479 lines
61 KiB
XML
479 lines
61 KiB
XML
<document id="A252837688D1A14782C7F550308AF93C" ID-DOI="10.5281/zenodo.14662346" ID-ISSN="1942-1354" ID-Zenodo-Dep="14662346" ID-ZooBank="D5B7D848-4BC0-4719-A8EA-086DD57232B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736984875840" checkinUser="felipe" docAuthor="Smith, Paul E. Skelley Andrew B. T." docDate="2024" docId="943B87C254293E735C8AE04831433CB8" docLanguage="en" docName="InsectaMundi.2024.1091.1-19.pdf" docOrigin="Insecta Mundi 2024 (91)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Rhyparus platypygus Skelley and Smith 2024, new species" docType="treatment" docVersion="2" lastPageNumber="14" masterDocId="6802FFBA54253E7C5C4AE12332363A33" masterDocTitle="Five unusual new species of Rhyparus Westwood, 1845 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="11" updateTime="1736986152089" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="BA99BF72D49300ABE6E1E86159B92464">Five unusual new species of Rhyparus Westwood, 1845 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini)</mods:title>
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<mods:name id="74153162991D0176EB571CE767A96EF8" type="personal">
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<mods:namePart id="CD43EF1BAD87A5BDD89B818C638979AB">Smith, Paul E. Skelley Andrew B. T.</mods:namePart>
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<mods:typeOfResource id="1175E93106A93C60FC14557A4FA63863">text</mods:typeOfResource>
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<mods:title id="72675146B9FB246D5A9195C9D74A872D">Insecta Mundi</mods:title>
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<mods:part id="CAC60145C3AB3AF8DD56DD395E6D3BD1">
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<mods:date id="584E47E97A6226436E847B28FD18F9E9">2024</mods:date>
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<mods:number id="CED24090F2671E7BD8AEC982A98C182E">2024-12-20</mods:number>
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<mods:number id="43597880BA273CFF093B6CCD78970FCB">2024</mods:number>
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<mods:detail id="81B61CFCFCFB769122E3C8463144E9EF" type="issue">
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<mods:number id="622442A65093D516DD1EFC795BBB36BE">91</mods:number>
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<mods:identifier id="ADE38EB14DAEB03DA42D822F96584949" type="DOI">10.5281/zenodo.14662346</mods:identifier>
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<mods:identifier id="D51BC6300C5A5BC43D0EF3343B467483" type="ISSN">1942-1354</mods:identifier>
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<treatment id="943B87C254293E735C8AE04831433CB8" ID-DOI="http://doi.org/10.5281/zenodo.14662372" ID-Zenodo-Dep="14662372" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8" httpUri="http://treatment.plazi.org/id/943B87C254293E735C8AE04831433CB8" lastPageId="15" lastPageNumber="14" pageId="12" pageNumber="11">
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<subSubSection id="5488655F54293E705C8AE048316F3BBA" box="[192,857,363,393]" pageId="12" pageNumber="11" type="nomenclature">
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<paragraph id="1C2D36D454293E705C8AE048316F3BBA" blockId="12.[192,857,363,425]" box="[192,857,363,393]" pageId="12" pageNumber="11">
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<heading id="476581B854293E705C8AE048316F3BBA" box="[192,857,363,393]" fontSize="12" level="2" pageId="12" pageNumber="11" reason="2">
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<emphasis id="2EE6EAC654293E705C8AE048316F3BBA" bold="true" box="[192,857,363,393]" pageId="12" pageNumber="11">
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<taxonomicName id="DB924D5754293E705C8AE04830813BBA" authority="Skelley and Smith" authorityName="Skelley and Smith" authorityYear="2024" box="[192,695,363,393]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="platypygus" status="sp. nov.">
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<emphasis id="2EE6EAC654293E705C8AE04833FF3BBB" bold="true" box="[192,457,363,392]" italics="true" pageId="12" pageNumber="11">Rhyparus platypygus</emphasis>
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Skelley and Smith
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</taxonomicName>
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,
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<taxonomicNameLabel id="35D557BD54293E705E8EE04F316F3BBA" box="[708,857,364,393]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5488655F54293E705C8AE0AC33B33B9A" box="[192,389,399,425]" pageId="12" pageNumber="11" type="description">
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<paragraph id="1C2D36D454293E705C8AE0AC33B33B9A" blockId="12.[192,857,363,425]" box="[192,389,399,425]" pageId="12" pageNumber="11">
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<figureCitation id="84A92A5154293E705C8AE0AC33563B9A" box="[192,352,399,425]" captionStart-0="Figures 18–25" captionStart-1="Figures 26–29" captionStartId-0="13.[192,271,1765,1789]" captionStartId-1="15.[194,273,1189,1213]" captionTargetBox-0="[193,1440,229,1736]" captionTargetBox-1="[192,1440,229,1159]" captionTargetId-0="figure-11@13.[192,1440,229,1736]" captionTargetId-1="figure-260@15.[192,1440,229,1159]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionText-0="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." captionText-1="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14662354" figureDoi-1="http://doi.org/10.5281/zenodo.14662356" httpUri-0="https://zenodo.org/record/14662354/files/figure.png" httpUri-1="https://zenodo.org/record/14662356/files/figure.png" pageId="12" pageNumber="11">Figures 18–28</figureCitation>
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,
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<figureCitation id="84A92A5154293E705D20E0AC33B33B9A" box="[362,389,399,425]" captionStart="Figure 41" captionStartId="18.[191,260,1014,1038]" captionTargetBox="[192,1441,229,984]" captionTargetId="figure-376@18.[192,1440,230,984]" captionTargetPageId="18" captionText="Figure 41. Distribution map of the new species of Rhyparus. Rhyparus cucullatus yellow circles, Rhyparus hexadentatus white stars, Rhyparus loughlini light blue squares, Rhyparus platypygus light green triangles, Rhyparus pteronellus red inverted triangle." figureDoi="http://doi.org/10.5281/zenodo.14662364" httpUri="https://zenodo.org/record/14662364/files/figure.png" pageId="12" pageNumber="11">41</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5488655F54293E705C8AE09A30543848" pageId="12" pageNumber="11" type="diagnosis">
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<paragraph id="1C2D36D454293E705C8AE09A30543848" blockId="12.[191,1441,441,635]" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705C8AE09A330A3BE0" bold="true" box="[192,316,441,467]" pageId="12" pageNumber="11">Diagnosis.</emphasis>
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A member of
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<taxonomicName id="DB924D5754293E705DAEE09A307D3BE0" authorityName="Westwood" authorityYear="1845" box="[484,587,441,467]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="2EE6EAC654293E705DAEE09A307D3BE0" box="[484,587,441,467]" italics="true" pageId="12" pageNumber="11">Rhyparus</emphasis>
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</taxonomicName>
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, distinguished from all other Western Hemisphere species by the following combination of characters: small size; pronotal intercostal surfaces not densely punctate; juxtasutural elytral costa interrupted by caudal bulb; pygidial juncture with abdominal ventrite 5 not perpendicular to body (45° angle in male,
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<figureCitation id="84A92A5154293E705D4EE33D3364380B" box="[260,338,542,568]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="12" pageNumber="11">Fig. 28</figureCitation>
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; much less in female,
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<figureCitation id="84A92A5154293E705E00E33D30AE380B" box="[586,664,542,568]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="12" pageNumber="11">Fig. 27</figureCitation>
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; compare with typically perpendicular juncture in
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<figureCitation id="84A92A5154293E70589FE33D3712380B" box="[1237,1316,542,568]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="12" pageNumber="11">Fig. 29</figureCitation>
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); male and female pygidia convex or flattened (not concave), with medial carina broad; and distribution in
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<collectingCountry id="6485764454293E7058A6E31C3753386A" box="[1260,1381,575,601]" name="Guatemala" pageId="12" pageNumber="11">Guatemala</collectingCountry>
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(
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<collectingRegion id="DE56F83654293E70593EE31C32D23848" country="Guatemala" name="Izabal" pageId="12" pageNumber="11">Izabal</collectingRegion>
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) and
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<collectingCountry id="6485764454293E705D68E34233A43848" box="[290,402,609,635]" name="Honduras" pageId="12" pageNumber="11">Honduras</collectingCountry>
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(
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<collectingRegion id="DE56F83654293E705DEBE34233D93848" box="[417,495,609,635]" country="Honduras" name="Cortes" pageId="12" pageNumber="11">Cortés</collectingRegion>
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) (
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<figureCitation id="84A92A5154293E705E4FE34230633848" box="[517,597,609,635]" captionStart="Figure 41" captionStartId="18.[191,260,1014,1038]" captionTargetBox="[192,1441,229,984]" captionTargetId="figure-376@18.[192,1440,230,984]" captionTargetPageId="18" captionText="Figure 41. Distribution map of the new species of Rhyparus. Rhyparus cucullatus yellow circles, Rhyparus hexadentatus white stars, Rhyparus loughlini light blue squares, Rhyparus platypygus light green triangles, Rhyparus pteronellus red inverted triangle." figureDoi="http://doi.org/10.5281/zenodo.14662364" httpUri="https://zenodo.org/record/14662364/files/figure.png" pageId="12" pageNumber="11">Fig. 41</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="5488655F54293E725C8AE3A9317F3C15" lastPageId="14" lastPageNumber="13" pageId="12" pageNumber="11" type="description">
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<paragraph id="1C2D36D454293E705C8AE3A93300393A" blockId="12.[191,1442,650,1920]" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705C8AE3A930163897" bold="true" box="[192,544,650,677]" pageId="12" pageNumber="11">
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Description.
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<typeStatus id="C329887654293E705D11E3A833FE3896" box="[347,456,651,677]" pageId="12" pageNumber="11" type="holotype">Holotype</typeStatus>
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female
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</emphasis>
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(
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<figureCitation id="84A92A5154293E705E7BE3A9309B3896" box="[561,685,650,677]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="12" pageNumber="11">Fig. 18–20</figureCitation>
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). Body length
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<quantity id="DB6A9B3154293E705F12E3A931F73896" box="[856,961,650,677]" metricMagnitude="-3" metricUnit="m" metricValue="3.63" pageId="12" pageNumber="11" unit="mm" value="3.63">3.63 mm</quantity>
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, width
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<quantity id="DB6A9B3154293E705850E3A836B53896" box="[1050,1155,651,677]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="12" pageNumber="11" unit="mm" value="1.19">1.19 mm</quantity>
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. Elongate, parallel-sided, flattened, dark brownish black; dorsal costae weakly alutaceous, somewhat glossy; costal intervals strongly alutaceous, dull; partly clothed with indistinct, fine, very short setae on head and all longitudinal costae on pronotum and elytra.
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</paragraph>
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<paragraph id="1C2D36D454293E705D4AE23231193E24" blockId="12.[191,1442,650,1920]" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705D4AE23233733918" bold="true" box="[256,325,785,811]" pageId="12" pageNumber="11">Head.</emphasis>
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Surface dull, alutaceous, transversely subhexagonal (
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<figureCitation id="84A92A5154293E705FD1E23231DB3918" box="[923,1005,785,811]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="12" pageNumber="11">Fig. 21</figureCitation>
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). Clypeus trapezoidal, anteriorly truncate in anterior view, concave between teeth in dorsal view; anterior margin narrowly upturned with sharp obtuse tooth on each side of medial third, lateral thirds concave to rounded obtuse tooth anterior of clypeo-genal junction. Gena moderately lobed laterally, distinctly excavate anterior of eyes. Clypeal disc convex with two indistinct longitudinal costae weakly evident in anterior view and dorsal profile; peridiscal impression strong basally, anteriorly a coarsely punctured, weak groove. Frons with four small, distinct frontodiscal costae, lateral costae longer than medial costae. Head with evenly scattered punctures; coarser punctures at base of head, smaller punctures on clypeal disc; punctures with a short, pale yellow seta.
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</paragraph>
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<paragraph id="1C2D36D454293E705D4AE53D301A3FBB" blockId="12.[191,1442,650,1920]" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705D4AE53D33B63E0B" bold="true" box="[256,384,1054,1080]" pageId="12" pageNumber="11">Pronotum.</emphasis>
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Surface alutaceous, dull; costae weakly alutaceous, glossy, widest in middle; with paramedian, discolateral, and submarginal costae distinct, all with anterior portions convex. Paramedian costae incomplete, prominent at anterior margin, obsolete at constriction near anterior third, distinct straight but diverging towards posterior margin. Discolateral costa nearly straight, distinctly interrupted by fovea on anterior third, present at anterior margin. Submarginal costa complete, sinuate; posterior lobe reduced, flattened. Median interval at constriction of paramedian costae with few small punctures. Other costal intervals impunctate except for four large pores at anterior margin and at basal third on each side of paramedian costae. Interval between discolateral and submarginal costae with posterior half convex medially, subcostate, ends at the posterior angle of the pronotum. Lateral margin deeply notched, with prominent anterior and intermediate lobes; anterior lobe slightly smaller than intermediate lobe, both angulate laterally; posterior lobe lacking. Posterior marginal groove broad, deep, and crenated by pronotal costae.
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</paragraph>
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<paragraph id="1C2D36D454293E705D4AE4AC31513F99" blockId="12.[191,1442,650,1920]" box="[256,871,1423,1450]" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705D4AE4AC33403F9A" bold="true" box="[256,374,1423,1449]" pageId="12" pageNumber="11">Scutellum</emphasis>
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. Scutellar shield minute, narrow, apex acute.
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</paragraph>
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<paragraph id="1C2D36D454293E725D4AE49233D43BB4" blockId="12.[191,1442,650,1920]" lastBlockId="14.[191,1442,230,1399]" lastPageId="14" lastPageNumber="13" pageId="12" pageNumber="11">
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<emphasis id="2EE6EAC654293E705D4AE49233793FF8" bold="true" box="[256,335,1457,1483]" pageId="12" pageNumber="11">Elytra.</emphasis>
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Surface of intervals alutaceous, dull; costae weakly alutaceous, glossy, widest in middle; flattened intervals with coarse, distinctly separated punctures (
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<figureCitation id="84A92A5154293E705F44E4F131B33FDE" box="[782,901,1490,1517]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="12" pageNumber="11">Fig. 18, 22</figureCitation>
|
||
). Juxtasutural costa interrupted caudally by bulb, not attaining apex; weakly convex. Discomedian costa straight from base to apex; weakly more prominent anteriorly, medially convex as discolateral costa; gradually swelling at apex to small postdiscal bulb; trichome of costa small, hidden in groove. Discolateral costa prominent, straight medially, weakly curving at caudal ends. Posthumeral costa distinct, complete to apex, narrower than discolateral costa. Submarginal costa narrow entire length; turning upward weakly and abruptly inward to caudal bulb, forming a small, tubercular angulation. Costal interval 1 (between juxtasutural and discomedian costae) flattened, with 18 coarse punctures in outer row. Costal interval 2 (between discomedian and discolateral costae) with two rows of coarse, rounded, distinctly separated punctures. Costal interval 3 (between discolateral and posthumeral costae) with three rows of coarse, rounded, distinctly separated punctures; punctures in median row smaller than outer rows. Costal interval 4 (between posthumeral and submarginal costae) with punctures somewhat transverse, not connected to adjacent row medially; rugose-washboard surface medially, decreasing basally. Caudal trichomes at end of costae small, hidden; trichome of caudal bulb narrowed, transversely elongate. Caudal bulbs moderately large, transverse; in dorsal view, weakly prominent laterally, curving inward and caudally to interrupt juxtasutural costa; anterior margin projecting, broadly rounded, dividing trichome opposite of discomedian costa; in caudal view, somewhat broadly rounded dorsally and laterally. Dorsal surface of caudal bulb weakly alutaceous, weakly glossy, apparently impunctate, with scattered minute setae; surface between bulb and caudal margin of elytra dull, concave with several foveate punctures.
|
||
</paragraph>
|
||
<caption id="48ED665C54283E715C8AE7C637093D2F" ID-DOI="http://doi.org/10.5281/zenodo.14662354" ID-Zenodo-Dep="14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="13" pageNumber="12" startId="13.[192,271,1765,1789]" targetBox="[193,1440,229,1736]" targetPageId="13" targetType="figure">
|
||
<paragraph id="1C2D36D454283E715C8AE7C637093D2F" blockId="13.[192,1440,1765,1820]" pageId="13" pageNumber="12">
|
||
<emphasis id="2EE6EAC654283E715C8AE7C6336D3CCE" bold="true" box="[192,347,1765,1789]" pageId="13" pageNumber="12">Figures 18–25.</emphasis>
|
||
<taxonomicName id="DB924D5754283E715D2BE7C630063CCE" authorityName="Skelley and Smith" authorityYear="2024" box="[353,560,1765,1789]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="platypygus" status="sp. nov.">
|
||
<emphasis id="2EE6EAC654283E715D2BE7C630063CCE" box="[353,560,1765,1789]" italics="true" pageId="13" pageNumber="12">Rhyparus platypygus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2EE6EAC654283E715E7CE7C530873CCE" bold="true" box="[566,689,1765,1790]" pageId="13" pageNumber="12">
|
||
<taxonomicNameLabel id="35D557BD54283E715E7CE7C530873CCE" box="[566,689,1765,1790]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male.
|
||
<emphasis id="2EE6EAC654283E715F12E7C631973CCE" bold="true" box="[856,929,1765,1789]" pageId="13" pageNumber="12">18–20)</emphasis>
|
||
Dorsal, ventral, lateral habitus.
|
||
<emphasis id="2EE6EAC654283E7158A9E7C637303CCE" bold="true" box="[1251,1286,1765,1789]" pageId="13" pageNumber="12">21)</emphasis>
|
||
Head and pronotum, oblique anterior view.
|
||
<emphasis id="2EE6EAC654283E715DBAE62730243D2F" bold="true" box="[496,530,1796,1820]" pageId="13" pageNumber="12">22)</emphasis>
|
||
Elytra, posterior view.
|
||
<emphasis id="2EE6EAC654283E715EB1E627312B3D2F" bold="true" box="[763,797,1796,1820]" pageId="13" pageNumber="12">23)</emphasis>
|
||
Pygidium.
|
||
<emphasis id="2EE6EAC654283E715FD8E62731EC3D2F" bold="true" box="[914,986,1796,1820]" pageId="13" pageNumber="12">24–25)</emphasis>
|
||
Genitalia, caudal and lateral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C2D36D4542B3E725D4AE0AD316E383F" blockId="14.[191,1442,230,1399]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725D4AE0AD33823B9B" bold="true" box="[256,436,398,424]" pageId="14" pageNumber="13">Ventral thorax.</emphasis>
|
||
Surface alutaceous, dull, punctures bearing a short seta. Metaventrite flattened; median longitudinal groove on posterior two-thirds of ventrite, width slightly less than mesotarsi, equally widened at ends; disc on either side of groove with minutely punctate area bounded laterally by short series of coarse punctures; punctures on remainder of disc fine, evenly scattered.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E725D4AE3373139390F" blockId="14.[191,1442,230,1399]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725D4AE3373342381D" bold="true" box="[256,372,532,558]" pageId="14" pageNumber="13">Abdomen</emphasis>
|
||
. Abdominal ventrites 2–4 with narrow, transverse, anterior groove fluted; posterior margin with row of small, shallow punctures; ventrites laterally with indistinct, shallow, triangular depression with coarse punctures, depression more distinct on posterior ventrites; rest of surface with scattered, fine punctures bearing a short seta. Abdominal ventrite 5 (last) as long as preceding two ventrites medially; surface evenly coarsely punctate, punctures separated by one puncture diameter; medially with cluster of punctures forming small fovea each side of midline; laterally with transversely elongate, triangular fovea; anterior margin with narrow, punctate groove extending to lateral margin. Pygidium greatly depressed against abdominal ventrite 5 (
|
||
<figureCitation id="84A92A51542B3E72588CE3FD370938CB" box="[1222,1343,734,760]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="14" pageNumber="13">Fig. 26–27</figureCitation>
|
||
), surface parallel with surface of ventrites; median carina broadly flattened, widening to a weakly bilobed projection that encroaches upon the two medial fossae on ventrite 5.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E725D4AE26036703E15" blockId="14.[191,1442,230,1399]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725D4AE260330A396E" bold="true" box="[256,316,835,861]" pageId="14" pageNumber="13">Legs.</emphasis>
|
||
All femora weakly glossy, with moderately deep, distinct punctures; all punctures bearing a very short seta. Profemoral punctures enlarged and prominent, mesofemoral and metafemoral punctures moderate in size and less distinct. Mesofemur robust medially; posterior margin weakly sinuate, with weakly convex swelling at middle and apical third. Metafemur narrow, moderately widened medially. Protibia with bidentate apex; inner tooth minute; medial margin weakly concave in apical third. Mesotibia and metatibia with apex truncate; lacking inner apical spine and concave inner margin. Mesotarsomere 1 as long as mesotarsomeres 2–4; ventral setae fine, glossy surface visible. Metatarsomere 1 as long as metatarsomeres 2–5 combined.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E725D4AE50E33703EFD" blockId="14.[191,1442,230,1399]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725D4AE50E339C3E7B" bold="true" box="[256,426,1069,1096]" pageId="14" pageNumber="13">
|
||
<typeStatus id="C3298876542B3E725D4AE50E33533E74" box="[256,357,1069,1095]" pageId="14" pageNumber="13" type="allotype">Allotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
Body length
|
||
<quantity id="DB6A9B31542B3E725E76E50D30963E7B" box="[572,672,1070,1096]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="13" unit="mm" value="3.8">3.80 mm</quantity>
|
||
, width
|
||
<quantity id="DB6A9B31542B3E725EBAE50D31633E7B" box="[752,853,1070,1096]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="13" unit="mm" value="1.2">1.20 mm</quantity>
|
||
. Similar to female in most characters. Pronotum with paramedian costae weakly curving inward while diverging to posterior margin. Mesotibia with apex truncate; large, medially projecting spine on inner apical angle; inner margin concave in apical quarter. Metatibia weakly widened to truncate apex; large, medially projecting spine on inner apical angle; inner margin weaky concave in apical third.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E725D4AE5F531AF3F45" blockId="14.[191,1442,230,1399]" pageId="14" pageNumber="13">
|
||
Apical half of pygidium with central carina basally split, Y-shaped; top of Y prominent, not tuberculate; median carina broad, flattened (
|
||
<figureCitation id="84A92A51542B3E725E63E5D430953F22" box="[553,675,1271,1297]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="14" pageNumber="13">Fig. 19, 23</figureCitation>
|
||
), surface convex, lacking concavity in lateral view (compare
|
||
<figureCitation id="84A92A51542B3E725907E5D437AA3F22" box="[1357,1436,1271,1297]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="14" pageNumber="13">Fig. 22</figureCitation>
|
||
,
|
||
<figureCitation id="84A92A51542B3E725C8BE43A32EA3F00" box="[193,220,1305,1331]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="14" pageNumber="13">28</figureCitation>
|
||
with
|
||
<figureCitation id="84A92A51542B3E725D56E43A335C3F00" box="[284,362,1305,1331]" captionStart="Figures 26–29" captionStartId="15.[194,273,1189,1213]" captionTargetBox="[192,1440,229,1159]" captionTargetId="figure-260@15.[192,1440,229,1159]" captionTargetPageId="15" captionText="Figures 26–29. Abdominal ventrites and pygidia. 26–27) Rhyparus platypygus new species female holotype, ventral and lateral views. 28) R. platypygus paratype male, lateral view. 29) R. loughlini new species holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="14" pageNumber="13">Fig. 29</figureCitation>
|
||
); each side of median carinae with alutaceous, triangular depression. Genitalia (
|
||
<figureCitation id="84A92A51542B3E7258ADE43A37693F07" box="[1255,1375,1305,1332]" captionStart="Figures 18–25" captionStartId="13.[192,271,1765,1789]" captionTargetBox="[193,1440,229,1736]" captionTargetId="figure-11@13.[192,1440,229,1736]" captionTargetPageId="13" captionText="Figures 18–25. Rhyparus platypygus new species, paratype male. 18–20) Dorsal, ventral, lateral habitus. 21) Head and pronotum, oblique anterior view. 22) Elytra, posterior view. 23) Pygidium. 24–25) Genitalia, caudal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.14662354" httpUri="https://zenodo.org/record/14662354/files/figure.png" pageId="14" pageNumber="13">Fig. 24–25</figureCitation>
|
||
) with phallobase arched, tubular; parameres short, less than one-quarter length of phallobase; parameres straight in lateral view, gradually narrowing to rounded apex in caudal view.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E725C8AE4A6317F3C15" blockId="14.[192,1441,1413,1575]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725C8AE4A633003FAC" bold="true" box="[192,310,1413,1439]" pageId="14" pageNumber="13">Variation.</emphasis>
|
||
Length
|
||
<quantity id="DB6A9B31542B3E725DD2E4A5300B3F93" box="[408,573,1414,1441]" metricMagnitude="-3" metricUnit="m" metricValue="3.78" metricValueMax="4.06" metricValueMin="3.5" pageId="14" pageNumber="13" unit="mm" value="3.78" valueMax="4.06" valueMin="3.5">3.50–4.06 mm</quantity>
|
||
, width
|
||
<quantity id="DB6A9B31542B3E725EDFE4A5310A3F93" box="[661,828,1414,1440]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" metricValueMax="1.31" metricValueMin="1.13" pageId="14" pageNumber="13" unit="mm" value="1.22" valueMax="1.31" valueMin="1.13">1.13–1.31 mm</quantity>
|
||
. Elytra with discolateral costal anterior and posterior ends weakly curved inward, appearing straight in some, curving more distinctly in others, slightly sinuate medially in a few specimens. Paramedian pronotal costae are posteriorly straight or weakly curving, and the degree of divergence posteriorly is somewhat sexually dimorphic. However, some specimens vary enough that gender cannot be confidently determined based on this character.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5488655F542B3E735C8AE71633983F7F" lastPageId="15" lastPageNumber="14" pageId="14" pageNumber="13" type="materials_examined">
|
||
<paragraph id="1C2D36D4542B3E725C8AE71636633CA0" blockId="14.[192,1442,1589,1919]" pageId="14" pageNumber="13">
|
||
<emphasis id="2EE6EAC6542B3E725C8AE71633953C7C" bold="true" box="[192,419,1589,1615]" pageId="14" pageNumber="13">Material examined.</emphasis>
|
||
<materialsCitation id="ACFA3C89542B3E725DE3E71533BE3CA0" collectingDate="2002-06-07" collectorName="R. E. Woodruff" country="Guatemala" location="Morales" pageId="14" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Izabal" typeStatus="holotype">
|
||
<typeStatus id="C3298876542B3E725DE3E71530273C63" box="[425,529,1590,1616]" pageId="14" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
female and
|
||
<typeStatus id="C3298876542B3E725ED2E71530C73C63" box="[664,753,1590,1616]" pageId="14" pageNumber="13" type="allotype">allotype</typeStatus>
|
||
male deposited in FSCA: “
|
||
<collectingCountry id="64857644542B3E725851E71536F13C63" box="[1051,1223,1590,1616]" name="Guatemala" pageId="14" pageNumber="13">GUATEMALA</collectingCountry>
|
||
: Dept.
|
||
<collectingRegion id="DE56F836542B3E72595FE715376C3C63" box="[1301,1370,1590,1616]" country="Guatemala" name="Izabal" pageId="14" pageNumber="13">Izabal</collectingRegion>
|
||
, / E of
|
||
<location id="194D600F542B3E725C8AE774332B3C42" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542B3E725C8AE774332B3C42" box="[192,285,1623,1649]" country="Guatemala" name="Morales" pageId="14" pageNumber="13" stateProvince="Izabal">Morales</location>
|
||
,
|
||
<date id="682C1014542B3E725D6DE77433AB3C42" box="[295,413,1623,1649]" pageId="14" pageNumber="13" value="2002-06-07">
|
||
<collectingDate id="7868E9FC542B3E725D6DE77433AB3C42" box="[295,413,1623,1649]" pageId="14" pageNumber="13" value="2002-06-07">7-VI-2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B1675302542B3E725DEDE77B307A3C42" box="[423,588,1623,1650]" pageId="14" pageNumber="13">R.E. Woodruff</collectorName>
|
||
,
|
||
<collectingMethod id="C5D34EC3542B3E725E1CE77430FB3C41" box="[598,717,1623,1650]" pageId="14" pageNumber="13">dung traps</collectingMethod>
|
||
// [red paper]
|
||
<typeStatus id="C3298876542B3E725F25E77436363C42" box="[879,1024,1623,1649]" pageId="14" pageNumber="13" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="0A94FD5D542B3E72584CE77A362D3C46" box="[1030,1051,1625,1653]" count="1" pageId="14" pageNumber="13" type="female">♀</specimenCount>
|
||
/
|
||
<taxonomicName id="DB924D57542B3E72587AE77436AF3C42" authorityName="Westwood" authorityYear="1845" box="[1072,1177,1623,1649]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2EE6EAC6542B3E72587AE77436AF3C42" box="[1072,1177,1623,1649]" italics="true" pageId="14" pageNumber="13">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="DB924D57542B3E7258E4E77437153C42" authorityName="Skelley and Smith" authorityYear="2024" box="[1198,1315,1623,1649]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="platypygus">
|
||
<emphasis id="2EE6EAC6542B3E7258E4E77437153C42" box="[1198,1315,1623,1649]" italics="true" pageId="14" pageNumber="13">platypygus</emphasis>
|
||
</taxonomicName>
|
||
/ des. 2024 / Skelley & Smith”
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="C3298876542B3E725DD8E75A33C43CA0" box="[402,498,1657,1683]" pageId="14" pageNumber="13" type="allotype">Allotype</typeStatus>
|
||
with similar label except blue, stating “
|
||
<typeStatus id="C3298876542B3E725FEEE75A361A3CA0" box="[932,1068,1657,1683]" pageId="14" pageNumber="13" type="allotype">ALLOTYPE</typeStatus>
|
||
<specimenCount id="0A94FD5D542B3E725878E75936713CA5" box="[1074,1095,1658,1686]" count="1" pageId="14" pageNumber="13" type="male">♂</specimenCount>
|
||
”.
|
||
</paragraph>
|
||
<paragraph id="1C2D36D4542B3E735D4AE7B833983F7F" blockId="14.[192,1442,1589,1919]" lastBlockId="15.[192,1440,1296,1356]" lastPageId="15" lastPageNumber="14" pageId="14" pageNumber="13">
|
||
<materialsCitation id="ACFA3C89542B3E725D4AE7B831CD3C86" box="[256,1019,1690,1717]" collectingDate="2002-06-07" collectionCode="FSCA, CMNC" collectorName="R. E. Woodruff" country="Guatemala" location="Morales" pageId="14" pageNumber="13" specimenCount="2" stateProvince="Izabal" typeStatus="paratype">
|
||
<emphasis id="2EE6EAC6542B3E725D4AE7B833443C86" bold="true" box="[256,370,1691,1717]" pageId="14" pageNumber="13">
|
||
<typeStatus id="C3298876542B3E725D4AE7B833443C86" box="[256,370,1691,1717]" pageId="14" pageNumber="13" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(13): same data as holotype (
|
||
<typeStatus id="C3298876542B3E725EFBE7B9313A3C87" box="[689,780,1690,1716]" pageId="14" pageNumber="13" type="allotype">allotype</typeStatus>
|
||
+ 2
|
||
<collectionCode id="7A83AE11542B3E725F71E7B831B63C86" box="[827,896,1691,1717]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="13" type="Museum">FSCA</collectionCode>
|
||
, 1
|
||
<collectionCode id="7A83AE11542B3E725FD6E7B831C53C86" box="[924,1011,1691,1717]" pageId="14" pageNumber="13">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<emphasis id="2EE6EAC6542B3E72584EE7B937A93C86" bold="true" box="[1028,1439,1690,1717]" pageId="14" pageNumber="13">
|
||
<materialsCitation id="ACFA3C89542B3E72584EE7B937AC3C86" box="[1028,1434,1690,1717]" collectorName="Department" country="Guatemala" location="Izabal" pageId="14" pageNumber="13" specimenCount="1" stateProvince="Izabal" typeStatus="paratype">
|
||
<collectingCountry id="64857644542B3E72584EE7B936843C87" box="[1028,1202,1690,1716]" name="Guatemala" pageId="14" pageNumber="13">GUATEMALA</collectingCountry>
|
||
:
|
||
<collectorName id="B1675302542B3E7258F1E7B8377D3C86" box="[1211,1355,1691,1717]" pageId="14" pageNumber="13">Department</collectorName>
|
||
<collectingRegion id="DE56F836542B3E72591AE7B837AC3C86" box="[1360,1434,1691,1717]" country="Guatemala" name="Izabal" pageId="14" pageNumber="13">Izabal</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
</emphasis>
|
||
<materialsCitation id="ACFA3C89542B3E725C8AE79E30F83CCB" collectingDate="2003-05-27" collectingDateMax="2003-05-28" collectingDateMin="2003-05-27" collectionCode="FSCA" collectorName="R Woodruff, J & Monzon" country="Guatemala" county="Morales" elevation="1250" location="Finca Firmeza" municipality="Sierra del Merendon" pageId="14" pageNumber="13" specimenCount="1" typeStatus="paratype">
|
||
E. of
|
||
<collectingCounty id="F54C4E58542B3E725CB6E79F336F3CE5" box="[252,345,1724,1750]" pageId="14" pageNumber="13">Morales</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="FC49ACAE542B3E725D2FE79F30663CE5" box="[357,592,1724,1750]" pageId="14" pageNumber="13">Sierra del Merendon</collectingMunicipality>
|
||
,
|
||
<location id="194D600F542B3E725E16E79F31363CE5" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542B3E725E16E79F31363CE5" box="[604,768,1724,1750]" country="Guatemala" county="Morales" municipality="Sierra del Merendon" name="Finca Firmeza" pageId="14" pageNumber="13">Finca Firmeza</location>
|
||
,
|
||
<locationDeviation id="0D9F2FF6542B3E725F46E79F31CF3CE5" box="[780,1017,1724,1751]" location="Honduras" pageId="14" pageNumber="13">
|
||
<quantity id="DB6A9B31542B3E725F46E79F31083CE5" box="[780,830,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="13" unit="km" value="1.0">1km</quantity>
|
||
W of Honduras
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="DB6A9B31542B3E72584CE79F366D3CE4" box="[1030,1115,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="13" unit="m" value="1250.0">
|
||
<elevation id="97BFD1E7542B3E72584CE79F366D3CE4" box="[1030,1115,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="13" unit="m" value="1250.0">1250 m</elevation>
|
||
</quantity>
|
||
elev.,
|
||
<date id="682C1014542B3E7258EBE79F37093CE5" box="[1185,1343,1724,1750]" pageId="14" pageNumber="13" value="2003-05-27" valueMax="2003-05-28" valueMin="2003-05-27">
|
||
<collectingDate id="7868E9FC542B3E7258EBE79F37093CE5" box="[1185,1343,1724,1750]" pageId="14" pageNumber="13" value="2003-05-27" valueMax="2003-05-28" valueMin="2003-05-27">27-28-V-2003</collectingDate>
|
||
</date>
|
||
, at light, cloud forest
|
||
<collectorName id="B1675302542B3E725D05E7FD33D93CCB" box="[335,495,1758,1784]" pageId="14" pageNumber="13">R Woodruff, J</collectorName>
|
||
.
|
||
<collectorName id="B1675302542B3E725DB0E7FD306E3CCB" box="[506,600,1758,1784]" pageId="14" pageNumber="13">Monzon</collectorName>
|
||
(3
|
||
<collectionCode id="7A83AE11542B3E725ECAE7FD30F13CCB" box="[640,711,1758,1784]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="13" type="Museum">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ACFA3C89542B3E725E90E7FD33D43D08" collectingDate="2001-06-21" collectingDateMax="2001-06-22" collectingDateMin="2001-06-21" collectionCode="WBWC" collectorName="W. B. Warner & J. Monzon-Sierra & R. Cunningham" country="Guatemala" elevation="1183" latitude="15.364167" location="Finca Firmesa" longLatPrecision="21" longitude="-88.69111" municipality="Morales" pageId="14" pageNumber="13" specimenCount="1" typeStatus="paratype">
|
||
SE of
|
||
<collectingMunicipality id="FC49ACAE542B3E725F54E7FD314D3CCB" box="[798,891,1758,1784]" pageId="14" pageNumber="13">Morales</collectingMunicipality>
|
||
,
|
||
<location id="194D600F542B3E725FCCE7FD36103CCB" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542B3E725FCCE7FD36103CCB" box="[902,1062,1758,1784]" country="Guatemala" latitude="15.364167" longLatPrecision="21" longitude="-88.69111" municipality="Morales" name="Finca Firmesa" pageId="14" pageNumber="13">Finca Firmesa</location>
|
||
2,
|
||
<geoCoordinate id="79A65013542B3E725800E7FD36FB3CCA" box="[1098,1229,1758,1785]" degrees="15" direction="north" minutes="21" orientation="latitude" pageId="14" pageNumber="13" precision="15" seconds="51" value="15.364167">15°21’51’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="79A65013542B3E725890E7FD37693CCB" box="[1242,1375,1758,1784]" degrees="88" direction="west" minutes="41" orientation="longitude" pageId="14" pageNumber="13" precision="15" seconds="28" value="-88.69111">88°41’28’’W</geoCoordinate>
|
||
,
|
||
<quantity id="DB6A9B31542B3E725926E7FD32ED3D2A" metricMagnitude="3" metricUnit="m" metricValue="1.182624" pageId="14" pageNumber="13" unit="ft" value="3880.0">
|
||
<elevation id="97BFD1E7542B3E725926E7FD32ED3D2A" metricMagnitude="3" metricUnit="m" metricValue="1.182624" pageId="14" pageNumber="13" unit="ft" value="3880.0">3880 ft.</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="682C1014542B3E725CADE7DC33A23D2A" box="[231,404,1791,1817]" pageId="14" pageNumber="13" value="2001-06-21" valueMax="2001-06-22" valueMin="2001-06-21">
|
||
<collectingDate id="7868E9FC542B3E725CADE7DC33A23D2A" box="[231,404,1791,1817]" pageId="14" pageNumber="13" value="2001-06-21" valueMax="2001-06-22" valueMin="2001-06-21">21–22-VI-2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B1675302542B3E725DEBE62330043D29" box="[417,562,1792,1818]" pageId="14" pageNumber="13">W.B. Warner</collectorName>
|
||
,
|
||
<collectorName id="B1675302542B3E725E75E62331353D2A" box="[575,771,1791,1818]" pageId="14" pageNumber="13">J. Monzón-Sierra</collectorName>
|
||
, &
|
||
<collectorName id="B1675302542B3E725F61E62331D53D2A" box="[811,995,1791,1818]" pageId="14" pageNumber="13">R. Cunningham</collectorName>
|
||
, ultraviolet, mercury vapor, & mercury halogen light (2
|
||
<collectionCode id="7A83AE11542B3E725D34E60233EC3D08" box="[382,474,1825,1851]" pageId="14" pageNumber="13">WBWC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ACFA3C89542B3E725DA5E602310F3D6E" collectingDate="2002-06-06" collectingDateMax="2002-06-07" collectingDateMin="2002-06-06" collectionCode="FSCA" collectorName="J. & C. Monzon & Davis & R. E. Woodruff" country="Guatemala" elevation="1150" location="Cerro Negro Norte" pageId="14" pageNumber="13" specimenCount="1" typeStatus="paratype">
|
||
near
|
||
<location id="194D600F542B3E725E63E60231323D08" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542B3E725E63E60231323D08" box="[553,772,1825,1851]" country="Guatemala" name="Cerro Negro Norte" pageId="14" pageNumber="13">Cerro Negro Norte</location>
|
||
,
|
||
<locationDeviation id="0D9F2FF6542B3E725F5BE60231DC3D08" box="[785,1002,1825,1851]" location="Honduras" pageId="14" pageNumber="13">
|
||
<quantity id="DB6A9B31542B3E725F5BE602317A3D08" box="[785,844,1825,1851]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="13" unit="km" value="1.0">1 km</quantity>
|
||
W Honduras
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="DB6A9B31542B3E725FBDE60236783D08" box="[1015,1102,1825,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="13" unit="m" value="1150.0">
|
||
<elevation id="97BFD1E7542B3E725FBDE60236783D08" box="[1015,1102,1825,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="13" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="682C1014542B3E725811E60236D83D08" box="[1115,1262,1825,1851]" pageId="14" pageNumber="13" value="2002-06-06" valueMax="2002-06-07" valueMin="2002-06-06">
|
||
<collectingDate id="7868E9FC542B3E725811E60236D83D08" box="[1115,1262,1825,1851]" pageId="14" pageNumber="13" value="2002-06-06" valueMax="2002-06-07" valueMin="2002-06-06">6–7-VI-2002</collectingDate>
|
||
</date>
|
||
, mercury vapor light,
|
||
<collectorName id="B1675302542B3E725D48E66033843D6E" box="[258,434,1859,1885]" pageId="14" pageNumber="13">J. & C. Monzon</collectorName>
|
||
, S.
|
||
<collectorName id="B1675302542B3E725D9DE661302F3D6F" box="[471,537,1858,1884]" pageId="14" pageNumber="13">Davis</collectorName>
|
||
,
|
||
<collectorName id="B1675302542B3E725E69E66030F03D6F" box="[547,710,1858,1885]" pageId="14" pageNumber="13">R.E. Woodruff</collectorName>
|
||
(2
|
||
<collectionCode id="7A83AE11542B3E725EA0E66031073D6E" box="[746,817,1859,1885]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="13" type="Museum">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="2EE6EAC6542B3E725F0BE66136E03D6E" bold="true" box="[833,1238,1858,1885]" pageId="14" pageNumber="13">
|
||
<materialsCitation id="ACFA3C89542B3E725F0BE66136E73D6E" box="[833,1233,1858,1885]" collectorName="Department" country="Honduras" location="Cortes" pageId="14" pageNumber="13" specimenCount="1" stateProvince="Cortes" typeStatus="paratype">
|
||
<collectingCountry id="64857644542B3E725F0BE66131D73D6F" box="[833,993,1858,1884]" name="Honduras" pageId="14" pageNumber="13">HONDURAS</collectingCountry>
|
||
:
|
||
<collectorName id="B1675302542B3E725FA6E660364A3D6E" box="[1004,1148,1859,1885]" pageId="14" pageNumber="13">Department</collectorName>
|
||
<collectingRegion id="DE56F836542B3E7258C8E66136E73D6E" box="[1154,1233,1858,1885]" country="Honduras" name="Cortes" pageId="14" pageNumber="13">Cortés</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
</emphasis>
|
||
<materialsCitation id="ACFA3C89542B3E735897E66033E13F18" collectingDate="1994-08-26" collectionCode="CMNC" collectorName="P. N. Cusuco & R. Anderson & Berlese" country="Honduras" elevation="1960" lastPageId="15" lastPageNumber="14" location="Cerro Jilinco" municipality="Buenos Aires" pageId="14" pageNumber="13" specimenCount="1" typeStatus="paratype">
|
||
<collectorName id="B1675302542B3E725897E66037513D6E" box="[1245,1383,1859,1885]" pageId="14" pageNumber="13">P.N. Cusuco</collectorName>
|
||
,
|
||
<locationDeviation id="0D9F2FF6542B3E72593BE66033463D4D" location="Cofradia" pageId="14" pageNumber="13">
|
||
<quantity id="DB6A9B31542B3E72593BE66032D33D4D" metricMagnitude="4" metricUnit="m" metricValue="1.8699999999999999" pageId="14" pageNumber="13" unit="km" value="18.7">18.7 km</quantity>
|
||
N Cofradía
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="DB6A9B31542B3E725D36E64733FD3D4D" box="[380,459,1892,1919]" metricMagnitude="3" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="13" unit="km" value="5.4">5.4 km</quantity>
|
||
W
|
||
<collectingMunicipality id="FC49ACAE542B3E725DBCE64630B83D4D" box="[502,654,1892,1919]" pageId="14" pageNumber="13">Buenos Aires</collectingMunicipality>
|
||
,
|
||
<location id="194D600F542B3E725ED1E64731183D4D" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542B3E725ED1E64731183D4D" box="[667,814,1892,1918]" country="Honduras" municipality="Buenos Aires" name="Cerro Jilinco" pageId="14" pageNumber="13">Cerro Jilinco</location>
|
||
,
|
||
<quantity id="DB6A9B31542B3E725F71E64731A43D4C" box="[827,914,1892,1919]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="14" pageNumber="13" unit="m" value="1960.0">
|
||
<elevation id="97BFD1E7542B3E725F71E64731A43D4C" box="[827,914,1892,1919]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="14" pageNumber="13" unit="m" value="1960.0">1960 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="682C1014542B3E725FD5E64736003D4D" box="[927,1078,1892,1918]" pageId="14" pageNumber="13" value="1994-08-26">
|
||
<collectingDate id="7868E9FC542B3E725FD5E64736003D4D" box="[927,1078,1892,1918]" pageId="14" pageNumber="13" value="1994-08-26">26-VIII-1994</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B1675302542B3E725808E64636E23D4D" box="[1090,1236,1892,1919]" pageId="14" pageNumber="13">R. Anderson</collectorName>
|
||
, pine/cloud forest litter,
|
||
<collectorName id="B1675302542A3E735D4BE43333643F19" box="[257,338,1296,1322]" pageId="15" pageNumber="14">Berlese</collectorName>
|
||
(1
|
||
<collectionCode id="7A83AE11542A3E735D32E43233F93F18" box="[376,463,1297,1323]" pageId="15" pageNumber="14">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ACFA3C89542A3E735DAEE43231303F19" box="[484,774,1296,1323]" country="Honduras" location="Department" pageId="15" pageNumber="14" specimenCount="1" stateProvince="Ocotepeque" typeStatus="paratype">
|
||
<emphasis id="2EE6EAC6542A3E735DAEE43231303F19" bold="true" box="[484,774,1296,1323]" pageId="15" pageNumber="14">
|
||
<location id="194D600F542A3E735DAEE43230423F18" LSID="urn:lsid:plazi:treatment:943B87C254293E735C8AE04831433CB8:194D600F542A3E735DAEE43230423F18" box="[484,628,1297,1323]" country="Honduras" name="Department" pageId="15" pageNumber="14" stateProvince="Ocotepeque">Department</location>
|
||
<collectingRegion id="DE56F836542A3E735E31E43331303F19" box="[635,774,1296,1322]" country="Honduras" name="Ocotepeque" pageId="15" pageNumber="14">Ocotepeque</collectingRegion>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ACFA3C89542A3E735F5EE432339D3F7F" collectingDate="1995-05-21" collectionCode="RHTC" collectorName="R. B. Guisayote & Turnbow" country="Honduras" pageId="15" pageNumber="14" specimenCount="1" typeStatus="paratype">
|
||
<collectorName id="B1675302542A3E735F5EE432318D3F19" box="[788,955,1296,1323]" pageId="15" pageNumber="14">R.B. Güisayote</collectorName>
|
||
,
|
||
<date id="682C1014542A3E735F8DE43336093F18" box="[967,1087,1296,1323]" pageId="15" pageNumber="14" value="1995-05-21">
|
||
<collectingDate id="7868E9FC542A3E735F8DE43336093F18" box="[967,1087,1296,1323]" pageId="15" pageNumber="14" value="1995-05-21">21-V-1995</collectingDate>
|
||
</date>
|
||
, R.
|
||
<collectorName id="B1675302542A3E735821E43336E43F19" box="[1131,1234,1296,1322]" pageId="15" pageNumber="14">Turnbow</collectorName>
|
||
, mercury vapor & blacklight (1
|
||
<collectionCode id="7A83AE11542A3E735D19E41133943F7F" box="[339,418,1330,1356]" pageId="15" pageNumber="14">RHTC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="48ED665C542A3E735C88E58637393EE8" ID-DOI="http://doi.org/10.5281/zenodo.14662356" ID-Zenodo-Dep="14662356" httpUri="https://zenodo.org/record/14662356/files/figure.png" pageId="15" pageNumber="14" startId="15.[194,273,1189,1213]" targetBox="[192,1440,229,1159]" targetPageId="15" targetType="figure">
|
||
<paragraph id="1C2D36D4542A3E735C88E58637393EE8" blockId="15.[192,1440,1189,1244]" pageId="15" pageNumber="14">
|
||
<emphasis id="2EE6EAC6542A3E735C88E58633693E8E" bold="true" box="[194,351,1189,1213]" pageId="15" pageNumber="14">Figures 26–29.</emphasis>
|
||
Abdominal ventrites and pygidia.
|
||
<emphasis id="2EE6EAC6542A3E735E82E58631263E8E" bold="true" box="[712,784,1189,1213]" pageId="15" pageNumber="14">26–27)</emphasis>
|
||
<taxonomicName id="DB924D57542A3E735F52E58631DC3E8E" authorityName="Skelley and Smith" authorityYear="2024" box="[792,1002,1189,1213]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="platypygus" status="sp. nov.">
|
||
<emphasis id="2EE6EAC6542A3E735F52E58631DC3E8E" box="[792,1002,1189,1213]" italics="true" pageId="15" pageNumber="14">Rhyparus platypygus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2EE6EAC6542A3E735FB8E58636593E8E" bold="true" box="[1010,1135,1189,1213]" pageId="15" pageNumber="14">
|
||
<taxonomicNameLabel id="35D557BD542A3E735FB8E58636593E8E" box="[1010,1135,1189,1213]" pageId="15" pageNumber="14" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
female holotype, ventral and lateral views.
|
||
<emphasis id="2EE6EAC6542A3E735D02E5E0335C3EE8" bold="true" box="[328,362,1219,1243]" pageId="15" pageNumber="14">28)</emphasis>
|
||
<taxonomicName id="DB924D57542A3E735D3AE5E733C33EE8" authorityName="Skelley and Smith" authorityYear="2024" box="[368,501,1219,1243]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="platypygus">
|
||
<emphasis id="2EE6EAC6542A3E735D3AE5E733C33EE8" box="[368,501,1219,1243]" italics="true" pageId="15" pageNumber="14">R. platypygus</emphasis>
|
||
</taxonomicName>
|
||
paratype male, lateral view.
|
||
<emphasis id="2EE6EAC6542A3E735F58E5E031023EE8" bold="true" box="[786,820,1219,1243]" pageId="15" pageNumber="14">29)</emphasis>
|
||
<taxonomicName id="DB924D57542A3E735F70E5E7319A3EE8" authorityName="Skelley and Smith" authorityYear="2024" box="[826,940,1219,1243]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="loughlini" status="new species">
|
||
<emphasis id="2EE6EAC6542A3E735F70E5E7319A3EE8" box="[826,940,1219,1243]" italics="true" pageId="15" pageNumber="14">R. loughlini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2EE6EAC6542A3E735FF8E5E0361A3EE8" bold="true" box="[946,1068,1219,1243]" pageId="15" pageNumber="14">
|
||
<taxonomicNameLabel id="35D557BD542A3E735FF8E5E0361A3EE8" box="[946,1068,1219,1243]" pageId="15" pageNumber="14" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="5488655F542A3E735C8AE47F33C53F8A" pageId="15" pageNumber="14" type="etymology">
|
||
<paragraph id="1C2D36D4542A3E735C8AE47F33C53F8A" blockId="15.[191,1440,1372,1465]" pageId="15" pageNumber="14">
|
||
<emphasis id="2EE6EAC6542A3E735C8AE47F33723F45" bold="true" box="[192,324,1372,1398]" pageId="15" pageNumber="14">Etymology.</emphasis>
|
||
Named
|
||
<taxonomicName id="DB924D57542A3E735DEAE47F30173F45" authorityName="Skelley and Smith" authorityYear="2024" box="[416,545,1372,1398]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="platypygus">
|
||
“
|
||
<emphasis id="2EE6EAC6542A3E735DE6E47F30173F45" box="[428,545,1372,1398]" italics="true" pageId="15" pageNumber="14">platypygus</emphasis>
|
||
</taxonomicName>
|
||
” for the highly modified pygidium of the females, which is also flattened in males when compared to other species. Derived from the Greek “
|
||
<emphasis id="2EE6EAC6542A3E735F1CE45E31A03FA4" box="[854,918,1405,1431]" italics="true" pageId="15" pageNumber="14">platys</emphasis>
|
||
” meaning broad or flat, combined with “
|
||
<emphasis id="2EE6EAC6542A3E73592EE45D37A33FA4" box="[1380,1429,1406,1431]" italics="true" pageId="15" pageNumber="14">pyge</emphasis>
|
||
” meaning rump or buttocks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5488655F542A3E735C8AE4EB31433CB8" pageId="15" pageNumber="14" type="discussion">
|
||
<paragraph id="1C2D36D4542A3E735C8AE4EB31433CB8" blockId="15.[192,1440,1480,1675]" pageId="15" pageNumber="14">
|
||
<emphasis id="2EE6EAC6542A3E735C8AE4EB337E3FD1" bold="true" box="[192,328,1480,1506]" pageId="15" pageNumber="14">Comments.</emphasis>
|
||
The modified and depressed female pygidium of
|
||
<taxonomicName id="DB924D57542A3E735F30E4EA363A3FD1" authorityName="Skelley and Smith" authorityYear="2024" box="[890,1036,1480,1507]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="platypygus">
|
||
<emphasis id="2EE6EAC6542A3E735F30E4EA31BC3FD1" box="[890,906,1481,1506]" italics="true" pageId="15" pageNumber="14">R</emphasis>
|
||
.
|
||
<emphasis id="2EE6EAC6542A3E735FDDE4EB363A3FD1" box="[919,1036,1480,1506]" italics="true" pageId="15" pageNumber="14">platypygus</emphasis>
|
||
</taxonomicName>
|
||
is presently unique for the Western Hemisphere members of
|
||
<taxonomicName id="DB924D57542A3E735D9FE4C930083C37" authorityName="Westwood" authorityYear="1845" box="[469,574,1514,1540]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2EE6EAC6542A3E735D9FE4C930083C37" box="[469,574,1514,1540]" italics="true" pageId="15" pageNumber="14">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
. However, this form of pygidium occurs in a group of species related to
|
||
<taxonomicName id="DB924D57542A3E735905E4C931683C15" authority="Minkina, Anichtchenko, Vasiljeva and Skelley" authorityName="Minkina, Anichtchenko, Vasiljeva and Skelley" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="stebnickae">
|
||
<emphasis id="2EE6EAC6542A3E735905E4C9336E3C15" italics="true" pageId="15" pageNumber="14">Rhyparus stebnickae</emphasis>
|
||
Minkina, Anichtchenko, Vasiljeva and Skelley
|
||
</taxonomicName>
|
||
found widely in
|
||
<collectingCountry id="64857644542A3E735850E72F36BE3C15" box="[1050,1160,1548,1574]" name="Indonesia" pageId="15" pageNumber="14">Indonesia</collectingCountry>
|
||
and oceanic countries to its east (e.g.,
|
||
<bibRefCitation id="78034B25542A3E735D18E70E30063C74" author="Minkina L. & Anichtchenko A & Vasiljeva A & Skelley PE" box="[338,560,1581,1608]" pageId="15" pageNumber="14" pagination="81 - 101" refId="ref11379" refString="Minkina L., Anichtchenko A, Vasiljeva A, Skelley PE. 2022. Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology 22: 81 - 101." type="journal article" year="2022">Minkina et al. 2022</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="78034B25542A3E735E71E70E30453C74" author="Minkina L & Jakl S & Bezdek A." box="[571,627,1581,1607]" pageId="15" pageNumber="14" pagination="1 - 5" refId="ref11466" refString="Minkina L, Jakl S, Bezdek A. 2023. A new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species with modified pygidium and last abdominal ventrite in females. Faunitaxys 11 (66): 1 - 5." type="journal article" year="2023">2023</bibRefCitation>
|
||
, 2024;
|
||
<bibRefCitation id="78034B25542A3E735E89E70E31FC3C74" author="Minkina L & Jakl S." box="[707,970,1581,1607]" pageId="15" pageNumber="14" pagination="131 - 152" refId="ref11431" refString="Minkina L, Jakl S. 2024. Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomical Series 20: 131 - 152." type="book chapter" year="2024">Minkina and Jákl 2024</bibRefCitation>
|
||
). All of these Eastern Hemisphere species differ from
|
||
<taxonomicName id="DB924D57542A3E735D0EE77333E03C5A" authorityName="Skelley and Smith" authorityYear="2024" box="[324,470,1615,1641]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="platypygus">
|
||
<emphasis id="2EE6EAC6542A3E735D0EE77333623C5A" box="[324,340,1616,1641]" italics="true" pageId="15" pageNumber="14">R</emphasis>
|
||
.
|
||
<emphasis id="2EE6EAC6542A3E735D2BE76C33E03C5A" box="[353,470,1615,1641]" italics="true" pageId="15" pageNumber="14">platypygus</emphasis>
|
||
</taxonomicName>
|
||
in several characters, most notably by being at least two times larger in body size and having densely punctate surfaces between the pronotal costae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |