279 lines
35 KiB
XML
279 lines
35 KiB
XML
<document id="9536074AB2B752CC427B0AB7A1183584" ID-CLB-Dataset="54646" ID-DOI="10.11646/zootaxa.5278.1.1" ID-GBIF-Dataset="1528757a-5499-40b9-a728-c5bf0c770b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFA7FFE3FF6D6A3ACCC1FDF4" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma longiclava Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docUuid="B8F5BDEE-1B98-4F5F-94FB-646D070B6153" docUuidSource="ZooBank" docVersion="9" lastPageNumber="15" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="13" updateTime="1728574925831" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="37C55A1A9286CC64EDA879C98B90F42E">Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:name id="0ED0F93D4DF937CC1A9A4EC27A3CEFCC" type="personal">
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<mods:namePart id="E6F05F1C520CA65B6CFE507BCE06ECF9">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="6EE2E3AC85EC556034281BAAA29769E6" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation id="1ADAD2B01CF375E7A746A5DFD9D49F74">Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier id="E8339829EE833C0AB955C3B48248D2CD" type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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<mods:namePart id="6E52DAE94DA2D3AE15693CC078A4B5F6">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="0C5F0DBCEF70F4CF3F3778AB3C553F46">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:title id="ADB42971B72318AAC8D517AF0D368E2A">Zootaxa</mods:title>
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<mods:date id="C61948D5C0B9B2F00559D90014D51000">2023</mods:date>
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<mods:number id="05846A4431E43225B38F36954DD4D6BE">2023-05-04</mods:number>
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<mods:number id="388FB32A0BAEE1FEEC3745E9C8F895A0">5278</mods:number>
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<treatment id="A148D509FFA7FFE3FF6D6A3ACCC1FDF4" ID-DOI="http://doi.org/10.5281/zenodo.7894862" ID-GBIF-Taxon="209583694" ID-Zenodo-Dep="7894862" LSID="urn:lsid:zoobank.org:act:B8F5BDEE-1B98-4F5F-94FB-646D070B6153" httpUri="http://treatment.plazi.org/id/A148D509FFA7FFE3FF6D6A3ACCC1FDF4" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="61FB3794FFA7FFE1FF6D6A3AC982FE40" box="[151,506,366,393]" pageId="12" pageNumber="13" type="nomenclature">
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<heading id="7216D373FFA7FFE1FF6D6A3AC982FE40" bold="true" box="[151,506,366,393]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<emphasis id="1B95B80DFFA7FFE1FF6D6A3AC982FE40" bold="true" box="[151,506,366,393]" pageId="12" pageNumber="13">
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<taxonomicName id="EEE11F9CFFA7FFE1FF6D6A3AC9C8FE40" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,432,366,393]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
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<emphasis id="1B95B80DFFA7FFE1FF6D6A3AC9C8FE40" bold="true" box="[151,432,366,393]" italics="true" pageId="12" pageNumber="13">Leptostigma longiclava</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00A60576FFA7FFE1FE426A3AC982FE40" box="[440,506,367,393]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="61FB3794FFA7FFE1FF6D6AC7C909FE18" pageId="12" pageNumber="13" type="description">
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<paragraph id="295E641FFFA7FFE1FF6D6AC7CCE2FE64" blockId="12.[151,1178,366,465]" box="[151,1178,402,429]" pageId="12" pageNumber="13">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="295E641FFFA7FFE1FF6D6AE2C909FE18" blockId="12.[151,1178,366,465]" box="[151,369,439,465]" pageId="12" pageNumber="13">
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<figureCitation id="B1DA789AFFA7FFE1FF6D6AE2C961FE18" box="[151,281,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="12" pageNumber="13">Figs. 3K–L</figureCitation>
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,
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<figureCitation id="B1DA789AFFA7FFE1FEDF6AE2C915FE18" box="[293,365,439,465]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="12.[151,250,1978,2002]" captionStartId-1="13.[151,250,1835,1859]" captionTargetBox-0="[167,1420,777,1941]" captionTargetBox-1="[165,1424,741,1799]" captionTargetId-0="figure-193@12.[167,1420,777,1954]" captionTargetId-1="figure-281@13.[159,1429,741,1811]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 11. Leptostigma longiclava holotype RBCM-EH2017.050.4085.001, A, photograph, B–E wing drawings: B, right forewing; C, left forewing; D, left hind wing; E, right hind wing. All to scale, 5 mm." captionText-1="FIGURE 12. F-848, tentative Leptostigma longiclava, A, photograph, B–C, drawings: B, right forewing; C, left forewing. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7894872" figureDoi-1="http://doi.org/10.5281/zenodo.7894878" httpUri-0="https://zenodo.org/record/7894872/files/figure.png" httpUri-1="https://zenodo.org/record/7894878/files/figure.png" pageId="12" pageNumber="13">11–12</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFA7FFE1FF6D6AAACDF9FDF4" pageId="12" pageNumber="13" type="materials_examined">
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<paragraph id="295E641FFFA7FFE1FF6D6AAACDF9FDF4" blockId="12.[151,1436,510,753]" pageId="12" pageNumber="13">
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<materialsCitation id="99896E42FFA7FFE1FF6D6AAACDF9FDF4" ID-GBIF-Occurrence="4100387313" collectionCode="RBCM" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
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<emphasis id="1B95B80DFFA7FFE1FF6D6AAAC973FDD0" bold="true" box="[151,267,511,537]" pageId="12" pageNumber="13">
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<typeStatus id="F65ADABDFFA7FFE1FF6D6AAAC97EFDD0" box="[151,262,511,537]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="4FF0FCDAFFA7FFE1FEEA6AABC91AFDD0" box="[272,354,510,537]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="12" pageNumber="13" type="Museum">RBCM</collectionCode>
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.EH2017.050.4085.0001a, b (part, counterpart) (
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<figureCitation id="B1DA789AFFA7FFE1FC8D6AAACB80FDD0" box="[887,1016,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="12" pageNumber="13">Figs. 3K–L</figureCitation>
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,
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<figureCitation id="B1DA789AFFA7FFE1FBF86AAACC67FDD0" box="[1026,1055,511,537]" captionStart="FIGURE 11" captionStartId="12.[151,250,1978,2002]" captionTargetBox="[167,1420,777,1941]" captionTargetId="figure-193@12.[167,1420,777,1954]" captionTargetPageId="12" captionText="FIGURE 11. Leptostigma longiclava holotype RBCM-EH2017.050.4085.001, A, photograph, B–E wing drawings: B, right forewing; C, left forewing; D, left hind wing; E, right hind wing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894872" httpUri="https://zenodo.org/record/7894872/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
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). Mostly complete in dorsal aspect but missing much of the abdomen; collected at McAbee by John Leahy; housed in the collections of the
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<collectionCode id="4FF0FCDAFFA7FFE1FAD36976CD06FDF4" box="[1321,1406,547,573]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="12" pageNumber="13" type="Museum">RBCM</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFA7FFE1FF3D6913CC11FD38" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="295E641FFFA7FFE1FF3D6913CC11FD38" blockId="12.[151,1436,510,753]" pageId="12" pageNumber="13">
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<emphasis id="1B95B80DFFA7FFE1FF3D6913C942FDA9" bold="true" box="[199,314,582,608]" pageId="12" pageNumber="13">Diagnosis</emphasis>
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. Distinguished from other species of
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<taxonomicName id="EEE11F9CFFA7FFE1FD036912CBFBFDA9" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[761,899,583,608]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="1B95B80DFFA7FFE1FD036912CBFBFDA9" box="[761,899,583,608]" italics="true" pageId="12" pageNumber="13">Leptostigma</emphasis>
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</taxonomicName>
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by a combination of antenna with club long, darkened, penultimate segment longer than ultimate, flagellar base formed by long, dark, apparently single (entire) segment. Forewing lightly, evenly infuscate throughout (no fascia or macula). Tentatively diagnostic: R+M shorter than 1-M, 2r-rs joins cell 3rm distinctly behind its middle, 2r-m joins M far behind 2m-cu, anal cells meet at crossvein or are at most minimally separated by a very short fusion of 1A with 2A+3A.
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</paragraph>
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</subSubSection>
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<caption id="7D9E3497FFA7FFE1FF6D6CEFCB96F83C" ID-DOI="http://doi.org/10.5281/zenodo.7894872" ID-Zenodo-Dep="7894872" httpUri="https://zenodo.org/record/7894872/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1978,2002]" targetBox="[167,1420,777,1941]" targetPageId="12" targetType="figure">
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<paragraph id="295E641FFFA7FFE1FF6D6CEFCB96F83C" blockId="12.[151,1436,1978,2038]" pageId="12" pageNumber="13">
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<emphasis id="1B95B80DFFA7FFE1FF6D6CEFC959F81B" bold="true" box="[151,289,1978,2002]" pageId="12" pageNumber="13">FIGURE 11.</emphasis>
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<taxonomicName id="EEE11F9CFFA7FFE1FED06CEECA63F81B" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[298,539,1978,2002]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="longiclava">
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<emphasis id="1B95B80DFFA7FFE1FED06CEECA63F81B" box="[298,539,1978,2002]" italics="true" pageId="12" pageNumber="13">Leptostigma longiclava</emphasis>
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</taxonomicName>
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holotype RBCM-EH2017.050.4085.001, A, photograph, B–E wing drawings: B, right forewing; C, left forewing; D, left hind wing; E, right hind wing. All to scale, 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="61FB3794FFA6FFE0FF3D6BC3CC89FD04" pageId="13" pageNumber="14" type="description">
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<paragraph id="295E641FFFA6FFE0FF3D6BC3CC89FD04" blockId="13.[151,1437,150,717]" pageId="13" pageNumber="14">
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<emphasis id="1B95B80DFFA6FFE0FF3D6BC3C921FF79" bold="true" box="[199,345,150,176]" pageId="13" pageNumber="14">Description.</emphasis>
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<typeStatus id="F65ADABDFFA6FFE0FE9E6BC2C9A8FF78" box="[356,464,151,177]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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. Body. Length not known by preservation, tentative specimen F-848
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<emphasis id="1B95B80DFFA6FFE0FB016BC2CD6DFF79" box="[1275,1301,151,176]" italics="true" pageId="13" pageNumber="14">ca</emphasis>
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.
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<quantity id="EE19C9FAFFA6FFE0FAD26BC2CD02FF78" box="[1320,1402,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="14" unit="mm" value="14.0">14 mm</quantity>
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as preserved, likely
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<quantity id="EE19C9FAFFA6FFE0FEA66BEFC9AFFF1C" box="[348,471,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="13" pageNumber="14" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
|
||
in life. Head, thorax, scape, pedicel, basal flagellomere, femora dark; antennal club, pterostigma, tibiae, metasoma darkened as preserved; flagellar segment 2 rather pale, wing membrane lightly infuscate throughout. Head as preserved difficult to interpret. Antenna apparently 7-segmented, with scape, particularly pedicel very short, first flagellomere longest (about one third of flagellum), those following gradually becoming shorter and, except apical one, wider, apical one half-oval. Thorax with distinct median longitudinal mesonotal sulcus, notauli complete, meeting well before transverse scutellum; metascutellum narrow (narrower than normal in
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FF6D6AC6C92AFE64" authorityName="Benson" authorityYear="1938" box="[151,338,403,429]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
|
||
, Corynidinae), cenchri not apparent. Forewing length 14 (
|
||
<typeStatus id="F65ADABDFFA6FFE0FC1A6AC6CC3EFE64" box="[992,1094,403,429]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
)–
|
||
<quantity id="EE19C9FAFFA6FFE0FBA76AC6CCD4FE64" box="[1117,1196,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="16.0">16 mm</quantity>
|
||
, maximum width 4.5 (
|
||
<typeStatus id="F65ADABDFFA6FFE0FF656AE2C97BFE18" box="[159,259,439,465]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
)–
|
||
<quantity id="EE19C9FAFFA6FFE0FEE16AE3C924FE18" box="[283,348,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="5.0">5 mm</quantity>
|
||
, length/width 3.1–3.2; pterostigma narrow, similar to width of C&R basal of pterostigma; R+M as long as 1-M in left wing, shorter in right; 2r-rs shorter than Rs between 2rm, 2r-rs, entering Rs behind middle of cell 3rm; 2r-m distinctly curved, joins Rs about same level as 2m-cu joining M, joins M behind 2m-cu for more than half length of 2r-m; 3r-m almost straight; 1-M curved; 1-M and 1m-cu slightly diverging toward wing anterior margin; cell 2mcu with M shorter than 1m-cu; 2m-cu slightly curved or angular mid-length; cu-a slightly postfurcal; anal cells minimally separated by a very short fusion of 1A with 2A+3A, no crossvein between them (in F-848, anal cell region not preserved in
|
||
<typeStatus id="F65ADABDFFA6FFE0FE5969DACA72FD60" box="[419,522,655,681]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
). Hind wing mostly indistinctly preserved, crossveins 3r-m, m-cu present, distant for less than length of 2r-m. Abdominal tergum 1 possibly split medially, with hind margin straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFA6FFE0FF6D6C7EC94AF8AF" ID-DOI="http://doi.org/10.5281/zenodo.7894878" ID-Zenodo-Dep="7894878" httpUri="https://zenodo.org/record/7894878/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1835,1859]" targetBox="[165,1424,741,1799]" targetPageId="13" targetType="figure">
|
||
<paragraph id="295E641FFFA6FFE0FF6D6C7EC94AF8AF" blockId="13.[151,1436,1835,1895]" pageId="13" pageNumber="14">
|
||
<emphasis id="1B95B80DFFA6FFE0FF6D6C7EC958F88A" bold="true" box="[151,288,1835,1859]" pageId="13" pageNumber="14">FIGURE 12.</emphasis>
|
||
F-848, tentative
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FE356C79CAC6F88A" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[463,702,1835,1859]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longiclava">
|
||
<emphasis id="1B95B80DFFA6FFE0FE356C79CAC6F88A" box="[463,702,1835,1859]" italics="true" pageId="13" pageNumber="14">Leptostigma longiclava</emphasis>
|
||
</taxonomicName>
|
||
, A, photograph, B–C, drawings: B, right forewing; C, left forewing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFA6FFE3FF3D6CC5C916FE40" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="295E641FFFA6FFE3FF3D6CC5C916FE40" blockId="13.[151,1436,1936,2034]" lastBlockId="14.[151,1437,151,573]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis id="1B95B80DFFA6FFE0FF3D6CC5C943F863" bold="true" box="[199,315,1936,1962]" pageId="13" pageNumber="14">Remarks.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FE226CC4CA17F863" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[472,623,1936,1962]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FE226CC4CA17F863" box="[472,623,1936,1962]" italics="true" pageId="13" pageNumber="14">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FD8D6CC5CACFF863" bold="true" box="[631,695,1936,1962]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FD8D6CC5CACFF863" box="[631,695,1936,1962]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FD0A6CC4CBD4F863" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[752,940,1936,1962]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FD0A6CC4CBD4F863" box="[752,940,1936,1962]" italics="true" pageId="13" pageNumber="14">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FC4E6CC5CB8CF863" bold="true" box="[948,1012,1936,1962]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FC4E6CC5CB8CF863" box="[948,1012,1936,1962]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its club long and darkened [short, wide, and dark in
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FEA06CE0C995F807" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[346,493,1972,1998]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FEA06CE0C995F807" box="[346,493,1972,1998]" italics="true" pageId="13" pageNumber="14">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FE096CE1CA48F807" bold="true" box="[499,560,1972,1998]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FE096CE1CA48F807" box="[499,560,1972,1998]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; long and pale in
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FD0B6CE0CBD2F807" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[753,938,1972,1998]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FD0B6CE0CBD2F807" box="[753,938,1972,1998]" italics="true" pageId="13" pageNumber="14">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FC556CE1CB94F807" bold="true" box="[943,1004,1972,1998]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FC556CE1CB94F807" box="[943,1004,1972,1998]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
]; differs venationally, tentatively, from both
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FF2A6C8CC91DF83B" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[208,357,2008,2034]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FF2A6C8CC91DF83B" box="[208,357,2008,2034]" italics="true" pageId="13" pageNumber="14">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FE966C8DC9D2F83B" bold="true" box="[364,426,2008,2034]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FE966C8DC9D2F83B" box="[364,426,2008,2034]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EEE11F9CFFA6FFE0FE1A6C8CCAE2F83B" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[480,666,2008,2034]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA6FFE0FE1A6C8CCAE2F83B" box="[480,666,2008,2034]" italics="true" pageId="13" pageNumber="14">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA6FFE0FD5B6C8DCAA7F83B" bold="true" box="[673,735,2008,2034]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="00A60576FFA6FFE0FD5B6C8DCAA7F83B" box="[673,735,2008,2034]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in 2r-rs joining cell 3rm distinctly distal to its middle [vs. at or before middle]. Differs from
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FE276BC2CACAFF78" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[477,690,151,177]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FE276BC2CACAFF78" box="[477,690,151,177]" italics="true" pageId="14" pageNumber="15">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FD436BC2CA8FFF78" bold="true" box="[697,759,151,177]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FD436BC2CA8FFF78" box="[697,759,151,177]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
that has a generally similar antenna (club long, darkened) in that its flagellar base is formed by one long, dark segment [vs. two pale, relatively short ones], and by its penultimate club segment longer than ultimate [vs. shorter].
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FD4D6B8ACB34FF30" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[695,844,223,249]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FD4D6B8ACB34FF30" box="[695,844,223,249]" italics="true" pageId="14" pageNumber="15">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FCAE6B8ACBECFF30" bold="true" box="[852,916,223,249]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FCAE6B8ACBECFF30" box="[852,916,223,249]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also differs from
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FB9A6B8ACD4FFF30" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1120,1335,223,249]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FB9A6B8ACD4FFF30" box="[1120,1335,223,249]" italics="true" pageId="14" pageNumber="15">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FAC46B8ACD06FF30" bold="true" box="[1342,1406,223,249]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FAC46B8ACD06FF30" box="[1342,1406,223,249]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having anal cells at most minimally separated [vs. distant for more than half width/height of cell 2a]. Differs from
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FA7C6A56C97EFE89" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="proxivena" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FA7C6A56C97EFE89" italics="true" pageId="14" pageNumber="15">L. proxivena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FEF46A72C935FE88" bold="true" box="[270,333,295,321]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FEF46A72C935FE88" box="[270,333,295,321]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FEA66A72C99FFE88" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[348,487,295,321]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fasciatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FEA66A72C99FFE88" box="[348,487,295,321]" italics="true" pageId="14" pageNumber="15">L. fasciatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FE146A72CA55FE88" bold="true" box="[494,557,295,321]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FE146A72CA55FE88" box="[494,557,295,321]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EEE11F9CFFA5FFE3FD966A72CB75FE88" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[620,781,295,321]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaemacula" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA5FFE3FD966A72CB75FE88" box="[620,781,295,321]" italics="true" pageId="14" pageNumber="15">L. alaemacula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA5FFE3FCEE6A72CB2BFE88" bold="true" box="[788,851,295,321]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="00A60576FFA5FFE3FCEE6A72CB2BFE88" box="[788,851,295,321]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its forewing lightly, evenly infuscate throughout [vs. with dark transverse fascia or macula at or basal to level of pterostigmal base] and 2r-m entering M far behind, not close to 2m-cu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFA5FFE3FF3D6AC7CACCFE18" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="295E641FFFA5FFE3FF3D6AC7CACCFE18" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="15">
|
||
<emphasis id="1B95B80DFFA5FFE3FF3D6AC7C94EFE65" bold="true" box="[199,310,402,428]" pageId="14" pageNumber="15">Material.</emphasis>
|
||
<materialsCitation id="99896E42FFA5FFE3FEB96AC6CACCFE18" ID-GBIF-Occurrence="4100387320" collectionCode="TRU" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="F65ADABDFFA5FFE3FEB96AC6C9D3FE64" box="[323,427,403,429]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
|
||
and tentatively (based on venational characters) specimen F-848 (
|
||
<figureCitation id="B1DA789AFFA5FFE3FB3D6AC6CD67FE64" box="[1223,1311,403,429]" captionStart="FIGURE 12" captionStartId="13.[151,250,1835,1859]" captionTargetBox="[165,1424,741,1799]" captionTargetId="figure-281@13.[159,1429,741,1811]" captionTargetPageId="13" captionText="FIGURE 12. F-848, tentative Leptostigma longiclava, A, photograph, B–C, drawings: B, right forewing; C, left forewing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894878" httpUri="https://zenodo.org/record/7894878/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
|
||
), collector unknown at McAbee, in the collections of
|
||
<collectionCode id="4FF0FCDAFFA5FFE3FD8E6AE2CAC8FE18" box="[628,688,439,465]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="14" pageNumber="15" type="Herbarium">TRU</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFA5FFE3FF3D6A8EC917FDD0" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="295E641FFFA5FFE3FF3D6A8EC917FDD0" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="15">
|
||
<emphasis id="1B95B80DFFA5FFE3FF3D6A8EC936FE3C" bold="true" box="[199,334,475,501]" pageId="14" pageNumber="15">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="1B95B80DFFA5FFE3FDF36A8ECA2CFE3C" box="[521,596,475,501]" italics="true" pageId="14" pageNumber="15">longus</emphasis>
|
||
, meaning long, and
|
||
<emphasis id="1B95B80DFFA5FFE3FCC76A8ECB02FE3C" box="[829,890,475,501]" italics="true" pageId="14" pageNumber="15">clava</emphasis>
|
||
, club, referring to the antennae. The epithet is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFA5FFE3FF3D6977CCC1FDF4" box="[199,1209,546,573]" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="295E641FFFA5FFE3FF3D6977CCC1FDF4" blockId="14.[151,1437,151,573]" box="[199,1209,546,573]" pageId="14" pageNumber="15">
|
||
<emphasis id="1B95B80DFFA5FFE3FF3D6977C9ECFDF4" bold="true" box="[199,404,546,573]" pageId="14" pageNumber="15">Locality and age.</emphasis>
|
||
McAbee,
|
||
<collectingRegion id="EB25AAFDFFA5FFE3FDF36976CAB5FDF4" box="[521,717,547,573]" country="Canada" name="British Columbia" pageId="14" pageNumber="15">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="51F6248FFFA5FFE3FD236976CB48FDF4" box="[729,816,547,573]" name="Canada" pageId="14" pageNumber="15">Canada</collectingCountry>
|
||
, Hoodoo Face beds; mid-Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |