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<document id="144E4B5A38AA8D8EBF245E406C9BD597" ID-DOI="10.1080/00222933.2019.1606953" ID-GBIF-Dataset="d2615e10-8703-400b-b571-6c431b0ed9a7" ID-ISSN="1464-5262" ID-Zenodo-Dep="3675571" ID-ZooBank="09AA5C86-5138-42A7-9646-9FF2A7F46767" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582134181895" checkinUser="valdenar" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="9F138799E555B06A9ABD5DC5D4A7F9B1" docLanguage="en" docName="JNATHIST.53.11–12.725–747.pdf.imf" docOrigin="Journal of Natural History 53 (11)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Aegidinus Arrow 1904" docType="key" docVersion="13" lastPageNumber="731" masterDocId="632AFFE1E556B06D9A1D5E52D024FFC2" masterDocTitle="Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature" masterLastPageNumber="747" masterPageNumber="725" pageNumber="727" updateTime="1698719875536" updateUser="plazi">
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<mods:title id="961C696C6FB972FBB49361C520F2E516">Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature</mods:title>
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<mods:namePart id="02C7AE147945369B3C1A2332EF3F61BF">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="D1CDE09D759A08F61D4B7D8027C54710">Akhmetova, Lilia A.</mods:namePart>
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<mods:namePart id="1D9AF7873F641625881BD9A2E16A6D03">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="296F22715461D3DEFF9276B44F589B3E">Journal of Natural History</mods:title>
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<mods:number id="0FDCDFC843190BA5682FF66265AEEC9B">53</mods:number>
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<treatment id="9F138799E555B06A9ABD5DC5D4A7F9B1" ID-DOI="http://doi.org/10.5281/zenodo.4561501" ID-GBIF-Taxon="162173913" ID-Zenodo-Dep="4561501" LSID="urn:lsid:plazi:treatment:9F138799E555B06A9ABD5DC5D4A7F9B1" httpUri="http://treatment.plazi.org/id/9F138799E555B06A9ABD5DC5D4A7F9B1" lastPageId="7" lastPageNumber="731" pageId="3" pageNumber="727">
<subSubSection id="5FA06504E555B06E9ABD5DC5D20BFC6D" box="[160,559,919,943]" pageId="3" pageNumber="727" type="nomenclature">
<paragraph id="1705368FE555B06E9ABD5DC5D20BFC6D" blockId="3.[160,559,919,943]" box="[160,559,919,943]" pageId="3" pageNumber="727">
<emphasis id="25CEEA9DE555B06E9ABD5DC5D20BFC6D" bold="true" box="[160,559,919,943]" pageId="3" pageNumber="727">
<heading id="4C4D81E3E555B06E9ABD5DC5D20CFC6D" box="[160,552,919,943]" pageId="3" pageNumber="727">
Key to
<taxonomicName id="D0BA4D0CE555B06E9AEA5DC5D14BFC6D" authorityName="Arrow" authorityYear="1904" box="[247,367,919,943]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus">Aegidinus</taxonomicName>
species (males
</heading>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E555B06A9ABD5D88D4A7F9B1" lastPageId="7" lastPageNumber="731" pageId="3" pageNumber="727" type="key">
<key id="632B61BEE555B06A9ABD5D88D4A7F9B1" lastPageId="7" lastPageNumber="731" pageId="3" pageNumber="727">
<keyStep id="AC4E2E2AE555B06E9ABD5D88D4A0FB98" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD5D88D4A7FBD6" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5D88D4A7FBD6" pageId="3" pageNumber="727">
1. Parameres separated into dorsomedial and ventrolateral lobes (
<figureCitation id="8F812A0AE555B06E99BA5D88D428FC30" box="[935,1036,986,1010]" captionStart="Figure 1" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[167,1148,143,1370]" captionTargetId="figure@4.[166,1149,142,1371]" captionTargetPageId="4" captionText="Figure 1. Aegidinus guianensis, holotype, female (a, b, d), female, paratype of Ae. oreibates (c), female from Villa Napoleão, Brazil (e), male, holotype of Ae. oreibates (f), male, paratype of Ae. oreibates (g–i). Habitus in dorsal view (a, c, f), labels (b), female external genitalia (d, e), aedeagus in lateral (h) and ventral view (g), parameres in dorsal view (i)." figureDoi="http://doi.org/10.5281/zenodo.3675573" httpUri="https://zenodo.org/record/3675573/files/figure.png" pageId="3" pageNumber="727">Figures 1</figureCitation>
(h,i), 2(e,f) and 3(b
<emphasis id="25CEEA9DE555B06E9AE55DAED121FBD6" box="[248,261,1020,1044]" italics="true" pageId="3" pageNumber="727">–</emphasis>
d))....................................................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5A4DD4A0FB98" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5A4DD4A0FB98" pageId="3" pageNumber="727">
- Parameres not separated into dorsomedial and ventrolateral lobes (
<figureCitation id="8F812A0AE555B06E99C85A4DD417FBF5" box="[981,1075,1055,1079]" captionStart="Figure 6" captionStartId="17.[160,225,1443,1466]" captionTargetBox="[190,1148,145,1407]" captionTargetId="figure@17.[166,1149,142,1411]" captionTargetPageId="17" captionText="Figure 6. Aegidinus cornutus, male (a, c, d, e), female (b, f). Habitus in dorsal view (a, b); female external genitalia (f); aedeagus in lateral (c) and ventral view (e); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675583" httpUri="https://zenodo.org/record/3675583/files/figure.png" pageId="3" pageNumber="727">Figure 6</figureCitation>
(c
<emphasis id="25CEEA9DE555B06E9E5B5A4DD477FBF5" box="[1094,1107,1055,1079]" italics="true" pageId="3" pageNumber="727">–</emphasis>
e))... ................................................................................................................................
<taxonomicName id="D0BA4D0CE555B06E99BF5A10D4A0FB98" authorityName="Colby" authorityYear="2009" box="[930,1156,1090,1114]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="25CEEA9DE555B06E99BF5A10D4A0FB98" bold="true" box="[930,1156,1090,1114]" italics="true" pageId="3" pageNumber="727">Aegidinus cornutus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE555B06E9ABD5A25D4A7FB70" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD5A25D4A7FB4D" blockId="3.[160,1156,986,1727]" box="[160,1155,1143,1167]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5A25D4A7FB4D" box="[160,1155,1143,1167]" pageId="3" pageNumber="727">
2. Phallobase with ventroapical plate (
<figureCitation id="8F812A0AE555B06E98495A25D29DFB4D" box="[596,697,1143,1167]" captionStart="Figure 1" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[167,1148,143,1370]" captionTargetId="figure@4.[166,1149,142,1371]" captionTargetPageId="4" captionText="Figure 1. Aegidinus guianensis, holotype, female (a, b, d), female, paratype of Ae. oreibates (c), female from Villa Napoleão, Brazil (e), male, holotype of Ae. oreibates (f), male, paratype of Ae. oreibates (g–i). Habitus in dorsal view (a, c, f), labels (b), female external genitalia (d, e), aedeagus in lateral (h) and ventral view (g), parameres in dorsal view (i)." figureDoi="http://doi.org/10.5281/zenodo.3675573" httpUri="https://zenodo.org/record/3675573/files/figure.png" pageId="3" pageNumber="727">Figures 1</figureCitation>
(g), 3(c) and 4(d)) ......................................... 3
</keyLead>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5AC8D4A7FB70" blockId="3.[160,1156,986,1727]" box="[160,1155,1178,1202]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5AC8D4A7FB70" box="[160,1155,1178,1202]" pageId="3" pageNumber="727">
- Phallobase without ventroapical plate (
<figureCitation id="8F812A0AE555B06E98935AC8D2DCFB70" box="[654,760,1178,1202]" captionStart="Figure 2" captionStartId="5.[160,225,1371,1394]" captionTargetBox="[189,1148,153,1339]" captionTargetId="figure@5.[166,1149,142,1340]" captionTargetPageId="5" captionText="Figure 2. Aegidinus steinheili, lectotype, male (a, d–f), male with erroneous label ‘Bresil Steinheil...’. (b), female, paralectotype (c, g). Habitus in dorsal view (a–c), female external genitalia (g), aedeagus in lateral (f) and ventral view (d), parameres in dorsal view (e)." figureDoi="http://doi.org/10.5281/zenodo.3675575" httpUri="https://zenodo.org/record/3675575/files/figure.png" pageId="3" pageNumber="727">Figures 2</figureCitation>
(d), 4(h) and 5(d)).............................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE555B06E9ABD5A9DD4A7FAC8" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD5A9DD4A7FB25" blockId="3.[160,1156,986,1727]" box="[160,1155,1231,1255]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5A9DD4A7FB25" box="[160,1155,1231,1255]" pageId="3" pageNumber="727">3. Parameres symmetrical .......................................................................................................................... 4</keyLead>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5AA0D4A7FAC8" blockId="3.[160,1156,986,1727]" box="[160,1155,1266,1290]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5AA0D4A7FAC8" box="[160,1155,1266,1290]" pageId="3" pageNumber="727">- Parameres asymmetrical .................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE555B06E9ABD5B75D4A7FA65" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD5B75D4A0FAA0" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5B75D4A0FAA0" pageId="3" pageNumber="727">
4. Ventrolateral lobe of paramere with subapical tooth (
<bibRefCitation id="732B4B7EE555B06E99015B75D3BAFAFD" author="Colby J" box="[796,926,1319,1343]" pageId="3" pageNumber="727" pagination="1 - 41" refId="ref9012" refString="Colby J. 2009. Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi. 76: 1 - 41." type="journal article" year="2009">
Colby
<date id="6304104FE555B06E997A5B75D3BAFAFD" box="[871,926,1319,1343]" pageId="3" pageNumber="727" value="2009-01">2009</date>
</bibRefCitation>
, fig. 55)........................ ...............................................................................................................................
<taxonomicName id="D0BA4D0CE555B06E99705B18D4A0FAA0" authorityName="Colby" authorityYear="2009" box="[877,1156,1354,1378]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="25CEEA9DE555B06E99705B18D4A0FAA0" bold="true" box="[877,1156,1354,1378]" italics="true" pageId="3" pageNumber="727">Aegidinus howdenorum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5B3FD4A7FA65" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5B3FD4A7FA65" pageId="3" pageNumber="727">
- Ventrolateral lobe of paramere without subapical tooth (
<figureCitation id="8F812A0AE555B06E99455B3FD38AFA47" box="[856,942,1389,1413]" captionStart="Figure 1" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[167,1148,143,1370]" captionTargetId="figure@4.[166,1149,142,1371]" captionTargetPageId="4" captionText="Figure 1. Aegidinus guianensis, holotype, female (a, b, d), female, paratype of Ae. oreibates (c), female from Villa Napoleão, Brazil (e), male, holotype of Ae. oreibates (f), male, paratype of Ae. oreibates (g–i). Habitus in dorsal view (a, c, f), labels (b), female external genitalia (d, e), aedeagus in lateral (h) and ventral view (g), parameres in dorsal view (i)." figureDoi="http://doi.org/10.5281/zenodo.3675573" httpUri="https://zenodo.org/record/3675573/files/figure.png" pageId="3" pageNumber="727">Figure 1</figureCitation>
(h)).............................. ............................................................................................................................
<taxonomicName id="D0BA4D0CE555B06E99965BDDD4A7FA65" baseAuthorityName="Westwood" baseAuthorityYear="1845" box="[907,1155,1423,1447]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="25CEEA9DE555B06E99965BDDD4A7FA65" bold="true" box="[907,1155,1423,1447]" italics="true" pageId="3" pageNumber="727">Aegidinus guianensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE555B06E9ABD5B97D4A7F9A5" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD5B97D4A0F9E0" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD5B97D4A0F9E0" pageId="3" pageNumber="727">
5. Parameres longer, more asymmetrical (
<figureCitation id="8F812A0AE555B06E986B5B97D2F7FA1F" box="[630,723,1477,1501]" captionStart="Figure 4" captionStartId="7.[160,225,1306,1329]" captionTargetBox="[167,1142,143,1271]" captionTargetId="figure@7.[166,1149,142,1274]" captionTargetPageId="7" captionText="Figure 4. Aegidinus spp. Ae. noriegai sp. nov., holotype (a–d), Ae. simulatus, male (e–h). Habitus in dorsal view (a, e); aedeagus in lateral (b, f) and ventral view (d, h); parameres in dorsal view (c, g)." figureDoi="http://doi.org/10.5281/zenodo.3675579" httpUri="https://zenodo.org/record/3675579/files/figure.png" pageId="3" pageNumber="727">Figure 4</figureCitation>
(b,c)); ventroapical plate of phallobase longer than wide (
<figureCitation id="8F812A0AE555B06E9B695BB5D1F7FA3D" box="[372,467,1511,1535]" captionStart="Figure 4" captionStartId="7.[160,225,1306,1329]" captionTargetBox="[167,1142,143,1271]" captionTargetId="figure@7.[166,1149,142,1274]" captionTargetPageId="7" captionText="Figure 4. Aegidinus spp. Ae. noriegai sp. nov., holotype (a–d), Ae. simulatus, male (e–h). Habitus in dorsal view (a, e); aedeagus in lateral (b, f) and ventral view (d, h); parameres in dorsal view (c, g)." figureDoi="http://doi.org/10.5281/zenodo.3675579" httpUri="https://zenodo.org/record/3675579/files/figure.png" pageId="3" pageNumber="727">Figure 4</figureCitation>
(d)); protibia without medioapical tooth.......................................... ..........................................................................................................................
<taxonomicName id="D0BA4D0CE555B06E99525858D40FF9E0" authority="Akhmetova &amp; Vaz-de-Mello, 2019" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2019" box="[847,1067,1546,1570]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="noriegai" status="sp. nov.">
<emphasis id="25CEEA9DE555B06E99525858D40FF9E0" bold="true" box="[847,1067,1546,1570]" italics="true" pageId="3" pageNumber="727">Aegidinus noriegai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EFD57E6E555B06E9E365858D4A0F9E0" box="[1067,1156,1546,1570]" pageId="3" pageNumber="727" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="1705368FE555B06E9ABD587FD4A7F9A5" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD587FD4A7F9A5" pageId="3" pageNumber="727">
- Parameres shorter, less asymmetrical (
<figureCitation id="8F812A0AE555B06E9867587FD2F1F987" box="[634,725,1581,1605]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (b–d), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (j–l); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="3" pageNumber="727">Figure 3</figureCitation>
(b,d)); ventroapical plate of phallobase wider than long (
<figureCitation id="8F812A0AE555B06E9B8A581DD1D1F9A5" box="[407,501,1615,1639]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (b–d), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (j–l); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="3" pageNumber="727">Figure 3</figureCitation>
(c)); protibia with medioapical tooth
<taxonomicName id="D0BA4D0CE555B06E99BC581DD4A7F9A5" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[929,1155,1615,1639]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE555B06E99BC581DD4A7F9A5" bold="true" box="[929,1155,1615,1639]" italics="true" pageId="3" pageNumber="727">Aegidinus candezei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE555B0699ABD58D7D4A7F9A5" lastPageId="4" lastPageNumber="728" pageId="3" pageNumber="727">
<paragraph id="1705368FE555B06E9ABD58D7D4A7F97D" blockId="3.[160,1156,986,1727]" pageId="3" pageNumber="727">
<keyLead id="AC4B95BAE555B06E9ABD58D7D4A7F97D" pageId="3" pageNumber="727">
6. Mediobasal margins of dorsomedial lobes of parameres feebly sclerotised, membranous (
<figureCitation id="8F812A0AE555B06E9AFA58F5D161F97D" box="[231,325,1703,1727]" captionStart="Figure 2" captionStartId="5.[160,225,1371,1394]" captionTargetBox="[189,1148,153,1339]" captionTargetId="figure@5.[166,1149,142,1340]" captionTargetPageId="5" captionText="Figure 2. Aegidinus steinheili, lectotype, male (a, d–f), male with erroneous label ‘Bresil Steinheil...’. (b), female, paralectotype (c, g). Habitus in dorsal view (a–c), female external genitalia (g), aedeagus in lateral (f) and ventral view (d), parameres in dorsal view (e)." figureDoi="http://doi.org/10.5281/zenodo.3675575" httpUri="https://zenodo.org/record/3675575/files/figure.png" pageId="3" pageNumber="727">Figure 2</figureCitation>
(e)); protibia with medioapical tooth...................................................................... 7
</keyLead>
</paragraph>
<caption id="43C56607E552B0699ABD5B28D277FA28" ID-DOI="http://doi.org/10.5281/zenodo.3675573" ID-Zenodo-Dep="3675573" httpUri="https://zenodo.org/record/3675573/files/figure.png" pageId="4" pageNumber="728" startId="4.[160,225,1402,1425]" targetBox="[167,1148,143,1370]" targetPageId="4">
<paragraph id="1705368FE552B0699ABD5B28D277FA28" blockId="4.[160,1155,1402,1514]" pageId="4" pageNumber="728">
<emphasis id="25CEEA9DE552B0699ABD5B28D0D9FA53" bold="true" box="[160,253,1402,1425]" pageId="4" pageNumber="728">Figure 1.</emphasis>
<taxonomicName id="D0BA4D0CE552B0699B1B5B28D1EEFA53" baseAuthorityName="Westwood" baseAuthorityYear="1845" box="[262,458,1402,1425]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="25CEEA9DE552B0699B1B5B28D1EEFA53" box="[262,458,1402,1425]" italics="true" pageId="4" pageNumber="728">Aegidinus guianensis</emphasis>
</taxonomicName>
, holotype, female (a, b, d), female, paratype of
<taxonomicName id="D0BA4D0CE552B06999835B28D433FA53" authorityName="J. Colby" authorityYear="2009" box="[926,1047,1402,1425]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="oreibates">
<emphasis id="25CEEA9DE552B06999835B28D433FA53" box="[926,1047,1402,1425]" italics="true" pageId="4" pageNumber="728">Ae. oreibates</emphasis>
</taxonomicName>
(c), female from Villa Napoleão, Brazil (e), male, holotype of
<taxonomicName id="D0BA4D0CE552B069989D5BC5D2DCFA6C" authorityName="J. Colby" authorityYear="2009" box="[640,760,1431,1454]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="oreibates">
<emphasis id="25CEEA9DE552B069989D5BC5D2DCFA6C" box="[640,760,1431,1454]" italics="true" pageId="4" pageNumber="728">Ae. oreibates</emphasis>
</taxonomicName>
(f), male, paratype of
<taxonomicName id="D0BA4D0CE552B06999CF5BC5D46FFA6C" authorityName="J. Colby" authorityYear="2009" box="[978,1099,1431,1454]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="oreibates">
<emphasis id="25CEEA9DE552B06999CF5BC5D46FFA6C" box="[978,1099,1431,1454]" italics="true" pageId="4" pageNumber="728">Ae. oreibates</emphasis>
</taxonomicName>
(g
<emphasis id="25CEEA9DE552B0699E7B5BCAD456FA6D" box="[1126,1138,1432,1455]" italics="true" pageId="4" pageNumber="728">–</emphasis>
i). Habitus in dorsal view (a, c, f), labels (b), female external genitalia (d, e), aedeagus in lateral (h) and ventral view (g), parameres in dorsal view (i).
</paragraph>
</caption>
<paragraph id="1705368FE552B0699ABD587FD4A7F9A5" blockId="4.[160,1156,1581,1639]" pageId="4" pageNumber="728">
<keyLead id="AC4B95BAE552B0699ABD587FD4A7F9A5" pageId="4" pageNumber="728">- Mediobasal margins of dorsomedial lobes of parameres strongly sclerotised (4G, 5E); protibia without medioapical tooth ............................................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE552B0689ABD58D7D4A7F987" lastPageId="5" lastPageNumber="729" pageId="4" pageNumber="728">
<paragraph id="1705368FE552B0699ABD58D7D4A7F97D" blockId="4.[160,1156,1669,1727]" pageId="4" pageNumber="728">
<keyLead id="AC4B95BAE552B0699ABD58D7D4A7F97D" pageId="4" pageNumber="728">
7. Ventrolateral lobes of parameres long and slender (in lateral view), noticeably longer than dorsomedial lobes (
<figureCitation id="8F812A0AE552B0699BA058F5D23CF97D" box="[445,536,1703,1727]" captionStart="Figure 2" captionStartId="5.[160,225,1371,1394]" captionTargetBox="[189,1148,153,1339]" captionTargetId="figure@5.[166,1149,142,1340]" captionTargetPageId="5" captionText="Figure 2. Aegidinus steinheili, lectotype, male (a, d–f), male with erroneous label ‘Bresil Steinheil...’. (b), female, paralectotype (c, g). Habitus in dorsal view (a–c), female external genitalia (g), aedeagus in lateral (f) and ventral view (d), parameres in dorsal view (e)." figureDoi="http://doi.org/10.5281/zenodo.3675575" httpUri="https://zenodo.org/record/3675575/files/figure.png" pageId="4" pageNumber="728">Figure 2</figureCitation>
(f)).......................................... ..........................................
<taxonomicName id="D0BA4D0CE552B0699E0758F5D4A7F97D" authorityName="R. Paulian" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[1050,1155,1703,1727]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="25CEEA9DE552B0699E0758F5D4A7F97D" bold="true" box="[1050,1155,1703,1727]" italics="true" pageId="4" pageNumber="728">steinheili</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<caption id="43C56607E553B0689ABD5B09D33EFA6F" ID-DOI="http://doi.org/10.5281/zenodo.3675575" ID-Zenodo-Dep="3675575" httpUri="https://zenodo.org/record/3675575/files/figure.png" pageId="5" pageNumber="729" startId="5.[160,225,1371,1394]" targetBox="[189,1148,153,1339]" targetPageId="5">
<paragraph id="1705368FE553B0689ABD5B09D33EFA6F" blockId="5.[160,1155,1371,1453]" pageId="5" pageNumber="729">
<emphasis id="25CEEA9DE553B0689ABD5B09D0DFFAB0" bold="true" box="[160,251,1371,1394]" pageId="5" pageNumber="729">Figure 2.</emphasis>
<taxonomicName id="D0BA4D0CE553B0689B1E5B09D198FAB0" authorityName="R. Paulian" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[259,444,1371,1394]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="729" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="25CEEA9DE553B0689B1E5B09D198FAB0" box="[259,444,1371,1394]" italics="true" pageId="5" pageNumber="729">Aegidinus steinheili</emphasis>
</taxonomicName>
, lectotype, male (a, d
<emphasis id="25CEEA9DE553B068988B5B09D287FAB0" box="[662,675,1371,1394]" italics="true" pageId="5" pageNumber="729">–</emphasis>
f), male with erroneous label
<emphasis id="25CEEA9DE553B06899D65B09D3F4FAB0" box="[971,976,1371,1394]" italics="true" pageId="5" pageNumber="729">‘</emphasis>
Bresil Steinheil...
<emphasis id="25CEEA9DE553B0689E645B09D45AFAB0" box="[1145,1150,1371,1394]" italics="true" pageId="5" pageNumber="729">’</emphasis>
. (b), female, paralectotype (c, g). Habitus in dorsal view (a
<emphasis id="25CEEA9DE553B06898C55B2AD2C1FA4D" box="[728,741,1400,1423]" italics="true" pageId="5" pageNumber="729">–</emphasis>
c), female external genitalia (g), aedeagus in lateral (f) and ventral view (d), parameres in dorsal view (e).
</paragraph>
</caption>
<paragraph id="1705368FE553B0689ABD5858D4A7F987" blockId="5.[160,1155,1546,1605]" pageId="5" pageNumber="729">
<keyLead id="AC4B95BAE553B0689ABD5858D4A7F987" pageId="5" pageNumber="729">- Ventrolateral lobes of parameres triangular and obtuse in lateral view, not longer than dorsomedial lobes ..................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<paragraph id="1705368FE553B0689ABD5830D4A7F9B8" blockId="5.[160,1155,1634,1727]" box="[160,1155,1634,1658]" pageId="5" pageNumber="729">
<keyStep id="AC4E2E2AE553B0689ABD5830D4A7F9B8" box="[160,1155,1634,1658]" pageId="5" pageNumber="729">
<keyLead id="AC4B95BAE553B0689ABD5830D4A7F9B8" box="[160,1155,1634,1658]" pageId="5" pageNumber="729">
8. Ventrolateral lobes of parameres as long as dorsomedial lobes......
<taxonomicName id="D0BA4D0CE553B06899A95830D4A7F9B8" authorityName="Colby" authorityYear="2009" box="[948,1155,1634,1658]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="729" phylum="Arthropoda" rank="species" species="petrovi">
<emphasis id="25CEEA9DE553B06899A95830D4A7F9B8" bold="true" box="[948,1155,1634,1658]" italics="true" pageId="5" pageNumber="729">Aegidinus petrovi</emphasis>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="1705368FE553B06A9ABD58D7D4A7FA63" blockId="5.[160,1155,1634,1727]" lastBlockId="7.[160,1155,1417,1441]" lastPageId="7" lastPageNumber="731" pageId="5" pageNumber="729">
<keyLead id="AC4B95BAE553B0689ABD58D7D4A7F97D" pageId="5" pageNumber="729">
- Ventrolateral lobes of parameres noticeably shorter than dorsomedial lobes............... ...............................................................................................................
<taxonomicName id="D0BA4D0CE553B068995C58F5D4A7F97D" authorityName="Colby" authorityYear="2009" box="[833,1155,1703,1727]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="729" phylum="Arthropoda" rank="species" species="teamscaraborum">
<emphasis id="25CEEA9DE553B068995C58F5D4A7F97D" bold="true" box="[833,1155,1703,1727]" italics="true" pageId="5" pageNumber="729">Aegidinus teamscaraborum</emphasis>
</taxonomicName>
</keyLead>
<keyStep id="AC4E2E2AE550B06A9ABD58D3D4A7FA63" lastPageId="7" lastPageNumber="731" pageId="6" pageNumber="730">
<keyLead id="AC4B95BAE550B06A9ABD58D3D4A7FA63" lastPageId="7" lastPageNumber="731" pageId="6" pageNumber="730">
9. Dorsal sides of parameres strongly overlapping and separated by slit (
<figureCitation id="8F812A0AE550B06B9E1B58D3D44CF95B" box="[1030,1128,1665,1689]" captionStart="Figure 4" captionStartId="7.[160,225,1306,1329]" captionTargetBox="[167,1142,143,1271]" captionTargetId="figure@7.[166,1149,142,1274]" captionTargetPageId="7" captionText="Figure 4. Aegidinus spp. Ae. noriegai sp. nov., holotype (a–d), Ae. simulatus, male (e–h). Habitus in dorsal view (a, e); aedeagus in lateral (b, f) and ventral view (d, h); parameres in dorsal view (c, g)." figureDoi="http://doi.org/10.5281/zenodo.3675579" httpUri="https://zenodo.org/record/3675579/files/figure.png" pageId="6" pageNumber="730">Figure 4</figureCitation>
(f), arrowed).......................................................... ..........................................................
<taxonomicName id="D0BA4D0CE550B06B998A58F1D4A7F979" authorityName="Colby" authorityYear="2009" box="[919,1155,1699,1723]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="species" species="simulatus">
<emphasis id="25CEEA9DE550B06B998A58F1D4A7F979" bold="true" box="[919,1155,1699,1723]" italics="true" pageId="6" pageNumber="730">Aegidinus simulatus</emphasis>
</taxonomicName>
- Dorsal sides of parameres less overlapping and not separated by slit (
<figureCitation id="8F812A0AE551B06A99FF5BDBD41BFA63" box="[994,1087,1417,1441]" captionStart="Figure 5" captionStartId="8.[160,225,1686,1709]" captionTargetBox="[170,1132,145,1646]" captionTargetId="figure@8.[166,1149,142,1654]" captionTargetPageId="8" captionText="Figure 5. Aegidinus colbyae, holotype male (a, e, d, f), paratype female (b, c). Habitus in dorsal view (a, b); female external genitalia (c); aedeagus in lateral (f) and ventral view (d); parameres in dorsal view (e)." figureDoi="http://doi.org/10.5281/zenodo.3675581" httpUri="https://zenodo.org/record/3675581/files/figure.png" pageId="7" pageNumber="731">Figure 5</figureCitation>
(f)) 10
</keyLead>
</keyStep>
</paragraph>
<caption id="43C56607E550B06B9ABD5B85D1CFF984" ID-DOI="http://doi.org/10.5281/zenodo.3675577" ID-Zenodo-Dep="3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="6" pageNumber="730" startId="6.[160,225,1495,1518]" targetBox="[167,1148,143,1463]" targetPageId="6">
<paragraph id="1705368FE550B06B9ABD5B85D1CFF984" blockId="6.[160,1157,1495,1606]" pageId="6" pageNumber="730">
<emphasis id="25CEEA9DE550B06B9ABD5B85D0DFFA2C" bold="true" box="[160,251,1495,1518]" pageId="6" pageNumber="730">Figure 3.</emphasis>
<taxonomicName id="D0BA4D0CE550B06B9B1E5B85D144FA2C" authorityName="Arrow" authorityYear="1904" box="[259,352,1495,1518]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE550B06B9B1E5B85D144FA2C" box="[259,352,1495,1518]" italics="true" pageId="6" pageNumber="730">Aegidinus</emphasis>
</taxonomicName>
spp.
<taxonomicName id="D0BA4D0CE550B06B9B8A5B85D26AFA2C" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[407,590,1495,1518]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE550B06B9B8A5B85D26AFA2C" box="[407,590,1495,1518]" italics="true" pageId="6" pageNumber="730">Aegidinus candezei</emphasis>
</taxonomicName>
, lectotype, male (a, e), male (b
<emphasis id="25CEEA9DE550B06B99985B85D3B6FA2C" box="[901,914,1495,1518]" italics="true" pageId="6" pageNumber="730">–</emphasis>
d), female (h, k), female, holotype of
<taxonomicName id="D0BA4D0CE550B06B9B065BA6D1BFF9C9" authorityName="Colby" authorityYear="2009" box="[283,411,1524,1547]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="species" species="sunidigea">
<emphasis id="25CEEA9DE550B06B9B065BA6D1BFF9C9" box="[283,411,1524,1547]" italics="true" pageId="6" pageNumber="730">Ae. sunidigea</emphasis>
</taxonomicName>
(g, j);
<taxonomicName id="D0BA4D0CE550B06B9BC05BA6D21EF9C9" authorityName="Arrow" authorityYear="1904" box="[477,570,1524,1547]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE550B06B9BC05BA6D21EF9C9" box="[477,570,1524,1547]" italics="true" pageId="6" pageNumber="730">Aegidinus</emphasis>
</taxonomicName>
sp., paralectotype of
<taxonomicName id="D0BA4D0CE550B06B99095BA6D3A8F9C9" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[788,908,1524,1547]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="730" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE550B06B99095BA6D3A8F9C9" box="[788,908,1524,1547]" italics="true" pageId="6" pageNumber="730">Ae. candezei</emphasis>
</taxonomicName>
(f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (j
<emphasis id="25CEEA9DE550B06B98D15840D2FDF9EB" box="[716,729,1554,1577]" italics="true" pageId="6" pageNumber="730">–</emphasis>
l); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d).
</paragraph>
</caption>
<caption id="43C56607E551B06A9ABD5B48D4A0FA8D" ID-DOI="http://doi.org/10.5281/zenodo.3675579" ID-Zenodo-Dep="3675579" httpUri="https://zenodo.org/record/3675579/files/figure.png" pageId="7" pageNumber="731" startId="7.[160,225,1306,1329]" targetBox="[167,1142,143,1271]" targetPageId="7">
<paragraph id="1705368FE551B06A9ABD5B48D4A0FA8D" blockId="7.[160,1156,1305,1359]" pageId="7" pageNumber="731">
<emphasis id="25CEEA9DE551B06A9ABD5B48D0D8FAF3" bold="true" box="[160,252,1306,1329]" pageId="7" pageNumber="731">Figure 4.</emphasis>
<taxonomicName id="D0BA4D0CE551B06A9B185B4BD146FAF2" authorityName="Arrow" authorityYear="1904" box="[261,354,1305,1328]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE551B06A9B185B4BD146FAF2" box="[261,354,1305,1328]" italics="true" pageId="7" pageNumber="731">Aegidinus</emphasis>
</taxonomicName>
spp.
<taxonomicName id="D0BA4D0CE551B06A9B865B4BD22BFAF2" authority="Akhmetova &amp; Vaz-de-Mello, 2019" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2019" box="[411,527,1305,1328]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="species" species="noriegai" status="sp. nov.">
<emphasis id="25CEEA9DE551B06A9B865B4BD22BFAF2" box="[411,527,1305,1328]" italics="true" pageId="7" pageNumber="731">Ae. noriegai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EFD57E6E551B06A98055B48D24DFAF3" box="[536,617,1306,1329]" pageId="7" pageNumber="731" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (a
<emphasis id="25CEEA9DE551B06A98FB5B48D2D7FAF3" box="[742,755,1306,1329]" italics="true" pageId="7" pageNumber="731">–</emphasis>
d),
<taxonomicName id="D0BA4D0CE551B06A99085B4BD3B1FAF2" authorityName="Colby" authorityYear="2009" box="[789,917,1305,1328]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="species" species="simulatus">
<emphasis id="25CEEA9DE551B06A99085B4BD3B1FAF2" box="[789,917,1305,1328]" italics="true" pageId="7" pageNumber="731">Ae. simulatus</emphasis>
</taxonomicName>
, male (e
<emphasis id="25CEEA9DE551B06A99F25B48D3D8FAF3" box="[1007,1020,1306,1329]" italics="true" pageId="7" pageNumber="731">–</emphasis>
h). Habitus in dorsal view (a, e); aedeagus in lateral (b, f) and ventral view (d, h); parameres in dorsal view (c, g).
</paragraph>
</caption>
<keyStep id="AC4E2E2AE551B06A9ABD5BECD4A7F9D9" pageId="7" pageNumber="731">
<paragraph id="1705368FE551B06A9ABD5BECD4A0FA3B" blockId="7.[160,1156,1470,1563]" pageId="7" pageNumber="731">
<keyLead id="AC4B95BAE551B06A9ABD5BECD4A0FA3B" pageId="7" pageNumber="731">
10. Dorsal processes of parameres carina shaped (
<figureCitation id="8F812A0AE551B06A98C65BECD31EFA14" box="[731,826,1470,1494]" captionStart="Figure 5" captionStartId="8.[160,225,1686,1709]" captionTargetBox="[170,1132,145,1646]" captionTargetId="figure@8.[166,1149,142,1654]" captionTargetPageId="8" captionText="Figure 5. Aegidinus colbyae, holotype male (a, e, d, f), paratype female (b, c). Habitus in dorsal view (a, b); female external genitalia (c); aedeagus in lateral (f) and ventral view (d); parameres in dorsal view (e)." figureDoi="http://doi.org/10.5281/zenodo.3675581" httpUri="https://zenodo.org/record/3675581/files/figure.png" pageId="7" pageNumber="731">Figure 5</figureCitation>
(e), arrowed)................................. ...........................................................................................................................
<taxonomicName id="D0BA4D0CE551B06A994B5BB3D40FFA3B" authority="Akhmetova &amp; Vaz-de-Mello, 2019" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2019" box="[854,1067,1505,1529]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="species" species="colbyae" status="sp. nov.">
<emphasis id="25CEEA9DE551B06A994B5BB3D40FFA3B" bold="true" box="[854,1067,1505,1529]" italics="true" pageId="7" pageNumber="731">Aegidinus colbyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EFD57E6E551B06A9E365BB3D4A0FA3B" box="[1067,1156,1505,1529]" pageId="7" pageNumber="731" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="1705368FE551B06A9ABD5851D4A7F9D9" blockId="7.[160,1156,1470,1563]" box="[160,1155,1539,1563]" pageId="7" pageNumber="731">
<keyLead id="AC4B95BAE551B06A9ABD5851D4A7F9D9" box="[160,1155,1539,1563]" pageId="7" pageNumber="731">- Dorsal processes of parameres tooth or spur shaped........................................................ 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="AC4E2E2AE551B06A9ABD586BD4A7F9B1" pageId="7" pageNumber="731">
<paragraph id="1705368FE551B06A9ABD586BD4A7F993" blockId="7.[160,1155,1593,1651]" box="[160,1155,1593,1617]" pageId="7" pageNumber="731">
<keyLead id="AC4B95BAE551B06A9ABD586BD4A7F993" box="[160,1155,1593,1617]" pageId="7" pageNumber="731">
11. Dorsal processes of parameres long, spur shaped........... ...........
<taxonomicName id="D0BA4D0CE551B06A999B586BD4A7F993" authorityName="Arrow" box="[902,1155,1593,1617]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="25CEEA9DE551B06A999B586BD4A7F993" bold="true" box="[902,1155,1593,1617]" italics="true" pageId="7" pageNumber="731">Aegidinus brasiliensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1705368FE551B06A9ABD5809D4A7F9B1" blockId="7.[160,1155,1593,1651]" box="[160,1155,1627,1651]" pageId="7" pageNumber="731">
<keyLead id="AC4B95BAE551B06A9ABD5809D4A7F9B1" box="[160,1155,1627,1651]" pageId="7" pageNumber="731">
- Dorsal processes of parameres short, tooth shaped............ ............
<taxonomicName id="D0BA4D0CE551B06A99B15809D4A7F9B1" authorityName="Colby" authorityYear="2009" box="[940,1155,1627,1651]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="731" phylum="Arthropoda" rank="species" species="howeae">
<emphasis id="25CEEA9DE551B06A99B15809D4A7F9B1" bold="true" box="[940,1155,1627,1651]" italics="true" pageId="7" pageNumber="731">Aegidinus howeae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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