297 lines
39 KiB
XML
297 lines
39 KiB
XML
<document id="EE61BF1045F5E1AD0A2F5A0EA25B2B9D" ID-CLB-Dataset="54646" ID-DOI="10.11646/zootaxa.5278.1.1" ID-GBIF-Dataset="1528757a-5499-40b9-a728-c5bf0c770b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFA5FFFDFF6D69DACCC1F831" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma longipallidum Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docUuid="EEEB4CA5-0EF0-4B45-985E-08D39CF2A996" docUuidSource="ZooBank" docVersion="9" lastPageNumber="17" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="15" updateTime="1728574925831" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="296ABC44CC2F93FAE9FAC0ED33F2E6BD">Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:namePart id="5D5269CD3BFCED4FD966627C6F870B57">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="75BE191CEFE43610CFFB4918C30ADF15">Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier id="93D41944E63C6D7FC5B0D7D2D25798BF" type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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<mods:namePart id="CE8957A7BE45712C3BC3B7F3573D383E">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="DF5017078E8605D0DFF93B23FCF1B6C6">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:title id="D79322E68DC7AF9DBB69B93D9C117645">Zootaxa</mods:title>
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<mods:date id="BC216D7B313FBCA06EA6708F1338AEE0">2023</mods:date>
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<mods:number id="8E589CFA40DE3BC7669DFAE59B953D43">2023-05-04</mods:number>
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<treatment id="A148D509FFA5FFFDFF6D69DACCC1F831" ID-DOI="http://doi.org/10.5281/zenodo.7894864" ID-GBIF-Taxon="209583695" ID-Zenodo-Dep="7894864" LSID="urn:lsid:zoobank.org:act:EEEB4CA5-0EF0-4B45-985E-08D39CF2A996" httpUri="http://treatment.plazi.org/id/A148D509FFA5FFFDFF6D69DACCC1F831" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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<subSubSection id="61FB3794FFA5FFE3FF6D69DACA5EFD60" box="[151,550,654,681]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="295E641FFFA5FFE3FF6D69DACA5EFD60" blockId="14.[151,1186,654,753]" box="[151,550,654,681]" pageId="14" pageNumber="15">
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<emphasis id="1B95B80DFFA5FFE3FF6D69DACA5EFD60" bold="true" box="[151,550,654,681]" pageId="14" pageNumber="15">
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<taxonomicName id="EEE11F9CFFA5FFE3FF6D69DAC9A5FD60" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,477,654,681]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
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<emphasis id="1B95B80DFFA5FFE3FF6D69DAC9A5FD60" bold="true" box="[151,477,654,681]" italics="true" pageId="14" pageNumber="15">Leptostigma longipallidum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00A60576FFA5FFE3FE1E69DACA5EFD60" box="[484,550,655,681]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="61FB3794FFA5FFE3FF6D69E7C90CFD38" pageId="14" pageNumber="15" type="description">
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<paragraph id="295E641FFFA5FFE3FF6D69E7CCDAFD04" blockId="14.[151,1186,654,753]" box="[151,1186,690,717]" pageId="14" pageNumber="15">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<uuid id="5D475ECAFFA5FFE3FD7969E7CCDAFD04" box="[643,1186,690,717]" pageId="14" pageNumber="15">EEEB4CA5-0EF0-4B45-985E-08D39CF2A996</uuid>
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<paragraph id="295E641FFFA5FFE3FF6D6982C90CFD38" blockId="14.[151,1186,654,753]" box="[151,372,726,753]" pageId="14" pageNumber="15">
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<figureCitation id="B1DA789AFFA5FFE3FF6D6982C962FD38" box="[151,282,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="14" pageNumber="15">Figs. 3C–D</figureCitation>
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,
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<figureCitation id="B1DA789AFFA5FFE3FEDC6983C917FD38" box="[294,367,726,753]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="14.[151,250,1956,1980]" captionStartId-1="15.[151,250,1982,2006]" captionStartId-2="16.[151,250,1877,1901]" captionTargetBox-0="[151,1436,777,1928]" captionTargetBox-1="[169,1418,417,1947]" captionTargetBox-2="[168,1420,1101,1813]" captionTargetId-0="figure-207@14.[151,1436,777,1933]" captionTargetId-1="figure-136@15.[169,1418,417,1958]" captionTargetId-2="figure-483@16.[168,1420,1101,1854]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURE 13. Leptostigma longipallidum, holotype GSC 142841, A, photograph, B–C, wing drawings: B, right forewing; C, left forewing. All to scale, 5 mm." captionText-1="FIGURE 14. F-771, tentative Leptostigma longipallidum, A, photograph, B–E, wing drawings: B, right forewing; C, left forewing; D, right hind wing; E. left wing. All to scale, 5 mm." captionText-2="FIGURE 15. F-1185 tentative Leptostigma longipallidum forewing: A, photograph; B, drawing. Scale is 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7894882" figureDoi-1="http://doi.org/10.5281/zenodo.7894886" figureDoi-2="http://doi.org/10.5281/zenodo.7894890" httpUri-0="https://zenodo.org/record/7894882/files/figure.png" httpUri-1="https://zenodo.org/record/7894886/files/figure.png" httpUri-2="https://zenodo.org/record/7894890/files/figure.png" pageId="14" pageNumber="15">13–15</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<caption id="7D9E3497FFA5FFE3FF6D6CF1C990F816" ID-DOI="http://doi.org/10.5281/zenodo.7894882" ID-Zenodo-Dep="7894882" httpUri="https://zenodo.org/record/7894882/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1956,1980]" targetBox="[151,1436,777,1928]" targetPageId="14" targetType="figure">
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<paragraph id="295E641FFFA5FFE3FF6D6CF1C990F816" blockId="14.[151,1436,1956,2016]" pageId="14" pageNumber="15">
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<emphasis id="1B95B80DFFA5FFE3FF6D6CF1C959F875" bold="true" box="[151,289,1956,1980]" pageId="14" pageNumber="15">FIGURE 13.</emphasis>
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<taxonomicName id="EEE11F9CFFA5FFE3FED26CF0CA43F875" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[296,571,1956,1980]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longipallidum">
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<emphasis id="1B95B80DFFA5FFE3FED26CF0CA43F875" box="[296,571,1956,1980]" italics="true" pageId="14" pageNumber="15">Leptostigma longipallidum</emphasis>
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</taxonomicName>
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, holotype GSC 142841, A, photograph, B–C, wing drawings: B, right forewing; C, left forewing. All to scale, 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="61FB3794FFA4FFE2FF6D6BC2CD2BFF30" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="295E641FFFA4FFE2FF6D6BC2CD2BFF30" blockId="15.[151,1437,151,393]" pageId="15" pageNumber="16">
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<materialsCitation id="99896E42FFA4FFE2FF6D6BC2CD2BFF30" ID-GBIF-Occurrence="4100387303" collectionCode="A, GSC" collectorName="McAbee & John Leahy" pageId="15" pageNumber="16" specimenCode="GSC 142841" specimenCount="1" typeStatus="holotype">
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<emphasis id="1B95B80DFFA4FFE2FF6D6BC2C973FF78" bold="true" box="[151,267,151,177]" pageId="15" pageNumber="16">
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<typeStatus id="F65ADABDFFA4FFE2FF6D6BC2C97EFF78" box="[151,262,151,177]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="7947CC64FFA4FFE2FEEE6BC2C9DFFF78" box="[276,423,151,177]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="15" pageNumber="16">GSC 142841</specimenCode>
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(part, counterpart) (
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<figureCitation id="B1DA789AFFA4FFE2FD746BC2CB6CFF78" box="[654,788,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="15" pageNumber="16">Figs. 3C–D</figureCitation>
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,
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<figureCitation id="B1DA789AFFA4FFE2FCDB6BC2CB38FF78" box="[801,832,151,177]" captionStart="FIGURE 13" captionStartId="14.[151,250,1956,1980]" captionTargetBox="[151,1436,777,1928]" captionTargetId="figure-207@14.[151,1436,777,1933]" captionTargetPageId="14" captionText="FIGURE 13. Leptostigma longipallidum, holotype GSC 142841, A, photograph, B–C, wing drawings: B, right forewing; C, left forewing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894882" httpUri="https://zenodo.org/record/7894882/files/figure.png" pageId="15" pageNumber="16">13</figureCitation>
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).
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<collectionCode id="4FF0FCDAFFA4FFE2FCAF6BC2CB10FF78" box="[853,872,151,177]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="15" pageNumber="16" type="Herbarium">A</collectionCode>
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very damaged head, thorax, partial abdomen and fragments of legs, both forewings only missing the apical regions overlaying indistinct portions of hind wings; from
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<collectorName id="841401C9FFA4FFE2FF6D6B8AC88FFF30" box="[151,247,223,249]" pageId="15" pageNumber="16">McAbee</collectorName>
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(
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<collectionCode id="4FF0FCDAFFA4FFE2FEFC6B8AC946FF30" box="[262,318,223,249]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="15" pageNumber="16">GSC</collectionCode>
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locality V-016800), collector
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<collectorName id="841401C9FFA4FFE2FD756B8ACB6DFF30" box="[655,789,223,249]" pageId="15" pageNumber="16">John Leahy</collectorName>
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||
; housed in the collections of the
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<collectionCode id="4FF0FCDAFFA4FFE2FB7E6B8ACCC2FF30" box="[1156,1210,223,249]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="15" pageNumber="16">GSC</collectionCode>
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(Vancouver).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFA4FFE2FF3D6A57CD64FE40" pageId="15" pageNumber="16" type="diagnosis">
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<paragraph id="295E641FFFA4FFE2FF3D6A57CD64FE40" blockId="15.[151,1437,151,393]" pageId="15" pageNumber="16">
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<emphasis id="1B95B80DFFA4FFE2FF3D6A57C942FED5" bold="true" box="[199,314,258,284]" pageId="15" pageNumber="16">Diagnosis</emphasis>
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. Distinguished from other species of
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<taxonomicName id="EEE11F9CFFA4FFE2FD1C6A56CB08FED5" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[742,880,259,284]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis id="1B95B80DFFA4FFE2FD1C6A56CB08FED5" box="[742,880,259,284]" italics="true" pageId="15" pageNumber="16">Leptostigma</emphasis>
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</taxonomicName>
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by antenna with a combination of club elongate, paler than flagellar base formed with two long darkened segments; forewing lightly, evenly infuscate throughout. Tentatively diagnostic: R+M longer than 1-M, 2r-rs joins cell 3rm at or before its middle. R+M longer than 1-M, 2r-rs joins cell 3rm at or before its middle, 2r-m joins M far behind 2m-cu, anal cells meet at crossvein.
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</paragraph>
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</subSubSection>
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<caption id="7D9E3497FFA4FFE2FF6D6CEBCB74F830" ID-DOI="http://doi.org/10.5281/zenodo.7894886" ID-Zenodo-Dep="7894886" httpUri="https://zenodo.org/record/7894886/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1982,2006]" targetBox="[169,1418,417,1947]" targetPageId="15" targetType="figure">
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<paragraph id="295E641FFFA4FFE2FF6D6CEBCB74F830" blockId="15.[151,1436,1982,2042]" pageId="15" pageNumber="16">
|
||
<emphasis id="1B95B80DFFA4FFE2FF6D6CEBC95CF81F" bold="true" box="[151,292,1982,2006]" pageId="15" pageNumber="16">FIGURE 14.</emphasis>
|
||
F-771, tentative
|
||
<taxonomicName id="EEE11F9CFFA4FFE2FE266CEACA8AF81F" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[476,754,1982,2006]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipallidum">
|
||
<emphasis id="1B95B80DFFA4FFE2FE266CEACA8AF81F" box="[476,754,1982,2006]" italics="true" pageId="15" pageNumber="16">Leptostigma longipallidum</emphasis>
|
||
</taxonomicName>
|
||
, A, photograph, B–E, wing drawings: B, right forewing; C, left forewing; D, right hind wing; E. left wing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFBBFFFDFF3D6BC3CB88FCF0" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="295E641FFFBBFFFDFF3D6BC3CB88FCF0" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF3D6BC3C929FF79" bold="true" box="[199,337,150,176]" pageId="16" pageNumber="17">Description</emphasis>
|
||
.
|
||
<typeStatus id="F65ADABDFFBBFFFDFE9F6BC2C9A9FF78" box="[357,465,151,177]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
. Body length as preserved (F-777, certainly incomplete)
|
||
<quantity id="EE19C9FAFFBBFFFDFB816BC2CCAAFF78" box="[1147,1234,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="mm" value="20.0">20 mm</quantity>
|
||
, dark throughout including small leg fragments, wing membrane lightly infuscate throughout, pterostigma dark; in F-1185 middle part of wing wide with long, dense pubescence (similar in some living cimbicids,
|
||
<emphasis id="1B95B80DFFBBFFFDFBC66B8ACC28FF30" box="[1084,1104,223,249]" italics="true" pageId="16" pageNumber="17">cf</emphasis>
|
||
.
|
||
<bibRefCitation id="4D7019EEFFBBFFFDFBA56B8ACD1CFF30" author="Vilhelmsen, L. & Smith, D. R. & Malagon-Aldana, L. A." box="[1119,1380,223,249]" pageId="16" pageNumber="17" pagination="1 - 36" refId="ref19867" refString="Vilhelmsen, L., Smith, D. R. & Malagon-Aldana, L. A. (2018) A review of the South American genera of Cimbicidae (Insecta, Hymenoptera). European Journal of Taxonomy, 482, 1 - 36. https: // doi. org / 10.5852 / ejt. 2018.482" type="journal article" year="2018">
|
||
Vilhelmsen
|
||
<emphasis id="1B95B80DFFBBFFFDFB1D6B8ACD59FF30" box="[1255,1313,223,249]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
,
|
||
<figureCitation id="B1DA789AFFBBFFFDFA8A6B8AC8A6FED4" captionStart="FIGURE 2" captionStartId="4.[151,250,1147,1171]" captionTargetBox="[151,1391,564,1120]" captionTargetId="figure-185@4.[151,1436,561,1124]" captionTargetPageId="4" captionText="FIGURE 2. Cenocimbex menatensis Nel, holotype MNHN-LP-R.11195: A, extract from a reflectance transformation imaging (RTI) file (Cultural Heritage Imaging 2012); B, drawing of left forewing; C, drawing of right forewing preserved portion; D, left hind wing. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894834" httpUri="https://zenodo.org/record/7894834/files/figure.png" pageId="16" pageNumber="17">Fig. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="B1DA789AFFBBFFFDFF116A56C968FED4" box="[235,272,259,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1927,1951]" captionTargetBox="[163,1426,705,1888]" captionTargetId="figure-293@9.[151,1436,705,1903]" captionTargetPageId="9" captionText="FIGURE 6. F-1182, tentative Leptostigma brevilatum, A, photograph, and B–E, drawings of: B, left forewing; C, right forewing; D, left hindwing, and; E, right hind wing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894854" httpUri="https://zenodo.org/record/7894854/files/figure.png" pageId="16" pageNumber="17">6D</figureCitation>
|
||
,
|
||
<figureCitation id="B1DA789AFFBBFFFDFEE76A56C928FED4" box="[285,336,259,285]" captionStart="FIGURE 13" captionStartId="14.[151,250,1956,1980]" captionTargetBox="[151,1436,777,1928]" captionTargetId="figure-207@14.[151,1436,777,1933]" captionTargetPageId="14" captionText="FIGURE 13. Leptostigma longipallidum, holotype GSC 142841, A, photograph, B–C, wing drawings: B, right forewing; C, left forewing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894882" httpUri="https://zenodo.org/record/7894882/files/figure.png" pageId="16" pageNumber="17">13A</figureCitation>
|
||
;
|
||
<bibRefCitation id="4D7019EEFFBBFFFDFEA76A56CA77FED4" author="Yan, Y. & Niu, G. & Zhang, Y. & Ren, Q. & Du, S. & Lan, B. & Wei, M." box="[349,527,259,285]" pageId="16" pageNumber="17" pagination="7853" refId="ref20304" refString="Yan, Y., Niu, G., Zhang, Y., Ren, Q., Du, S., Lan, B. & Wei, M. (2019) Complete mitochondrial genome sequence of Labriocimbex sinicus, a new genus and new species of Cimbicidae (Hymenoptera) from China. PeerJ, 7, e 7853 https: // doi. org / 10.7717 / peerj. 7853" type="journal article" year="2019">
|
||
Yan
|
||
<emphasis id="1B95B80DFFBBFFFDFE686A56C9B4FED4" box="[402,460,259,285]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
,
|
||
<figureCitation id="B1DA789AFFBBFFFDFDE66A56CA18FED4" box="[540,608,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1979,2003]" captionTargetBox="[159,1427,1145,1945]" captionTargetId="figure-357@1.[151,1436,1137,1956]" captionTargetPageId="1" captionText="FIGURE 1. Forewing and hind wing of Corynis crassicornis (Rossi) (Corynidinae) with veins (roman font) and cells (italic font) labelled. Drawn from Vilhelmsen (2019) figure 15B. Scale = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894832" httpUri="https://zenodo.org/record/7894832/files/figure.png" pageId="16" pageNumber="17">Fig. 1</figureCitation>
|
||
; may be absent in other
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFC866A56CC4AFED4" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[892,1074,259,285]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Cenocimbicinae">Cenocimbicinae</taxonomicName>
|
||
by preservation). Head (F-777) roundish, slightly transverse, with eyes long, strongly converging forward, reaching rather close to both clypeus and occiput, ocelli in orthogonal triangle, posterior ones somewhat diminished, distant from each other for more than their greatest diameter; toruli touching clypeus, distant for more than their diameter; temples inflated, in front view toward clypeus as wide as eye; clypeus wide, moderately short. Antennal club elongate, paler than funicle, funicle 2-segmented, cylindrical, somewhat darkened. Lateral lobes of pronotum wide; median longitudinal mesonotal sulcus, notauli well developed, joined well before small transverse scutellum, metanotum short. Forewing length
|
||
<typeStatus id="F65ADABDFFBBFFFDFF6D6AAAC881FDD0" box="[151,249,511,537]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
<quantity id="EE19C9FAFFBBFFFDFF056AAAC919FDD0" box="[255,353,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="16" pageNumber="17" unit="mm" value="17.8">17.8 mm</quantity>
|
||
(F-1185,
|
||
<quantity id="EE19C9FAFFBBFFFDFE316AABCA64FDD0" box="[459,540,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="15.0">15 mm</quantity>
|
||
; F-777,
|
||
<quantity id="EE19C9FAFFBBFFFDFD8F6AABCAA3FDD0" box="[629,731,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="16" pageNumber="17" unit="mm" value="16.5">16.5 mm</quantity>
|
||
), maximum width
|
||
<quantity id="EE19C9FAFFBBFFFDFC526AABCC63FDD0" box="[936,1051,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.3" metricValueMin="4.0" pageId="16" pageNumber="17" unit="mm" value="4.65" valueMax="5.3" valueMin="4.0">4–5.3 mm</quantity>
|
||
, length/width 3.4–3.7, pterostigma narrow, tapering, similar to width of C-R basal to pterostigma, R+M longer than 1-M, maximum length of cell 2r shorter than 3r, 2r-rs same length or shorter than Rs between 2r-m and 2r-rs, joins Rs before middle of cell 3rm, 2r-rm curved (more or less sigmoidal), joins Rs similar level to 2m-cu joins M, joins M behind 2m-cu for more than half (for half only in F-1185) of 2r-m length; 1-M more or less curved, cell 1mcu with 1-M, 1m-cu sub-parallel to slightly diverging, 3r-m almost straight, cell 2mcu with M shorter than 1m-cu or (in F-1185) of similar length, 2m-cu slightly bent toward base, cu-a postfurcal (scarcely so if at all in F-777), anal cells meet at crossvein. Hind wing with 3r-m, m-cu, cu-a present, m-cu for its length from 3r-m, cu-a before middle of cell mcu, for twice its length basal to m-cu. Abdominal tergum 1 entire, with hind margin weakly convex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFBBFFFDFF3D6816CD79FC24" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="295E641FFFBBFFFDFF3D6816CD79FC24" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF3D6816C94AFC94" bold="true" box="[199,306,835,861]" pageId="16" pageNumber="17">Remarks</emphasis>
|
||
. Differs from all congeners with known antenna (
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFC8B6816CC70FC94" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[881,1032,835,861]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFC8B6816CC70FC94" box="[881,1032,835,861]" italics="true" pageId="16" pageNumber="17">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFBE86816CC2BFC94" bold="true" box="[1042,1107,835,861]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFBE86816CC2BFC94" box="[1042,1107,835,861]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFB9E6816CC80FC94" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1124,1272,835,861]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFB9E6816CC80FC94" box="[1124,1272,835,861]" italics="true" pageId="16" pageNumber="17">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFAF86816CD3BFC94" bold="true" box="[1282,1347,835,861]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFAF86816CD3BFC94" box="[1282,1347,835,861]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFA7C6816C933FC48" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFA7C6816C933FC48" italics="true" pageId="16" pageNumber="17">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFEAA6832C9F3FC48" bold="true" box="[336,395,871,897]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFEAA6832C9F3FC48" box="[336,395,871,897]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) by its club paler than its funicle. Differs venationally from
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFBF16832CCE4FC48" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1035,1180,871,897]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFBF16832CCE4FC48" box="[1035,1180,871,897]" italics="true" pageId="16" pageNumber="17">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFB5B6832CCA4FC48" bold="true" box="[1185,1244,871,897]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFB5B6832CCA4FC48" box="[1185,1244,871,897]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFAF76832CDE4FC48" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1293,1436,871,897]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFAF76832CDE4FC48" box="[1293,1436,871,897]" italics="true" pageId="16" pageNumber="17">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFF6D68DEC8ACFC6C" bold="true" box="[151,212,907,933]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFF6D68DEC8ACFC6C" box="[151,212,907,933]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by R+M longer than 1-M, from
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFDCE68DECB7CFC6C" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[564,772,907,933]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFDCE68DECB7CFC6C" box="[564,772,907,933]" italics="true" pageId="16" pageNumber="17">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFCF068DECB3EFC6C" bold="true" box="[778,838,907,933]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFCF068DECB3EFC6C" box="[778,838,907,933]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its anal cells meeting at a crossvein. Differs from
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFA7C68DEC97CFC01" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="proxivena" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFA7C68DEC97CFC01" italics="true" pageId="16" pageNumber="17">L. proxivena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFEF168FAC931FC00" bold="true" box="[267,329,943,969]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFEF168FAC931FC00" box="[267,329,943,969]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFEAD68FAC9A7FC00" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[343,479,943,969]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="fasciatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFEAD68FAC9A7FC00" box="[343,479,943,969]" italics="true" pageId="16" pageNumber="17">L. fasciatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFE1C68FACA5CFC00" bold="true" box="[486,548,943,969]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFE1C68FACA5CFC00" box="[486,548,943,969]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFDC868FACAB7FC00" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[562,719,943,969]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaemacula" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFBBFFFDFDC868FACAB7FC00" box="[562,719,943,969]" italics="true" pageId="16" pageNumber="17">L. alaemacula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFBBFFFDFD2C68FACB6CFC00" bold="true" box="[726,788,943,969]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="00A60576FFBBFFFDFD2C68FACB6CFC00" box="[726,788,943,969]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, with unknown antenna by wing membrane lightly, evenly infuscate throughout (without fascia, macula) and 2r-m entering M far behind, and not close to, 2m-cu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFBBFFFDFF3D68A3CBF9FBFC" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="295E641FFFBBFFFDFF3D68A3CBF9FBFC" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF3D68A3C957FBD9" bold="true" box="[199,303,1014,1040]" pageId="16" pageNumber="17">Material</emphasis>
|
||
.
|
||
<materialsCitation id="99896E42FFBBFFFDFEC468A2CBF9FBFC" ID-GBIF-Occurrence="4100387317" collectionCode="TRU" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="F65ADABDFFBBFFFDFEC468A2C9DEFBD8" box="[318,422,1015,1041]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
and tentatively (by venation) specimen F-777, F-771 (part, counterpart) (
|
||
<figureCitation id="B1DA789AFFBBFFFDFB1368A2CD44FBD8" box="[1257,1340,1015,1041]" captionStart="FIGURE 14" captionStartId="15.[151,250,1982,2006]" captionTargetBox="[169,1418,417,1947]" captionTargetId="figure-136@15.[169,1418,417,1958]" captionTargetPageId="15" captionText="FIGURE 14. F-771, tentative Leptostigma longipallidum, A, photograph, B–E, wing drawings: B, right forewing; C, left forewing; D, right hind wing; E. left wing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894886" httpUri="https://zenodo.org/record/7894886/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||
), and F-1185 (
|
||
<figureCitation id="B1DA789AFFBBFFFDFF276F4EC948FBFC" box="[221,304,1050,1077]" captionStart="FIGURE 15" captionStartId="16.[151,250,1877,1901]" captionTargetBox="[168,1420,1101,1813]" captionTargetId="figure-483@16.[168,1420,1101,1854]" captionTargetPageId="16" captionText="FIGURE 15. F-1185 tentative Leptostigma longipallidum forewing: A, photograph; B, drawing. Scale is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894890" httpUri="https://zenodo.org/record/7894890/files/figure.png" pageId="16" pageNumber="17">Fig. 15</figureCitation>
|
||
), both collector unknown, in the collections of
|
||
<collectionCode id="4FF0FCDAFFBBFFFDFCBB6F4ECB05FBFC" box="[833,893,1051,1077]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="16" pageNumber="17" type="Herbarium">TRU</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFBBFFFDFF6D6C00CD7DF8A4" ID-DOI="http://doi.org/10.5281/zenodo.7894890" ID-Zenodo-Dep="7894890" box="[151,1285,1877,1902]" httpUri="https://zenodo.org/record/7894890/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1877,1901]" targetBox="[168,1420,1101,1813]" targetPageId="16" targetType="figure">
|
||
<paragraph id="295E641FFFBBFFFDFF6D6C00CD7DF8A4" blockId="16.[151,1285,1877,1902]" box="[151,1285,1877,1902]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF6D6C00C958F8A4" bold="true" box="[151,288,1877,1901]" pageId="16" pageNumber="17">FIGURE 15.</emphasis>
|
||
F-1185 tentative
|
||
<taxonomicName id="EEE11F9CFFBBFFFDFE286C03CA9CF8A7" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[466,740,1878,1902]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="longipallidum">
|
||
<emphasis id="1B95B80DFFBBFFFDFE286C03CA9CF8A7" box="[466,740,1878,1902]" italics="true" pageId="16" pageNumber="17">Leptostigma longipallidum</emphasis>
|
||
</taxonomicName>
|
||
forewing: A, photograph; B, drawing. Scale is 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFBBFFFDFF3D6CC2C947F81D" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="295E641FFFBBFFFDFF3D6CC2C947F81D" blockId="16.[151,1436,1942,2041]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF3D6CC2C936F878" bold="true" box="[199,334,1943,1969]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="1B95B80DFFBBFFFDFDFB6CC3CA34F879" box="[513,588,1942,1968]" italics="true" pageId="16" pageNumber="17">longus</emphasis>
|
||
, meaning long, and
|
||
<emphasis id="1B95B80DFFBBFFFDFCD76CC3CBFFF879" box="[813,903,1942,1968]" italics="true" pageId="16" pageNumber="17">pallidus</emphasis>
|
||
, pale, referring to the antennal club. The epithet is an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFBBFFFDFF3D6C8BCCC1F831" box="[199,1209,2014,2041]" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="295E641FFFBBFFFDFF3D6C8BCCC1F831" blockId="16.[151,1436,1942,2041]" box="[199,1209,2014,2041]" pageId="16" pageNumber="17">
|
||
<emphasis id="1B95B80DFFBBFFFDFF3D6C8BC9ECF830" bold="true" box="[199,404,2014,2041]" pageId="16" pageNumber="17">Locality and age.</emphasis>
|
||
McAbee,
|
||
<collectingRegion id="EB25AAFDFFBBFFFDFDF36C8BCAB5F831" box="[521,717,2014,2040]" country="Canada" name="British Columbia" pageId="16" pageNumber="17">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="51F6248FFFBBFFFDFD236C8BCB48F831" box="[729,816,2014,2040]" name="Canada" pageId="16" pageNumber="17">Canada</collectingCountry>
|
||
, Hoodoo Face beds; mid-Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |