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<document id="10F46A51FCC3B2E946D4FAA83E3CD1DC" ID-CLB-Dataset="31625" ID-DOI="10.11646/zootaxa.4344.1.11" ID-GBIF-Dataset="ac5a7c87-f97d-468e-b6cf-288ce1960482" ID-ISSN="1175-5326" ID-Zenodo-Dep="1042227" ID-ZooBank="633EE23D-F2A3-41E2-8594-D5908B0B60D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509954947573" checkinUser="plazi" docAuthor="Song, Xiao-Bin, Maruyama, Munetoshi &amp; Peng, Zhong" docDate="2017" docId="963F87AFFFD2FFF5FF0575A176336FF9" docLanguage="en" docName="zootaxa.4344.1.11.pdf" docOrigin="Zootaxa 4344 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebioderus maolanus Song &amp; Maruyama &amp; Peng 2017" docType="treatment" docVersion="7" masterDocId="6A06FFD7FFD1FFF1FF92716B753D6962" masterDocTitle="Discovery of the genus Lebioderus (Coleoptera, Carabidae, Paussinae) from China, with description of two new species" updateTime="1698480712914" updateUser="ExternalLinkService">
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<mods:title id="8B18BE470E93AE5D4D5788FD6BE7DDAD">Discovery of the genus Lebioderus (Coleoptera, Carabidae, Paussinae) from China, with description of two new species</mods:title>
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<mods:namePart id="8792498E27C79BE20B2290B5052841FE">Song, Xiao-Bin</mods:namePart>
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<mods:namePart id="AC1D1F7ED24C636A75B5EA7E6F901A41">Maruyama, Munetoshi</mods:namePart>
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<mods:namePart id="0F5FAA8C693C9058C1BCC964BD2B90F5">Peng, Zhong</mods:namePart>
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<mods:date id="6C20BEE701685ACDA1230703DAECDC70">2017</mods:date>
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<treatment id="963F87AFFFD2FFF5FF0575A176336FF9" ID-DOI="http://doi.org/10.5281/zenodo.6038749" ID-GBIF-Taxon="137239271" ID-Zenodo-Dep="6038749" LSID="urn:lsid:plazi:treatment:963F87AFFFD2FFF5FF0575A176336FF9" httpUri="http://treatment.plazi.org/id/963F87AFFFD2FFF5FF0575A176336FF9" lastPageId="4" pageId="3">
<subSubSection id="568C6532FFD2FFF2FF0575A175C76C66" pageId="3" type="nomenclature">
<paragraph id="1E2936B9FFD2FFF2FF0575A176776D87" blockId="3.[151,1436,1226,2015]" box="[151,842,1226,1253]" pageId="3">
<heading id="456181D5FFD2FFF2FF0575A176776D87" bold="true" box="[151,842,1226,1253]" fontSize="11" level="1" pageId="3" reason="1">
<emphasis id="2CE2EAABFFD2FFF2FF0575A176776D87" bold="true" box="[151,842,1226,1253]" pageId="3">
<taxonomicName id="D9964D3AFFD2FFF2FF0575A174A16D86" ID-CoL="8MKRM" authorityName="Song &amp; Maruyama &amp; Peng" authorityYear="2017" box="[151,412,1226,1252]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="maolanus">
<emphasis id="2CE2EAABFFD2FFF2FF0575A174A16D86" bold="true" box="[151,412,1226,1252]" italics="true" pageId="3">Lebioderus maolanus</emphasis>
</taxonomicName>
Song and Maruyama, new species
</emphasis>
</heading>
</paragraph>
<paragraph id="1E2936B9FFD2FFF2FF05758675C76C66" blockId="3.[151,1436,1226,2015]" box="[151,250,1261,1284]" pageId="3">
(
<figureCitation id="86AD2A3CFFD2FFF2FF0D758675CF6C66" box="[159,242,1261,1284]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="3">Fig. 3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="568C6532FFD2FFF2FF05747877E06C2F" pageId="3" type="materials_examined">
<paragraph id="1E2936B9FFD2FFF2FF05747877E06C2F" blockId="3.[151,1436,1226,2015]" pageId="3">
<materialsCitation id="AEFE3CE4FFD2FFF2FF05747877E06C2F" ID-GBIF-Occurrence="1699341402" collectingDate="2015-07-19" collectionCode="SNUC" collectorName="Maolan N. R. &amp; Jia-Yao Hu" country="China" county="Libo County" elevation="750" latitude="25.277222" location="Libo County" longLatPrecision="21" longitude="107.92472" pageId="3" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="2CE2EAABFFD2FFF2FF057478747D6C48" bold="true" box="[151,320,1299,1322]" pageId="3">
<typeStatus id="C12D881BFFD2FFF2FF05747875F26C48" box="[151,207,1299,1322]" pageId="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="C12D881BFFD2FFF2FEDB7478748D6C48" box="[329,432,1299,1322]" pageId="3" type="holotype">Holotype</typeStatus>
:
<specimenCount id="0890FD30FFD2FFF2FE2C747874EF6C48" box="[446,466,1299,1322]" pageId="3" type="female"></specimenCount>
(
<collectionCode id="7887AE7CFFD2FFF2FE71747877126C48" box="[483,559,1299,1322]" pageId="3">SNUC</collectionCode>
), labeled
<collectingCountry id="66817629FFD2FFF2FD32747877C06C48" box="[672,765,1299,1322]" name="China" pageId="3">CHINA</collectingCountry>
:
<collectingRegion id="DC52F85BFFD2FFF2FC99747876576C48" box="[779,874,1299,1322]" country="China" name="Guizhou" pageId="3">Guizhou</collectingRegion>
,
<collectingCounty id="F7484E35FFD2FFF2FCE57478713E6C48" box="[887,1027,1299,1322]" pageId="3">Libo County</collectingCounty>
,
<collectorName id="B363536FFFD2FFF2FB83747871A26C48" box="[1041,1183,1299,1322]" pageId="3">Maolan N.R.</collectorName>
(″Ễ),
<geoCoordinate id="7BA2507EFFD2FFF2FA84747870AA6C48" box="[1302,1431,1299,1322]" direction="north" orientation="latitude" pageId="3" precision="15" value="25.277222">25°1638″N</geoCoordinate>
,
<geoCoordinate id="7BA2507EFFD2FFF2FF05745D741E6C2F" box="[151,291,1334,1357]" direction="east" orientation="longitude" pageId="3" precision="15" value="107.92472">107°5529″E</geoCoordinate>
, alt.
<elevation id="95BBD18AFFD2FFF2FEC7745D74A86C2F" box="[341,405,1334,1357]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" unit="m" value="750.0">
<quantity id="D96E9B5CFFD2FFF2FEC7745D74A86C2F" box="[341,405,1334,1357]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" unit="m" value="750.0">750m</quantity>
</elevation>
,
<collectingDate id="7A6CE991FFD2FFF2FE0D745D77196C2F" box="[415,548,1334,1357]" pageId="3" value="2015-07-19">
<date id="6A281079FFD2FFF2FE0D745D77196C2F" box="[415,548,1334,1357]" pageId="3" value="2015-07-19">19.VII.2015</date>
</collectingDate>
,
<collectorName id="B363536FFFD2FFF2FDBD745D779A6C2F" box="[559,679,1334,1357]" pageId="3">Jia-Yao Hu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="568C6532FFD2FFF2FF55743376D86C98" pageId="3" type="diagnosis">
<paragraph id="1E2936B9FFD2FFF2FF55743376D86C98" blockId="3.[151,1436,1226,2015]" pageId="3">
<emphasis id="2CE2EAABFFD2FFF2FF55743374016C0D" bold="true" box="[199,316,1368,1391]" pageId="3">Diagnosis.</emphasis>
The new species is very similar to
<taxonomicName id="D9964D3AFFD2FFF2FD23743276FA6C0D" authority="Nagel, 2009" authorityName="Nagel" authorityYear="2009" box="[689,967,1367,1392]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="2CE2EAABFFD2FFF2FD23743276026C0D" box="[689,831,1367,1392]" italics="true" pageId="3">L. brancuccii</emphasis>
<bibRefCitation id="7A074B48FFD2FFF2FCD7743376FA6C0D" author="Nagel" box="[837,967,1368,1391]" pageId="3" refString="Nagel, P. (2009) Flanged bombardier beetles from Laos (Carabidae, Paussinae). Entomologica Basiliensia et Collectionis Frey, 31, 101 - 113." type="journal article" year="2009">Nagel, 2009</bibRefCitation>
</taxonomicName>
(
<figureCitation id="86AD2A3CFFD2FFF2FC447433710E6C0D" box="[982,1075,1368,1391]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="3">Figs. 3B</figureCitation>
,
<figureCitation id="86AD2A3CFFD2FFF2FBAC743371416C0D" box="[1086,1148,1368,1391]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">4DF</figureCitation>
) in the shapes of antennae, pronotum and legs, but can be readily separated from it by the 1
<taxonomicName id="D9964D3AFFD2FFF2FCD27410769B6CF0" box="[832,934,1403,1426]" pageId="3" rank="subSpecies" subSpecies="ventral">st ventral</taxonomicName>
projection rounded at apex (
<figureCitation id="86AD2A3CFFD2FFF2FB4B7410700D6CF0" box="[1241,1328,1403,1426]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4A</figureCitation>
: arrow; in
<taxonomicName id="D9964D3AFFD2FFF2FF0574F574156CD7" authorityName="Nagel" authorityYear="2009" box="[151,296,1437,1461]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="2CE2EAABFFD2FFF2FF0574F574156CD7" box="[151,296,1437,1461]" italics="true" pageId="3">L. brancuccii</emphasis>
</taxonomicName>
, see
<figureCitation id="86AD2A3CFFD2FFF2FEF774F574806CD7" box="[357,445,1438,1461]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4D</figureCitation>
), longitudinal groove much wider near the posterior margin of head (
<figureCitation id="86AD2A3CFFD2FFF2FB5174F570266CD7" box="[1219,1307,1438,1461]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4B</figureCitation>
; head of
<taxonomicName id="D9964D3AFFD2FFF2FA1474F574346CB5" authorityName="Nagel" authorityYear="2009" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="2CE2EAABFFD2FFF2FA1474F574346CB5" italics="true" pageId="3">L. brancuccii</emphasis>
</taxonomicName>
, see
<figureCitation id="86AD2A3CFFD2FFF2FEAC74AB74B26CB5" box="[318,399,1472,1495]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4E</figureCitation>
), lateral projection of pronotum more developed (
<figureCitation id="86AD2A3CFFD2FFF2FC3074AB76C86CB5" box="[930,1013,1472,1495]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4C</figureCitation>
; pronotum of
<taxonomicName id="D9964D3AFFD2FFF2FB1E74AA70246CB5" authorityName="Nagel" authorityYear="2009" box="[1164,1305,1471,1496]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="2CE2EAABFFD2FFF2FB1E74AA70246CB5" box="[1164,1305,1471,1496]" italics="true" pageId="3">L. brancuccii</emphasis>
</taxonomicName>
, see
<figureCitation id="86AD2A3CFFD2FFF2FADD74AB70A16CB5" box="[1359,1436,1472,1495]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4F</figureCitation>
), and the glandular openings of pygidium not shaped as pronounced triangular.
</paragraph>
</subSubSection>
<subSubSection id="568C6532FFD2FFF5FF55776D712A6F51" lastPageId="4" pageId="3" type="description">
<paragraph id="1E2936B9FFD2FFF2FF55776D718E6F7F" blockId="3.[151,1436,1226,2015]" box="[199,1203,1542,1565]" pageId="3">
<emphasis id="2CE2EAABFFD2FFF2FF55776D748F6F7F" bold="true" box="[199,434,1542,1565]" pageId="3">Description. Female.</emphasis>
Body (
<figureCitation id="86AD2A3CFFD2FFF2FD93776D77626F7F" box="[513,607,1542,1565]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="3">Figs. 3A</figureCitation>
,
<figureCitation id="86AD2A3CFFD2FFF2FDF8776D77B76F7F" box="[618,650,1542,1565]" captionStart="FIGURE 5" captionStartId="6.[151,255,1073,1096]" captionTargetBox="[151,1436,194,1049]" captionTargetId="figure@6.[151,1436,194,1049]" captionTargetPageId="6" captionText="FIGURE 5. A. Lebioderus sinicus with Carebara ants. B. Wutongshan, Shenzhen, Guangdong, China. C. L. maolanus, sifted from leave litter. D. Type locality of L. maolanus (Maolan, Guizhou)." httpUri="https://zenodo.org/record/1042237/files/figure.png" pageId="3">5C</figureCitation>
)
<quantity id="D96E9B5CFFD2FFF2FD0B776D77C66F7F" box="[665,763,1542,1565]" metricMagnitude="-3" metricUnit="m" metricValue="7.84" pageId="3" unit="mm" value="7.84">7.84 mm</quantity>
; chestnut brown, but tibiae slightly paler.
</paragraph>
<paragraph id="1E2936B9FFD2FFF2FF557743718C6FE7" blockId="3.[151,1436,1226,2015]" pageId="3">
Head (
<figureCitation id="86AD2A3CFFD2FFF2FE9C774374516F5D" box="[270,364,1576,1599]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="3">Figs. 3A</figureCitation>
,
<figureCitation id="86AD2A3CFFD2FFF2FEEA774374A56F5D" box="[376,408,1576,1599]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">4B</figureCitation>
) with distinct longitudinal groove along the midline; disc with shallow depressions at both sides of midline; temples well protruded laterad; dorsal surface matte, finely, densely punctate; sparsely covered with minute, recumbent setae; eyes rather large, less prominent than temples; neck narrow, strongly constricted.
</paragraph>
<paragraph id="1E2936B9FFD2FFF2FF5577FB719D6E6D" blockId="3.[151,1436,1226,2015]" pageId="3">
Antennae (
<figureCitation id="86AD2A3CFFD2FFF2FEAD77FB74A86FC5" box="[319,405,1680,1703]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4A</figureCitation>
) with segment I parallel-sided; Antennal club with four distinct transverse furrows; posterior margin with five produced denticles, 2nd dorsal denticles short; 3rd dorsal denticles almost as long as 4th and 5th; 2nd to 5th dorsal denticles apically with short trichome brush; anterior margin with 1st to 5th projections rounded at apex; each projection with a few short setae at apex; surface sparsely covered with minute, recumbent setae.
</paragraph>
<paragraph id="1E2936B9FFD2FFF2FF55767075E96EF8" blockId="3.[151,1436,1226,2015]" pageId="3">
Pronotum (
<figureCitation id="86AD2A3CFFD2FFF2FEAC767074B26E50" box="[318,399,1819,1842]" captionStart="FIGURE 4" captionStartId="5.[151,255,1556,1579]" captionTargetBox="[165,1424,273,1517]" captionTargetId="figure@5.[151,1436,263,1531]" captionTargetPageId="5" captionText="FIGURE 4. Lebioderus maolanus sp. nov. A. Antennal club B. Head C. Pronotum. L. brancuccii D. Antennal club E. Head F. Pronotum. Scales: A F = 0.5 mm." httpUri="https://zenodo.org/record/1042235/files/figure.png" pageId="3">Fig. 4C</figureCitation>
) broad, about 1.75 times as wide as long; distinctly wider than head; divided by a transverse cleft; fore part about 0.92 times as long as the hind part; both ends of longitudinal groove indistinct around 1/2 of anterior part and anterior 1/3 of posterior part; dorsal surface matte, finely, densely punctate; sparsely covered with minute, recumbent setae.
</paragraph>
<paragraph id="1E2936B9FFD2FFF2FF5576CD71D16EBD" blockId="3.[151,1436,1226,2015]" pageId="3">
Elytra (
<figureCitation id="86AD2A3CFFD2FFF2FE8576CD74576EDF" box="[279,362,1958,1981]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="3">Fig. 3A</figureCitation>
) with 7 to 8 pairs of long setae on lateral margins and 2 pairs of long setae along the anterior part of midline; shoulders well protruded; dorsal surface matte; sparsely covered with minute, recumbent setae.
</paragraph>
<caption id="4AE96631FFD5FFF5FF057441740F6C01" httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="4" targetBox="[161,1421,206,1282]" targetPageId="4">
<paragraph id="1E2936B9FFD5FFF5FF057441740F6C01" blockId="4.[151,1437,1321,1379]" pageId="4">
<emphasis id="2CE2EAABFFD5FFF5FF05744174256C23" bold="true" box="[151,280,1322,1345]" pageId="4">FIGURE 3.</emphasis>
Dorsal habitus of
<taxonomicName id="D9964D3AFFD5FFF5FE4F744277686C23" authorityName="Westwood" authorityYear="1838" box="[477,597,1321,1345]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="2CE2EAABFFD5FFF5FE4F744277686C23" box="[477,597,1321,1345]" italics="true" pageId="4">Lebioderus</emphasis>
</taxonomicName>
species. A.
<taxonomicName id="D9964D3AFFD5FFF5FD44744176646C23" authority="Song &amp; Maruyama &amp; Peng, 2017" authorityName="Song &amp; Maruyama &amp; Peng" authorityYear="2017" box="[726,857,1321,1345]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="maolanus" status="sp. nov.">
<emphasis id="2CE2EAABFFD5FFF5FD44744176646C23" box="[726,857,1321,1345]" italics="true" pageId="4">L. maolanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37D157D0FFD5FFF5FCF2744176886C23" box="[864,949,1322,1345]" pageId="4" rank="species">
<emphasis id="2CE2EAABFFD5FFF5FCF2744176886C23" bold="true" box="[864,949,1322,1345]" pageId="4">sp. nov.</emphasis>
</taxonomicNameLabel>
, female (Holotype). B.
<taxonomicName id="D9964D3AFFD5FFF5FB20744170026C23" authorityName="Nagel" authorityYear="2009" box="[1202,1343,1321,1345]" class="Insecta" family="Carabidae" genus="Lebioderus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="2CE2EAABFFD5FFF5FB20744170026C23" box="[1202,1343,1321,1345]" italics="true" pageId="4">L. brancuccii</emphasis>
</taxonomicName>
, female. Scale = 2 mm.
</paragraph>
</caption>
<paragraph id="1E2936B9FFD5FFF5FF5574F977AD6CA9" blockId="4.[151,1435,1426,1691]" pageId="4">
Pygidium (
<figureCitation id="86AD2A3CFFD5FFF5FED374F974A76CCB" box="[321,410,1426,1449]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="4">Fig. 3A</figureCitation>
) with sharp edge; glandular openings situated proximally at the left and right margin; disc smooth, moderately covered with minute setae.
</paragraph>
<paragraph id="1E2936B9FFD5FFF5FF5574BC71C76C8C" blockId="4.[151,1435,1426,1691]" box="[199,1274,1495,1518]" pageId="4">
Legs (
<figureCitation id="86AD2A3CFFD5FFF5FE9B74BC74606C8C" box="[265,349,1495,1518]" captionStart="FIGURE 3" captionStartId="4.[151,255,1322,1345]" captionTargetBox="[161,1421,206,1282]" captionTargetId="figure@4.[151,1436,194,1297]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Lebioderus species. A. L. maolanus sp. nov., female (Holotype). B. L. brancuccii, female. Scale = 2 mm." httpUri="https://zenodo.org/record/1042233/files/figure.png" pageId="4">Fig. 3A</figureCitation>
) robust; tibiae almost straight, strongly compressed; sparsely covered with short setae.
</paragraph>
<paragraph id="1E2936B9FFD5FFF5FF55749174416F73" blockId="4.[151,1435,1426,1691]" box="[199,380,1530,1553]" pageId="4">Male. Unknown.</paragraph>
<paragraph id="1E2936B9FFD5FFF5FF557777712A6F51" blockId="4.[151,1435,1426,1691]" box="[199,1047,1564,1587]" pageId="4">
<emphasis id="2CE2EAABFFD5FFF5FF55777774486F51" bold="true" box="[199,373,1564,1587]" pageId="4">Measurements.</emphasis>
BL, 7.84; HW, 1.59; ACL, 1.75; PL, 1.28; PW, 2.24; EL, 5.23.
</paragraph>
</subSubSection>
<subSubSection id="568C6532FFD5FFF5FF557754771F6F1B" pageId="4" type="etymology">
<paragraph id="1E2936B9FFD5FFF5FF55775477B76F34" blockId="4.[151,1435,1426,1691]" box="[199,650,1599,1622]" pageId="4">
<emphasis id="2CE2EAABFFD5FFF5FF557754747A6F34" bold="true" box="[199,327,1599,1622]" pageId="4">Etymology.</emphasis>
Named after the
<typeStatus id="C12D881BFFD5FFF5FD92775477136F34" box="[512,558,1599,1622]" pageId="4">type</typeStatus>
locality.
</paragraph>
<paragraph id="1E2936B9FFD5FFF5FF557709771F6F1B" blockId="4.[151,1435,1426,1691]" box="[199,546,1634,1657]" pageId="4">
<materialsCitation id="AEFE3CE4FFD5FFF5FF557709771F6F1B" ID-GBIF-Occurrence="1699341401" box="[199,546,1634,1657]" country="China" location="Guizhou" pageId="4" specimenCount="1" stateProvince="Guizhou">
<emphasis id="2CE2EAABFFD5FFF5FF557709745D6F1B" bold="true" box="[199,352,1634,1657]" pageId="4">Distrubution.</emphasis>
<collectingCountry id="66817629FFD5FFF5FEF57709749A6F1B" box="[359,423,1634,1657]" name="China" pageId="4">China</collectingCountry>
(
<collectingRegion id="DC52F85BFFD5FFF5FE277709772A6F1B" box="[437,535,1634,1657]" country="China" name="Guizhou" pageId="4">Guizhou</collectingRegion>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="568C6532FFD5FFF5FF5577EF76336FF9" box="[199,782,1668,1691]" pageId="4" type="biology_ecology">
<paragraph id="1E2936B9FFD5FFF5FF5577EF76336FF9" blockId="4.[151,1435,1426,1691]" box="[199,782,1668,1691]" pageId="4">
<emphasis id="2CE2EAABFFD5FFF5FF5577EF74126FF9" bold="true" box="[199,303,1668,1691]" pageId="4">Host ant.</emphasis>
Unknown. Sifted from leave litter (
<figureCitation id="86AD2A3CFFD5FFF5FD3E77EF763C6FF9" box="[684,769,1668,1691]" captionStart="FIGURE 5" captionStartId="6.[151,255,1073,1096]" captionTargetBox="[151,1436,194,1049]" captionTargetId="figure@6.[151,1436,194,1049]" captionTargetPageId="6" captionText="FIGURE 5. A. Lebioderus sinicus with Carebara ants. B. Wutongshan, Shenzhen, Guangdong, China. C. L. maolanus, sifted from leave litter. D. Type locality of L. maolanus (Maolan, Guizhou)." httpUri="https://zenodo.org/record/1042237/files/figure.png" pageId="4">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>