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<document id="74F8D05FAC5F5BF6940D79B6D5BD2B71" ID-CLB-Dataset="21546" ID-DOI="10.11646/zootaxa.4809.2.8" ID-GBIF-Dataset="0e06c542-08b9-4f44-81b5-a36a6c0561fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="3934243" ID-ZooBank="A8FECC8E-8BAC-4AF5-B695-C27A947A52AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594126326473" checkinUser="plazi" docAuthor="Lin, Yu-Chi, Braby, M. F. &amp; Hsu, Yu-Feng" docDate="2020" docId="9C078A1B2503FFA340CEA9FAFD94FACD" docLanguage="en" docName="zootaxa.4809.2.8.pdf" docOrigin="Zootaxa 4809 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epsteinius Lin, Braby &amp; Hsu 2020, gen. nov." docType="treatment" docUuid="48BE2E0B-CBFE-4F8E-8B77-D64C375B8BD7" docUuidSource="ZooBank" docVersion="8" lastPageNumber="376" masterDocId="603EF2632501FFA14059A91BFFC5FF9F" masterDocTitle="A new genus and species of slug caterpillar (Lepidoptera: Limacodidae) from Taiwan" masterLastPageNumber="382" masterPageNumber="374" pageNumber="376" updateTime="1698843161627" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E3506E78A90FB15B4E0BA97BBE6AFC80" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD4665C7672D84D8C94587FBEF047931">
<mods:title id="760094A06F3B15FF498DEF13012E2EDB">A new genus and species of slug caterpillar (Lepidoptera: Limacodidae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="A8C9709680C9C95417DE4D480586B581" type="personal">
<mods:role id="1130335E0068BFA0E0D4BA1E9D56F220">
<mods:roleTerm id="EC2A982FC739792EAFFFDE177D80E419">Author</mods:roleTerm>
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<mods:namePart id="4AC7286018A196792F51970D8ED0567D">Lin, Yu-Chi</mods:namePart>
</mods:name>
<mods:name id="26506C43BBFFA829CD02BC406D417219" type="personal">
<mods:role id="74C333855981A8678DA0E745FCD6198C">
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<mods:namePart id="215E0083BE967100156468CEA1894D6B">Braby, M. F.</mods:namePart>
</mods:name>
<mods:name id="39D8E87DD16FE45642D5D50CAE0082FB" type="personal">
<mods:role id="2FE01D2D2B9198F3050D50F194A3274B">
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<mods:namePart id="F72A4D59315185A3E8987AAE6640DE91">Hsu, Yu-Feng</mods:namePart>
</mods:name>
<mods:typeOfResource id="4530B0C1AF322ED65E051187FE5E06BA">text</mods:typeOfResource>
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<mods:title id="18A654B3F2880A00FE1825AAB87603F8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B4B38836A31EE5D354373A6107302FE9">
<mods:date id="F92F67E0228D4065A3BD264D492E9EEC">2020</mods:date>
<mods:detail id="1F619D5E4015F49228C3F17043EFF551" type="pubDate">
<mods:number id="942ACDE22E505A18A8CCDF838EBDB2B8">2020-07-07</mods:number>
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<mods:detail id="415743EED4E96DB6C4B4A380A545DD51" type="volume">
<mods:number id="620276D1811825C28B6DA046456D0C3A">4809</mods:number>
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<mods:detail id="CC42AD3BF134418D526589F14065726D" type="issue">
<mods:number id="D9B366BD28F39A21CBCACDDF2C3187EC">2</mods:number>
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<mods:classification id="07B4F1A1A1CCA959E9BB926E96F6D69F">journal article</mods:classification>
<mods:identifier id="BC05328AC1C6CE7B00DF1776E5ADB519" type="CLB-Dataset">21546</mods:identifier>
<mods:identifier id="D0536053AB6975A5EA1533635D5ED89D" type="DOI">10.11646/zootaxa.4809.2.8</mods:identifier>
<mods:identifier id="5F8F11E3034E3BC18B975250FAAE81D4" type="GBIF-Dataset">0e06c542-08b9-4f44-81b5-a36a6c0561fe</mods:identifier>
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<treatment id="9C078A1B2503FFA340CEA9FAFD94FACD" ID-DOI="http://doi.org/10.5281/zenodo.4324051" ID-GBIF-Taxon="164971402" ID-Zenodo-Dep="4324051" LSID="urn:lsid:zoobank.org:act:48BE2E0B-CBFE-4F8E-8B77-D64C375B8BD7" httpUri="http://treatment.plazi.org/id/9C078A1B2503FFA340CEA9FAFD94FACD" lastPageNumber="376" pageId="2" pageNumber="376">
<subSubSection id="5CB468862503FFA340CEA9FAFDB8FF63" box="[151,637,225,252]" pageId="2" pageNumber="376" type="nomenclature">
<paragraph id="14113B0D2503FFA340CEA9FAFDB8FF63" blockId="2.[151,686,225,288]" box="[151,637,225,252]" pageId="2" pageNumber="376">
<heading id="4F598C612503FFA340CEA9FAFDB8FF63" bold="true" box="[151,637,225,252]" fontSize="11" level="1" pageId="2" pageNumber="376" reason="1">
<emphasis id="26DAE71F2503FFA340CEA9FAFDB8FF63" bold="true" box="[151,637,225,252]" pageId="2" pageNumber="376">
<taxonomicName id="D3AE408E2503FFA340CEA9FAFDC7FF63" ID-CoL="9BDKF" authority="Lin, Braby &amp; Hsu" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[151,514,225,252]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="26DAE71F2503FFA340CEA9FAFED3FF64" bold="true" box="[151,278,225,251]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
Lin, Braby &amp; Hsu
</taxonomicName>
,
<taxonomicNameLabel id="3DE95A642503FFA34256A9F9FDB8FF63" box="[527,637,226,252]" pageId="2" pageNumber="376" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CB468862503FFA340CEA81EFC91FEFD" pageId="2" pageNumber="376" type="description">
<paragraph id="14113B0D2503FFA340CEA81EFD6BFEBF" blockId="2.[151,686,225,288]" box="[151,686,261,288]" pageId="2" pageNumber="376">
(
<figureCitation id="8C9527882503FFA340C6A81EFEE8FE80" box="[159,301,261,287]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,853,878]" captionTargetBox="[163,1433,188,827]" captionTargetId="figure@3.[154,1434,181,827]" captionTargetPageId="3" captionText="FIGURES 1–12. Adults of Epsteinius translucidus and Microleon longipalpis: 1, 2, E. translucidus Lin, sp. nov. holotype male upper side and underside; 3, 4, E. translucidus Lin, sp. nov. paratype female upper side and underside; 5, 6, M. longipalpis male upper side and underside; 7, 8, M. longipalpis female upper side and underside; 9, 10, M. longipalpis holotype male upper side and label data (Yokohama, Japan) (NHM); 11, 12, male heads of E. translucidus Lin, sp. nov. and M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934245" httpUri="https://zenodo.org/record/3934245/files/figure.png" pageId="2" pageNumber="376">Figs 1–4, 11</figureCitation>
,
<figureCitation id="8C9527882503FFA34160A81EFE44FE80" box="[313,385,261,287]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1067,1092]" captionTargetBox="[207,1429,181,1031]" captionTargetId="figure@4.[158,1429,181,1042]" captionTargetPageId="4" captionText="FIGURES 13–16. Male genitalia of Epsteinius translucidus and Microleon longipalpis: 13, 14, genitalia (ventral) and phallus (lateral) of E. translucidus Lin, sp. nov.; 15, 16, genitalia (ventral) and phallus (lateral) of M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934247" httpUri="https://zenodo.org/record/3934247/files/figure.png" pageId="2" pageNumber="376">13, 14</figureCitation>
,
<figureCitation id="8C9527882503FFA341D4A81EFE6EFE80" box="[397,427,261,287]" captionStart="FIGURES 17–18" captionStartId="5.[151,264,1902,1927]" captionTargetBox="[154,1434,418,1876]" captionTargetId="figure@5.[153,1434,417,1877]" captionTargetPageId="5" captionText="FIGURES 17–18. Female genitalia of Epsteinius translucidus and Microleon longipalpis: 17, E. translucidus Lin, sp. nov.; 18, M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934249" httpUri="https://zenodo.org/record/3934249/files/figure.png" pageId="2" pageNumber="376">17</figureCitation>
,
<figureCitation id="8C9527882503FFA341EEA81EFE3AFE80" box="[439,511,261,287]" captionStart="FIGURES 19–33" captionStartId="6.[151,264,1647,1672]" captionTargetBox="[152,1436,182,1621]" captionTargetId="figure@6.[151,1436,181,1622]" captionTargetPageId="6" captionText="FIGURES 19–33. Life history of Epsteinius translucidus and Microleon longipalpis: 19–23, M. longipalpis, showing dorsal and subdorsal views of early instar larva, dorsal view and subdorsal views of final instar larva, and cocoon without reverse neat and circular cut after adult emergence; 24–33, E. translucidus Lin, sp. nov.; 24–25, early instar larva, showing dorsal and subdorsal views; 26–27, final instar larva with brown lateral patch, showing dorsal and subdorsal views; 28, cocoon, without reverse neat and circular cut after adult emergence; 29–30, adult female, showing lateral and ventral views; 31–32, final instar larva without brown lateral patch, showing dorsal and subdorsal views; 33, final instar larva on host plant, showing transparent edge of larval body from abaxial leaf surface under sunshine." figureDoi="http://doi.org/10.5281/zenodo.3934251" httpUri="https://zenodo.org/record/3934251/files/figure.png" pageId="2" pageNumber="376">29, 30</figureCitation>
,
<figureCitation id="8C9527882503FFA34252A81EFD63FEBF" box="[523,678,261,288]" captionStart="FIGURES 34–46" captionStartId="7.[151,264,931,956]" captionTargetBox="[152,1436,181,906]" captionTargetId="figure@7.[152,1436,181,906]" captionTargetPageId="7" captionText="FIGURES 34–46. Head, pupa, and female 5th tarsomere of Epsteinius translucidus and Microleon longipalpis: 34–35, head with right labial palpus and galeae of M. longipalpis; 36–37, head with right labial palpus and galeae of E. translucidus Lin, sp. nov.; 38–39, eclosed pupa and ruptured pupal eye of M. longipalpis (arrow denotes fracture line); 40–42, uneclosed pupa, uneclosed pupal eye, and eclosed ruptured pupal eye of E. translucidus Lin, sp. nov. (arrow denotes fracture line); 43–44, female 5th tarsomere and sensilla of M.longipalpis; 45–46, female 5th tarsomere and sensilla of E. translucidus Lin, sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.3934253" httpUri="https://zenodo.org/record/3934253/files/figure.png" pageId="2" pageNumber="376">36, 37, 45, 46</figureCitation>
)
</paragraph>
<paragraph id="14113B0D2503FFA340CEA851FC91FEFD" blockId="2.[151,852,329,354]" box="[151,852,329,354]" pageId="2" pageNumber="376">
<typeStatus id="CB1585AF2503FFA340CEA851FF0CFEFD" box="[151,201,330,354]" pageId="2" pageNumber="376">Type</typeStatus>
species.
<taxonomicName id="D3AE408E2503FFA3417FA851FDFAFEFD" authority="Lin" authorityName="Lin" authorityYear="2020" box="[294,575,329,354]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="species" species="translucidus" status="sp. nov.">
<emphasis id="26DAE71F2503FFA3417FA851FDD6FEFD" box="[294,531,330,354]" italics="true" pageId="2" pageNumber="376">Epsteinius translucidus</emphasis>
Lin
</taxonomicName>
,
<taxonomicNameLabel id="3DE95A642503FFA34210A851FD5DFEFE" box="[585,664,330,353]" pageId="2" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="5CB468862503FFA340CEA894FF2DFA95" pageId="2" pageNumber="376" type="diagnosis">
<paragraph id="14113B0D2503FFA340CEA894FC10FD54" blockId="2.[151,1437,399,1363]" pageId="2" pageNumber="376">
<emphasis id="26DAE71F2503FFA340CEA894FED7FE35" bold="true" box="[151,274,399,426]" pageId="2" pageNumber="376">Diagnosis.</emphasis>
<taxonomicName id="D3AE408E2503FFA34142A88AFE55FE35" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[283,400,401,426]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34142A88AFE55FE35" box="[283,400,401,426]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
differs from
<taxonomicName id="D3AE408E2503FFA34272A88BFD5AFE35" authorityName="Butler" authorityYear="1885" box="[555,671,400,426]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34272A88BFD5AFE35" box="[555,671,400,426]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA342E9A88BFC81FE35" box="[688,836,399,426]" captionStart="FIGURES 1–12" captionStartId="3.[151,264,853,878]" captionTargetBox="[163,1433,188,827]" captionTargetId="figure@3.[154,1434,181,827]" captionTargetPageId="3" captionText="FIGURES 1–12. Adults of Epsteinius translucidus and Microleon longipalpis: 1, 2, E. translucidus Lin, sp. nov. holotype male upper side and underside; 3, 4, E. translucidus Lin, sp. nov. paratype female upper side and underside; 5, 6, M. longipalpis male upper side and underside; 7, 8, M. longipalpis female upper side and underside; 9, 10, M. longipalpis holotype male upper side and label data (Yokohama, Japan) (NHM); 11, 12, male heads of E. translucidus Lin, sp. nov. and M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934245" httpUri="https://zenodo.org/record/3934245/files/figure.png" pageId="2" pageNumber="376">Figs 5–9, 12</figureCitation>
,
<figureCitation id="8C9527882503FFA3430BA894FC58FE35" box="[850,925,399,426]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1067,1092]" captionTargetBox="[207,1429,181,1031]" captionTargetId="figure@4.[158,1429,181,1042]" captionTargetPageId="4" captionText="FIGURES 13–16. Male genitalia of Epsteinius translucidus and Microleon longipalpis: 13, 14, genitalia (ventral) and phallus (lateral) of E. translucidus Lin, sp. nov.; 15, 16, genitalia (ventral) and phallus (lateral) of M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934247" httpUri="https://zenodo.org/record/3934247/files/figure.png" pageId="2" pageNumber="376">15, 16</figureCitation>
,
<figureCitation id="8C9527882503FFA343F2A88BFC0CFE35" box="[939,969,400,426]" captionStart="FIGURES 17–18" captionStartId="5.[151,264,1902,1927]" captionTargetBox="[154,1434,418,1876]" captionTargetId="figure@5.[153,1434,417,1877]" captionTargetPageId="5" captionText="FIGURES 17–18. Female genitalia of Epsteinius translucidus and Microleon longipalpis: 17, E. translucidus Lin, sp. nov.; 18, M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934249" httpUri="https://zenodo.org/record/3934249/files/figure.png" pageId="2" pageNumber="376">18</figureCitation>
,
<figureCitation id="8C9527882503FFA34381A88BFBB9FE35" box="[984,1148,399,426]" captionStart="FIGURES 34–46" captionStartId="7.[151,264,931,956]" captionTargetBox="[152,1436,181,906]" captionTargetId="figure@7.[152,1436,181,906]" captionTargetPageId="7" captionText="FIGURES 34–46. Head, pupa, and female 5th tarsomere of Epsteinius translucidus and Microleon longipalpis: 34–35, head with right labial palpus and galeae of M. longipalpis; 36–37, head with right labial palpus and galeae of E. translucidus Lin, sp. nov.; 38–39, eclosed pupa and ruptured pupal eye of M. longipalpis (arrow denotes fracture line); 40–42, uneclosed pupa, uneclosed pupal eye, and eclosed ruptured pupal eye of E. translucidus Lin, sp. nov. (arrow denotes fracture line); 43–44, female 5th tarsomere and sensilla of M.longipalpis; 45–46, female 5th tarsomere and sensilla of E. translucidus Lin, sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.3934253" httpUri="https://zenodo.org/record/3934253/files/figure.png" pageId="2" pageNumber="376">34, 35, 43, 44</figureCitation>
;
<bibRefCitation id="703F46FC2503FFA344D3A88BFAFBFE35" author="Solovyev, A. V." box="[1162,1342,400,426]" pageId="2" pageNumber="376" pagination="17 - 43" refId="ref5247" refString="Solovyev, A. V. (2008) The limacodid moths (Lepidoptera, Limacodidae) of Russia. Eversmannia, 15 - 16, 17 - 43." type="journal article" year="2008">Solovyev, 2008</bibRefCitation>
;
<bibRefCitation id="703F46FC2503FFA34515A88BFF16FE51" author="Sasaki, A." pageId="2" pageNumber="376" pagination="99 - 106" refId="ref5146" refString="Sasaki, A. (2016) A taxonomic note of genus Microleon (Limacodidae) in Japan, with descriptions of two new species. Japan Heterocerists's Journal, 279, 99 - 106." type="journal article" year="2016">Sasaki, 2016</bibRefCitation>
;
<bibRefCitation id="703F46FC2503FFA34086A8AFFE61FE51" author="Sohn, J. - C. &amp; Kim, S. - S. &amp; Choi, S. - W." box="[223,420,436,463]" pageId="2" pageNumber="376" pagination="391 - 394" refId="ref5182" refString="Sohn, J. - C., Kim, S. - S. &amp; Choi, S. - W. (2018) Three species of Limacodidae (Lepidoptera, Zygaenoidea), new to Korea. Journal of Asia-Pacific Biodiversity, 11, 391 - 394. https: // doi. org / 10.1016 / j. japb. 2018.07.006" type="journal article" year="2018">
Sohn
<emphasis id="26DAE71F2503FFA34178A8AEFE91FE51" box="[289,340,436,462]" italics="true" pageId="2" pageNumber="376">et al</emphasis>
., 2018
</bibRefCitation>
) by the following characters: (1) adults are smaller in size—male forewing length of
<taxonomicName id="D3AE408E2503FFA3452DA8AEFF28FE6D" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3452DA8AEFF28FE6D" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
is
<quantity id="D35696E82503FFA34155A8CCFE57FE6C" box="[268,402,471,499]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="2" pageNumber="376" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
compared with
<quantity id="D35696E82503FFA3421CA8C3FD0EFE6C" box="[581,715,472,499]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.9" metricValueMin="6.3" pageId="2" pageNumber="376" unit="mm" value="7.1" valueMax="7.9" valueMin="6.3">6.3–7.9 mm</quantity>
for that of
<taxonomicName id="D3AE408E2503FFA3431FA8C3FC7FFE6D" authorityName="Butler" authorityYear="1885" box="[838,954,472,498]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3431FA8C3FC7FFE6D" box="[838,954,472,498]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
; (2) in
<taxonomicName id="D3AE408E2503FFA34452A8C2FB45FE6D" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[1035,1152,473,498]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34452A8C2FB45FE6D" box="[1035,1152,473,498]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
the hindwing is narrower with the termen rounded, whereas in
<taxonomicName id="D3AE408E2503FFA3426DA8E7FD6DFD89" authorityName="Butler" authorityYear="1885" box="[564,680,508,534]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3426DA8E7FD6DFD89" box="[564,680,508,534]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
it is broader with the termen distinctly angular; (3) the labial palpus of
<taxonomicName id="D3AE408E2503FFA340EFAB3AFEEEFDA5" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[182,299,545,570]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA340EFAB3AFEEEFDA5" box="[182,299,545,570]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
is shorter (approx. 2.0–2.5 times the diameter of the eye) than that of
<taxonomicName id="D3AE408E2503FFA34419AB3BFB71FDA5" authorityName="Butler" authorityYear="1885" box="[1088,1204,544,570]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34419AB3BFB71FDA5" box="[1088,1204,544,570]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
(approx. 3 times the diameter of the eye); (4) in
<taxonomicName id="D3AE408E2503FFA3418CAB5EFD8FFDC1" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[469,586,581,606]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3418CAB5EFD8FFDC1" box="[469,586,581,606]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
the upperside of the forewing has the inner half dark brown with a purple iridescent suffusion, whereas in
<taxonomicName id="D3AE408E2503FFA34259AB73FDB1FD1D" authorityName="Butler" authorityYear="1885" box="[512,628,616,642]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34259AB73FDB1FD1D" box="[512,628,616,642]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
there are usually two large irregular orange patches, one at base and the other in the median area of dorsum, and it lacks the purple suffusion; (5) in
<taxonomicName id="D3AE408E2503FFA343B3AB96FB9AFD39" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[1002,1119,653,678]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA343B3AB96FB9AFD39" box="[1002,1119,653,678]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
the scale fringe of the hindwing is grey, whereas in
<taxonomicName id="D3AE408E2503FFA341F4ABABFDE4FD55" authorityName="Butler" authorityYear="1885" box="[429,545,688,714]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA341F4ABABFDE4FD55" box="[429,545,688,714]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
it is conspicuously black at the tornus.
</paragraph>
<paragraph id="14113B0D2503FFA3409EABCEFAA3FC75" blockId="2.[151,1437,399,1363]" pageId="2" pageNumber="376">
In addition, the male genitalia of
<taxonomicName id="D3AE408E2503FFA34264ABCEFD77FD71" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[573,690,725,750]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34264ABCEFD77FD71" box="[573,690,725,750]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA34299ABCFFC84FD71" box="[704,833,724,750]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1067,1092]" captionTargetBox="[207,1429,181,1031]" captionTargetId="figure@4.[158,1429,181,1042]" captionTargetPageId="4" captionText="FIGURES 13–16. Male genitalia of Epsteinius translucidus and Microleon longipalpis: 13, 14, genitalia (ventral) and phallus (lateral) of E. translucidus Lin, sp. nov.; 15, 16, genitalia (ventral) and phallus (lateral) of M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934247" httpUri="https://zenodo.org/record/3934247/files/figure.png" pageId="2" pageNumber="376">Figs 13, 14</figureCitation>
) differs from that of
<taxonomicName id="D3AE408E2503FFA34477ABCFFB66FD71" authorityName="Butler" authorityYear="1885" box="[1070,1187,724,750]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34477ABCFFB66FD71" box="[1070,1187,724,750]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA344E8ABCFFAF7FD70" box="[1201,1330,723,751]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1067,1092]" captionTargetBox="[207,1429,181,1031]" captionTargetId="figure@4.[158,1429,181,1042]" captionTargetPageId="4" captionText="FIGURES 13–16. Male genitalia of Epsteinius translucidus and Microleon longipalpis: 13, 14, genitalia (ventral) and phallus (lateral) of E. translucidus Lin, sp. nov.; 15, 16, genitalia (ventral) and phallus (lateral) of M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934247" httpUri="https://zenodo.org/record/3934247/files/figure.png" pageId="2" pageNumber="376">Figs 15, 16</figureCitation>
) in many characters, including the form of the gnathos which has a sharply down curved tip, the valva which has two long processes along the dorsal margin, and the phallus which is shorter and basally enlarged and rounded with two conspicuous cornuti. In contrast, the genitalia of
<taxonomicName id="D3AE408E2503FFA342E7AA5BFCF7FCC5" authorityName="Butler" authorityYear="1885" box="[702,818,832,858]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA342E7AA5BFCF7FCC5" box="[702,818,832,858]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
has the gnathos with a T-shaped end, no process along the dorsal margin of the valva, and a relatively straight phallus without cornuti. The female genitalia of
<taxonomicName id="D3AE408E2503FFA34571AA7EFA58FCE1" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[1320,1437,869,894]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34571AA7EFA58FCE1" box="[1320,1437,869,894]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA340F9AA93FF36FC3D" box="[160,243,904,930]" captionStart="FIGURES 17–18" captionStartId="5.[151,264,1902,1927]" captionTargetBox="[154,1434,418,1876]" captionTargetId="figure@5.[153,1434,417,1877]" captionTargetPageId="5" captionText="FIGURES 17–18. Female genitalia of Epsteinius translucidus and Microleon longipalpis: 17, E. translucidus Lin, sp. nov.; 18, M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934249" httpUri="https://zenodo.org/record/3934249/files/figure.png" pageId="2" pageNumber="376">Fig. 17</figureCitation>
) differs from
<taxonomicName id="D3AE408E2503FFA341CCAA93FDCCFC3D" authorityName="Butler" authorityYear="1885" box="[405,521,904,930]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA341CCAA93FDCCFC3D" box="[405,521,904,930]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA34243AA93FDA8FC3D" box="[538,621,904,930]" captionStart="FIGURES 17–18" captionStartId="5.[151,264,1902,1927]" captionTargetBox="[154,1434,418,1876]" captionTargetId="figure@5.[153,1434,417,1877]" captionTargetPageId="5" captionText="FIGURES 17–18. Female genitalia of Epsteinius translucidus and Microleon longipalpis: 17, E. translucidus Lin, sp. nov.; 18, M. longipalpis." figureDoi="http://doi.org/10.5281/zenodo.3934249" httpUri="https://zenodo.org/record/3934249/files/figure.png" pageId="2" pageNumber="376">Fig. 18</figureCitation>
) in having a short and wide ductus bursae, and an ovate corpus bursae containing two signa each with a row of flat complex teeth. In
<taxonomicName id="D3AE408E2503FFA3430CAAB7FC0CFC59" authorityName="Butler" authorityYear="1885" box="[853,969,940,966]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3430CAAB7FC0CFC59" box="[853,969,940,966]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
, the ductus bursae is long and coiled, and the corpus bursae is ovate, containing only one signum with a set of sharp complex processes over its surface.
</paragraph>
<paragraph id="14113B0D2503FFA3409EAAEFFF2DFA95" blockId="2.[151,1437,399,1363]" pageId="2" pageNumber="376">
Although the early instar larvae of
<taxonomicName id="D3AE408E2503FFA3420DAAEEFD0CFB91" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[596,713,1013,1038]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3420DAAEEFD0CFB91" box="[596,713,1013,1038]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA34281AAEFFC9FFB91" box="[728,858,1011,1038]" captionStart="FIGURES 19–33" captionStartId="6.[151,264,1647,1672]" captionTargetBox="[152,1436,182,1621]" captionTargetId="figure@6.[151,1436,181,1622]" captionTargetPageId="6" captionText="FIGURES 19–33. Life history of Epsteinius translucidus and Microleon longipalpis: 19–23, M. longipalpis, showing dorsal and subdorsal views of early instar larva, dorsal view and subdorsal views of final instar larva, and cocoon without reverse neat and circular cut after adult emergence; 24–33, E. translucidus Lin, sp. nov.; 24–25, early instar larva, showing dorsal and subdorsal views; 26–27, final instar larva with brown lateral patch, showing dorsal and subdorsal views; 28, cocoon, without reverse neat and circular cut after adult emergence; 29–30, adult female, showing lateral and ventral views; 31–32, final instar larva without brown lateral patch, showing dorsal and subdorsal views; 33, final instar larva on host plant, showing transparent edge of larval body from abaxial leaf surface under sunshine." figureDoi="http://doi.org/10.5281/zenodo.3934251" httpUri="https://zenodo.org/record/3934251/files/figure.png" pageId="2" pageNumber="376">Figs 24, 25</figureCitation>
) and
<taxonomicName id="D3AE408E2503FFA343C3AAEFFBCBFB91" authorityName="Butler" authorityYear="1885" box="[922,1038,1012,1038]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA343C3AAEFFBCBFB91" box="[922,1038,1012,1038]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
spp. (
<figureCitation id="8C9527882503FFA3440AAAEFFB10FB91" box="[1107,1237,1012,1038]" captionStart="FIGURES 19–33" captionStartId="6.[151,264,1647,1672]" captionTargetBox="[152,1436,182,1621]" captionTargetId="figure@6.[151,1436,181,1622]" captionTargetPageId="6" captionText="FIGURES 19–33. Life history of Epsteinius translucidus and Microleon longipalpis: 19–23, M. longipalpis, showing dorsal and subdorsal views of early instar larva, dorsal view and subdorsal views of final instar larva, and cocoon without reverse neat and circular cut after adult emergence; 24–33, E. translucidus Lin, sp. nov.; 24–25, early instar larva, showing dorsal and subdorsal views; 26–27, final instar larva with brown lateral patch, showing dorsal and subdorsal views; 28, cocoon, without reverse neat and circular cut after adult emergence; 29–30, adult female, showing lateral and ventral views; 31–32, final instar larva without brown lateral patch, showing dorsal and subdorsal views; 33, final instar larva on host plant, showing transparent edge of larval body from abaxial leaf surface under sunshine." figureDoi="http://doi.org/10.5281/zenodo.3934251" httpUri="https://zenodo.org/record/3934251/files/figure.png" pageId="2" pageNumber="376">Figs 19, 20</figureCitation>
) are very similar, they diverge considerably in color pattern and shape in the final instar larvae, as follows: (1) in
<taxonomicName id="D3AE408E2503FFA344E7AD02FAF6FBAD" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[1214,1331,1049,1074]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA344E7AD02FAF6FBAD" box="[1214,1331,1049,1074]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
(
<figureCitation id="8C9527882503FFA34518AD03FECDFBC9" captionStart="FIGURES 19–33" captionStartId="6.[151,264,1647,1672]" captionTargetBox="[152,1436,182,1621]" captionTargetId="figure@6.[151,1436,181,1622]" captionTargetPageId="6" captionText="FIGURES 19–33. Life history of Epsteinius translucidus and Microleon longipalpis: 19–23, M. longipalpis, showing dorsal and subdorsal views of early instar larva, dorsal view and subdorsal views of final instar larva, and cocoon without reverse neat and circular cut after adult emergence; 24–33, E. translucidus Lin, sp. nov.; 24–25, early instar larva, showing dorsal and subdorsal views; 26–27, final instar larva with brown lateral patch, showing dorsal and subdorsal views; 28, cocoon, without reverse neat and circular cut after adult emergence; 29–30, adult female, showing lateral and ventral views; 31–32, final instar larva without brown lateral patch, showing dorsal and subdorsal views; 33, final instar larva on host plant, showing transparent edge of larval body from abaxial leaf surface under sunshine." figureDoi="http://doi.org/10.5281/zenodo.3934251" httpUri="https://zenodo.org/record/3934251/files/figure.png" pageId="2" pageNumber="376">Figs 26, 27, 31, 32</figureCitation>
) the shape of the body is elliptical, whereas in
<taxonomicName id="D3AE408E2503FFA3434CAD27FC4CFBC9" authorityName="Butler" authorityYear="1885" box="[789,905,1084,1110]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3434CAD27FC4CFBC9" box="[789,905,1084,1110]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
spp. (
<figureCitation id="8C9527882503FFA34395AD27FB8EFBC9" box="[972,1099,1084,1110]" captionStart="FIGURES 19–33" captionStartId="6.[151,264,1647,1672]" captionTargetBox="[152,1436,182,1621]" captionTargetId="figure@6.[151,1436,181,1622]" captionTargetPageId="6" captionText="FIGURES 19–33. Life history of Epsteinius translucidus and Microleon longipalpis: 19–23, M. longipalpis, showing dorsal and subdorsal views of early instar larva, dorsal view and subdorsal views of final instar larva, and cocoon without reverse neat and circular cut after adult emergence; 24–33, E. translucidus Lin, sp. nov.; 24–25, early instar larva, showing dorsal and subdorsal views; 26–27, final instar larva with brown lateral patch, showing dorsal and subdorsal views; 28, cocoon, without reverse neat and circular cut after adult emergence; 29–30, adult female, showing lateral and ventral views; 31–32, final instar larva without brown lateral patch, showing dorsal and subdorsal views; 33, final instar larva on host plant, showing transparent edge of larval body from abaxial leaf surface under sunshine." figureDoi="http://doi.org/10.5281/zenodo.3934251" httpUri="https://zenodo.org/record/3934251/files/figure.png" pageId="2" pageNumber="376">Figs 21, 22</figureCitation>
) it is narrower with a humped dorsal surface; (2) in
<taxonomicName id="D3AE408E2503FFA341DCAD7AFE3FFBE5" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[389,506,1121,1146]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA341DCAD7AFE3FFBE5" box="[389,506,1121,1146]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
there is usually an indistinct brown lateral patch (which varies among individuals), whereas in
<taxonomicName id="D3AE408E2503FFA34142AD9FFE4AFB01" authorityName="Butler" authorityYear="1885" box="[283,399,1156,1182]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA34142AD9FFE4AFB01" box="[283,399,1156,1182]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
spp. there is a distinct red subdorsal patch; (3) in
<taxonomicName id="D3AE408E2503FFA3438FAD9EFB8EFB01" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[982,1099,1157,1182]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3438FAD9EFB8EFB01" box="[982,1099,1157,1182]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
the body is semi-translucent, whereas in
<taxonomicName id="D3AE408E2503FFA3414EADB3FE4EFB5D" authorityName="Butler" authorityYear="1885" box="[279,395,1192,1218]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA3414EADB3FE4EFB5D" box="[279,395,1192,1218]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
spp. it is opaque; (4) in
<taxonomicName id="D3AE408E2503FFA342C5ADB2FCD4FB5D" authorityName="Lin, Braby &amp; Hsu" authorityYear="2020" box="[668,785,1193,1218]" class="Insecta" family="Limacodidae" genus="Epsteinius" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA342C5ADB2FCD4FB5D" box="[668,785,1193,1218]" italics="true" pageId="2" pageNumber="376">Epsteinius</emphasis>
</taxonomicName>
the spine-bearing protuberances are similar to those of its early instars, whereas in
<taxonomicName id="D3AE408E2503FFA341F0ADD7FDD8FB79" authorityName="Butler" authorityYear="1885" box="[425,541,1228,1254]" class="Insecta" family="Limacodidae" genus="Microleon" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="26DAE71F2503FFA341F0ADD7FDD8FB79" box="[425,541,1228,1254]" italics="true" pageId="2" pageNumber="376">Microleon</emphasis>
</taxonomicName>
spp. the protuberances in the final instar diminish in size compared with its early instars.
</paragraph>
</subSubSection>
<subSubSection id="5CB468862503FFA3409EAC0FFD94FACD" pageId="2" pageNumber="376" type="etymology">
<paragraph id="14113B0D2503FFA3409EAC0FFD94FACD" blockId="2.[151,1437,399,1363]" pageId="2" pageNumber="376">
<emphasis id="26DAE71F2503FFA3409EAC0FFE8BFAB1" bold="true" box="[199,334,1300,1326]" pageId="2" pageNumber="376">Etymology.</emphasis>
The generic name is named in honor of Marc E. Epstein for his enormous and longstanding contribution to the study of limacodid moths.
</paragraph>
</subSubSection>
</treatment>
</document>
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