. Body: length
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<emphasis box="[303,329,1707,1732]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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.
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<quantity box="[344,433,1706,1732]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="26" pageNumber="27" unit="mm" value="9.2">9.2 mm</quantity>
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; head, antenna dark; thorax dark, wings including pterostigma light; abdomen light. Head difficult to interpret as preserved; moderately transverse, occiput long, three distinct ocelli in low triangle, compound eyes not preserved, sole funicular segment thin, slightly more than half length of club; club expands gradually over three segments (“drop-like”). Thoracic dorsum only partially visible as preserved, median longitudinal mesonotal sulcus and notauli present; mesopleuron with no mesopleural sulcus visible. Forewing: membrane hyaline, length
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<emphasis box="[151,177,1887,1912]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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.
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<quantity box="[193,284,1886,1913]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="26" pageNumber="27" unit="mm" value="6.5">6.5 mm</quantity>
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; maximum width estimated
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<emphasis box="[607,633,1887,1912]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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.
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<quantity box="[649,739,1886,1913]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="mm" value="2.5">2.5 mm</quantity>
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, length/width:
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<emphasis box="[910,936,1887,1912]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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. 2.6; pterostigma wide,
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<emphasis box="[1214,1240,1887,1912]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
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. as cell 2r; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r; 2r-rs longer than Rs between 2r-m, 2r-rs, joining Rs near middle of cell 3rm; 2r-rm straight, short, joins M at 2m-cu; 1-M curved; cell 1mcu with M, 1m-cu sub-parallel; 3r-m straight; cell 2mcu with M slightly longer than 1m-cu; 2m-cu straight; cu-a postfurcal; anal cell separated by 1A fusion with 2A+3A (no crossvein) for length of about a third length of cell 2a. Abdomen: long, wide; ovipositor thick, not exceeding abdomen apex.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="materials_examined">
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<paragraph blockId="27.[151,1437,151,321]" pageId="27" pageNumber="28">
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<emphasis bold="true" box="[199,310,222,248]" pageId="27" pageNumber="28">Material.</emphasis>
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<materialsCitation collectionCode="UCM" pageId="27" pageNumber="28" specimenCode="UCM 4570" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[319,423,223,249]" pageId="27" pageNumber="28">Holotype</typeStatus>
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and
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<specimenCode box="[480,611,222,249]" collectionCode="UCM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="27" pageNumber="28" type="University or college">UCM 4570</specimenCode>
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, a rather complete specimen missing only the legs and much of the hind wings.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="27.[151,1437,151,321]" box="[199,788,294,321]" pageId="27" pageNumber="28">
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<emphasis bold="true" box="[199,404,294,321]" pageId="27" pageNumber="28">Locality and age.</emphasis>
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Florissant Formation, Priabonian.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |