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<document id="AC20CF1FCC76AAFD77EA21115E9EC306" ID-CLB-Dataset="58359" ID-DOI="10.11646/zootaxa.5285.1.7" ID-GBIF-Dataset="77e7fae7-a1e0-43cb-af62-d4e9597f329f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7935889" ID-ZooBank="F2895A5A-A06E-480F-A6C8-061150B57041" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684131653704" checkinUser="plazi" docAuthor="Pati, Sameer Kumar, Bajantri, Parashuram Prabhu &amp; Hegde, Gopalkrishna Dattatraya" docDate="2023" docId="9F5287886A17FF8B94EA6A18AD4CF8A0" docLanguage="en" docName="zootaxa.5285.1.7.pdf" docOrigin="Zootaxa 5285 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5285.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aradhya Pati &amp; Bajantri &amp; Hegde 2023, gen. nov." docType="treatment" docUuid="DC7F9116-1CA9-4423-9C31-ECDE89DD0C71" docUuidSource="ZooBank" docVersion="7" lastPageNumber="170" masterDocId="636BFFF06A15FF82947D685EA945FFA5" masterDocTitle="A new genus and new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India" masterLastPageNumber="175" masterPageNumber="161" pageNumber="163" updateTime="1699505753996" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="869EA88FE7450842818B33E4500D1DC6">A new genus and new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India</mods:title>
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<mods:namePart id="E444784A62F4750B3DC518A4BCEE4BC6">Pati, Sameer Kumar</mods:namePart>
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<mods:namePart id="975189302F03CA94C5999F5C6814079C">Bajantri, Parashuram Prabhu</mods:namePart>
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<mods:namePart id="0D7612C73C200E6D3A8C2C1808A18D9A">Hegde, Gopalkrishna Dattatraya</mods:namePart>
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<paragraph id="1744369E6A17FF8094EA6A18A830FDC4" blockId="2.[151,373,582,609]" box="[151,373,582,609]" pageId="2" pageNumber="163">
<heading id="4C0C81F26A17FF8094EA6A18A830FDC4" bold="true" box="[151,373,582,609]" fontSize="11" level="1" pageId="2" pageNumber="163" reason="1">
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<taxonomicName id="D0FB4D1D6A17FF8094EA6A18A9BAFDC4" ID-CoL="8b32c181-d229-4ef4-b844-751e7d68e1a8" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[151,255,582,609]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A17FF8094EA6A18A9BAFDC4" bold="true" box="[151,255,582,609]" italics="true" pageId="2" pageNumber="163">Aradhya</emphasis>
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<taxonomicNameLabel id="3EBC57F76A17FF80957A6A19A830FDC4" box="[263,373,583,609]" pageId="2" pageNumber="163" rank="genus">gen. nov.</taxonomicNameLabel>
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<typeStatus id="C840883C6A17FF8094EA6AD1A997FD0C" box="[151,210,655,681]" pageId="2" pageNumber="163">Type</typeStatus>
species.
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<taxonomicName id="D0FB4D1D6A17FF8095446AD1A8B0FD0C" ID-CoL="90e8146e-992e-427d-923d-7cc05de0c133" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[313,501,655,681]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A17FF8095446AD1A8B0FD0C" box="[313,501,655,681]" italics="true" pageId="2" pageNumber="163">Aradhya placida</emphasis>
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<taxonomicNameLabel id="3EBC57F76A17FF8095816AD1AB10FD0C" box="[508,597,655,681]" pageId="2" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation; gender feminine.
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<emphasis id="258FEA8C6A17FF8094BA6AECA807FD69" bold="true" box="[199,322,690,716]" pageId="2" pageNumber="163">Diagnosis.</emphasis>
Carapace transversely ovate, broader than long (CW/CL = 1.21.3), strongly inflated, deep (
<collectingCountry id="6FEC760E6A17FF8091116AEDACD1FD68" box="[1388,1428,691,717]" name="Switzerland" pageId="2" pageNumber="163">CH</collectingCountry>
/ CW = 0.60.7); dorsal surface generally smooth, glabrous, strongly arched; anterolateral margins cristate, lacking distinct serrations; front strongly deflexed, with narrow anterior margin (FW/CW = 0.25); frontal medial triangle incomplete, lateral margins indiscernible; epigastric and postorbital cristae poorly developed; external orbital angle indistinct; epibranchial tooth very low; branchial regions strongly inflated; cervical grooves relatively shallow, not reaching level of postorbital cristae; epistome posterior margin with distinct, triangular medial lobe and strongly sinuous lateral lobes (
<figureCitation id="8FC02A1B6A17FF8095F16BD5AB54FC00" box="[396,529,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2A, B</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096616BD5ABC4FC00" box="[540,641,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096F06BD5AB91FC00" box="[653,724,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">4AC</figureCitation>
). First, second maxillipeds each with long flagellum on exopod (
<figureCitation id="8FC02A1B6A17FF8094E26BF1A9BFFC6C" box="[159,250,942,969]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">Fig. 5A</figureCitation>
). TME lacking flagellum or with relatively short flagellum, reaching about half width of merus (
<figureCitation id="8FC02A1B6A17FF8091496BF1ACD2FC6C" box="[1332,1431,943,969]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">Figs. 3G</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8094EA6B8DA9F9FC48" box="[151,188,979,1005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">4D</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8094B46B8CA9A8FC48" box="[201,237,978,1005]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5B</figureCitation>
). Chelipeds relatively smooth, unequal in adult males (
<figureCitation id="8FC02A1B6A17FF80971F6B8DAAAFFC48" box="[866,1002,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF80978A6B8DAD7DFC48" box="[1015,1080,979,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3AF</figureCitation>
) and subequal in adult females (
<figureCitation id="8FC02A1B6A17FF8094E26BA9A864FBB4" box="[159,289,1015,1041]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">Fig. 4A, C</figureCitation>
). Ambulatory legs slender, long; dactylus (P2P5) recurved, longer than propodus, distal chitinous part very short or reduced (
<figureCitation id="8FC02A1B6A17FF8095AB6C45AB25FB90" box="[470,608,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096136C45ABEAFB90" box="[622,687,1051,1077]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3AF</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096C06C45AA41FB90" box="[701,772,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">4AC</figureCitation>
). Male s2/s3 cristate, not reaching edge of sternum; male s3/s4 deep, broad, reaching edge of sternum (
<figureCitation id="8FC02A1B6A17FF8096E06C61AA45FBFC" box="[669,768,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097736C61AA34FBFC" box="[782,881,1087,1113]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097036C60AAE7FBFC" box="[894,930,1086,1113]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5C</figureCitation>
). Male pleonal locking mechanism with low tubercles on submedial part of s5 (
<figureCitation id="8FC02A1B6A17FF8096616C3DAB30FBD8" box="[540,629,1122,1149]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">Fig. 5C</figureCitation>
). Male sternopleonal cavity relatively short, reaching anteriorly to level of mid-length of cheliped coxae (
<figureCitation id="8FC02A1B6A17FF80966D6CD9AB36FB04" box="[528,627,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096026CD9ABA4FB04" box="[639,737,1159,1185]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096936CD8AA57FB04" box="[750,786,1158,1185]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5C</figureCitation>
). Male pleon relatively narrow, triangular; pleonal somite 6 trapezoidal, relatively narrow (proximal width ca. 1.3× medial length), shorter than telson, with gently concave lateral margins (
<figureCitation id="8FC02A1B6A17FF8095326C91A8F7FB4C" box="[335,434,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8095C26C91AB67FB4C" box="[447,546,1231,1257]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8096536C90AB16FB4C" box="[558,595,1230,1257]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF80961D6C91ABC0FB4C" box="[608,645,1231,1257]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="2" pageNumber="163">6A</figureCitation>
). Male telson narrow, elongated (medial length ca. 1.31.4× proximal width), with gently concave lateral margins (
<figureCitation id="8FC02A1B6A17FF8096E76CADABBAFAA8" box="[666,767,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097706CADAA37FAA8" box="[781,882,1267,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097FD6CACAAE1FAA8" box="[896,932,1266,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097CF6CADAA9DFAA8" box="[946,984,1267,1293]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="2" pageNumber="163">6A</figureCitation>
). G1 relatively stout, long, tip reaching pleonal locking structure
<emphasis id="258FEA8C6A17FF8095C46D49A8BAFA95" box="[441,511,1303,1328]" italics="true" pageId="2" pageNumber="163">in situ</emphasis>
; flexible zone small; ultimate article relatively stout, conical, relatively short, ca. 0.3× length of penultimate article, gently curved outwards at angle of ca. 15° from longitudinal axis, tip broad, tubular, dorsal flap absent; penultimate article relatively stout, outer margin strongly convex, shelf-like at basal half, inner margin characteristically convex distally (
<figureCitation id="8FC02A1B6A17FF8096D06DDDAA54FA38" box="[685,785,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097606DDCAA7BFA38" box="[797,830,1410,1437]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8097376DDDAACBFA38" box="[842,910,1411,1437]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="2" pageNumber="163">6BE</figureCitation>
). G2 distinctly shorter than G1, ca. 0.6× length of G1; ultimate article very short, ca. 0.2× length of penultimate article (
<figureCitation id="8FC02A1B6A17FF8097BB6DF9AD5AFA64" box="[966,1055,1447,1473]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">Figs. 3I</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8090566DF8AD0CFA64" box="[1067,1097,1446,1473]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8090286DF9AD33FA64" box="[1109,1142,1447,1473]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="2" pageNumber="163">6F</figureCitation>
). Female pleon and telson ovate in outline; pleonal somite 6 subequal in length to telson; telson subtriangular, with broad, straight apex (
<figureCitation id="8FC02A1B6A17FF8091126D95A9FEF9AC" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">Fig. 4E</figureCitation>
). Female pleopods 25 endopods slender and longer than exopods; pleopods 35 exopods conspicuously stouter than endopods (
<figureCitation id="8FC02A1B6A17FF8095306E4DA8E1F988" box="[333,420,1555,1581]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">Fig. 4F</figureCitation>
). Vulvae on s6 close to each other (VD/SW = ca. 0.1), opening inwards, subovate, large, touching s5/s6 (
<figureCitation id="8FC02A1B6A17FF8095346E69A8E1F9F4" box="[329,420,1591,1617]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">Fig. 4G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5FE165156A17FF8094BA6E05AD84F918" pageId="2" pageNumber="163" type="etymology">
<paragraph id="1744369E6A17FF8094BA6E05AD84F918" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="163">
<emphasis id="258FEA8C6A17FF8094BA6E05A80BF9D0" bold="true" box="[199,334,1627,1653]" pageId="2" pageNumber="163">Etymology.</emphasis>
The genus is named after Miss Aaradhya Bajantri, the only daughter of the second author of the present paper. “
<taxonomicName id="D0FB4D1D6A17FF80953B6E21A8EEF93C" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[326,427,1663,1689]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">Aradhya</taxonomicName>
” also means “the first one” in Sanskrit. Crabs of this new genus are the first ones among the Indian gecarcinucid crabs observed to be non-aggressive and quite calm. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="5FE165156A17FF8B94BA6E99AD4CF8A0" lastPageId="9" lastPageNumber="170" pageId="2" pageNumber="163" type="discussion">
<paragraph id="1744369E6A17FF8594BA6E99AD83FE2C" blockId="2.[151,1437,654,2013]" lastBlockId="7.[151,1437,150,393]" lastPageId="7" lastPageNumber="168" pageId="2" pageNumber="163">
<emphasis id="258FEA8C6A17FF8094BA6E99A87EF944" bold="true" box="[199,315,1735,1761]" pageId="2" pageNumber="163">Remarks.</emphasis>
Among the Indian gecarcinucid genera,
<taxonomicName id="D0FB4D1D6A17FF8097716E99AA29F944" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[780,876,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A17FF8097716E99AA29F944" box="[780,876,1735,1761]" italics="true" pageId="2" pageNumber="163">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A17FF80970B6E99AAA5F944" bold="true" box="[886,992,1735,1761]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="3EBC57F76A17FF80970B6E99AAA5F944" box="[886,992,1735,1761]" pageId="2" pageNumber="163" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
most closely resembles
<taxonomicName id="D0FB4D1D6A17FF8090846E99ACD0F944" authorityName="Pati &amp; Sudha Devi" authorityYear="2015" box="[1273,1429,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A17FF8090846E99ACD0F944" box="[1273,1429,1735,1761]" italics="true" pageId="2" pageNumber="163">Arcithelphusa</emphasis>
</taxonomicName>
,
<taxonomicName id="D0FB4D1D6A17FF8094EA6EB5ABFFF8A0" authority=", Rajathelphusa, and Pavizham" authorityName="Rajathelphusa, and Pavizham" box="[151,698,1771,1797]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A17FF8094EA6EB5A81DF8A0" box="[151,344,1771,1797]" italics="true" pageId="2" pageNumber="163">Cylindrotelphusa</emphasis>
,
<emphasis id="258FEA8C6A17FF8095146EB5AB4EF8A0" box="[361,523,1771,1797]" italics="true" pageId="2" pageNumber="163">Rajathelphusa</emphasis>
, and
<emphasis id="258FEA8C6A17FF8096316EB5ABFFF8A0" box="[588,698,1771,1797]" italics="true" pageId="2" pageNumber="163">Pavizham</emphasis>
</taxonomicName>
, because all possess an ovate, relatively narrow (CW/CL = 1.2 1.4) and deep carapace (
<collectingCountry id="6FEC760E6A17FF8095DE6F51A88FF88C" box="[419,458,1807,1833]" name="Switzerland" pageId="2" pageNumber="163">CH</collectingCountry>
/CW = 0.50.7), with a narrow front (FW/CW = 0.25) and an incomplete frontal median triangle (
<figureCitation id="8FC02A1B6A17FF8094816F6DA8C7F8E8" box="[252,386,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2A, B</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8095F36F6DA8B6F8E8" box="[398,499,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8095826F6DAB03F8E8" box="[511,582,1843,1869]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="2" pageNumber="163">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A17FF8096016F6DAA6EF8E8" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[636,811,1843,1869]" pageId="2" pageNumber="163" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A17FF8096D26F6DABADF8E8" box="[687,744,1843,1869]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, B, D, E;
<bibRefCitation id="736A4B6F6A17FF8097846F6DAD8FF8E8" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[1017,1226,1843,1869]" pageId="2" pageNumber="163" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5A, B;
<bibRefCitation id="736A4B6F6A17FF8091286F6DA9B2F8D4" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." pageId="2" pageNumber="163" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A17FF8091F56F6DA9F1F8D4" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11a, b, 13a, b;
<bibRefCitation id="736A4B6F6A17FF8095A36F09ABC2F8D4" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[478,647,1879,1905]" pageId="2" pageNumber="163" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A17FF8096716F09AB00F8D4" box="[524,581,1879,1905]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, B, 7A, C;
<bibRefCitation id="736A4B6F6A17FF8097106F09AD53F8D4" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[877,1046,1879,1905]" pageId="2" pageNumber="163" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A17FF8097E66F09AA96F8D4" box="[923,979,1879,1905]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
2022
</bibRefCitation>
: figs. 3A, C, 8A, C).
<taxonomicName id="D0FB4D1D6A17FF8091796F09AC21F8D4" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1284,1380,1879,1905]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A17FF8091796F09AC21F8D4" box="[1284,1380,1879,1905]" italics="true" pageId="2" pageNumber="163">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A17FF8091166F09A98DF830" bold="true" pageId="2" pageNumber="163">
<taxonomicNameLabel id="3EBC57F76A17FF8091166F09A98DF830" pageId="2" pageNumber="163" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is nevertheless distinguished from
<taxonomicName id="D0FB4D1D6A17FF8096256F25ABB1F830" authorityName="Pati &amp; Sudha Devi" authorityYear="2015" box="[600,756,1915,1941]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A17FF8096256F25ABB1F830" box="[600,756,1915,1941]" italics="true" pageId="2" pageNumber="163">Arcithelphusa</emphasis>
</taxonomicName>
,
<taxonomicName id="D0FB4D1D6A17FF80977E6F25AC67F830" authority=", Rajathelphusa, and Pavizham" authorityName="Rajathelphusa, and Pavizham" box="[771,1314,1915,1941]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A17FF80977E6F25AA81F830" box="[771,964,1915,1941]" italics="true" pageId="2" pageNumber="163">Cylindrotelphusa</emphasis>
,
<emphasis id="258FEA8C6A17FF8097AE6F25AD30F830" box="[979,1141,1915,1941]" italics="true" pageId="2" pageNumber="163">Rajathelphusa</emphasis>
, and
<emphasis id="258FEA8C6A17FF8090C96F25AC67F830" box="[1204,1314,1915,1941]" italics="true" pageId="2" pageNumber="163">Pavizham</emphasis>
</taxonomicName>
mainly by its narrow and elongated male telson, the medial length ca. 1.31.4 times the proximal width (
<figureCitation id="8FC02A1B6A17FF8090B86FC1AC6DF81C" box="[1221,1320,1951,1977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="2" pageNumber="163">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8091486FC1ACDDF81C" box="[1333,1432,1951,1977]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="2" pageNumber="163">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8094EA6F9CA9F9F878" box="[151,188,1986,2013]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="2" pageNumber="163">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A17FF8094B76F9DA9A8F878" box="[202,237,1987,2013]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="2" pageNumber="163">6A</figureCitation>
) (vs. male telson relatively broader and shorter, the medial length ca. 0.91.1 times the proximal width; see
<bibRefCitation id="736A4B6F6A12FF8594B968C9A832FF14" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[196,375,151,177]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF85948568C9A877FF14" box="[248,306,151,177]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2C, F;
<bibRefCitation id="736A4B6F6A12FF85967F68C9AB9DFF14" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[514,728,151,177]" pageId="7" pageNumber="168" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5C;
<bibRefCitation id="736A4B6F6A12FF85973968C9AAB2FF14" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[836,1015,151,177]" pageId="7" pageNumber="168" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A12FF85970568C9AAF6FF14" box="[888,947,151,177]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11h, 13h;
<bibRefCitation id="736A4B6F6A12FF8590CC68C9AC1AFF14" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1201,1375,151,177]" pageId="7" pageNumber="168" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A12FF85909D68C9AC5FFF14" box="[1248,1306,151,177]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3D, 7E;
<bibRefCitation id="736A4B6F6A12FF85948868E5A8DBFF70" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[245,414,187,213]" pageId="7" pageNumber="168" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A12FF85955E68E5A81EFF70" box="[291,347,187,213]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2022
</bibRefCitation>
: figs. 4A, 9C), the relatively stouter ultimate article of the G1 (
<figureCitation id="8FC02A1B6A12FF85902368E5AD84FF70" box="[1118,1217,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8590B068E4ADAAFF70" box="[1229,1263,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85908768E5AC7EFF70" box="[1274,1339,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="7" pageNumber="168">6BE</figureCitation>
) (vs. G1 ultimate article relatively slender; see
<bibRefCitation id="736A4B6F6A12FF8596416881ABA3FF5C" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[572,742,223,249]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8596136881ABE0FF5C" box="[622,677,223,249]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3E, F, J, O;
<bibRefCitation id="736A4B6F6A12FF8597E16881AD2DFF5C" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[924,1128,223,249]" pageId="7" pageNumber="168" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 4AC;
<bibRefCitation id="736A4B6F6A12FF8590936881ACDDFF5C" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[1262,1432,223,249]" pageId="7" pageNumber="168" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A12FF85915D6881AC12FF5C" box="[1312,1367,223,249]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2017
</bibRefCitation>
: figs. 12ac, 14ac;
<bibRefCitation id="736A4B6F6A12FF85950C695DAB5FFEB8" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[369,538,259,285]" pageId="7" pageNumber="168" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A12FF8595E2695DA89DFEB8" box="[415,472,259,285]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2021
</bibRefCitation>
: fig. 9BD, FH;
<bibRefCitation id="736A4B6F6A12FF85969E695DAAC9FEB8" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[739,908,259,285]" pageId="7" pageNumber="168" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A12FF85976C695DAA0CFEB8" box="[785,841,259,285]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2022
</bibRefCitation>
: figs. 5CE, 10AC), and the characteristically convex distal inner margin of the penultimate article of the G1 (
<figureCitation id="8FC02A1B6A12FF8597286979AAF2FEE4" box="[853,951,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597BF6978AAA1FEE4" box="[962,996,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597936979AD75FEE4" box="[1006,1072,295,321]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="7" pageNumber="168">6B, C</figureCitation>
) (vs. G1 penultimate article with an almost straight distal inner margin; see
<bibRefCitation id="736A4B6F6A12FF85960C6915AA5BFEC0" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[625,798,331,357]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8596DE6915AB9EFEC0" box="[675,731,331,357]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3E, F, J, O;
<bibRefCitation id="736A4B6F6A12FF8597A46915ADECFEC0" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[985,1193,331,357]" pageId="7" pageNumber="168" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 4A, C;
<bibRefCitation id="736A4B6F6A12FF85914F6915A996FE2C" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." pageId="7" pageNumber="168" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A12FF8591196915ACD8FEC0" box="[1380,1437,331,357]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2017
</bibRefCitation>
: figs. 12a, b, 13a, b;
<bibRefCitation id="736A4B6F6A12FF8595C16931AB21FE2C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[444,612,367,393]" pageId="7" pageNumber="168" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A12FF8595946931AB67FE2C" box="[489,546,367,393]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2021
</bibRefCitation>
: fig. 9B, D, F, H;
<bibRefCitation id="736A4B6F6A12FF8597516931AA91FE2C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[812,980,367,393]" pageId="7" pageNumber="168" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A12FF8597246931AAD7FE2C" box="[857,914,367,393]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2022
</bibRefCitation>
: figs. 5C, E, 10A, C).
</paragraph>
<caption id="438466166A16FF8194EA6FC9AAEDF876" ID-DOI="http://doi.org/10.5281/zenodo.7935895" ID-Zenodo-Dep="7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="3" pageNumber="164" startId="3.[151,250,1943,1967]" targetBox="[192,1395,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="1744369E6A16FF8194EA6FC9AAEDF876" blockId="3.[151,1436,1943,2003]" pageId="3" pageNumber="164">
<emphasis id="258FEA8C6A16FF8194EA6FC9A856F80A" bold="true" box="[151,275,1943,1967]" pageId="3" pageNumber="164">FIGURE 2.</emphasis>
<taxonomicName id="D0FB4D1D6A16FF8195676FC9A881F80A" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[282,452,1943,1967]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A16FF8195676FC9A881F80A" box="[282,452,1943,1967]" italics="true" pageId="3" pageNumber="164">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A16FF8195B76FC9AB2DF80A" bold="true" box="[458,616,1943,1967]" pageId="3" pageNumber="164">
<taxonomicNameLabel id="3EBC57F76A16FF8195B76FC9AB2DF80A" box="[458,616,1943,1967]" pageId="3" pageNumber="164" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="438466166A11FF8694EA6F1CAB45F863" ID-DOI="http://doi.org/10.5281/zenodo.7935897" ID-Zenodo-Dep="7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="4" pageNumber="165" startId="4.[151,250,1858,1882]" targetBox="[151,1435,181,1834]" targetPageId="4" targetType="figure">
<paragraph id="1744369E6A11FF8694EA6F1CAB45F863" blockId="4.[151,1436,1858,1990]" pageId="4" pageNumber="165">
<emphasis id="258FEA8C6A11FF8694EA6F1CA851F8FF" bold="true" box="[151,276,1858,1882]" pageId="4" pageNumber="165">FIGURE 3.</emphasis>
<taxonomicName id="D0FB4D1D6A11FF8695616F1CA883F8FF" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[284,454,1858,1882]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A11FF8695616F1CA883F8FF" box="[284,454,1858,1882]" italics="true" pageId="4" pageNumber="165">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A11FF8695B36F1CAB2BF8FF" bold="true" box="[462,622,1858,1882]" pageId="4" pageNumber="165">
<taxonomicNameLabel id="3EBC57F76A11FF8695B36F1CAB2BF8FF" box="[462,622,1858,1882]" pageId="4" pageNumber="165" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I).
</paragraph>
</caption>
<caption id="438466166A10FF8794EA6F28ADD6F85F" ID-DOI="http://doi.org/10.5281/zenodo.7935901" ID-Zenodo-Dep="7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="5" pageNumber="166" startId="5.[151,250,1910,1934]" targetBox="[214,1372,181,1886]" targetPageId="5" targetType="figure">
<paragraph id="1744369E6A10FF8794EA6F28ADD6F85F" blockId="5.[151,1436,1910,2042]" pageId="5" pageNumber="166">
<emphasis id="258FEA8C6A10FF8794EA6F28A851F82B" bold="true" box="[151,276,1910,1934]" pageId="5" pageNumber="166">FIGURE 4.</emphasis>
<taxonomicName id="D0FB4D1D6A10FF8795666F28A880F82B" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[283,453,1910,1934]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A10FF8795666F28A880F82B" box="[283,453,1910,1934]" italics="true" pageId="5" pageNumber="166">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A10FF8795B16F28AB2EF82B" bold="true" box="[460,619,1910,1934]" pageId="5" pageNumber="166">
<taxonomicNameLabel id="3EBC57F76A10FF8795B16F28AB2EF82B" box="[460,619,1910,1934]" pageId="5" pageNumber="166" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D).
</paragraph>
</caption>
<caption id="438466166A13FF8494EA6FC9AB0EF852" ID-DOI="http://doi.org/10.5281/zenodo.7935903" ID-Zenodo-Dep="7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="6" pageNumber="167" startId="6.[151,250,1943,1967]" targetBox="[203,1384,181,1919]" targetPageId="6" targetType="figure">
<paragraph id="1744369E6A13FF8494EA6FC9AB0EF852" blockId="6.[151,1436,1943,2039]" pageId="6" pageNumber="167">
<emphasis id="258FEA8C6A13FF8494EA6FC9A857F80A" bold="true" box="[151,274,1943,1967]" pageId="6" pageNumber="167">FIGURE 5.</emphasis>
<taxonomicName id="D0FB4D1D6A13FF8495656FC9A884F80A" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[280,449,1943,1967]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A13FF8495656FC9A884F80A" box="[280,449,1943,1967]" italics="true" pageId="6" pageNumber="167">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A13FF8495BA6FC9AB27F80A" bold="true" box="[455,610,1943,1967]" pageId="6" pageNumber="167">
<taxonomicNameLabel id="3EBC57F76A13FF8495BA6FC9AB27F80A" box="[455,610,1943,1967]" pageId="6" pageNumber="167" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F).
</paragraph>
</caption>
<caption id="438466166A12FF8594EA6D16AA37FA0D" ID-DOI="http://doi.org/10.5281/zenodo.7935907" ID-Zenodo-Dep="7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="7" pageNumber="168" startId="7.[151,250,1352,1376]" targetBox="[153,1434,419,1327]" targetPageId="7" targetType="figure">
<paragraph id="1744369E6A12FF8594EA6D16AA37FA0D" blockId="7.[151,1437,1352,1448]" pageId="7" pageNumber="168">
<emphasis id="258FEA8C6A12FF8594EA6D16A856FAC5" bold="true" box="[151,275,1352,1376]" pageId="7" pageNumber="168">FIGURE 6.</emphasis>
<taxonomicName id="D0FB4D1D6A12FF8595676D16A886FAC5" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[282,451,1352,1376]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A12FF8595676D16A886FAC5" box="[282,451,1352,1376]" italics="true" pageId="7" pageNumber="168">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A12FF8595B76D16AB22FAFA" bold="true" box="[458,615,1352,1376]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="3EBC57F76A12FF8595B76D16AB22FAFA" box="[458,615,1352,1376]" pageId="7" pageNumber="168" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F).
</paragraph>
</caption>
<paragraph id="1744369E6A12FF8A94BA6D88ADCCFDC4" blockId="7.[151,1437,1494,2025]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="169" pageId="7" pageNumber="168">
<taxonomicName id="D0FB4D1D6A12FF8594BA6D88A862FA55" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[199,295,1494,1520]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A12FF8594BA6D88A862FA55" box="[199,295,1494,1520]" italics="true" pageId="7" pageNumber="168">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A12FF85954C6D88A8D8FA55" bold="true" box="[305,413,1494,1520]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="3EBC57F76A12FF85954C6D88A8D8FA55" box="[305,413,1494,1520]" pageId="7" pageNumber="168" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is morphologically closer to
<taxonomicName id="D0FB4D1D6A12FF85968F6D88AACBFA55" authorityName="Pati &amp; Sudha Devi" authorityYear="2015" box="[754,910,1494,1520]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A12FF85968F6D88AACBFA55" box="[754,910,1494,1520]" italics="true" pageId="7" pageNumber="168">Arcithelphusa</emphasis>
</taxonomicName>
than any other Indian genera of gecarcinucid crabs as both genera share several features in common, including a strongly arched and generally smooth dorsal surface of the carapace (
<figureCitation id="8FC02A1B6A12FF8595DA6E40AB69F99C" box="[423,556,1566,1593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A, B</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85964A6E40ABDEF99C" box="[567,667,1566,1593]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8596DA6E40ABA8F99D" box="[679,749,1566,1592]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF85975E6E40AA95F99D" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[803,976,1566,1592]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8597286E41AACBF99D" box="[853,910,1566,1592]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, B, D, E), the absence of distinct serrations on the anterolateral margins of the carapace (
<figureCitation id="8FC02A1B6A12FF85976B6E1CAA39F9F9" box="[790,892,1602,1628]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597F76E1CAAB6F9F8" box="[906,1011,1602,1629]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85907C6E1CAD02F9F9" box="[1025,1095,1602,1628]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8590FF6E1CAC72F9F9" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[1154,1335,1602,1628]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8590CA6E1DADB7F9F9" box="[1207,1266,1602,1628]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, D), the poorly developed epigastric and postorbital cristae (
<figureCitation id="8FC02A1B6A12FF85974C6E38AAD0F925" box="[817,917,1638,1664]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597DC6E38AD40F924" box="[929,1029,1638,1665]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85906C6E38AD12F925" box="[1041,1111,1638,1664]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8590F06E38AC7EF925" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[1165,1339,1638,1664]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8590BD6E39ADBDF925" box="[1216,1272,1638,1664]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, D), an indistinct external orbital angle (
<figureCitation id="8FC02A1B6A12FF8596256ED4ABF8F901" box="[600,701,1674,1700]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8596B66ED4AA74F900" box="[715,817,1674,1701]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597436ED4AAC0F901" box="[830,901,1674,1700]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8597C36ED4AD2AF901" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[958,1135,1674,1700]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF85978C6ED5AD6EF901" box="[1009,1067,1674,1700]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, D), the relatively shallow and short cervical grooves (
<figureCitation id="8FC02A1B6A12FF85964E6EF0ABDDF96D" box="[563,664,1710,1736]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8596D86EF0AA49F96C" box="[677,780,1710,1737]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597646EF0AA25F96D" box="[793,864,1710,1736]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8597E46EF0AD0EF96D" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[921,1099,1710,1736]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8597B06EF1AD42F96D" box="[973,1031,1710,1736]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2A, D), the absence of a flagellum or with a relatively short flagellum on the TME (
<figureCitation id="8FC02A1B6A12FF8597486E8CAADFF949" box="[821,922,1746,1772]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">Figs. 3G</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597DA6E8CAA89F949" box="[935,972,1746,1772]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4D</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8597A56E8CAAB9F948" box="[984,1020,1746,1773]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5B</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8590496E8CADA1F949" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[1076,1252,1746,1772]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF85901A6E8DADE4F949" box="[1127,1185,1746,1772]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3B, N), the relatively smooth chelipeds (
<figureCitation id="8FC02A1B6A12FF8595A16EA8AB24F8B5" box="[476,609,1782,1808]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8596106EA8ABE8F8B5" box="[621,685,1782,1808]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3AF</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8596C46EA8AA45F8B5" box="[697,768,1782,1808]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="7" pageNumber="168">4AC</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF85974A6EA8AAA0F8B5" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[823,997,1782,1808]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8597176EA9AAE7F8B5" box="[874,930,1782,1808]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2AF), the relatively shorter male sternopleonal cavity reaching anteriorly to the level of mid-length of the cheliped coxae (
<figureCitation id="8FC02A1B6A12FF8590EF6F44ADB0F891" box="[1170,1269,1818,1844]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="7" pageNumber="168">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85917E6F44AC22F890" box="[1283,1383,1818,1845]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8591096F44ACD2F890" box="[1396,1431,1818,1845]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5C</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8594BE6F60A831F8FD" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[195,372,1854,1880]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF85948A6F61A875F8FD" box="[247,304,1854,1880]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 2C, F), the relatively stout G1 with a relatively shorter ultimate article, ca. 0.30.4 times the length of the penultimate article and the strongly convex outer margin of the penultimate article at the basal half (
<figureCitation id="8FC02A1B6A12FF8594E26FD8A847F805" box="[159,258,1926,1952]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8595736FD8A86AF804" box="[270,303,1926,1953]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8595476FD8A83BF805" box="[314,382,1926,1952]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="7" pageNumber="168">6BE</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8595CF6FD8AB1BF805" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[434,606,1926,1952]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8595996FD9AB59F805" box="[484,540,1926,1952]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3E, F, J, O), and the relatively shorter G2, ca. 0.60.7 times the length of the G1, with a very short ultimate article, ca. 0.2 times the length of the penultimate article (
<figureCitation id="8FC02A1B6A12FF8590C36FF4AC52F861" box="[1214,1303,1962,1988]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="7" pageNumber="168">Figs. 3I</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF85915E6FF4AC04F860" box="[1315,1345,1962,1989]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="7" pageNumber="168">5F</figureCitation>
,
<figureCitation id="8FC02A1B6A12FF8591316FF4AC28F861" box="[1356,1389,1962,1988]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="7" pageNumber="168">6F</figureCitation>
; see
<bibRefCitation id="736A4B6F6A12FF8594EA6F90A803F84D" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[151,326,1998,2024]" pageId="7" pageNumber="168" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A12FF8594B76F91A846F84D" box="[202,259,1998,2024]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3H, K, P). In addition to the shorter, elongated male telson and the stouter G1 ultimate article of the new genus,
<taxonomicName id="D0FB4D1D6A1DFF8A951C68C9A884FF14" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[353,449,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A951C68C9A884FF14" box="[353,449,151,177]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A95BA68C9AB75FF14" bold="true" box="[455,560,151,177]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A95BA68C9AB75FF14" box="[455,560,151,177]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is differentiated from
<taxonomicName id="D0FB4D1D6A1DFF8A975568C9AA81FF14" authorityName="Pati &amp; Sudha Devi" authorityYear="2015" box="[808,964,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A975568C9AA81FF14" box="[808,964,151,177]" italics="true" pageId="8" pageNumber="169">Arcithelphusa</emphasis>
</taxonomicName>
by the relatively narrow male pleon (
<figureCitation id="8FC02A1B6A1DFF8A911968C9A9FFFF70" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A94BB68E5A86CFF70" box="[198,297,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A954868E4A81CFF70" box="[309,345,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A951868E5A8CCFF70" box="[357,393,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6A</figureCitation>
) (vs. male pleon relatively broad; see
<bibRefCitation id="736A4B6F6A1DFF8A974468E5AAADFF70" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[825,1000,187,213]" pageId="8" pageNumber="169" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A1DFF8A971168E5AAE0FF70" box="[876,933,187,213]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2019
</bibRefCitation>
: figs. 2C, F, 3D), the relatively narrow male pleonal somite 6, the proximal width ca. 1.3 times the medial length (
<figureCitation id="8FC02A1B6A1DFF8A97936881AD17FF5C" box="[1006,1106,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90226881AD87FF5C" box="[1119,1218,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90B26880ADB1FF5C" box="[1231,1268,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A917D6881AC61FF5C" box="[1280,1316,223,249]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6A</figureCitation>
) (vs. male pleonal somite 6 relatively broad, the proximal width ca. 1.71.8 times the medial length; see
<bibRefCitation id="736A4B6F6A1DFF8A90CE695DAC25FEB8" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[1203,1376,259,285]" pageId="8" pageNumber="169" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A1DFF8A909B695DAC5BFEB8" box="[1254,1310,259,285]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2019
</bibRefCitation>
: figs. 2C, F, 3D), the relatively small flexible zone of the G1 (
<figureCitation id="8FC02A1B6A1DFF8A97766979AA2AFEE4" box="[779,879,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A97066978AAD9FEE4" box="[891,924,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A97D56979AAAEFEE4" box="[936,1003,295,321]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6B, C</figureCitation>
) (vs. G1 flexible zone relatively large; see
<bibRefCitation id="736A4B6F6A1DFF8A94BE6915A830FEC0" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[195,373,331,357]" pageId="8" pageNumber="169" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A1DFF8A948A6915A874FEC0" box="[247,305,331,357]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3E, J, O), and the gently curved ultimate article of the G1 (
<figureCitation id="8FC02A1B6A1DFF8A902F6915ADF2FEC0" box="[1106,1207,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90B96914ADA3FEC0" box="[1220,1254,330,357]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A908E6915AC7DFEC0" box="[1267,1336,331,357]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6B, D</figureCitation>
) (vs. G1 ultimate article strongly bent; see
<bibRefCitation id="736A4B6F6A1DFF8A96696931AB84FE2C" author="Pati, S. K. &amp; Sujila, P. S. &amp; Sudha Devi, A. R." box="[532,705,367,393]" pageId="8" pageNumber="169" pagination="203 - 214" refId="ref10745" refString="Pati, S. K., Sujila, P. S. &amp; Sudha Devi, A. R. (2019) Description of a new species of freshwater crab of the genus Arcithelphusa Pati &amp; Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India. Zootaxa, 4674 (2), 203 - 214. https: // doi. org / 10.11646 / zootaxa. 4674.2.2" type="journal article" year="2019">
Pati
<emphasis id="258FEA8C6A1DFF8A963B6931AB3AFE2C" box="[582,639,367,393]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2019
</bibRefCitation>
: fig. 3E, F, J, O).
<taxonomicName id="D0FB4D1D6A1DFF8A97FB6931AAA3FE2C" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[902,998,367,393]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A97FB6931AAA3FE2C" box="[902,998,367,393]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A97936931AD12FE2C" bold="true" box="[1006,1111,367,393]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A97936931AD12FE2C" box="[1006,1111,367,393]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is found at a lower elevation (
<quantity id="D0039B7B6A1DFF8A94E269CDA9ADFE08" box="[159,232,403,429]" metricMagnitude="2" metricUnit="m" metricValue="6.29" pageId="8" pageNumber="169" unit="m" value="629.0">629 m</quantity>
) on an isolated mountain, which is surrounded by the Kali and Gangavali rivers, with deep valleys (
<figureCitation id="8FC02A1B6A1DFF8A913769CDACCEFE08" box="[1354,1419,403,429]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="8" pageNumber="169">Fig. 1</figureCitation>
). The known congeners of
<taxonomicName id="D0FB4D1D6A1DFF8A95CB69E9AB17FE74" authorityName="Pati &amp; Sudha Devi" authorityYear="2015" box="[438,594,439,465]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A95CB69E9AB17FE74" box="[438,594,439,465]" italics="true" pageId="8" pageNumber="169">Arcithelphusa</emphasis>
</taxonomicName>
have been recorded from the Wayanad mountain plateau at slightly higher elevations (
<quantity id="D0039B7B6A1DFF8A956B6985A8D3FE50" box="[278,406,475,501]" metricMagnitude="2" metricUnit="m" metricValue="7.865" metricValueMax="8.64" metricValueMin="7.09" pageId="8" pageNumber="169" unit="m" value="786.5" valueMax="864.0" valueMin="709.0">709864 m</quantity>
) (
<figureCitation id="8FC02A1B6A1DFF8A95D16985A8A9FE50" box="[428,492,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="8" pageNumber="169">Fig. 1</figureCitation>
). The Wayanad plateau is some
<quantity id="D0039B7B6A1DFF8A97336984AAE7FE50" box="[846,930,474,501]" metricMagnitude="5" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="169" unit="km" value="350.0">350 km</quantity>
away from the Bare where
<taxonomicName id="D0FB4D1D6A1DFF8A90B26985AC6AFE50" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1231,1327,475,501]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A90B26985AC6AFE50" box="[1231,1327,475,501]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A91486985ACD9FE50" bold="true" box="[1333,1436,475,501]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A91486985ACD9FE50" box="[1333,1436,475,501]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
occurs. The Western Ghats between the Wayanad plateau and the Bare has some mountain peaks and deep valleys (
<figureCitation id="8FC02A1B6A1DFF8A94E26A7DA9A4FD98" box="[159,225,547,573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="8" pageNumber="169">Fig. 1</figureCitation>
), which form “sky islands” acting as barriers between these two genera. The morphological differences and the geographical isolation between them corroborate the recognition of
<taxonomicName id="D0FB4D1D6A1DFF8A97C46A19AD5CFDC4" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[953,1049,583,609]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A97C46A19AD5CFDC4" box="[953,1049,583,609]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A905D6A19ADCCFDC4" bold="true" box="[1056,1161,583,609]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A905D6A19ADCCFDC4" box="[1056,1161,583,609]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="1744369E6A1DFF8A94BA6A35A88CF8E8" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="169">
<taxonomicName id="D0FB4D1D6A1DFF8A94BA6A35A862FD20" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[199,295,619,645]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A94BA6A35A862FD20" box="[199,295,619,645]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A954E6A35A8E5FD20" bold="true" box="[307,416,619,645]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A954E6A35A8E5FD20" box="[307,416,619,645]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is further separated from
<taxonomicName id="D0FB4D1D6A1DFF8A96AB6A35AAD2FD20" box="[726,919,619,645]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A96AB6A35AAD2FD20" box="[726,919,619,645]" italics="true" pageId="8" pageNumber="169">Cylindrotelphusa</emphasis>
</taxonomicName>
and
<emphasis id="258FEA8C6A1DFF8A97AB6A35AD3DFD20" box="[982,1144,619,645]" italics="true" pageId="8" pageNumber="169">Rajathelphusa</emphasis>
by the generally smooth dorsal surface of the carapace (
<figureCitation id="8FC02A1B6A1DFF8A95886AD1AB1CFD0C" box="[501,601,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96186AD1AB8FFD0C" box="[613,714,655,681]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96AB6AD1AA5EFD0C" box="[726,795,655,681]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
) (vs. carapace dorsal surface relatively rugose; see
<bibRefCitation id="736A4B6F6A1DFF8A91236AD1A860FD68" author="Bahir, M. M. &amp; Yeo, D. C. J." pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5A;
<bibRefCitation id="736A4B6F6A1DFF8A95EE6AEDAB03FD68" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[403,582,691,717]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A95BA6AEDAB44FD68" box="[455,513,691,717]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11a, 13a;
<bibRefCitation id="736A4B6F6A1DFF8A96816AEDAAEFFD68" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[764,938,691,717]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A97516AEDAA23FD68" box="[812,870,691,717]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, 7A;
<bibRefCitation id="736A4B6F6A1DFF8A90286AEDAC46FD68" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1109,1283,691,717]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A90F96AEDADFBFD68" box="[1156,1214,691,717]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 8A); the relatively shallow cervical grooves (
<figureCitation id="8FC02A1B6A1DFF8A964F6A89ABD2FD54" box="[562,663,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96D96A89AA4FFD54" box="[676,778,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A976A6A89AA1EFD54" box="[791,859,727,753]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
) (vs. cervical grooves relatively deep; see
<bibRefCitation id="736A4B6F6A1DFF8A913D6A89A840FCB0" author="Bahir, M. M. &amp; Yeo, D. C. J." pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5A;
<bibRefCitation id="736A4B6F6A1DFF8A95106AA5AB5CFCB0" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[365,537,763,789]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A95E26AA5A892FCB0" box="[415,471,763,789]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11a, 13a;
<bibRefCitation id="736A4B6F6A1DFF8A96BB6AA5AA28FCB0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[710,877,763,789]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A968E6AA5AA6EFCB0" box="[755,811,763,789]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, 7A;
<bibRefCitation id="736A4B6F6A1DFF8A90726AA5ADF3FCB0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1039,1206,763,789]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A90406AA5AD31FCB0" box="[1085,1140,763,789]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 8A); the lack of a flagellum or at most with a shorter flagellum on the TME, which is about half the width of the merus (
<figureCitation id="8FC02A1B6A1DFF8A914E6B41ACD2FC9C" box="[1331,1431,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3G</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A94EA6B1DA9F9FCF8" box="[151,188,835,861]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4D</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A94BB6B1CA9ADFCF8" box="[198,232,834,861]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5B</figureCitation>
) (vs. flagellum on the TME relatively long, reaching beyond half the width of the merus; see
<bibRefCitation id="736A4B6F6A1DFF8A90936B1DACDDFCF8" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[1262,1432,835,861]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A915D6B1DAC12FCF8" box="[1312,1367,835,861]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11i, 13i;
<bibRefCitation id="736A4B6F6A1DFF8A95496B39A89AFC24" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[308,479,871,897]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A951E6B39A8D9FC24" box="[355,412,871,897]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3C, 7D;
<bibRefCitation id="736A4B6F6A1DFF8A96F96B39AA6AFC24" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[644,815,871,897]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A96CE6B39ABAEFC24" box="[691,747,871,897]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 9A); the relatively smooth chelipeds (
<figureCitation id="8FC02A1B6A1DFF8A916F6B39ACD2FC24" box="[1298,1431,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A94EA6BD5A992FC00" box="[151,215,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3AF</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A949F6BD5A863FC00" box="[226,294,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
) (vs. chelipeds relatively rugose; see
<bibRefCitation id="736A4B6F6A1DFF8A96B86BD5AAD7FC00" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[709,914,907,933]" pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5AC;
<bibRefCitation id="736A4B6F6A1DFF8A90656BD5AD86FC00" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[1048,1219,907,933]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A90376BD5ADC4FC00" box="[1098,1153,907,933]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11a, c, d, 13a, c, d;
<bibRefCitation id="736A4B6F6A1DFF8A94AD6BF1A83EFC6C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[208,379,943,969]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A94836BF1A872FC6C" box="[254,311,943,969]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, 4G, 7A, G;
<bibRefCitation id="736A4B6F6A1DFF8A960B6BF1AA64FC6C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[630,801,943,969]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A96D96BF1AB98FC6C" box="[676,733,943,969]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: figs. 8A, 9D); the gently concave lateral margins of the male telson (
<figureCitation id="8FC02A1B6A1DFF8A95546B8DA8C9FC48" box="[297,396,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A95E46B8DA8B9FC48" box="[409,508,979,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96746B8CAB68FC48" box="[521,557,978,1005]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96476B8DAB1BFC48" box="[570,606,979,1005]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6A</figureCitation>
) (vs. male telson with strongly concave lateral margins; see
<bibRefCitation id="736A4B6F6A1DFF8A91706B8DA996FBB4" author="Bahir, M. M. &amp; Yeo, D. C. J." pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5C;
<bibRefCitation id="736A4B6F6A1DFF8A95416BA9A8AEFBB4" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[316,491,1015,1041]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A95126BA9A8EDFBB4" box="[367,424,1015,1041]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11h, 13h;
<bibRefCitation id="736A4B6F6A1DFF8A96E26BA9AA0FFBB4" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[671,842,1015,1041]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A96B36BA9AA43FBB4" box="[718,774,1015,1041]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3D, 7E;
<bibRefCitation id="736A4B6F6A1DFF8A97916BA9ADD2FBB4" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1004,1175,1015,1041]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A90666BA9AD11FBB4" box="[1051,1108,1015,1041]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 9C); the relatively stout G1 (
<figureCitation id="8FC02A1B6A1DFF8A957A6C45A829FB90" box="[263,364,1051,1077]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A95056C44A8DFFB90" box="[376,410,1050,1077]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A95DB6C45A8AFFB90" box="[422,490,1051,1077]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6B, D</figureCitation>
) (vs. G1 relatively slender; see
<bibRefCitation id="736A4B6F6A1DFF8A972F6C45AD61FB90" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[850,1060,1051,1077]" pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 4A, B;
<bibRefCitation id="736A4B6F6A1DFF8A90CD6C45AC25FB90" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[1200,1376,1051,1077]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A909E6C45AC59FB90" box="[1251,1308,1051,1077]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 12a, c, 14a, c;
<bibRefCitation id="736A4B6F6A1DFF8A95406C61A8ACFBFC" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[317,489,1087,1113]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A95116C61A8E0FBFC" box="[364,421,1087,1113]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 9B, F;
<bibRefCitation id="736A4B6F6A1DFF8A96016C61AA62FBFC" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[636,807,1087,1113]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A96D66C61ABA1FBFC" box="[683,740,1087,1113]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 10A); and the relatively stouter penultimate article of the G1, with the outer margin being strongly convex and shelf-like at the basal half (
<figureCitation id="8FC02A1B6A1DFF8A90016C3DADA4FBD8" box="[1148,1249,1123,1149]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90926C3CAC55FBD8" box="[1263,1296,1122,1149]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A91606C3DAC27FBD8" box="[1309,1378,1123,1149]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6B, D</figureCitation>
) (vs. G1 penultimate article relatively slenderer, with the outer margin straight to relatively less convex at the basal half; see
<bibRefCitation id="736A4B6F6A1DFF8A94BF6CF5A8D6FB60" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[194,403,1195,1221]" pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 4A, B;
<bibRefCitation id="736A4B6F6A1DFF8A96636CF5AB89FB60" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[542,716,1195,1221]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A962C6CF5ABCCFB60" box="[593,649,1195,1221]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 12a, c, 14a, c;
<bibRefCitation id="736A4B6F6A1DFF8A97C96CF5AD18FB60" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[948,1117,1195,1221]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A979F6CF5AD5FFB60" box="[994,1050,1195,1221]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 9B, F;
<bibRefCitation id="736A4B6F6A1DFF8A90926CF5ACDDFB60" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1263,1432,1195,1221]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A91606CF5AC10FB60" box="[1309,1365,1195,1221]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 10A). The anterolateral margins of the carapace lack distinct serrations (
<figureCitation id="8FC02A1B6A1DFF8A97976C91AD08FB4C" box="[1002,1101,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90246C91ADF9FB4C" box="[1113,1212,1231,1257]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90B56C91AC4AFB4C" box="[1224,1295,1231,1257]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
), and the G1 ultimate article is relatively shorter, ca. 0.3 times the length of the penultimate article (
<figureCitation id="8FC02A1B6A1DFF8A90286CADADFDFAA8" box="[1109,1208,1267,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90BE6CACADA0FAA8" box="[1219,1253,1266,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A90926CADAC54FAA8" box="[1263,1297,1267,1293]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6B</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1DFF8A91416CADACD9FAA8" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1340,1436,1267,1293]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A91416CADACD9FAA8" box="[1340,1436,1267,1293]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A94EA6D49A9BAFA94" bold="true" box="[151,255,1303,1329]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A94EA6D49A9BAFA94" box="[151,255,1303,1329]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
; whereas the anterolateral margins have distinct serrations (see
<bibRefCitation id="736A4B6F6A1DFF8A97C36D49ADCFFA94" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[958,1162,1303,1329]" pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 5A;
<bibRefCitation id="736A4B6F6A1DFF8A90926D49ACDDFA94" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[1263,1432,1303,1329]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A915D6D49AC12FA94" box="[1312,1367,1303,1329]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 11a, 13a), and the G1 ultimate article is relatively longer, ca. 0.40.6 times the length of the penultimate article (see
<bibRefCitation id="736A4B6F6A1DFF8A94B66D01A8DEFADC" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[203,411,1375,1401]" pageId="8" pageNumber="169" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 4A;
<bibRefCitation id="736A4B6F6A1DFF8A96796D01ABF4FADC" author="Pati, S. K. &amp; Rajesh, L. &amp; Raj, S. &amp; Sheeja, V. U. &amp; Kumar, A. B. &amp; Sureshan, P. M." box="[516,689,1375,1401]" pageId="8" pageNumber="169" pagination="1295 - 1330" refId="ref10644" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. &amp; Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51, 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
Pati
<emphasis id="258FEA8C6A1DFF8A964B6D01AB2AFADC" box="[566,623,1375,1401]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2017
</bibRefCitation>
: figs. 12a, 14a) in
<taxonomicName id="D0FB4D1D6A1DFF8A97FD6D01AD04FADC" box="[896,1089,1375,1401]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A97FD6D01AD04FADC" box="[896,1089,1375,1401]" italics="true" pageId="8" pageNumber="169">Cylindrotelphusa</emphasis>
</taxonomicName>
. The epigastric and postorbital cristae are poorly developed (
<figureCitation id="8FC02A1B6A1DFF8A95956DDDAB08FA38" box="[488,589,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96276DDDAB85FA38" box="[602,704,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A96B06DDDAA51FA38" box="[717,788,1411,1437]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
), the external orbital angle is indistinct (
<figureCitation id="8FC02A1B6A1DFF8A90A26DDDAC01FA38" box="[1247,1348,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A912C6DDDA9EFFA64" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A94C56DF9A9BBFA64" box="[184,254,1447,1473]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
), and the G2 is relatively shorter, ca. 0.6 times the length of G1, with the ultimate article very short, ca. 0.2 times the length of the penultimate article (
<figureCitation id="8FC02A1B6A1DFF8A96D36D95AA42FA40" box="[686,775,1483,1509]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">Figs. 3I</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A97686D94AA76FA40" box="[789,819,1482,1509]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="8" pageNumber="169">5F</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A97426D95AA1BFA40" box="[831,862,1483,1509]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="8" pageNumber="169">6F</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1DFF8A97F06D95AAA8FA40" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[909,1005,1483,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A97F06D95AAA8FA40" box="[909,1005,1483,1509]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A978B6D95AD25FA40" bold="true" box="[1014,1120,1483,1509]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A978B6D95AD25FA40" box="[1014,1120,1483,1509]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
; whereas the epigastric and postorbital cristae are well-developed (see
<bibRefCitation id="736A4B6F6A1DFF8A96046DB1AA66F9AC" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[633,803,1519,1545]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A96DA6DB1ABA5F9AC" box="[679,736,1519,1545]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, 4A, B, 7A, B;
<bibRefCitation id="736A4B6F6A1DFF8A90416DB1ADA3F9AC" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1084,1254,1519,1545]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A90176DB1ADE6F9AC" box="[1130,1187,1519,1545]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 8A, B), the external orbital angle is distinct (see
<bibRefCitation id="736A4B6F6A1DFF8A96466E4DABA2F988" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[571,743,1555,1581]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A96176E4DABE6F988" box="[618,675,1555,1581]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: figs. 3A, 4A, B, 7A, B;
<bibRefCitation id="736A4B6F6A1DFF8A907B6E4DADF7F988" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1030,1202,1555,1581]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A90486E4DAD2BF988" box="[1077,1134,1555,1581]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 8A, B), and the G2 is relatively longer, ca. 1.0 times the length of G1, with the ultimate article long, ca. 0.5 times the length of the penultimate article (see
<bibRefCitation id="736A4B6F6A1DFF8A95D46E05AB1DF9D0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[425,600,1627,1653]" pageId="8" pageNumber="169" pagination="14" refId="ref10903" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2021) Description of a new genus with two new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India. Zoological Studies, 60, 14. https: // doi. org / 10.6620 / ZS. 2021.60 - 14" type="journal article" year="2021">
Raj
<emphasis id="258FEA8C6A1DFF8A95A46E05AB56F9D0" box="[473,531,1627,1653]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
: fig. 9E, I;
<bibRefCitation id="736A4B6F6A1DFF8A96A06E05AAC9F9D0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[733,908,1627,1653]" pageId="8" pageNumber="169" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1DFF8A97706E05AA02F9D0" box="[781,839,1627,1653]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
2022
</bibRefCitation>
: fig. 10D) in
<emphasis id="258FEA8C6A1DFF8A90546E05AD8EF9D0" box="[1065,1227,1627,1653]" italics="true" pageId="8" pageNumber="169">Rajathelphusa</emphasis>
.
<taxonomicName id="D0FB4D1D6A1DFF8A90A66E05ACD9F9D0" box="[1243,1436,1627,1653]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A90A66E05ACD9F9D0" box="[1243,1436,1627,1653]" italics="true" pageId="8" pageNumber="169">Cylindrotelphusa</emphasis>
</taxonomicName>
is known to dwell in both lower and higher elevations (
<quantity id="D0039B7B6A1DFF8A97736E21AA32F93C" box="[782,887,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="4.915" metricValueMax="9.8" metricValueMin="0.03" pageId="8" pageNumber="169" unit="m" value="491.5" valueMax="980.0" valueMin="3.0">3980 m</quantity>
), but it is restricted to the southern Indian states of
<collectingRegion id="D53FF87C6A1DFF8A94CB6EFDA844F918" box="[182,257,1699,1725]" country="India" name="Kerala" pageId="8" pageNumber="169">Kerala</collectingRegion>
and
<collectingRegion id="D53FF87C6A1DFF8A95456EFDA8FBF918" box="[312,446,1699,1725]" country="India" name="Tamil Nadu" pageId="8" pageNumber="169">Tamil Nadu</collectingRegion>
(
<figureCitation id="8FC02A1B6A1DFF8A95B36EFDAB55F918" box="[462,528,1699,1725]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="8" pageNumber="169">Fig. 1</figureCitation>
).
<taxonomicName id="D0FB4D1D6A1DFF8A96546EFDABAFF918" box="[553,746,1699,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A96546EFDABAFF918" box="[553,746,1699,1725]" italics="true" pageId="8" pageNumber="169">Cylindrotelphusa</emphasis>
</taxonomicName>
is probably not known beyond these two states, and this fact along with the morphological differences between
<taxonomicName id="D0FB4D1D6A1DFF8A96A76E99AADEF944" box="[730,923,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1DFF8A96A76E99AADEF944" box="[730,923,1735,1761]" italics="true" pageId="8" pageNumber="169">Cylindrotelphusa</emphasis>
</taxonomicName>
and
<taxonomicName id="D0FB4D1D6A1DFF8A97A86E99AD70F944" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[981,1077,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A97A86E99AD70F944" box="[981,1077,1735,1761]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A90436E99ADECF944" bold="true" box="[1086,1193,1735,1761]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A90436E99ADECF944" box="[1086,1193,1735,1761]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
support their generic separation. On the other hand,
<emphasis id="258FEA8C6A1DFF8A967F6EB5ABE1F8A0" box="[514,676,1771,1797]" italics="true" pageId="8" pageNumber="169">Rajathelphusa</emphasis>
occurs at high mountains (
<quantity id="D0039B7B6A1DFF8A97996EB4AD33F8A0" box="[996,1142,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.1865" metricValueMax="1.623" metricValueMin="0.75" pageId="8" pageNumber="169" unit="m" value="1186.5" valueMax="1623.0" valueMin="750.0">7501623 m</quantity>
altitude) of the Southern Western Ghats and is geographically clearly isolated from
<taxonomicName id="D0FB4D1D6A1DFF8A97546F51AACCF88C" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[809,905,1807,1833]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A97546F51AACCF88C" box="[809,905,1807,1833]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A97EC6F51AABEF88C" bold="true" box="[913,1019,1807,1833]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A97EC6F51AABEF88C" box="[913,1019,1807,1833]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
of the Central Western Ghats mainly by the Palghat gap (
<figureCitation id="8FC02A1B6A1DFF8A950B6F6DA8FDF8E8" box="[374,440,1843,1869]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="8" pageNumber="169">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1DFF8B94BA6F09AD69FCF8" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,150,1797]" lastPageId="9" lastPageNumber="170" pageId="8" pageNumber="169">
The superficial resemblance of
<taxonomicName id="D0FB4D1D6A1DFF8A96526F09ABCAF8D4" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[559,655,1879,1905]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A96526F09ABCAF8D4" box="[559,655,1879,1905]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A96E56F09AA46F8D4" bold="true" box="[664,771,1879,1905]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A96E56F09AA46F8D4" box="[664,771,1879,1905]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
with
<emphasis id="258FEA8C6A1DFF8A973B6F09AAF1F8D4" box="[838,948,1879,1905]" italics="true" pageId="8" pageNumber="169">Pavizham</emphasis>
notwithstanding, the new genus is distinct from the latter genus mainly by the elongated male telson and the stouter G1 ultimate article. Other important differences between them are as follows: the carapace is deeper and strongly arched dorsally,
<collectingCountry id="6FEC760E6A1DFF8A90B66FC1ADB4F81C" box="[1227,1265,1951,1977]" name="Switzerland" pageId="8" pageNumber="169">CH</collectingCountry>
/CW = 0.60.7 (
<figureCitation id="8FC02A1B6A1DFF8A94E26F9DA860F878" box="[159,293,1987,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="8" pageNumber="169">Figs. 2A, B</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A954C6F9DA8D3F878" box="[305,406,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="8" pageNumber="169">3A, C, E</figureCitation>
,
<figureCitation id="8FC02A1B6A1DFF8A95DF6F9DA8A3F878" box="[418,486,1987,2013]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="8" pageNumber="169">4AC</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1DFF8A966E6F9DAB36F878" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[531,627,1987,2013]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1DFF8A966E6F9DAB36F878" box="[531,627,1987,2013]" italics="true" pageId="8" pageNumber="169">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1DFF8A96066F9DABA1F878" bold="true" box="[635,740,1987,2013]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="3EBC57F76A1DFF8A96066F9DABA1F878" box="[635,740,1987,2013]" pageId="8" pageNumber="169" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. carapace less deep and gently arched dorsally,
<collectingCountry id="6FEC760E6A1DFF8A91576F9DAC15F878" box="[1322,1360,1987,2013]" name="Switzerland" pageId="8" pageNumber="169">CH</collectingCountry>
/CW =
<quantity id="D0039B7B6A1CFF8B94EA68C8A99DFF14" box="[151,216,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="170" unit="in" value="0.5">0.5 in</quantity>
<emphasis id="258FEA8C6A1CFF8B94A268C9A808FF14" box="[223,333,151,177]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B95FA68C9AB74FF14" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[391,561,151,177]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B95CB68C9A8ABFF14" box="[438,494,151,177]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: fig. 3A, C); the flagellum on the TME is missing or with a shorter flagellum, which reaches about half the width of the merus (
<figureCitation id="8FC02A1B6A1CFF8B96C468E5AA5EFF70" box="[697,795,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">Figs. 3G</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B975B68E5AA0EFF70" box="[806,843,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="9" pageNumber="170">4D</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B972B68E4AA32FF70" box="[854,887,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5B</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B97DF68E5AD47FF70" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[930,1026,187,213]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B97DF68E5AD47FF70" box="[930,1026,187,213]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B907568E5AD35FF70" bold="true" box="[1032,1136,187,213]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B907568E5AD35FF70" box="[1032,1136,187,213]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. flagellum on the TME almost as long as the width of the merus in
<emphasis id="258FEA8C6A1CFF8B96E96881AA47FF5C" box="[660,770,223,249]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B973D6881AAB5FF5C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[832,1008,223,249]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B970C6881AAEEFF5C" box="[881,939,223,249]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: fig. 5A); the chelipeds are relatively smooth (
<figureCitation id="8FC02A1B6A1CFF8B9487695DA8C4FEB8" box="[250,385,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B95F3695DA88BFEB8" box="[398,462,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">3AF</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B95A6695DAB5AFEB8" box="[475,543,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="9" pageNumber="170">4AC</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B9630695DABE8FEB8" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[589,685,259,285]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B9630695DABE8FEB8" box="[589,685,259,285]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B96CB695DAA65FEB8" bold="true" box="[694,800,259,285]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B96CB695DAA65FEB8" box="[694,800,259,285]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. chelipeds relatively rugose in
<emphasis id="258FEA8C6A1CFF8B90D2695DAC58FEB8" box="[1199,1309,259,285]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B9124695DA9B2FEE4" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B91F5695DA9F1FEE4" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: figs. 3A, 4D); the male s2/s3 does not reach the edges of the sternum (
<figureCitation id="8FC02A1B6A1CFF8B90696979AD33FEE4" box="[1044,1142,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B90FF6979ADA6FEE4" box="[1154,1251,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B90936978AC55FEE4" box="[1262,1296,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5C</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B91416979ACD9FEE4" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1340,1436,295,321]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B91416979ACD9FEE4" box="[1340,1436,295,321]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B94EA6915A845FEC0" bold="true" box="[151,256,331,357]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B94EA6915A845FEC0" box="[151,256,331,357]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. male s2/s3 reaching the edges of the sternum in
<emphasis id="258FEA8C6A1CFF8B972D6915AAFBFEC0" box="[848,958,331,357]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B97856915ADE5FEC0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1016,1184,331,357]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B905B6915AD1BFEC0" box="[1062,1118,331,357]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: figs. 3D, 4B, 5B); the male sternopleonal cavity is relatively shorter, reaching anteriorly to the level of mid-length of the cheliped coxae (
<figureCitation id="8FC02A1B6A1CFF8B94E269CDA844FE08" box="[159,257,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B957169CDA82EFE08" box="[268,363,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B950869CCA8D2FE08" box="[373,407,402,429]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5C</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B95BC69CDAB64FE08" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[449,545,403,429]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B95BC69CDAB64FE08" box="[449,545,403,429]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B965569CDABD5FE08" bold="true" box="[552,656,403,429]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B965569CDABD5FE08" box="[552,656,403,429]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. male sternopleonal cavity relatively longer, reaching anteriorly to the level of the anterior margin of the cheliped coxae in
<emphasis id="258FEA8C6A1CFF8B977E69E9AA34FE74" box="[771,881,439,465]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B97D769E9AD15FE74" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[938,1104,439,465]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B97AA69E9AD4BFE74" box="[983,1038,439,465]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: figs. 3D, 4B); the male pleon is relatively narrow, with the pleonal somite 6 being relatively narrow, the proximal width ca. 1.3 times the medial length (
<figureCitation id="8FC02A1B6A1CFF8B949369A1A817FDBC" box="[238,338,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B951D69A1A880FDBC" box="[352,453,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">3B, D, F</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B95AF69A0A8B2FDBC" box="[466,503,510,537]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B967969A1AB6DFDBC" box="[516,552,511,537]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">6A</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B962A69A1ABF2FDBC" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[599,695,511,537]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B962A69A1ABF2FDBC" box="[599,695,511,537]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B96BC69A1AA69FDBC" bold="true" box="[705,812,511,537]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B96BC69A1AA69FDBC" box="[705,812,511,537]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. male pleon relatively broad, with relatively broad pleonal somite 6, the proximal width ca. 1.7 times the medial length in
<emphasis id="258FEA8C6A1CFF8B97D66A7DAD5CFD98" box="[939,1049,547,573]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B902F6A7DADBDFD98" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[1106,1272,547,573]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B90026A7DADF3FD98" box="[1151,1206,547,573]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: figs. 3D, 4A); the outer margin of the G1 penultimate article is strongly convex and shelf-like at the basal half (
<figureCitation id="8FC02A1B6A1CFF8B90AA6A19AC7EFDC4" box="[1239,1339,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">Figs. 3H</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B913A6A18AC2CFDC4" box="[1351,1385,582,609]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B91096A19A9EEFD20" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">6B, C</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B94A56A35A87DFD20" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[216,312,619,645]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B94A56A35A87DFD20" box="[216,312,619,645]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B95426A35A8ECFD20" bold="true" box="[319,425,619,645]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B95426A35A8ECFD20" box="[319,425,619,645]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. G1 penultimate article with a less convex outer margin at the basal half in
<emphasis id="258FEA8C6A1CFF8B915B6A35ACD1FD20" box="[1318,1428,619,645]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B94BE6AD1A834FD0C" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[195,369,655,681]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B948E6AD1A868FD0C" box="[243,301,655,681]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: fig. 5CE); and the G2 is short, ca. 0.6 times the length of G1, with the ultimate article very short, ca. 0.2 times the length of the penultimate article (
<figureCitation id="8FC02A1B6A1CFF8B97706AEDAA20FD68" box="[781,869,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">Figs. 3I</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B970C6AECAACAFD68" box="[881,911,690,717]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5F</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B97E76AEDAAFCFD68" box="[922,953,691,717]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">6F</figureCitation>
) in
<taxonomicName id="D0FB4D1D6A1CFF8B97996AEDAD01FD68" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[996,1092,691,717]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B97996AEDAD01FD68" box="[996,1092,691,717]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B90366AEDADF1FD68" bold="true" box="[1099,1204,691,717]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B90366AEDADF1FD68" box="[1099,1204,691,717]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(vs. G2 long, ca. 1.4 times the length of G1, with the ultimate article long, ca. 0.6 times the length of the penultimate article in
<emphasis id="258FEA8C6A1CFF8B915B6A89ACD1FD54" box="[1318,1428,727,753]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B94BF6AA5A829FCB0" author="Raj, S. &amp; Kumar, A. B. &amp; Ng, P. K. L." box="[194,364,763,789]" pageId="9" pageNumber="170" pagination="49" refId="ref10973" refString="Raj, S., Kumar, A. B. &amp; Ng, P. K. L. (2022) New genus and two new species of semiterrestrial crabs (Decapoda: Brachyura: Gecarcinucidae) from the southern Western Ghats, India. Zoological Studies, 61, 49. https: // doi. org / 10.6620 / ZS. 2022.61 - 49" type="journal article" year="2022">
Raj
<emphasis id="258FEA8C6A1CFF8B948D6AA5A86CFCB0" box="[240,297,763,789]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
: fig. 5F).
<taxonomicName id="D0FB4D1D6A1CFF8B95A46AA5AB7CFCB0" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[473,569,763,789]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B95A46AA5AB7CFCB0" box="[473,569,763,789]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B963C6AA5ABEFFCB0" bold="true" box="[577,682,763,789]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B963C6AA5ABEFFCB0" box="[577,682,763,789]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is also separated from
<emphasis id="258FEA8C6A1CFF8B97D36AA5AD59FCB0" box="[942,1052,763,789]" italics="true" pageId="9" pageNumber="170">Pavizham</emphasis>
geographically as both genera are apart from each other by a distance of about
<quantity id="D0039B7B6A1CFF8B96E96B40ABAFFC9C" box="[660,746,798,825]" metricMagnitude="5" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="170" unit="km" value="650.0">650 km</quantity>
with several mountain peaks and deep valleys, including the prominent Palghat gap between the Central- and Southern Western Ghats (
<figureCitation id="8FC02A1B6A1CFF8B97A76B1DAD5EFCF8" box="[986,1051,835,861]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="9" pageNumber="170">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1CFF8B94BA6B39AA68F918" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="170">
Crabs of the following genera are likely to coexist with
<taxonomicName id="D0FB4D1D6A1CFF8B97096B39AA91FC24" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[884,980,871,897]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B97096B39AA91FC24" box="[884,980,871,897]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B979C6B39AD15FC24" bold="true" box="[993,1104,871,897]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B979C6B39AD15FC24" box="[993,1104,871,897]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
:
<taxonomicName id="D0FB4D1D6A1CFF8B90186B39ACDDFC24" authority="Pati &amp; Yeo, 2022" authorityName="Pati &amp; Yeo" authorityYear="2022" box="[1125,1432,871,897]" class="Malacostraca" family="Gecarcinucidae" genus="Barusa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1CFF8B90186B39ADF3FC25" box="[1125,1206,871,896]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
<bibRefCitation id="736A4B6F6A1CFF8B90BE6B39ACDDFC24" author="Pati, S. K. &amp; Yeo, D. C. J." box="[1219,1432,871,897]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo, 2022</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D0FB4D1D6A1CFF8B94EA6BD5A88EFC00" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[151,459,907,933]" class="Malacostraca" family="Gecarcinucidae" genus="Barytelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1CFF8B94EA6BD5A869FC00" box="[151,300,907,933]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
Alcock, 1909
</taxonomicName>
;
<taxonomicName id="D0FB4D1D6A1CFF8B95AB6BD5AA71FC00" authority="Pati &amp; Sharma, 2014" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[470,820,907,933]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1CFF8B95AB6BD5AB05FC00" box="[470,576,907,933]" italics="true" pageId="9" pageNumber="170">Ghatiana</emphasis>
Pati &amp; Sharma, 2014
</taxonomicName>
;
<taxonomicName id="D0FB4D1D6A1CFF8B97426BD5AD1FFC00" authority="Bahir &amp; Yeo, 2007" authorityName="Bahir &amp; Yeo" authorityYear="2007" box="[831,1114,907,933]" class="Malacostraca" family="Gecarcinucidae" genus="Vanni" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1CFF8B97426BD5AA3AFC01" box="[831,895,907,932]" italics="true" pageId="9" pageNumber="170">Vanni</emphasis>
<bibRefCitation id="736A4B6F6A1CFF8B97F86BD5AD1FFC00" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[901,1114,907,933]" pageId="9" pageNumber="170" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo, 2007</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="D0FB4D1D6A1CFF8B90E96BD5ACD9FC00" authority="Bahir &amp; Yeo, 2007" authorityName="Bahir &amp; Yeo" authorityYear="2007" box="[1172,1436,907,933]" class="Malacostraca" family="Gecarcinucidae" genus="Vela" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="258FEA8C6A1CFF8B90E96BD5AD81FC00" box="[1172,1220,907,933]" italics="true" pageId="9" pageNumber="170">Vela</emphasis>
<bibRefCitation id="736A4B6F6A1CFF8B90B66BD5ACD9FC00" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[1227,1436,907,933]" pageId="9" pageNumber="170" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo, 2007</bibRefCitation>
</taxonomicName>
(cf.
<bibRefCitation id="736A4B6F6A1CFF8B94BE6BF1A883FC6C" author="Pati, S. K. &amp; Thackeray, T." box="[195,454,943,969]" pageId="9" pageNumber="170" pagination="627 - 647" refId="ref10357" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
;
<bibRefCitation id="736A4B6F6A1CFF8B95AD6BF1ABCFFC6C" author="Pati, S. K. &amp; Yeo, D. C. J." box="[464,650,943,969]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
;
<bibRefCitation id="736A4B6F6A1CFF8B96EE6BF1AA0DFC6C" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[659,840,943,969]" pageId="9" pageNumber="170" pagination="2022019" refId="ref10826" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="258FEA8C6A1CFF8B96B96BF1ABBEFC6C" box="[708,763,943,969]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022a
</bibRefCitation>
,
<bibRefCitation id="736A4B6F6A1CFF8B972F6BF1AAC8FC6C" author="Pati, S. K. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[850,909,943,969]" pageId="9" pageNumber="170" pagination="112 - 126" refId="ref10499" refString="Pati, S. K., Bajantri, P. P. &amp; Hegde, G. D. (2023) A new freshwater crab species of the genus Vela Bahir &amp; Yeo, 2007 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats of Karnataka, India. Zootaxa, 5239, 112 - 126. https: // doi. org / 10.11646 / zootaxa. 5239.1.5" type="journal article" year="2023">2023</bibRefCitation>
; unpublished data).
<taxonomicName id="D0FB4D1D6A1CFF8B90176BF1AD8FFC6C" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1130,1226,943,969]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B90176BF1AD8FFC6C" box="[1130,1226,943,969]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B90AC6BF1AC7DFC6C" bold="true" box="[1233,1336,943,969]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B90AC6BF1AC7DFC6C" box="[1233,1336,943,969]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
need not be confused with those genera because its carapace is relatively deep, with the front relatively narrow (
<figureCitation id="8FC02A1B6A1CFF8B91726B8DAC21FC48" box="[1295,1380,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Fig. 2B</figureCitation>
) (vs. carapace relatively low, with the front relatively wider; see
<bibRefCitation id="736A4B6F6A1CFF8B97556BA9AAB0FBB4" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[808,1013,1015,1041]" pageId="9" pageNumber="170" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 32B;
<bibRefCitation id="736A4B6F6A1CFF8B901A6BA9AC29FBB4" author="Pati, S. K. &amp; Thackeray, T." box="[1127,1388,1015,1041]" pageId="9" pageNumber="170" pagination="627 - 647" refId="ref10357" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
: fig. 4C;
<bibRefCitation id="736A4B6F6A1CFF8B94BA6C45A8C3FB90" author="Pati, S. K. &amp; Yeo, D. C. J." box="[199,390,1051,1077]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: figs. 1B, 10B;
<bibRefCitation id="736A4B6F6A1CFF8B964A6C45ABB6FB90" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[567,755,1051,1077]" pageId="9" pageNumber="170" pagination="2022019" refId="ref10826" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="258FEA8C6A1CFF8B96176C45ABE7FB90" box="[618,674,1051,1077]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022a
</bibRefCitation>
: fig. 3B;
<bibRefCitation id="736A4B6F6A1CFF8B97266C45AD4FFB90" author="Pati, S. K. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[859,1034,1051,1077]" pageId="9" pageNumber="170" pagination="112 - 126" refId="ref10499" refString="Pati, S. K., Bajantri, P. P. &amp; Hegde, G. D. (2023) A new freshwater crab species of the genus Vela Bahir &amp; Yeo, 2007 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats of Karnataka, India. Zootaxa, 5239, 112 - 126. https: // doi. org / 10.11646 / zootaxa. 5239.1.5" type="journal article" year="2023">
Pati
<emphasis id="258FEA8C6A1CFF8B97F36C45AA82FB90" box="[910,967,1051,1077]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2023
</bibRefCitation>
: fig. 2B). In
<taxonomicName id="D0FB4D1D6A1CFF8B90E76C45ADBFFB90" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[1178,1274,1051,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B90E76C45ADBFFB90" box="[1178,1274,1051,1077]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B917F6C45AC29FB90" bold="true" box="[1282,1388,1051,1077]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B917F6C45AC29FB90" box="[1282,1388,1051,1077]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, the epigastric and postorbital cristae are poorly developed (
<figureCitation id="8FC02A1B6A1CFF8B96856C61AA0BFBFC" box="[760,846,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Fig. 2A</figureCitation>
) (vs. epigastric and postorbital cristae well developed in
<emphasis id="258FEA8C6A1CFF8B94C86C3DA843FBD9" box="[181,262,1123,1148]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B95406C3DA897FBD8" box="[317,466,1123,1149]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B96706C3DAB89FBD8" author="Pati, S. K. &amp; Yeo, D. C. J." box="[525,716,1123,1149]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: figs. 1A, 10A), the external orbital angle is indistinct (
<figureCitation id="8FC02A1B6A1CFF8B91416C3DACD0FBD8" box="[1340,1429,1123,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="9" pageNumber="170">Fig. 2A</figureCitation>
) (vs. external orbital angle distinct in
<emphasis id="258FEA8C6A1CFF8B96486CD9ABC3FB05" box="[565,646,1159,1184]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B96C06CD9AA17FB04" box="[701,850,1159,1185]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B97F06CD9AD0EFB04" author="Pati, S. K. &amp; Yeo, D. C. J." box="[909,1099,1159,1185]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: figs. 1A, 10A), the flagellum on the TME is absent or relatively short (
<figureCitation id="8FC02A1B6A1CFF8B960C6CF5AB93FB60" box="[625,726,1195,1221]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="9" pageNumber="170">Figs. 3G</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B969E6CF5AA4DFB60" box="[739,776,1195,1221]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="9" pageNumber="170">4D</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B97686CF4AA72FB60" box="[789,823,1194,1221]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">5B</figureCitation>
) (vs. flagellum on the TME relatively long in
<emphasis id="258FEA8C6A1CFF8B91366CF5ACD9FB61" box="[1355,1436,1195,1220]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B94B56C91A818FB4C" box="[200,349,1231,1257]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B95E56C91AB1DFB4C" author="Pati, S. K. &amp; Yeo, D. C. J." box="[408,600,1231,1257]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: fig. 1D), the male s3/s4 is distinct (
<figureCitation id="8FC02A1B6A1CFF8B978E6C91AD0FFB4C" box="[1011,1098,1230,1257]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">Fig. 5C</figureCitation>
) (vs. male s3/s4 indiscernible in
<emphasis id="258FEA8C6A1CFF8B94C96CADA840FAA9" box="[180,261,1267,1292]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B95466CADA895FAA8" box="[315,464,1267,1293]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B96776CADAB8DFAA8" author="Pati, S. K. &amp; Yeo, D. C. J." box="[522,712,1267,1293]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: figs. 1F, 10C), and the G1 ultimate article is relatively stout and short, ca. 0.3 times the length of the penultimate article (
<figureCitation id="8FC02A1B6A1CFF8B977B6D49AA1EFA94" box="[774,859,1303,1329]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">Fig. 6B</figureCitation>
) (vs. G1 ultimate article relatively slender and long, ca. 0.60.8 times the length of the penultimate article in
<emphasis id="258FEA8C6A1CFF8B976B6D65AA22FAF1" box="[790,871,1339,1364]" italics="true" pageId="9" pageNumber="170">Barusa</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B97E26D65AD71FAF0" box="[927,1076,1339,1365]" italics="true" pageId="9" pageNumber="170">Barytelphusa</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B90126D65AC6AFAF0" author="Pati, S. K. &amp; Yeo, D. C. J." box="[1135,1327,1339,1365]" pageId="9" pageNumber="170" pagination="1 - 27" refId="ref10426" refString="Pati, S. K. &amp; Yeo, D. C. J. (2022) A taxonomic revision of the freshwater crab genus Barytelphusa Alcock, 1909 (Decapoda: Brachyura: Gecarcinucidae), with descriptions of a new genus and three new species. Journal of Crustacean Biology, 42, 1 - 27. https: // doi. org / 10.1093 / jcbiol / ruab 083" type="journal article" year="2022">Pati &amp; Yeo 2022</bibRefCitation>
: figs. 3A, 7I). The G2 is shorter than the G1, with the ultimate article very short, ca. 0.2 times the length of the penultimate article in
<taxonomicName id="D0FB4D1D6A1CFF8B94826DDDA81AFA38" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[255,351,1411,1437]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B94826DDDA81AFA38" box="[255,351,1411,1437]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B95186DDDA888FA38" bold="true" box="[357,461,1411,1437]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B95186DDDA888FA38" box="[357,461,1411,1437]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="8FC02A1B6A1CFF8B95A66DDDAB0BFA38" box="[475,590,1411,1437]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">Fig. 6B, F</figureCitation>
); whereas the G2 is as long as or longer than the G1, with the ultimate article relatively long, ca. 0.30.5 times the length of the penultimate article in
<emphasis id="258FEA8C6A1CFF8B97C16DF9AAB9FA65" box="[956,1020,1447,1472]" italics="true" pageId="9" pageNumber="170">Vanni</emphasis>
and
<emphasis id="258FEA8C6A1CFF8B904F6DF9AD27FA64" box="[1074,1122,1447,1473]" italics="true" pageId="9" pageNumber="170">Vela</emphasis>
(see
<bibRefCitation id="736A4B6F6A1CFF8B90E06DF9AC29FA64" author="Bahir, M. M. &amp; Yeo, D. C. J." box="[1181,1388,1447,1473]" pageId="9" pageNumber="170" pagination="309 - 354" refId="ref9793" refString="Bahir, M. M. &amp; Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 309 - 354." type="journal article" year="2007">Bahir &amp; Yeo 2007</bibRefCitation>
: fig. 31C, G;
<bibRefCitation id="736A4B6F6A1CFF8B948B6D95A8E4FA40" author="Pati, S. K. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[246,417,1483,1509]" pageId="9" pageNumber="170" pagination="112 - 126" refId="ref10499" refString="Pati, S. K., Bajantri, P. P. &amp; Hegde, G. D. (2023) A new freshwater crab species of the genus Vela Bahir &amp; Yeo, 2007 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats of Karnataka, India. Zootaxa, 5239, 112 - 126. https: // doi. org / 10.11646 / zootaxa. 5239.1.5" type="journal article" year="2023">
Pati
<emphasis id="258FEA8C6A1CFF8B955A6D95A81AFA40" box="[295,351,1483,1509]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2023
</bibRefCitation>
: fig. 4E, I).
<taxonomicName id="D0FB4D1D6A1CFF8B96596D95ABC1FA40" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[548,644,1483,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B96596D95ABC1FA40" box="[548,644,1483,1509]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B96F66D95ABB6FA40" bold="true" box="[651,755,1483,1509]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B96F66D95ABB6FA40" box="[651,755,1483,1509]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is further separated from
<emphasis id="258FEA8C6A1CFF8B906F6D95AD39FA40" box="[1042,1148,1483,1509]" italics="true" pageId="9" pageNumber="170">Ghatiana</emphasis>
by the distinct male s2/s3 and s3/s4 (
<figureCitation id="8FC02A1B6A1CFF8B95726DB1A820F9AC" box="[271,357,1518,1545]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">Fig. 5C</figureCitation>
) (vs. male s2/s3 and s3/s4 indistinct in
<emphasis id="258FEA8C6A1CFF8B97626DB1AACCF9AC" box="[799,905,1519,1545]" italics="true" pageId="9" pageNumber="170">Ghatiana</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B97BF6DB1AD8FF9AC" author="Pati, S. K. &amp; Thackeray, T." box="[962,1226,1519,1545]" pageId="9" pageNumber="170" pagination="627 - 647" refId="ref10357" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
: fig. 5C;
<bibRefCitation id="736A4B6F6A1CFF8B914F6DB1A9A5F988" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." pageId="9" pageNumber="170" pagination="2022019" refId="ref10826" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="258FEA8C6A1CFF8B91196DB1ACD8F9AC" box="[1380,1437,1519,1545]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022a
</bibRefCitation>
: fig. 3C); the relatively short male sternopleonal cavity, which reaches anteriorly to the level of mid-length of the cheliped coxae (
<figureCitation id="8FC02A1B6A1CFF8B95056E69A88BF9F4" box="[376,462,1590,1617]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="9" pageNumber="170">Fig. 5C</figureCitation>
) (vs. male sternopleonal cavity relatively long, reaching anteriorly beyond the level of the bases of the third maxillipeds in
<emphasis id="258FEA8C6A1CFF8B964C6E05ABDEF9D0" box="[561,667,1627,1653]" italics="true" pageId="9" pageNumber="170">Ghatiana</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B96A86E05AA9BF9D0" author="Pati, S. K. &amp; Thackeray, T." box="[725,990,1627,1653]" pageId="9" pageNumber="170" pagination="627 - 647" refId="ref10357" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
: fig. 5C;
<bibRefCitation id="736A4B6F6A1CFF8B903A6E05AC46F9D0" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[1095,1283,1627,1653]" pageId="9" pageNumber="170" pagination="2022019" refId="ref10826" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="258FEA8C6A1CFF8B90076E05ADF7F9D0" box="[1146,1202,1627,1653]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022a
</bibRefCitation>
: fig. 3C); and the relatively stouter ultimate article of the G1 (
<figureCitation id="8FC02A1B6A1CFF8B96D06E21AA47F93C" box="[685,770,1663,1689]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">Fig. 6B</figureCitation>
) (vs. G1 ultimate article relatively slender in
<emphasis id="258FEA8C6A1CFF8B917D6E21AC2FF93C" box="[1280,1386,1663,1689]" italics="true" pageId="9" pageNumber="170">Ghatiana</emphasis>
; see
<bibRefCitation id="736A4B6F6A1CFF8B94EA6EFDA8DAF918" author="Pati, S. K. &amp; Thackeray, T." box="[151,415,1699,1725]" pageId="9" pageNumber="170" pagination="627 - 647" refId="ref10357" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
: fig. 5D;
<bibRefCitation id="736A4B6F6A1CFF8B96756EFDAB86F918" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[520,707,1699,1725]" pageId="9" pageNumber="170" pagination="2022019" refId="ref10826" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="258FEA8C6A1CFF8B96466EFDAB36F918" box="[571,627,1699,1725]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2022a
</bibRefCitation>
: fig. 4D).
</paragraph>
<paragraph id="1744369E6A1CFF8B94BA6E98AD4CF8A0" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="170">
<emphasis id="258FEA8C6A1CFF8B94BA6E98A8ABF945" bold="true" box="[199,494,1734,1760]" pageId="9" pageNumber="170">Geographic distribution.</emphasis>
<taxonomicName id="D0FB4D1D6A1CFF8B95896E99AB11F944" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[500,596,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="258FEA8C6A1CFF8B95896E99AB11F944" box="[500,596,1735,1761]" italics="true" pageId="9" pageNumber="170">Aradhya</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1CFF8B96266E99AB81F944" bold="true" box="[603,708,1735,1761]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="3EBC57F76A1CFF8B96266E99AB81F944" box="[603,708,1735,1761]" pageId="9" pageNumber="170" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is currently known only from an isolated mountain of the Central Western Ghats in the Uttara Kannada district of
<collectingRegion id="D53FF87C6A1CFF8B96CF6EB5AA27F8A0" box="[690,866,1771,1797]" country="India" name="Karnataka" pageId="9" pageNumber="170">Karnataka state</collectingRegion>
,
<collectingCountry id="6FEC760E6A1CFF8B97136EB5AAEDF8A0" box="[878,936,1771,1797]" name="India" pageId="9" pageNumber="170">India</collectingCountry>
(
<figureCitation id="8FC02A1B6A1CFF8B97CB6EB5AABDF8A0" box="[950,1016,1771,1797]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="9" pageNumber="170">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>