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<document id="063746B240270592E313D02459ADDA61" ID-CLB-Dataset="269410" ID-DOI="10.5852/ejt.2023.891.2275" ID-GBIF-Dataset="ffda19c0-6b47-42e0-9959-4cb763e4ec4d" ID-ISSN="2118-9773" ID-Zenodo-Dep="8367911" ID-ZooBank="63FFD632-0F96-47B3-A42E-EA30DA7F766E" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1695315726654" checkinUser="plazi" docAuthor="Grosser, Clemens, Barjadze, Shalva, Maghradze, Eter, Shavadze, Lado, Pešić, Vladimir &amp; Faille, Arnaud" docDate="2023" docId="9E7B87ACFF852A50FE032CB13A5FBAB6" docLanguage="en" docName="EJT.2023.891.110-127.pdf" docOrigin="European Journal of Taxonomy 891" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2275/9799" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dina samegreloensis Grosser, Barjadze &amp; Shavadze 2023, sp. nov." docType="treatment" docUuid="4E387A11-FFF9-4ACF-99BA-2F1EC712BE79" docUuidSource="ZooBank" docVersion="4" lastPageNumber="121" masterDocId="6242FFD4FF832A5BFFB72F663F33BE3F" masterDocTitle="On the taxonomic status of Dina ratschaensis Kobakhidze, 1958 with a description of two new species - Dina imeretiensis sp. nov. and D. samegreloensis sp. nov. (Annelida, Hirudinida: Erpobdellidae)" masterLastPageNumber="127" masterPageNumber="110" pageNumber="116" updateTime="1699689684381" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F87A5E68774D72FA35AD076DE5181FA4">On the taxonomic status of Dina ratschaensis Kobakhidze, 1958 with a description of two new species - Dina imeretiensis sp. nov. and D. samegreloensis sp. nov. (Annelida, Hirudinida: Erpobdellidae)</mods:title>
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<mods:namePart id="C5B00ACBC071437CB7178165025D19E7">Grosser, Clemens</mods:namePart>
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<mods:affiliation id="A407A0EDC0DC913E64B386413DB31327">Independent researcher, Am Wasserturm 20, 04523 Elstertrebnitz, Germany. Ilia State University, Institute of Zoology, Giorgi Tsereteli 3, 0162 Tbilisi, Georgia. Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro. Department of Entomology, Coleoptera, Stuttgart State Museum of Natural History, Rosenstein 1, 70191 Stuttgart, Germany.</mods:affiliation>
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<mods:namePart id="711C64BF13115FE7A518399832C2849A">Maghradze, Eter</mods:namePart>
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<mods:affiliation id="C3794AB95A715751AC51B3A1A8EACF70">Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro.</mods:affiliation>
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<emphasis id="24A6EAA8FF852A5DFE032CB13D83BDCD" bold="true" box="[436,688,983,1010]" italics="true" pageId="6" pageNumber="116">Dina samegreloensis</emphasis>
Grosser, Barjadze &amp; Shavadze
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<paragraph id="166D36BAFF852A5DFD502B7B3C7EBA07" blockId="6.[365,1223,983,1080]" box="[743,845,1053,1080]" pageId="6" pageNumber="116">
<figureCitation id="8EE92A3FFF852A5DFD502B7B3C7EBA07" box="[743,845,1053,1080]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1892,1918]" captionStartId-1="8.[189,232,1841,1867]" captionTargetBox-0="[190,1398,862,1850]" captionTargetBox-1="[231,1398,306,1781]" captionTargetId-0="figure-213@7.[189,1398,862,1850]" captionTargetId-1="figure-11@8.[189,1398,266,1781]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." captionText-1="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi-0="http://doi.org/10.5281/zenodo.8367919" figureDoi-1="http://doi.org/10.5281/zenodo.8367921" httpUri-0="https://zenodo.org/record/8367919/files/figure.png" httpUri-1="https://zenodo.org/record/8367921/files/figure.png" pageId="6" pageNumber="116">Figs 4–5</figureCitation>
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<emphasis id="24A6EAA8FF852A5DFF0A2B393E05BA46" bold="true" box="[189,310,1119,1145]" pageId="6" pageNumber="116">Diagnosis</emphasis>
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Small-sized erpobdellids with a 
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<emphasis id="24A6EAA8FF852A5DFD9F2BE83D50BA98" box="[552,611,1166,1191]" italics="true" pageId="6" pageNumber="116">Dina</emphasis>
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-like heteronomous annulation.The midbody somites are subdivided into annuli b1, b2, a2, b5 and the broadened annulus b6 (
<figureCitation id="8EE92A3FFF852A5DFCC42BD73CE7BAF4" box="[883,980,1201,1227]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4F</figureCitation>
). The male genital pore is situated in furrow b2/a2 and the female one in furrow b6/b1. The genital pores are separated by three annuli (
<figureCitation id="8EE92A3FFF852A5DFF712B913E17BB2E" box="[198,292,1271,1297]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4G</figureCitation>
). Preserved leeches show numerous small and inconspicuous papillae on the dorsal and ventral sides. The cornua of the genital atrium are short, straight and directed slightly laterally (
<figureCitation id="8EE92A3FFF852A5DFB502A7C3A5ABB0A" box="[1255,1385,1306,1333]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="6" pageNumber="116">Fig. 5B–C</figureCitation>
). The vasa deferentia are very slightly curled up to the third ganglion behind the female genital pore. The ovisacs extend to the end of the second or the beginning of the third somite behind the female gonopore. They are unwinded to the end of the first somite and then coiled to the end (
<figureCitation id="8EE92A3FFF852A5DFB832AE23BA7BBA0" box="[1076,1172,1412,1439]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="6" pageNumber="116">Fig. 5A</figureCitation>
).
</paragraph>
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<emphasis id="24A6EAA8FF852A5DFF0A2AA33E77BBE0" bold="true" box="[189,324,1477,1503]" pageId="6" pageNumber="116">Etymology</emphasis>
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<paragraph id="166D36BAFF852A5DFF0A2A933A44B831" blockId="6.[189,1399,1524,1550]" box="[189,1399,1524,1550]" pageId="6" pageNumber="116">
<taxonomicName id="D1D24D39FF852A5DFF0A2A933E82B831" authority="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille, 2023" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[189,433,1524,1550]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="6" pageNumber="116" phylum="Annelida" rank="species" species="samegreloensis" status="sp. nov.">
<emphasis id="24A6EAA8FF852A5DFF0A2A933E82B831" box="[189,433,1524,1550]" italics="true" pageId="6" pageNumber="116">Dina samegreloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF852A5DFE012A923D3DB831" box="[438,526,1524,1550]" pageId="6" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
is named after the region of 
<collectingCountry id="6EC5762AFF852A5DFCE72A923C9DB831" box="[848,942,1524,1550]" name="Georgia" pageId="6" pageNumber="116">Georgia</collectingCountry>
from which the 
<typeStatus id="C9698818FF852A5DFBD02A923BFDB831" box="[1127,1230,1524,1550]" pageId="6" pageNumber="116" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="5EC86531FF852A5DFF0A29533A68B8D7" pageId="6" pageNumber="116" type="materials_examined">
<paragraph id="166D36BAFF852A5DFF0A29533E5FB870" blockId="6.[189,364,1589,1615]" box="[189,364,1589,1615]" pageId="6" pageNumber="116">
<heading id="4D2581D6FF852A5DFF0A29533E5FB870" bold="true" box="[189,364,1589,1615]" fontSize="11" level="3" pageId="6" pageNumber="116" reason="3">
<emphasis id="24A6EAA8FF852A5DFF0A29533E5FB870" bold="true" box="[189,364,1589,1615]" pageId="6" pageNumber="116">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF852A5DFF5D29023A68B8D7" blockId="6.[189,1398,1636,1768]" pageId="6" pageNumber="116">
<materialsCitation id="A6BA3CE7FF852A5DFF5D29023A68B8D7" ID-GBIF-Occurrence="4415817305" collectingDate="2021-10-07" collectionCode="IZISU" collectorName="Sh. Barjadze, L &amp; E. Maghradze" country="Georgia" elevation="492" latitude="42.476593" location="village Pirveli Balda" longLatPrecision="1" longitude="42.391182" municipality="Martvili Municipality" pageId="6" pageNumber="116" specimenCount="1" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="holotype">
<emphasis id="24A6EAA8FF852A5DFF5D29023E6FB841" bold="true" box="[234,348,1636,1662]" pageId="6" pageNumber="116">
<typeStatus id="C9698818FF852A5DFF5D29023E6FB841" box="[234,348,1636,1662]" pageId="6" pageNumber="116" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="6EC5762AFF852A5DFF0A29E13E0CB89E" box="[189,319,1671,1697]" name="Georgia" pageId="6" pageNumber="116">GEORGIA</collectingCountry>
• body length 
<quantity id="D12A9B5FFF852A5DFE4429E13D78B89D" box="[499,587,1671,1698]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="116" unit="mm" value="32.0">32 mm</quantity>
, width 
<quantity id="D12A9B5FFF852A5DFD1329E13DDEB89D" box="[676,749,1671,1698]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="116" unit="mm" value="6.0">6 mm</quantity>
, caudal sucker width 
<quantity id="D12A9B5FFF852A5DFC4529E13B08B89D" box="[1010,1083,1671,1698]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="116" unit="mm" value="4.0">4 mm</quantity>
; 
<collectingRegion id="D416F858FF852A5DFBFE29E13A45B89E" box="[1097,1398,1671,1697]" pageId="6" pageNumber="116">Samegrelo-Zemo Svaneti</collectingRegion>
region, Martvili Municipality, 
<location id="130D6061FF852A5DFD9B29CC3C14B8FB" LSID="urn:lsid:plazi:treatment:9E7B87ACFF852A50FE032CB13A5FBAB6:130D6061FF852A5DFD9B29CC3C14B8FB" box="[556,807,1706,1732]" country="Georgia" latitude="42.476593" longLatPrecision="1" longitude="42.391182" municipality="Martvili Municipality" name="village Pirveli Balda" pageId="6" pageNumber="116" stateProvince="Samegrelo-Zemo Svaneti">village Pirveli Balda</location>
, 
<taxonomicName id="D1D24D39FF852A5DFC8129CD3CE8B8FB" authority="Cave" authorityName="Cave" box="[822,987,1706,1733]" class="Malacostraca" family="Atyidae" genus="Xiphocaridinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="116" phylum="Arthropoda" rank="species" species="motena">
<location id="130D6061FF852A5DFC8129CD3CE8B8FB" LSID="urn:lsid:plazi:treatment:9E7B87ACFF852A50FE032CB13A5FBAB6:130D6061FF852A5DFC8129CD3CE8B8FB" box="[822,987,1706,1733]" country="Georgia" latitude="42.476593" longLatPrecision="1" longitude="42.391182" municipality="Martvili Municipality" name="Motena Cave" pageId="6" pageNumber="116" stateProvince="Samegrelo-Zemo Svaneti">Motena Cave</location>
</taxonomicName>
; 
<geoCoordinate id="73E6507DFF852A5DFC5D29CF3B94B8FA" box="[1002,1191,1705,1733]" degrees="42" direction="north" minutes="28" orientation="latitude" pageId="6" pageNumber="116" precision="1" seconds="35.73" value="42.476593">42°28′35.73″ N</geoCoordinate>
, 
<geoCoordinate id="73E6507DFF852A5DFB0029CF3A42B8FA" box="[1207,1393,1705,1733]" degrees="42" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="116" precision="1" seconds="28.25" value="42.391182">42°23′28.25″ E</geoCoordinate>
; altitude 
<quantity id="D12A9B5FFF852A5DFEAB29A83E59B8D7" box="[284,362,1742,1768]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="6" pageNumber="116" unit="m" value="492.0">
<elevation id="9DFFD189FF852A5DFEAB29A83E59B8D7" box="[284,362,1742,1768]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="6" pageNumber="116" unit="m" value="492.0">492 m</elevation>
</quantity>
; 
<date id="626C107AFF852A5DFEC129A83D30B8D7" box="[374,515,1742,1768]" pageId="6" pageNumber="116" value="2021-10-07">
<collectingDate id="7228E992FF852A5DFEC129A83D30B8D7" box="[374,515,1742,1768]" pageId="6" pageNumber="116" value="2021-10-07">7 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="BB27536CFF852A5DFDB829AB3DFAB8D7" box="[527,713,1741,1768]" pageId="6" pageNumber="116">Sh. Barjadze, L</collectorName>
. Shavadze and 
<collectorName id="BB27536CFF852A5DFCC929A83B11B8D8" box="[894,1058,1741,1768]" pageId="6" pageNumber="116">E. Maghradze</collectorName>
leg.; 
<collectionCode id="70C3AE7FFF852A5DFBD329A83B87B8D7" box="[1124,1204,1742,1768]" pageId="6" pageNumber="116">IZISU</collectionCode>
: AL-T 
<emphasis id="24A6EAA8FF852A5DFAB729A83A39B8D7" bold="true" box="[1280,1290,1742,1768]" pageId="6" pageNumber="116">-</emphasis>
00004.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5EC86531FF852A52FF0A28693B17BFC3" lastPageId="9" lastPageNumber="119" pageId="6" pageNumber="116" type="description">
<paragraph id="166D36BAFF852A5DFF0A28693E7DB916" blockId="6.[189,334,1807,1833]" box="[189,334,1807,1833]" pageId="6" pageNumber="116">
<heading id="4D2581D6FF852A5DFF0A28693E7DB916" bold="true" box="[189,334,1807,1833]" fontSize="11" level="3" pageId="6" pageNumber="116" reason="3">
<emphasis id="24A6EAA8FF852A5DFF0A28693E7DB916" bold="true" box="[189,334,1807,1833]" pageId="6" pageNumber="116">Description</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF852A5DFF0A28583BFAB9DB" blockId="6.[187,1399,1853,2021]" pageId="6" pageNumber="116">
<smallCapsWord id="108BA066FF852A5DFF0A28583E2EB969" baselines="1873,1873" box="[189,285,1854,1880]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="6" pageNumber="116">HABITUS</smallCapsWord>
. Small-sized erpobdellid. Preserved and contracted individuals reach a body length up to 
<quantity id="D12A9B5FFF852A5DFA9128583A44B967" box="[1318,1399,1854,1880]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="116" unit="mm" value="32.0">32 mm</quantity>
and a width up to 
<quantity id="D12A9B5FFF852A5DFE3A28063EE2B944" box="[397,465,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="116" unit="mm" value="6.0">6 mm</quantity>
(
<typeStatus id="C9698818FF852A5DFE5728073D78B944" box="[480,587,1889,1915]" pageId="6" pageNumber="116" type="holotype">holotype</typeStatus>
, 
<figureCitation id="8EE92A3FFF852A5DFDE028073DE4B944" box="[599,727,1889,1915]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4A–B</figureCitation>
). The body dorso-ventrally flattened in the posterior part, the first third (preclitellar and clitellar regions) cylindrical (
<figureCitation id="8EE92A3FFF852A5DFC3328E23CD7B9A1" box="[900,996,1924,1950]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4C</figureCitation>
). The mouth opening is wide and the upper lip barely noticeably elongated (
<figureCitation id="8EE92A3FFF852A5DFD7728C13C2CB9FE" box="[704,799,1959,1985]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4E</figureCitation>
). The caudal sucker is slightly wider than half of maximum body width (
<figureCitation id="8EE92A3FFF852A5DFE6728AC3D1CB9DB" box="[464,559,1994,2020]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="6" pageNumber="116">Fig. 4D</figureCitation>
). Small papillae numerous on dorsal and ventral surface.
</paragraph>
<paragraph id="166D36BAFF842A5CFF0A2E6F3E5EBFC9" blockId="7.[188,1400,265,502]" pageId="7" pageNumber="117">
<smallCapsWord id="108BA066FF842A5CFF0A2E6F3E79BF1D" baselines="284,285" box="[189,330,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Annulation" pageId="7" pageNumber="117">ANNULATION</smallCapsWord>
. The annulation is typical of the genus 
<taxonomicName id="D1D24D39FF842A5CFC992E6C3C5ABF1C" authorityName="R. Blanchard" authorityYear="1892" box="[814,873,266,291]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="7" pageNumber="117" phylum="Annelida" rank="genus">
<emphasis id="24A6EAA8FF842A5CFC992E6C3C5ABF1C" box="[814,873,266,291]" italics="true" pageId="7" pageNumber="117">Dina</emphasis>
</taxonomicName>
. The midbody somites are quinqueannulate and heteronomousely subdivided by clear furrows into annuli b1, b2, a2, b5 and the clearly broadened annulus b6. Annulus b1 is sometimes also slightly broadened, especially in the posterior part of the body (
<figureCitation id="8EE92A3FFF842A5CFF722E143E2DBFB2" box="[197,286,370,397]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="7" pageNumber="117">Fig. 4F</figureCitation>
). A tendency to split into further annuli is not visible. The male genital pore is situated in furrow b2/a2 and the female one in furrow b6/b1. The genital pores are separated by three annuli (
<figureCitation id="8EE92A3FFF842A5CFABF2EF03A59BF8F" box="[1288,1386,406,432]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="7" pageNumber="117">Fig. 4G</figureCitation>
). The dorsal and ventral sides are roughened by numerouse papillae. The papillae are very small and inconspicuous.
</paragraph>
<paragraph id="166D36BAFF842A5CFF0A2D783DD7BC64" blockId="7.[189,1398,542,603]" pageId="7" pageNumber="117">
<smallCapsWord id="108BA066FF842A5CFF0A2D783E64BC09" baselines="561,561" box="[189,343,542,568]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colouration" pageId="7" pageNumber="117">COLOURATION</smallCapsWord>
. The colouration of living specimen is pale pink. Preserved specimens are unicolored greyish without any dark patterns (
<figureCitation id="8EE92A3FFF842A5CFDE12D273DE4BC64" box="[598,727,577,603]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[190,1398,862,1850]" captionTargetId="figure-213@7.[189,1398,862,1850]" captionTargetPageId="7" captionText="Fig. 4. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), external morphology.A. Dorsal view. B. Ventral view. C. Lateral view. D. Caudal sucker. E. Oral sucker. F. Annulation of five midbody somites. G. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore." figureDoi="http://doi.org/10.5281/zenodo.8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="7" pageNumber="117">Fig. 4A–C</figureCitation>
).
</paragraph>
<paragraph id="166D36BAFF842A5CFF0A2DE53EDEBCA3" blockId="7.[189,493,642,669]" box="[189,493,642,669]" pageId="7" pageNumber="117">
<smallCapsWord id="108BA066FF842A5CFF0A2DE53FC5BCA4" baselines="662,662" box="[189,246,643,669]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Eyes" pageId="7" pageNumber="117">EYES</smallCapsWord>
. No eyes are visible.
</paragraph>
<paragraph id="166D36BAFF842A52FF0A2DA23A45BFB2" blockId="7.[187,1399,708,805]" lastBlockId="9.[189,1398,265,397]" lastPageId="9" lastPageNumber="119" pageId="7" pageNumber="117">
<smallCapsWord id="108BA066FF842A5CFF0A2DA23E20BCE3" baselines="727,727" box="[189,275,708,734]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sexual" pageId="7" pageNumber="117">SEXUAL</smallCapsWord>
<smallCapsWord id="108BA066FF842A5CFEAF2DAF3E5FBCE3" baselines="727" box="[280,364,713,732]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="7" pageNumber="117">ORGANS</smallCapsWord>
. The body of the genital atrium is large. The cornua are short, strong, straight and directed slightly laterally (
<figureCitation id="8EE92A3FFF842A5CFE3E2D813D3BBD3D" box="[393,520,743,770]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="7" pageNumber="117">Fig. 5B–C</figureCitation>
). The vasa deferentia are clearly offset from the cornua. They run relatively straight and only very slightly curled up to the third ganglion behind the female genital pore, then more coiled up to the end of the fifth somite behind the female genital pore (
<figureCitation id="8EE92A3FFF8A2A52FC582E6F3B7FBF1B" box="[1007,1100,265,292]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="9" pageNumber="119">Fig. 5A</figureCitation>
). The ovisacs run straight and unwinded to the end of the first somite behind the female genital pore. The right ovisac is strongly coiled and reaches the end of the second somite behind the female gonopore. The left ovisac has a slightly coiled end and reaches the annulus b2 of the third somite behind the female gonopore (
<figureCitation id="8EE92A3FFF8A2A52FABC2E143A5ABFB2" box="[1291,1385,370,397]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="9" pageNumber="119">Fig. 5A</figureCitation>
).
</paragraph>
<caption id="42AD6632FF842A5CFF0A28023A68B9D6" ID-DOI="http://doi.org/10.5281/zenodo.8367919" ID-Zenodo-Dep="8367919" httpUri="https://zenodo.org/record/8367919/files/figure.png" pageId="7" pageNumber="117" startId="7.[189,232,1892,1918]" targetBox="[190,1398,862,1850]" targetPageId="7">
<paragraph id="166D36BAFF842A5CFF0A28023A68B9D6" blockId="7.[187,1401,1892,2025]" pageId="7" pageNumber="117">
<emphasis id="24A6EAA8FF842A5CFF0A28023E3FB940" bold="true" box="[189,268,1892,1919]" pageId="7" pageNumber="117">Fig. 4.</emphasis>
<taxonomicName id="D1D24D39FF842A5CFEA128003CB6B940" authority="Grosser, Barjadze &amp; Shavadze" authorityName="Grosser, Barjadze &amp; Shavadze" authorityYear="2023" box="[278,901,1893,1919]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="7" pageNumber="117" phylum="Annelida" rank="species" species="samegreloensis" status="sp. nov.">
<emphasis id="24A6EAA8FF842A5CFEA128003D3CB940" box="[278,527,1893,1919]" italics="true" pageId="7" pageNumber="117">Dina samegreloensis</emphasis>
Grosser, Barjadze &amp; Shavadze
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF842A5CFC2528033CC2B940" box="[914,1009,1893,1919]" pageId="7" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IZISU: AL-T-00004), external morphology. 
<emphasis id="24A6EAA8FF842A5CFE0C28EE3EE3B99D" bold="true" box="[443,464,1928,1954]" pageId="7" pageNumber="117">A</emphasis>
. Dorsal view. 
<emphasis id="24A6EAA8FF842A5CFDC028EE3DB8B99D" bold="true" box="[631,651,1928,1954]" pageId="7" pageNumber="117">B</emphasis>
. Ventral view. 
<emphasis id="24A6EAA8FF842A5CFC8128EE3C78B99D" bold="true" box="[822,843,1928,1954]" pageId="7" pageNumber="117">C</emphasis>
. Lateral view. 
<emphasis id="24A6EAA8FF842A5CFC4128EE3B38B99D" bold="true" box="[1014,1035,1928,1954]" pageId="7" pageNumber="117">D</emphasis>
. Caudal sucker. 
<emphasis id="24A6EAA8FF842A5CFB7E28EE3BEEB99D" bold="true" box="[1225,1245,1928,1954]" pageId="7" pageNumber="117">E</emphasis>
. Oral sucker. 
<emphasis id="24A6EAA8FF842A5CFF0A28CD3FFCB9FA" bold="true" box="[189,207,1963,1989]" pageId="7" pageNumber="117">F</emphasis>
. Annulation of five midbody somites. 
<emphasis id="24A6EAA8FF842A5CFD3E28CD3D93B9FA" bold="true" box="[649,672,1963,1989]" pageId="7" pageNumber="117">G</emphasis>
. Position of genital pores and annulation in the clitellar region. Abbreviations: a2, b1, b2, b5, b6 = annuli of somites; f = female genital pore; m = male genital pore.
</paragraph>
</caption>
<caption id="42AD6632FF8B2A53FF0A28573A29B9AE" ID-DOI="http://doi.org/10.5281/zenodo.8367921" ID-Zenodo-Dep="8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="8" pageNumber="118" startId="8.[189,232,1841,1867]" targetBox="[231,1398,306,1781]" targetPageId="8">
<paragraph id="166D36BAFF8B2A53FF0A28573A29B9AE" blockId="8.[189,1398,1841,1938]" pageId="8" pageNumber="118">
<emphasis id="24A6EAA8FF8B2A53FF0A28573E3FB974" bold="true" box="[189,268,1841,1867]" pageId="8" pageNumber="118">Fig. 5.</emphasis>
<taxonomicName id="D1D24D39FF8B2A53FEA128543CB6B974" authority="Grosser, Barjadze &amp; Shavadze" authorityName="Grosser, Barjadze &amp; Shavadze" authorityYear="2023" box="[278,901,1841,1867]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="8" pageNumber="118" phylum="Annelida" rank="species" species="samegreloensis" status="sp. nov.">
<emphasis id="24A6EAA8FF8B2A53FEA128543D3CB974" box="[278,527,1841,1867]" italics="true" pageId="8" pageNumber="118">Dina samegreloensis</emphasis>
Grosser, Barjadze &amp; Shavadze
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF8B2A53FC3828543CDCB973" box="[911,1007,1842,1868]" pageId="8" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IZISU: AL-T-00004), sexual organs. 
<emphasis id="24A6EAA8FF8B2A53FED928323EB1B951" bold="true" box="[366,386,1876,1902]" pageId="8" pageNumber="118">A</emphasis>
. Schematic drawing of reproductive system. 
<emphasis id="24A6EAA8FF8B2A53FC2B28323C83B951" bold="true" box="[924,944,1876,1902]" pageId="8" pageNumber="118">B</emphasis>
. Schematic drawing of genital atrium. 
<emphasis id="24A6EAA8FF8B2A53FF0A28113FE1B9AE" bold="true" box="[189,210,1911,1937]" pageId="8" pageNumber="118">C</emphasis>
. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs.
</paragraph>
</caption>
<paragraph id="166D36BAFF8A2A52FF0B2ED23E71BFF1" blockId="9.[188,322,436,462]" box="[188,322,436,462]" pageId="9" pageNumber="119">
<heading id="4D2581D6FF8A2A52FF0B2ED23E71BFF1" bold="true" box="[188,322,436,462]" fontSize="11" level="3" pageId="9" pageNumber="119" reason="3">
<emphasis id="24A6EAA8FF8A2A52FF0B2ED23E71BFF1" bold="true" box="[188,322,436,462]" pageId="9" pageNumber="119">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF8A2A52FF0A2E843B17BFC3" blockId="9.[189,1060,482,509]" box="[189,1060,482,509]" pageId="9" pageNumber="119">
Information on variability is not yet possible. Only the 
<typeStatus id="C9698818FF8A2A52FCF32E843C98BFC3" box="[836,939,482,508]" pageId="9" pageNumber="119" type="holotype">holotype</typeStatus>
is known.
</paragraph>
</subSubSection>
<subSubSection id="5EC86531FF8A2A52FF0A2D423D77BCAF" pageId="9" pageNumber="119" type="biology_ecology">
<paragraph id="166D36BAFF8A2A52FF0A2D423E2EBC01" blockId="9.[189,285,548,574]" box="[189,285,548,574]" pageId="9" pageNumber="119">
<heading id="4D2581D6FF8A2A52FF0A2D423E2EBC01" bold="true" box="[189,285,548,574]" fontSize="11" level="3" pageId="9" pageNumber="119" reason="3">
<emphasis id="24A6EAA8FF8A2A52FF0A2D423E2EBC01" bold="true" box="[189,285,548,574]" pageId="9" pageNumber="119">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF8A2A52FF0A2D343D77BCAF" blockId="9.[188,1398,594,656]" pageId="9" pageNumber="119">
The single individual of this new leech species was found under a stone in the subterranean water stream in the dark zone of 
<taxonomicName id="D1D24D39FF8A2A52FE282D103D77BCAF" authority="Cave." authorityName="Cave." box="[415,580,630,656]" class="Malacostraca" family="Atyidae" genus="Xiphocaridinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="119" phylum="Arthropoda" rank="species" species="motena">Motena Cave.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5EC86531FF8A2A52FF0A2DD13C73BD3F" box="[189,832,695,768]" pageId="9" pageNumber="119" type="distribution">
<paragraph id="166D36BAFF8A2A52FF0A2DD13E65BCEE" blockId="9.[189,342,695,721]" box="[189,342,695,721]" pageId="9" pageNumber="119">
<heading id="4D2581D6FF8A2A52FF0A2DD13E65BCEE" bold="true" box="[189,342,695,721]" fontSize="11" level="3" pageId="9" pageNumber="119" reason="3">
<emphasis id="24A6EAA8FF8A2A52FF0A2DD13E65BCEE" bold="true" box="[189,342,695,721]" pageId="9" pageNumber="119">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF8A2A52FF0A2D803C73BD3F" blockId="9.[189,832,742,768]" box="[189,832,742,768]" pageId="9" pageNumber="119">
The new species is only known from the 
<typeStatus id="C9698818FF8A2A52FD282D803DE1BD3F" box="[671,722,742,768]" pageId="9" pageNumber="119">type</typeStatus>
location.
</paragraph>
</subSubSection>
<subSubSection id="5EC86531FF8A2A50FF0A2C413A5FBAB6" lastPageId="11" lastPageNumber="121" pageId="9" pageNumber="119" type="diagnosis">
<paragraph id="166D36BAFF8A2A52FF0A2C413EFDBD7E" blockId="9.[189,462,807,833]" box="[189,462,807,833]" pageId="9" pageNumber="119">
<heading id="4D2581D6FF8A2A52FF0A2C413EFDBD7E" bold="true" box="[189,462,807,833]" fontSize="11" level="3" pageId="9" pageNumber="119" reason="3">
<emphasis id="24A6EAA8FF8A2A52FF0A2C413EFDBD7E" bold="true" box="[189,462,807,833]" pageId="9" pageNumber="119">Differential diagnoses</emphasis>
</heading>
</paragraph>
<paragraph id="166D36BAFF8A2A52FF0A2C303A07BA7C" blockId="9.[187,1400,854,1091]" pageId="9" pageNumber="119">
<taxonomicName id="D1D24D39FF8A2A52FF0A2C303E56BD4F" authorityName="Johansson" authorityYear="1913" box="[189,357,854,880]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="species" species="absoloni">
<emphasis id="24A6EAA8FF8A2A52FF0A2C303E56BD4F" box="[189,357,854,880]" italics="true" pageId="9" pageNumber="119">Dina absoloni</emphasis>
</taxonomicName>
, a southeastern European cave leech was reported from 
<collectingCountry id="6EC5762AFF8A2A52FBBE2C303B55BD4F" box="[1033,1126,854,880]" name="Georgia" pageId="9" pageNumber="119">Georgia</collectingCountry>
by 
<bibRefCitation id="72434B4BFF8A2A52FB212C303A05BD4F" author="Lukin E. I." box="[1174,1334,854,880]" firstAuthor="Lukin" journalOrPublisher="Nauka, Leningrad" pageId="9" pageNumber="119" refId="ref7763" refString="Lukin E. I. 1976. Fauna USSR. Leeches: Leeches of Fresh and Brackish Waters. Nauka, Leningrad. [In Russian.]" title="Fauna USSR. Leeches: Leeches of Fresh and Brackish Waters" type="book" year="1976">Lukin (1976)</bibRefCitation>
. This species was not really found in 
<collectingCountry id="6EC5762AFF8A2A52FD952C1F3D4CBDAC" box="[546,639,889,915]" name="Georgia" pageId="9" pageNumber="119">Georgia</collectingCountry>
but was confused with other cave leeches (with 
<taxonomicName id="D1D24D39FF8A2A52FB132C1C3A64BDAC" authorityName="Kobakhidze" authorityYear="1958" box="[1188,1367,889,915]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF8A2A52FB132C1C3A64BDAC" box="[1188,1367,889,915]" italics="true" pageId="9" pageNumber="119">D. ratschaensis</emphasis>
</taxonomicName>
or an other similar species of 
<taxonomicName id="D1D24D39FF8A2A52FE502CFB3D12BD89" authorityName="R. Blanchard" authorityYear="1892" box="[487,545,925,950]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="genus">
<emphasis id="24A6EAA8FF8A2A52FE502CFB3D12BD89" box="[487,545,925,950]" italics="true" pageId="9" pageNumber="119">Dina</emphasis>
</taxonomicName>
, undescribed or here described; see also Discussion). 
<taxonomicName id="D1D24D39FF8A2A52FB342CFB3A15BD89" authorityName="Johansson" authorityYear="1913" box="[1155,1318,924,950]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="species" species="absoloni">
<emphasis id="24A6EAA8FF8A2A52FB342CFB3A15BD89" box="[1155,1318,924,950]" italics="true" pageId="9" pageNumber="119">Dina absoloni</emphasis>
</taxonomicName>
differs from species of 
<taxonomicName id="D1D24D39FF8A2A52FECA2CA63E84BDE6" authorityName="R. Blanchard" authorityYear="1892" box="[381,439,960,985]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="genus">
<emphasis id="24A6EAA8FF8A2A52FECA2CA63E84BDE6" box="[381,439,960,985]" italics="true" pageId="9" pageNumber="119">Dina</emphasis>
</taxonomicName>
living in Georgian karst caves by the position of the gonopores (the male genital pore is situated in furrow b1/b2 and the female one in furrow b5/b6) and differences in the reproductive system (ovisacs long, reach up to the eight ganglion behind the female gonopore; from the third ganglion onward, the ovisacs extend strong coiled alongside the nervous system to the caudal end; 
<figureCitation id="8EE92A3FFF8A2A52FB7B2B4F3A15BA7C" box="[1228,1318,1065,1091]" captionStart="Fig" captionStartId="10.[189,232,1727,1753]" captionTargetBox="[206,1397,278,1625]" captionTargetId="figure-11@10.[189,1398,265,1628]" captionTargetPageId="10" captionText="Fig. 7. Sexual organs of selected Dina spp.; schematic drawings and photograph. A. Dina ratschaensis from Sakishore Cave (Georgia). B. Dina cf. ratschaensis Kobakhidze, 1958 from Western Georgia (unknown cave). C. Dina absoloni Johansson, 1913 from Brestice (Herzegovina). A, C. Dissected by Grosser. B. Dissected by Epshtein (in Lukin 1976: fig. 263a–d; length × width of dissected specimens. A. 50 × 9 mm. B. b1 38 × 5 mm, b2 50 × 6 mm. C. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="9" pageNumber="119">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="166D36BAFF8A2A50FF0A2B0C3E9BBFC8" blockId="9.[189,1400,1130,1192]" lastBlockId="11.[189,1400,265,503]" lastPageId="11" lastPageNumber="121" pageId="9" pageNumber="119">
The cave dwelling leech 
<taxonomicName id="D1D24D39FF8A2A52FE5D2B0C3B39BABA" authority="Cichocka &amp; Bielecki, 2015" authorityName="Cichocka &amp; Bielecki" authorityYear="2015" box="[490,1034,1130,1157]" class="Clitellata" family="Erpobdellidae" genus="Erpobdella" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="species" species="borisi">
<emphasis id="24A6EAA8FF8A2A52FE5D2B0C3D88BABB" box="[490,699,1130,1156]" italics="true" pageId="9" pageNumber="119">Erpobdella borisi</emphasis>
<bibRefCitation id="72434B4BFF8A2A52FD742B0C3B39BABA" author="Cichocka J. M. &amp; Bielecki A. &amp; Kur J. &amp; Pikula D. &amp; Kilikowska A. &amp; Biernacka B." box="[707,1034,1130,1157]" firstAuthor="Cichocka" journalOrPublisher="Zootaxa" pageId="9" pageNumber="119" pagination="413 - 427" part="4013" refId="ref7219" refString="Cichocka J. M., Bielecki A., Kur J., Pikula D., Kilikowska A. &amp; Biernacka B. 2015. A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran. Zootaxa 4013 (3): 413 - 427. https: // doi. org / 10.11646 / zootaxa. 4013.3.5" title="A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran" type="journal article" year="2015">Cichocka &amp; Bielecki, 2015</bibRefCitation>
</taxonomicName>
, a species originally described from the Sahoolan Cave in northern 
<collectingCountry id="6EC5762AFF8A2A52FDDF2BE83DA4BA97" box="[616,663,1166,1192]" name="Iran" pageId="9" pageNumber="119">Iran</collectingCountry>
(
<collectingRegion id="D416F858FF8A2A52FD112BE83CEABA98" box="[678,985,1165,1192]" country="Iran" name="Azarbayjan-e Gharbi" pageId="9" pageNumber="119">West Azerbaijan Province</collectingRegion>
) shows a similar gonopore position and also a 
<taxonomicName id="D1D24D39FF882A50FEF72E6C3E4ABF1C" authorityName="R. Blanchard" authorityYear="1892" box="[320,377,266,291]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="genus">
<emphasis id="24A6EAA8FF882A50FEF72E6C3E4ABF1C" box="[320,377,266,291]" italics="true" pageId="11" pageNumber="121">Dina</emphasis>
</taxonomicName>
-like annulation with ring b6 divided into c11, c12. Therefore, the female gonopore is located in furrow b5/c11 (
<bibRefCitation id="72434B4BFF882A50FE422E4A3DC1BF78" author="Cichocka J. M. &amp; Bielecki A. &amp; Kur J. &amp; Pikula D. &amp; Kilikowska A. &amp; Biernacka B." box="[501,754,300,327]" etAl="et al." firstAuthor="Cichocka" journalOrPublisher="Zootaxa" pageId="11" pageNumber="121" pagination="413 - 427" part="4013" refId="ref7219" refString="Cichocka J. M., Bielecki A., Kur J., Pikula D., Kilikowska A. &amp; Biernacka B. 2015. A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran. Zootaxa 4013 (3): 413 - 427. https: // doi. org / 10.11646 / zootaxa. 4013.3.5" title="A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran" type="journal article" year="2015">
Cichocka 
<emphasis id="24A6EAA8FF882A50FDC72E4B3D90BF79" box="[624,675,300,326]" italics="true" pageId="11" pageNumber="121">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 4a–b). However, in 
<taxonomicName id="D1D24D39FF882A50FBAA2E4B3BA1BF78" box="[1053,1170,301,327]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="24A6EAA8FF882A50FBAA2E4B3B6BBF79" box="[1053,1112,301,326]" italics="true" pageId="11" pageNumber="121">Dina</emphasis>
spp.
</taxonomicName>
from the Georgian karst caves, the pore of the male genital organ is located in furrow b2/a2, and the female one in b6/b1. Further diffences are found in the shape of the reproductive system. The ovisacs of 
<taxonomicName id="D1D24D39FF882A50FB112E153A3FBFB3" authorityName="Cichocka &amp; Bielecki" authorityYear="2015" box="[1190,1292,370,396]" class="Clitellata" family="Erpobdellidae" genus="Erpobdella" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="borisi">
<emphasis id="24A6EAA8FF882A50FB112E153A3FBFB3" box="[1190,1292,370,396]" italics="true" pageId="11" pageNumber="121">E. borisi</emphasis>
</taxonomicName>
are long and extend to the seventh ganglion behind the female gonopore (
<bibRefCitation id="72434B4BFF882A50FC032EF03B98BF8E" author="Cichocka J. M. &amp; Bielecki A. &amp; Kur J. &amp; Pikula D. &amp; Kilikowska A. &amp; Biernacka B." box="[948,1195,406,433]" etAl="et al." firstAuthor="Cichocka" journalOrPublisher="Zootaxa" pageId="11" pageNumber="121" pagination="413 - 427" part="4013" refId="ref7219" refString="Cichocka J. M., Bielecki A., Kur J., Pikula D., Kilikowska A. &amp; Biernacka B. 2015. A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran. Zootaxa 4013 (3): 413 - 427. https: // doi. org / 10.11646 / zootaxa. 4013.3.5" title="A new leech species (Hirudinida: Erpobdellidae: Erpobdella) from a cave in the West Azerbaijan province of Iran" type="journal article" year="2015">
Cichocka 
<emphasis id="24A6EAA8FF882A50FB9D2EF13B6EBF8F" box="[1066,1117,406,432]" italics="true" pageId="11" pageNumber="121">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 5a), while in 
<taxonomicName id="D1D24D39FF882A50FF0A2EDC3E03BFEB" box="[189,304,442,468]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="24A6EAA8FF882A50FF0A2EDC3FCBBFEC" box="[189,248,442,467]" italics="true" pageId="11" pageNumber="121">Dina</emphasis>
spp.
</taxonomicName>
from Georgian karst caves they are short and extends at most to the third somite behind the female genital pore.
</paragraph>
<caption id="42AD6632FF8A2A52FF0B28A83BACB9D7" ID-DOI="http://doi.org/10.5281/zenodo.8367923" ID-Zenodo-Dep="8367923" box="[188,1183,1998,2025]" httpUri="https://zenodo.org/record/8367923/files/figure.png" pageId="9" pageNumber="119" startId="9.[188,231,1998,2024]" targetBox="[189,1398,1244,1959]" targetPageId="9">
<paragraph id="166D36BAFF8A2A52FF0B28A83BACB9D7" blockId="9.[188,1183,1998,2025]" box="[188,1183,1998,2025]" pageId="9" pageNumber="119">
<emphasis id="24A6EAA8FF8A2A52FF0B28A83E39B9D7" bold="true" box="[188,266,1998,2024]" pageId="9" pageNumber="119">Fig. 6.</emphasis>
<taxonomicName id="D1D24D39FF8A2A52FEA528A93DFBB9D6" authority="Kobakhidze, 1958" authorityName="Kobakhidze" authorityYear="1958" box="[274,712,1998,2025]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="119" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF8A2A52FEA528A93ED4B9D7" box="[274,487,1998,2024]" italics="true" pageId="9" pageNumber="119">Dina ratschaensis</emphasis>
<bibRefCitation id="72434B4BFF8A2A52FE5928A83DFBB9D6" author="Kobakhidze D. N." box="[494,712,1998,2025]" firstAuthor="Kobakhidze" journalOrPublisher="Bulletin of the Georgian SSR Academy of Sciences" pageId="9" pageNumber="119" pagination="591 - 592" part="21" refId="ref7696" refString="Kobakhidze D. N. 1958 New subspecies of the cave dwelling leech from Georgian SSR. Bulletin of the Georgian SSR Academy of Sciences 21 (5): 591 - 592. [In Russian.]" title="New subspecies of the cave dwelling leech from Georgian SSR" type="journal article" year="1958">Kobakhidze, 1958</bibRefCitation>
</taxonomicName>
. Living specimen from Sakishore Cave.
</paragraph>
</caption>
<caption id="42AD6632FF892A51FF0A29D93EF4B9B5" ID-DOI="http://doi.org/10.5281/zenodo.8367925" ID-Zenodo-Dep="8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="10" pageNumber="120" startId="10.[189,232,1727,1753]" targetBox="[206,1397,278,1625]" targetPageId="10">
<paragraph id="166D36BAFF892A51FF0A29D93EF4B9B5" blockId="10.[189,1399,1727,1930]" pageId="10" pageNumber="120">
<emphasis id="24A6EAA8FF892A51FF0A29D93E39B8E5" bold="true" box="[189,266,1727,1754]" pageId="10" pageNumber="120">Fig. 7.</emphasis>
Sexual organs of selected 
<taxonomicName id="D1D24D39FF892A51FDF229A63D89B8E5" box="[581,698,1728,1754]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="10" pageNumber="120" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="24A6EAA8FF892A51FDF229A63DB3B8E6" box="[581,640,1728,1753]" italics="true" pageId="10" pageNumber="120">Dina</emphasis>
spp.
</taxonomicName>
; schematic drawings and photograph. 
<emphasis id="24A6EAA8FF892A51FBCA29D93BA1B8E6" bold="true" box="[1149,1170,1727,1753]" pageId="10" pageNumber="120">A</emphasis>
. 
<taxonomicName id="D1D24D39FF892A51FB1529A63A44B8E5" authorityName="Kobakhidze" authorityYear="1958" box="[1186,1399,1728,1754]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="10" pageNumber="120" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF892A51FB1529A63A44B8E5" box="[1186,1399,1728,1754]" italics="true" pageId="10" pageNumber="120">Dina ratschaensis</emphasis>
</taxonomicName>
from Sakishore Cave (Georgia). 
<emphasis id="24A6EAA8FF892A51FDFA29853D52B8C2" bold="true" box="[589,609,1763,1789]" pageId="10" pageNumber="120">B</emphasis>
. 
<taxonomicName id="D1D24D39FF892A51FDC329823B6DB8C1" authority="Kobakhidze, 1958" authorityName="Kobakhidze" authorityYear="1958" box="[628,1118,1763,1790]" class="Clitellata" family="Erpobdellidae" genus="Dina" isUncertain="true" kingdom="Animalia" order="Arhynchobdellida" pageId="10" pageNumber="120" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF892A51FDC329823D9CB8C2" box="[628,687,1764,1789]" italics="true" pageId="10" pageNumber="120">Dina</emphasis>
cf. 
<emphasis id="24A6EAA8FF892A51FD5329853C44B8C2" box="[740,887,1763,1789]" italics="true" pageId="10" pageNumber="120">ratschaensis</emphasis>
<bibRefCitation id="72434B4BFF892A51FC3529853B6DB8C1" author="Kobakhidze D. N." box="[898,1118,1763,1790]" firstAuthor="Kobakhidze" journalOrPublisher="Bulletin of the Georgian SSR Academy of Sciences" pageId="10" pageNumber="120" pagination="591 - 592" part="21" refId="ref7696" refString="Kobakhidze D. N. 1958 New subspecies of the cave dwelling leech from Georgian SSR. Bulletin of the Georgian SSR Academy of Sciences 21 (5): 591 - 592. [In Russian.]" title="New subspecies of the cave dwelling leech from Georgian SSR" type="journal article" year="1958">Kobakhidze, 1958</bibRefCitation>
</taxonomicName>
from Western Georgia (unknown cave). 
<emphasis id="24A6EAA8FF892A51FE3B28603E92B91F" bold="true" box="[396,417,1798,1824]" pageId="10" pageNumber="120">C</emphasis>
. 
<taxonomicName id="D1D24D39FF892A51FE0528613C14B91F" authority="Johansson, 1913" authorityName="Johansson" authorityYear="1913" box="[434,807,1798,1824]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="10" pageNumber="120" phylum="Annelida" rank="species" species="absoloni">
<emphasis id="24A6EAA8FF892A51FE0528613D69B91F" box="[434,602,1798,1824]" italics="true" pageId="10" pageNumber="120">Dina absoloni</emphasis>
Johansson, 1913
</taxonomicName>
from Brestice (Herzegovina). 
<emphasis id="24A6EAA8FF892A51FB2328603BFDB91F" bold="true" box="[1172,1230,1798,1824]" pageId="10" pageNumber="120">A, C</emphasis>
. Dissected by Grosser. 
<emphasis id="24A6EAA8FF892A51FE91284F3E09B97C" bold="true" box="[294,314,1833,1859]" pageId="10" pageNumber="120">B</emphasis>
. Dissected by Epshtein (in 
<bibRefCitation id="72434B4BFF892A51FDC8284F3C3FB97C" author="Lukin E. I." box="[639,780,1833,1859]" firstAuthor="Lukin" journalOrPublisher="Nauka, Leningrad" pageId="10" pageNumber="120" refId="ref7763" refString="Lukin E. I. 1976. Fauna USSR. Leeches: Leeches of Fresh and Brackish Waters. Nauka, Leningrad. [In Russian.]" title="Fauna USSR. Leeches: Leeches of Fresh and Brackish Waters" type="book" year="1976">Lukin 1976</bibRefCitation>
: fig. 263a–d; length × width of dissected specimens. 
<emphasis id="24A6EAA8FF892A51FF0A282A3FE1B959" bold="true" box="[189,210,1868,1894]" pageId="10" pageNumber="120">A</emphasis>
. 50 × 9 mm. 
<emphasis id="24A6EAA8FF892A51FEC7282B3EB7B958" bold="true" box="[368,388,1869,1895]" pageId="10" pageNumber="120">B</emphasis>
. b1 38 × 5 mm, b2 50 × 6 mm. 
<emphasis id="24A6EAA8FF892A51FD4C282A3C23B959" bold="true" box="[763,784,1868,1894]" pageId="10" pageNumber="120">C</emphasis>
. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs.
</paragraph>
</caption>
<paragraph id="166D36BAFF882A50FF0A2D783A3BBCD7" blockId="11.[188,1399,541,744]" pageId="11" pageNumber="121">
However, with regard to the shape of the vasa deferentia and ovisacs (
<figureCitation id="8EE92A3FFF882A50FC562D783B72BC07" box="[993,1089,542,568]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[312,1343,310,1882]" captionTargetId="figure-18@5.[245,1343,265,1886]" captionTargetPageId="5" captionText="Fig. 3. Dina imeretiensis Grosser, Barjadze &amp; Maghradze sp. nov., holotype (IZISU: Al-T-00002), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367917" httpUri="https://zenodo.org/record/8367917/files/figure.png" pageId="11" pageNumber="121">Figs 3A</figureCitation>
, 
<figureCitation id="8EE92A3FFF882A50FBFB2D7B3B41BC07" box="[1100,1138,541,568]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="11" pageNumber="121">5A</figureCitation>
, 
<figureCitation id="8EE92A3FFF882A50FBCA2D783B90BC07" box="[1149,1187,542,568]" captionStart="Fig" captionStartId="10.[189,232,1727,1753]" captionTargetBox="[206,1397,278,1625]" captionTargetId="figure-11@10.[189,1398,265,1628]" captionTargetPageId="10" captionText="Fig. 7. Sexual organs of selected Dina spp.; schematic drawings and photograph. A. Dina ratschaensis from Sakishore Cave (Georgia). B. Dina cf. ratschaensis Kobakhidze, 1958 from Western Georgia (unknown cave). C. Dina absoloni Johansson, 1913 from Brestice (Herzegovina). A, C. Dissected by Grosser. B. Dissected by Epshtein (in Lukin 1976: fig. 263a–d; length × width of dissected specimens. A. 50 × 9 mm. B. b1 38 × 5 mm, b2 50 × 6 mm. C. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="11" pageNumber="121">7A</figureCitation>
) both new species show the greatest similarity with 
<taxonomicName id="D1D24D39FF882A50FDF52D243DCABC64" authorityName="Kobakhidze" authorityYear="1958" box="[578,761,577,603]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FDF52D243DCABC64" box="[578,761,577,603]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
. The vasa deferentia of 
<taxonomicName id="D1D24D39FF882A50FBA42D243BD9BC64" authority="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille, 2023" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[1043,1258,577,603]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="samegreloensis" status="sp. nov.">
<emphasis id="24A6EAA8FF882A50FBA42D243BD9BC64" box="[1043,1258,577,603]" italics="true" pageId="11" pageNumber="121">D. samegreloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF882A50FB462D273A7FBC64" box="[1265,1356,577,603]" pageId="11" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
are only very slightly curled up to the third ganglion behind the female genital pore. The vasa deferentia of 
<taxonomicName id="D1D24D39FF882A50FF0A2DEE3E5EBC9E" authority="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille, 2023" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[189,365,648,673]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis" status="sp. nov.">
<emphasis id="24A6EAA8FF882A50FF0A2DEE3E5EBC9E" box="[189,365,648,673]" italics="true" pageId="11" pageNumber="121">D. imeretiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF882A50FEC32DEE3EFDBC9D" box="[372,462,648,674]" pageId="11" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="D1D24D39FF882A50FDB22DEE3D88BC9E" authorityName="Kobakhidze" authorityYear="1958" box="[517,699,647,673]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FDB22DEE3D88BC9E" box="[517,699,647,673]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
are strongly curled along their entire course. The ovisacs of 
<taxonomicName id="D1D24D39FF882A50FF0A2DCD3E5DBCFB" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[189,366,683,708]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis">
<emphasis id="24A6EAA8FF882A50FF0A2DCD3E5DBCFB" box="[189,366,683,708]" italics="true" pageId="11" pageNumber="121">D. imeretiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="D1D24D39FF882A50FE112DCD3D6EBCFB" authorityName="Kobakhidze" authorityYear="1958" box="[422,605,682,708]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FE112DCD3D6EBCFB" box="[422,605,682,708]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
are strongly winded along their entire caudal course (
<figureCitation id="8EE92A3FFF882A50FB632DCC3A06BCFA" box="[1236,1333,682,709]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[312,1343,310,1882]" captionTargetId="figure-18@5.[245,1343,265,1886]" captionTargetPageId="5" captionText="Fig. 3. Dina imeretiensis Grosser, Barjadze &amp; Maghradze sp. nov., holotype (IZISU: Al-T-00002), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367917" httpUri="https://zenodo.org/record/8367917/files/figure.png" pageId="11" pageNumber="121">Figs 3A</figureCitation>
, 
<figureCitation id="8EE92A3FFF882A50FAF62DCD3A59BCFA" box="[1345,1386,683,709]" captionStart="Fig" captionStartId="10.[189,232,1727,1753]" captionTargetBox="[206,1397,278,1625]" captionTargetId="figure-11@10.[189,1398,265,1628]" captionTargetPageId="10" captionText="Fig. 7. Sexual organs of selected Dina spp.; schematic drawings and photograph. A. Dina ratschaensis from Sakishore Cave (Georgia). B. Dina cf. ratschaensis Kobakhidze, 1958 from Western Georgia (unknown cave). C. Dina absoloni Johansson, 1913 from Brestice (Herzegovina). A, C. Dissected by Grosser. B. Dissected by Epshtein (in Lukin 1976: fig. 263a–d; length × width of dissected specimens. A. 50 × 9 mm. B. b1 38 × 5 mm, b2 50 × 6 mm. C. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="11" pageNumber="121">7A</figureCitation>
). In contrast, the ovisacs of 
<taxonomicName id="D1D24D39FF882A50FE462DA83DFABCD7" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[497,713,718,744]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="samegreloensis">
<emphasis id="24A6EAA8FF882A50FE462DA83DFABCD7" box="[497,713,718,744]" italics="true" pageId="11" pageNumber="121">D. samegreloensis</emphasis>
</taxonomicName>
are only winded in their posterior half (
<figureCitation id="8EE92A3FFF882A50FB2A2DA83BC8BCD7" box="[1181,1275,717,744]" captionStart="Fig" captionStartId="8.[189,232,1841,1867]" captionTargetBox="[231,1398,306,1781]" captionTargetId="figure-11@8.[189,1398,266,1781]" captionTargetPageId="8" captionText="Fig. 5. Dina samegreloensis Grosser, Barjadze &amp; Shavadze sp. nov., holotype (IZISU: AL-T-00004), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367921" httpUri="https://zenodo.org/record/8367921/files/figure.png" pageId="11" pageNumber="121">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="166D36BAFF882A50FF0A2C763A5FBAB6" blockId="11.[188,1399,783,1162]" pageId="11" pageNumber="121">
<taxonomicName id="D1D24D39FF882A50FF0A2C763EA3BD16" authority="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille, 2023" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[189,400,784,809]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis" status="sp. nov.">
<emphasis id="24A6EAA8FF882A50FF0A2C763EA3BD16" box="[189,400,784,809]" italics="true" pageId="11" pageNumber="121">Dina imeretiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F9557D3FF882A50FE2C2C763ECABD15" box="[411,505,784,810]" pageId="11" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="D1D24D39FF882A50FD8E2C763DC7BD16" authorityName="Kobakhidze" authorityYear="1958" box="[569,756,783,809]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FD8E2C763DC7BD16" box="[569,756,783,809]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
can be clearly separated by the shape of the genital atrium. The cornua of the atrium in 
<taxonomicName id="D1D24D39FF882A50FDEB2C553C3EBD73" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[604,781,819,844]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis">
<emphasis id="24A6EAA8FF882A50FDEB2C553C3EBD73" box="[604,781,819,844]" italics="true" pageId="11" pageNumber="121">D. imeretiensis</emphasis>
</taxonomicName>
are nearly parallel and curved ventrally with straight ends (
<figureCitation id="8EE92A3FFF882A50FEB32C333EB5BD4F" box="[260,390,853,880]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[312,1343,310,1882]" captionTargetId="figure-18@5.[245,1343,265,1886]" captionTargetPageId="5" captionText="Fig. 3. Dina imeretiensis Grosser, Barjadze &amp; Maghradze sp. nov., holotype (IZISU: Al-T-00002), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367917" httpUri="https://zenodo.org/record/8367917/files/figure.png" pageId="11" pageNumber="121">Fig. 3B–C</figureCitation>
). In 
<taxonomicName id="D1D24D39FF882A50FE762C303D48BD50" authorityName="Kobakhidze" authorityYear="1958" box="[449,635,853,879]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FE762C303D48BD50" box="[449,635,853,879]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
the cornua are curved first laterally and then sharply to median and only slightly ventrally. The ends of the cornua are strongly kinked ventrally and not clearly offset from the vasa deferentia (
<figureCitation id="8EE92A3FFF882A50FE5F2CFA3D74BD89" box="[488,583,924,950]" captionStart="Fig" captionStartId="10.[189,232,1727,1753]" captionTargetBox="[206,1397,278,1625]" captionTargetId="figure-11@10.[189,1398,265,1628]" captionTargetPageId="10" captionText="Fig. 7. Sexual organs of selected Dina spp.; schematic drawings and photograph. A. Dina ratschaensis from Sakishore Cave (Georgia). B. Dina cf. ratschaensis Kobakhidze, 1958 from Western Georgia (unknown cave). C. Dina absoloni Johansson, 1913 from Brestice (Herzegovina). A, C. Dissected by Grosser. B. Dissected by Epshtein (in Lukin 1976: fig. 263a–d; length × width of dissected specimens. A. 50 × 9 mm. B. b1 38 × 5 mm, b2 50 × 6 mm. C. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="11" pageNumber="121">Fig. 7A</figureCitation>
). Further differences occur in the length of the ovisacs. The ovisacs of 
<taxonomicName id="D1D24D39FF882A50FF0B2CA63E5DBDE6" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[188,366,960,985]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis">
<emphasis id="24A6EAA8FF882A50FF0B2CA63E5DBDE6" box="[188,366,960,985]" italics="true" pageId="11" pageNumber="121">D. imeretiensis</emphasis>
</taxonomicName>
extend up to the second ganglion behind the female genital pore (
<figureCitation id="8EE92A3FFF882A50FBCF2CD93BE5BDE6" box="[1144,1238,959,985]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[312,1343,310,1882]" captionTargetId="figure-18@5.[245,1343,265,1886]" captionTargetPageId="5" captionText="Fig. 3. Dina imeretiensis Grosser, Barjadze &amp; Maghradze sp. nov., holotype (IZISU: Al-T-00002), sexual organs. A. Schematic drawing of reproductive system. B. Schematic drawing of genital atrium. C. Photograph of genital atrium. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367917" httpUri="https://zenodo.org/record/8367917/files/figure.png" pageId="11" pageNumber="121">Fig. 3A</figureCitation>
). The ovisacs of 
<taxonomicName id="D1D24D39FF882A50FF572C853EA9BDC3" authorityName="Kobakhidze" authorityYear="1958" box="[224,410,994,1020]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FF572C853EA9BDC3" box="[224,410,994,1020]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
are longer and extend up to the end of the second somite or up to the beginning of the third somite (on annulus b1) behind the female gonopore (
<figureCitation id="8EE92A3FFF882A50FC682B633B71BA1F" box="[991,1090,1029,1056]" captionStart="Fig" captionStartId="10.[189,232,1727,1753]" captionTargetBox="[206,1397,278,1625]" captionTargetId="figure-11@10.[189,1398,265,1628]" captionTargetPageId="10" captionText="Fig. 7. Sexual organs of selected Dina spp.; schematic drawings and photograph. A. Dina ratschaensis from Sakishore Cave (Georgia). B. Dina cf. ratschaensis Kobakhidze, 1958 from Western Georgia (unknown cave). C. Dina absoloni Johansson, 1913 from Brestice (Herzegovina). A, C. Dissected by Grosser. B. Dissected by Epshtein (in Lukin 1976: fig. 263a–d; length × width of dissected specimens. A. 50 × 9 mm. B. b1 38 × 5 mm, b2 50 × 6 mm. C. 30 × 6 mm. a = genital atrium; b = ovisacs; c = vas deferens; d = testisacs." figureDoi="http://doi.org/10.5281/zenodo.8367925" httpUri="https://zenodo.org/record/8367925/files/figure.png" pageId="11" pageNumber="121">Fig. 7A</figureCitation>
). The colouring of living specimens differs as well. The head, preclitellar region and caudal sucker of 
<taxonomicName id="D1D24D39FF882A50FB852B4F3BDBBA7C" authorityName="Kobakhidze" authorityYear="1958" box="[1074,1256,1065,1091]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="ratschaensis">
<emphasis id="24A6EAA8FF882A50FB852B4F3BDBBA7C" box="[1074,1256,1065,1091]" italics="true" pageId="11" pageNumber="121">D. ratschaensis</emphasis>
</taxonomicName>
are whitish, the clitellar and postclitellar regions are light brownish (
<figureCitation id="8EE92A3FFF882A50FCE32B2A3CAEBA59" box="[852,925,1100,1126]" captionStart="Fig" captionStartId="9.[188,231,1998,2024]" captionTargetBox="[189,1398,1244,1959]" captionTargetId="figure-298@9.[189,1398,1242,1963]" captionTargetPageId="9" captionText="Fig. 6. Dina ratschaensis Kobakhidze, 1958. Living specimen from Sakishore Cave." figureDoi="http://doi.org/10.5281/zenodo.8367923" httpUri="https://zenodo.org/record/8367923/files/figure.png" pageId="11" pageNumber="121">Fig. 6</figureCitation>
). 
<taxonomicName id="D1D24D39FF882A50FC022B2B3BB6BA59" authorityName="Grosser &amp; Barjadze &amp; Maghradze &amp; Shavadze &amp; Pešić &amp; Faille" authorityYear="2023" box="[949,1157,1101,1126]" class="Clitellata" family="Erpobdellidae" genus="Dina" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="121" phylum="Annelida" rank="species" species="imeretiensis">
<emphasis id="24A6EAA8FF882A50FC022B2B3BB6BA59" box="[949,1157,1101,1126]" italics="true" pageId="11" pageNumber="121">Dina imeretiensis</emphasis>
</taxonomicName>
shows a dark bluish postclitellar region, the caudal sucker is whitish and the other body regions are flesh coloured (
<figureCitation id="8EE92A3FFF882A50FAA22B093A69BAB6" box="[1301,1370,1135,1161]" captionStart="Fig" captionStartId="3.[188,232,1963,1989]" captionTargetBox="[189,1398,1104,1924]" captionTargetId="figure-282@3.[189,1398,1104,1924]" captionTargetPageId="3" captionText="Fig. 1. Dina imeretiensis Grosser, Barjadze &amp; Maghradze sp. nov. Living holotype and paratype from Prometheus Cave." figureDoi="http://doi.org/10.5281/zenodo.8367913" httpUri="https://zenodo.org/record/8367913/files/figure.png" pageId="11" pageNumber="121">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>