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<document ID-DOI="http://dx.doi.org/10.3897/jhr.90.81095" ID-GBIF-Dataset="17f3d5ae-6dad-47ac-b4ea-2fd05fcfbfb2" ID-Pensoft-Pub="1314-2607-90-23" ID-Pensoft-UUID="3DC37DEB2F815E458BCE0B7DB8077814" ID-Zenodo-Dep="6508928" ID-ZooBank="EA0491B03B624360B9F79A24E3693248" ModsDocID="1314-2607-90-23" checkinTime="1651307064127" checkinUser="pensoft" docAuthor="Bordera, Santiago &amp; Santos, Bernardo F." docDate="2022" docId="97DCB175866359F9AFCB3BC6FD7A1A6A" docLanguage="en" docName="JourHymenoptRes 90: 23-57" docOrigin="Journal of Hymenoptera Research 90" docPubDate="2022-04-29" docSource="http://dx.doi.org/10.3897/jhr.90.81095" docTitle="Piasites nigricollis Bordera &amp; Santos 2022, sp. nov." docType="treatment" docUuid="623E4B24-7B9D-4B64-9907-325576744831" docUuidSource="ZooBank" docVersion="4" id="3DC37DEB2F815E458BCE0B7DB8077814" lastPageNumber="23" masterDocId="3DC37DEB2F815E458BCE0B7DB8077814" masterDocTitle="A review of Piasites Seyrig (Hymenoptera, Ichneumonidae, Cryptinae), with description of seven new species" masterLastPageNumber="57" masterPageNumber="23" pageNumber="23" updateTime="1678757756438" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>A review of Piasites Seyrig (Hymenoptera, Ichneumonidae, Cryptinae), with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bordera, Santiago</mods:namePart>
<mods:affiliation>Departamento de Ciencias Ambientales y Recursos Naturales, University of Alicante, E- 03690 Alicante, Spain</mods:affiliation>
<mods:nameIdentifier type="email">s.bordera@ua.es</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Bernardo F.</mods:namePart>
<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier CP 50, 75231, Paris Cedex 05, France</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>90</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>23</mods:start>
<mods:end>57</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.90.81095</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.90.81095</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-90-23</mods:identifier>
<mods:identifier type="ZooBank">EA0491B03B624360B9F79A24E3693248</mods:identifier>
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<treatment ID-GBIF-Taxon="194907331" LSID="urn:lsid:zoobank.org:act:623E4B24-7B9D-4B64-9907-325576744831" httpUri="http://treatment.plazi.org/id/97DCB175866359F9AFCB3BC6FD7A1A6A" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection pageId="0" pageNumber="23" type="nomenclature">
<paragraph pageId="0" pageNumber="23">
<taxonomicName LSID="http://zoobank.org/623E4B24-7B9D-4B64-9907-325576744831" authority="Bordera &amp; Santos, 2022" authorityName="Bordera &amp; Santos" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Piasites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piasites nigricollis" order="Hymenoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="nigricollis" status="sp. nov.">Piasites nigricollis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Propodeum of Piasites, dorsal view, females (II): A P. nigricollis sp. nov. (holotype) B P. orbitalis sp. nov. (holotype) C P. lineatus sp. nov. (paratype) D P. seyrigi sp. nov. (paratype). Scale bars: 0.3 mm." figureDoi="10.3897/jhr.90.81095.figure2" httpUri="https://binary.pensoft.net/fig/677619" pageId="0" pageNumber="23">Figs 2A</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Piasites nigricollis sp. nov., female (holotype): A habitus, lateral view B head, front view C head and mesosoma, lateral view D head and mesoscutum, dorsal view E metasoma, dorsal view F front wing, areolet and vein 2 m-cu. Scale bars: 1 mm (A); 0.5 mm (C, E); 0.3 mm (B, D, F)." figureDoi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23">, 6</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Piasites species, habitus of males, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. perinetensis sp. nov. (paratype) E P. nigricollis sp. nov. (paratype). Scale bars: 1 mm." figureDoi="10.3897/jhr.90.81095.figure12" httpUri="https://binary.pensoft.net/fig/677630" pageId="0" pageNumber="23">, 12E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">, 13D, F 14C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="diagnosis">
<paragraph pageId="0" pageNumber="23">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="23">
<taxonomicName authorityName="Bordera &amp; Santos" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Piasites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piasites nigricollis" order="Hymenoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis italics="true" pageId="0" pageNumber="23">Piasites nigricollis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from all other
<taxonomicName authorityName="Seyrig" authorityYear="1952" class="Insecta" family="Ichneumonidae" genus="Piasites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piasites" order="Hymenoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="23">Piasites</emphasis>
</taxonomicName>
species by the combination of the following characters: pronotum shallowly and longitudinally strigose, ventrally smooth (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Piasites nigricollis sp. nov., female (holotype): A habitus, lateral view B head, front view C head and mesosoma, lateral view D head and mesoscutum, dorsal view E metasoma, dorsal view F front wing, areolet and vein 2 m-cu. Scale bars: 1 mm (A); 0.5 mm (C, E); 0.3 mm (B, D, F)." figureDoi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23">6C</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13D</figureCitation>
); mesoscutum evenly convex, mostly smooth and shiny with fine punctures on median lobe (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Piasites nigricollis sp. nov., female (holotype): A habitus, lateral view B head, front view C head and mesosoma, lateral view D head and mesoscutum, dorsal view E metasoma, dorsal view F front wing, areolet and vein 2 m-cu. Scale bars: 1 mm (A); 0.5 mm (C, E); 0.3 mm (B, D, F)." figureDoi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23">6C, D</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13D</figureCitation>
); mesopleuron mostly smooth and shiny, shallowly rugulose punctate at dorsal anterior corner, ventral anterior part shallowly punctate (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Piasites nigricollis sp. nov., female (holotype): A habitus, lateral view B head, front view C head and mesosoma, lateral view D head and mesoscutum, dorsal view E metasoma, dorsal view F front wing, areolet and vein 2 m-cu. Scale bars: 1 mm (A); 0.5 mm (C, E); 0.3 mm (B, D, F)." figureDoi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23">6C</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13D</figureCitation>
); propodeum strongly rugose punctate (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Propodeum of Piasites, dorsal view, females (II): A P. nigricollis sp. nov. (holotype) B P. orbitalis sp. nov. (holotype) C P. lineatus sp. nov. (paratype) D P. seyrigi sp. nov. (paratype). Scale bars: 0.3 mm." figureDoi="10.3897/jhr.90.81095.figure2" httpUri="https://binary.pensoft.net/fig/677619" pageId="0" pageNumber="23">2A</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13D</figureCitation>
); propleuron, anterior part of pronotum and mesoscutum brownish black, ocular orbits completely white cream (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Piasites nigricollis sp. nov., female (holotype): A habitus, lateral view B head, front view C head and mesosoma, lateral view D head and mesoscutum, dorsal view E metasoma, dorsal view F front wing, areolet and vein 2 m-cu. Scale bars: 1 mm (A); 0.5 mm (C, E); 0.3 mm (B, D, F)." figureDoi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23">6C, D</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13D</figureCitation>
). Additionally, male with metapleuron smooth to shallowly rugulose punctate (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Piasites species, males A-E head and mesosoma, lateral view: A P. carinatus B P. quasimodus sp. nov. (paratype) C P. seyrigi sp. nov. (paratype) D P. nigricollis sp. nov. (paratype) E P. perinetensis sp. nov. (paratype) F-G metapleuron: F P. nigricollis sp. nov. (paratype) G P. perinetensis sp. nov. (paratype). Scale bars: 0.5 mm (A-E); 0.2 mm (F, G)." figureDoi="10.3897/jhr.90.81095.figure13" httpUri="https://binary.pensoft.net/fig/677631" pageId="0" pageNumber="23">13F</figureCitation>
).
</paragraph>
<caption doi="10.3897/jhr.90.81095.figure6" httpUri="https://binary.pensoft.net/fig/677624" pageId="0" pageNumber="23" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Figure 6.</emphasis>
<taxonomicName authorityName="Bordera &amp; Santos" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Piasites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piasites nigricollis" order="Hymenoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis italics="true" pageId="0" pageNumber="23">Piasites nigricollis</emphasis>
</taxonomicName>
sp. nov., female (holotype):
<emphasis bold="true" pageId="0" pageNumber="23">A</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="23">B</emphasis>
head, front view
<emphasis bold="true" pageId="0" pageNumber="23">C</emphasis>
head and mesosoma, lateral view
<emphasis bold="true" pageId="0" pageNumber="23">D</emphasis>
head and mesoscutum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="23">E</emphasis>
metasoma, dorsal view
<emphasis bold="true" pageId="0" pageNumber="23">F</emphasis>
front wing, areolet and vein 2m-cu. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="23">A</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="23">C, E</emphasis>
); 0.3 mm (
<emphasis bold="true" pageId="0" pageNumber="23">B, D, F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">Description.</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Female.</emphasis>
Fore wing length about 6.8 mm.
</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Head</emphasis>
.
</emphasis>
In dorsal view, moderately narrowed behind eyes, about 0.55
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Gena in dorsal view rounded, weakly swollen. Posterior ocellus separated from eye about 1.55
<normalizedToken originalValue="×">x</normalizedToken>
its maximum diameter. Distance between posterior ocelli 1.1
<normalizedToken originalValue="×">x</normalizedToken>
their diameter. Occipital carina complete, evenly curved dorsally. Face smooth and shiny, slightly convex in the central part with very fine setiferous punctures. Clypeal suture weak, only present laterally. Clypeus about 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as broad as medially long, very convex with relatively long setae in the central part, prominent in lateral view, ventral margin rounded, with a very weak median denticle. Malar space about 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width. Mandible 1.60
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal width; dorsal tooth distinctly longer than ventral one. Antenna with 28 flagellomeres, strongly tapered towards apex; flagellomeres from f11 to apex conspicuously flattened ventrally; f1 about 8.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as its maximum width.
</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Mesosoma</emphasis>
.
</emphasis>
Pronotum shallow and longitudinally strigose, ventrally becoming smooth; epomia strong and relatively long, straigth, crossing the anterior depression of pronotum. Mesoscutum 1.05
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, evenly convex, predominantly smooth and shiny, with fine and dense setiferous punctures on median lobe, lateral parts smooth; notaulus deep and narrow, with very small and inconspicuous transverse keels. Scutellum rugulose anteriorly, smooth posteriorly, lateral carinae strong, reaching 0.7 its anterior length. Mesopleuron predominantly smooth and shiny, shallowly rugulose punctate at dorsal anterior corner, ventral anterior part shallowly punctate; sternaulus deep, with very small transverse keels; epicnemial carina very weak, reaching the subtegular ridge. Metapleuron shallowly transverse strigose punctate; juxtacoxal and submetapleural carinae strong and complete. Propodeum strongly rugose punctate; anterior and posterior transverse carinae complete, centrally angled towards anterior part, parallel; posterior transverse carina forming two low lateral crests. Hind leg with femur about 6.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as high. Areolet pentagonal, about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as high as wide, relatively small, 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as high as abscissa of vein 2m-cu above bulla. Hind wing with vein cu-a about 0.5
<normalizedToken originalValue="×">x</normalizedToken>
length of first abscissa of CU.
</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Metasoma</emphasis>
.
</emphasis>
T1 about 2.45
<normalizedToken originalValue="×">x</normalizedToken>
as long as posteriorly broad, granulate, strongly curved dorsally in lateral view, dorso-lateral carina very weak, only present at postpetiole. T2 about 0.85
<normalizedToken originalValue="×">x</normalizedToken>
as long as posteriorly broad, finely granulate as the following tergites. Ovipositor sheath about 0.55
<normalizedToken originalValue="×">x</normalizedToken>
as long as hind tibia.
</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Colour</emphasis>
.
</emphasis>
Body mostly orange. Head yellow cream; mandibular teeth, ocellar triangle and central part of occiput, scape and pedicel brownish black; f1-3 orange, following flagellomeres dark brown, f5-10 white. Mesosoma with propleuron, dorso-lateral part of pronotum and anterior part of mesoscutum, brownish black; subventral spot on pronotum and subtegular ridge yellow cream. T4 dorsally, and anterior part of T5 and T6, black; posterior part of tergites T5 and T6 and following tergites yellow cream. Wings hyaline, slightly yellowish, pterostigma yellow.
</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Male.</emphasis>
Fore wing length about 7.7 mm.
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Head</emphasis>
.
</emphasis>
Posterior ocellus separated from eye about 1.45
<normalizedToken originalValue="×">x</normalizedToken>
its maximum diameter. Distance between hind ocelli about 1.15
<normalizedToken originalValue="×">x</normalizedToken>
maximum diameter of posterior ocellus. Clypeus about 1.65
<normalizedToken originalValue="×">x</normalizedToken>
as broad as medially long. Malar space about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width. Antenna with at least 24 flagellomeres, flagellum not enlarged subapically; flagellomeres from f17 to apex flattened ventrally; f1 about 6.0
<normalizedToken originalValue="×">x</normalizedToken>
long as its maximum width.
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Mesosoma</emphasis>
.
</emphasis>
Scutellum anteriorly punctate. Metapleuron shallowly rugolose punctate. Posterior transverse carina of propodeum evenly curved centrally. Hind leg with femur about 5.45
<normalizedToken originalValue="×">x</normalizedToken>
as long as high. Areolet about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as high as wide, 0.75
<normalizedToken originalValue="×">x</normalizedToken>
as high as abscissa of vein 2m-cu above bulla. Hind wing with vein cu-a 0.55
<normalizedToken originalValue="×">x</normalizedToken>
as long as first abscissa of CU.
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Metasoma</emphasis>
.
</emphasis>
T1 about 3.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as posteriorly broad. T2 about 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as posteriorly broad.
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">Colour</emphasis>
.
</emphasis>
As in female, but antenna with white band from apex of f7 to f15. Other features as in female.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="etymology">
<paragraph pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="23">
From the Latin
<emphasis italics="true" pageId="0" pageNumber="23">collum</emphasis>
, meaning
<normalizedToken originalValue="“neck”">&quot;neck&quot;</normalizedToken>
, in reference to the black-coloured pronotum and anterior part of mesoscutum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="materials_examined">
<paragraph pageId="0" pageNumber="23">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="23">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Ivondro, Museum Paris, A Seyrig" country="Madagascar" location="Madagascar" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="Holotype">
<specimenCount type="female">1 ♀</specimenCount>
<specimenCount type="male">1 ♂</specimenCount>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<specimenCount></specimenCount>
;
<collectorName>Ivondro</collectorName>
;
<collectorName>Museum Paris</collectorName>
; VII-40;
<collectorName>A Seyrig</collectorName>
; MNHN
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Ivondro &amp; Museum Paris &amp; A Seyrig" country="Madagascar" location="Madagascar" specimenCount="1" specimenCount-male="1" typeStatus="Paratype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">
<typeStatus>Paratype</typeStatus>
</emphasis>
:
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<specimenCount type="male">1♂</specimenCount>
; same collection data as for holotype; VIII-40; MNHN
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>