160 lines
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160 lines
15 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFADFFEAFF6D6AC7CB13FDF4" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Allenbycimbex morrisae Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="7" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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<mods:number>1</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894850" ID-Zenodo-Dep="7894850" LSID="urn:lsid:plazi:treatment:A148D509FFADFFEAFF6D6AC7CB13FDF4" httpUri="http://treatment.plazi.org/id/A148D509FFADFFEAFF6D6AC7CB13FDF4" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<paragraph blockId="6.[151,1198,402,501]" box="[151,519,402,429]" pageId="6" pageNumber="7">
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,446,402,429]" class="Insecta" family="Cimbicidae" genus="Allenbycimbex" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="morrisae" status="n. sp.">
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<emphasis bold="true" box="[151,446,402,429]" italics="true" pageId="6" pageNumber="7">Allenbycimbex morrisae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[453,519,403,429]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="6.[151,1198,402,501]" box="[151,1198,438,465]" pageId="6" pageNumber="7">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,223,475,501]" pageId="6" pageNumber="7" type="description">
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<paragraph blockId="6.[151,1198,402,501]" box="[151,223,475,501]" pageId="6" pageNumber="7">
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<figureCitation box="[151,220,475,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1327,1351]" captionTargetBox="[155,1430,525,1300]" captionTargetId="figure-85@6.[151,1436,525,1304]" captionTargetPageId="6" captionText="FIGURE 4. Allenbycimbex morrisae, holotype RBCM.EH2007.002.0001: A, photograph; B, drawing; C, drawing of right wing; D, drawing of left wing. Scale bar for A and B is 5 mm (as the fossil has been stretched by geologic shear forces, C and D are altered to the presumed unstretched configuration)." figureDoi="http://doi.org/10.5281/zenodo.7894842" httpUri="https://zenodo.org/record/7894842/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7894842" ID-Zenodo-Dep="7894842" httpUri="https://zenodo.org/record/7894842/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1327,1351]" targetBox="[155,1430,525,1300]" targetPageId="6" targetType="figure">
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<paragraph blockId="6.[151,1436,1327,1423]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[151,278,1327,1351]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
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<taxonomicName authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[288,538,1327,1351]" class="Insecta" family="Cimbicidae" genus="Allenbycimbex" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="morrisae">
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<emphasis box="[288,538,1327,1351]" italics="true" pageId="6" pageNumber="7">Allenbycimbex morrisae</emphasis>
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</taxonomicName>
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, holotype RBCM.EH2007.002.0001: A, photograph; B, drawing; C, drawing of right wing; D, drawing of left wing. Scale bar for A and B is 5 mm (as the fossil has been stretched by geologic shear forces, C and D are altered to the presumed unstretched configuration).
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</paragraph>
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</caption>
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<subSubSection pageId="6" pageNumber="7" type="materials_examined">
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<paragraph blockId="6.[151,1437,1469,2036]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[151,262,1469,1495]" pageId="6" pageNumber="7">Material.</emphasis>
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<materialsCitation collectionCode="RBCM" collectorName="Mostly & Muriel Morris & Allenby Formation" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[272,376,1469,1495]" pageId="6" pageNumber="7">Holotype</typeStatus>
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<collectionCode box="[387,470,1469,1495]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="6" pageNumber="7" type="Museum">RBCM</collectionCode>
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.EH2007.002.0001a, b (part, counterpart).
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<collectorName box="[957,1036,1469,1495]" pageId="6" pageNumber="7">Mostly</collectorName>
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complete but somewhat deformed by geological shear forces, lacking head, fore and mid legs, hind tarsi, apical half of left wings, and apical half of abdomen; hind wings fragmentary and poorly preserved; wings lack visible colouration either originally or by preservation; collected by
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<collectorName box="[447,610,1577,1603]" pageId="6" pageNumber="7">Muriel Morris</collectorName>
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,
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<collectorName box="[619,837,1577,1603]" pageId="6" pageNumber="7">Allenby Formation</collectorName>
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; housed in the collections of the
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<collectionCode box="[1204,1289,1577,1603]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="6" pageNumber="7" type="Museum">RBCM</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,479,1613,1640]" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph blockId="6.[151,1437,1469,2036]" box="[199,479,1613,1640]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[199,322,1613,1639]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
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<paragraph blockId="6.[151,1437,1469,2036]" lastBlockId="7.[151,1437,151,573]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[199,345,1649,1675]" pageId="6" pageNumber="7">Description.</emphasis>
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Length as preserved (stretched, see below; incomplete head and abdomen)
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<quantity box="[1214,1304,1649,1676]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="6" pageNumber="7" unit="mm" value="8.9">8.9 mm</quantity>
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, apparently dark (preserved parts: hind tibiae, metasoma possibly paler). Temples somewhat inflated. Antennal club if correctly identified apparently short with truncated apex. Lateral pronotal lobes very wide, notauli apparently widely separated before scutellum (interpretation tentative). Hind coxae large, contiguous; femora not inflated, almost parallel-sided, shorter than abdomen wide; tibia thinner and much longer than femur. Forewing: length
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<emphasis box="[1148,1174,1794,1819]" italics="true" pageId="6" pageNumber="7">ca</emphasis>
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.
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<quantity box="[1189,1290,1793,1820]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="7" unit="mm" value="13.5">13.5 mm</quantity>
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as preserved (distorted), the same in life if it was compressed in width,
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<emphasis box="[804,830,1830,1855]" italics="true" pageId="6" pageNumber="7">ca</emphasis>
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.
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<quantity box="[845,924,1829,1856]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="mm" value="10.0">10 mm</quantity>
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in life if it was stretched in length; maximum width:
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<emphasis box="[232,258,1866,1891]" italics="true" pageId="6" pageNumber="7">ca</emphasis>
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.
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<quantity box="[275,363,1865,1892]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="3.5">3.5 mm</quantity>
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as preserved, about the same in life if it was stretched in length or
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<emphasis box="[1146,1172,1866,1891]" italics="true" pageId="6" pageNumber="7">ca</emphasis>
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. 4.25 if compressed in width; length/width: probably
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<emphasis box="[500,526,1902,1927]" italics="true" pageId="6" pageNumber="7">ca</emphasis>
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. 3.0; no membrane colouration detected (could be an artefact of preservation); pterostigma broad, distinctly wider than C-R basal to it; R+M longer than 1-M; 2r-rs not preserved; 2r-rm curved, joins Rs about same level as 2m-cu joins M, joins M behind 2m-cu for more than half length of 2r-m; 1-M almost straight; 1-M and 1m-cu diverging toward anterior; 3r-m distinctly bowed toward base; cell 2mcu with M shorter than 1m-cu; 2m-cu curved distinctly toward base; cu-a postfurcal; anal cell contracted for distance subequal to width (height) of 1a cell, with no crossvein; basal portion of 2A+3A sharply bent toward 1A, meets 1A at steep angle. Hind wing with m-cu not preserved, r-m and cu-a far distant (for much longer than in
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<taxonomicName authorityName="Nel" authorityYear="2004" box="[1186,1324,223,249]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis box="[1186,1324,223,249]" italics="true" pageId="7" pageNumber="8">Cenocimbex</emphasis>
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</taxonomicName>
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and more similar to that characteristic of
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[510,648,259,284]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus" status="n. gen.">
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<emphasis box="[510,648,259,284]" italics="true" pageId="7" pageNumber="8">Leptostigma</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[657,737,259,285]" pageId="7" pageNumber="8">
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<taxonomicNameLabel box="[657,737,259,285]" pageId="7" pageNumber="8" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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); other details obscure. Abdomen with 3 segments preserved (apparently sterna 2–4) each narrower than thorax and some 3.5× as wide as long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="discussion">
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<paragraph blockId="7.[151,1437,151,573]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[199,315,331,357]" pageId="7" pageNumber="8">Remarks.</emphasis>
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The fossil was distorted by geological shear forces during diagenesis, either stretching it along the plane of the length of its right forewing or compressing it 90° to that. These possibilities cannot be distinguished. As the forewings are about 90° to each other, we estimated its life shape by both graphically compressing the image along the length of the right forewing and by stretching it’s width so that elements of its venation are of equal length to those of its left wing.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="etymology">
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<paragraph blockId="7.[151,1437,151,573]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[199,334,511,537]" pageId="7" pageNumber="8">Etymology.</emphasis>
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The specific epithet is a patronymic formed from the surname of Muriel Morris, recognizing her contribution as collector and donor of the only known specimen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |