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<document id="5083997C6014B6A591FDC44F2C8BDFE9" ID-CLB-Dataset="305520" ID-DOI="10.5852/cr-palevol2023v22a20" ID-GBIF-Dataset="7dfd23ea-32a7-4f34-a894-8ed01d6e3638" ID-ISSN="1777-571X" ID-Zenodo-Dep="14224818" ID-ZooBank="urn:lsid:zoobank.org:pub:ECFD1C38-6AD9-4558-B16B-2CC573012BFF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1732636787430" checkinUser="felipe" docAuthor="Moreau, Jean-David &amp; Néraudeau, Didier" docDate="2023" docId="923B87E9FF9BFFF0FF05FC681FD0F840" docLanguage="en" docName="CRPalevol.22.20.455-466.pdf" docOrigin="Comptes Rendus Palevol 22 (20)" docSource="http://dx.doi.org/10.5852/cr-palevol2023v22a20" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Weichselia reticulata Fontaine emend. Alvin 1971" docType="treatment" docVersion="2" lastPageNumber="460" masterDocId="6E02FF91FF9CFFF7FFCAFF9C1E4FFFAC" masterDocTitle="Amber and plants from the Upper Cretaceous of La Gripperie-Saint-Symphorien (Charente-Maritime, Western France)" masterLastPageNumber="466" masterPageNumber="455" pageNumber="460" updateTime="1732751150832" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="78FB5FF3F550FCD88BFEA6C24DE8B8F5">Amber and plants from the Upper Cretaceous of La Gripperie-Saint-Symphorien (Charente-Maritime, Western France)</mods:title>
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<mods:namePart id="56E7C9022B737EEABCF76F64A51BB629">Moreau, Jean-David</mods:namePart>
<mods:affiliation id="032B4237240A4603DADC304B261FB5A6">Biogéosciences, CNRS, université de Bourgogne, 6 boulevard Gabriel, F- 21000 Dijon (France) jean. david. moreau @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="45E8649B8A75FD1D212A51786347053F">Néraudeau, Didier</mods:namePart>
<mods:affiliation id="400A3AA86F41C2FEAB840336901A8F45">Géosciences, CNRS, Université Rennes I, Campus de Beaulieu, 263 avenue du général Leclerc, F- 35042 Rennes (France) didier. neraudeau @ univ-rennes 1. fr</mods:affiliation>
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<treatment id="923B87E9FF9BFFF0FF05FC681FD0F840" ID-DOI="http://doi.org/10.5281/zenodo.14226110" ID-GBIF-Taxon="247601340" ID-Zenodo-Dep="14226110" LSID="urn:lsid:plazi:treatment:923B87E9FF9BFFF0FF05FC681FD0F840" httpUri="http://treatment.plazi.org/id/923B87E9FF9BFFF0FF05FC681FD0F840" lastPageNumber="460" pageId="7" pageNumber="460" scope_family="Matoniaceae" scope_order="Gleicheniales">
<subSubSection id="52886574FF9BFFF0FF05FC681C27FB83" pageId="7" pageNumber="460" type="nomenclature">
<paragraph id="1A2D36FFFF9BFFF0FF05FC681C27FB83" blockId="7.[207,701,1012,1103]" pageId="7" pageNumber="460">
<taxonomicName id="DD924D7CFF9BFFF0FF05FC681C27FB83" authority="(Stokes &amp; Webb, 1824) Fontaine emend. Alvin, 1971" authorityName="Fontaine emend. Alvin" authorityYear="1971" baseAuthorityName="Stokes &amp; Webb" baseAuthorityYear="1824" class="Polypodiopsida" family="Weichseliaceae" genus="Weichselia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="7" pageNumber="460" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="28E6EAEDFF9BFFF0FF05FC681FFEFBA3" bold="true" box="[207,433,1012,1039]" italics="true" pageId="7" pageNumber="460">Weichselia reticulata</emphasis>
(Stokes &amp; Webb, 1824) Fontaine emend.
<bibRefCitation id="7E034B0EFF9BFFF0FE22FB891C27FB83" author="ALVIN K. L." box="[488,616,1045,1071]" pageId="7" pageNumber="460" pagination="1 - 33" refId="ref6305" refString="ALVIN K. L. 1971. - Weichselia reticulata (Stokes et Webb) Fontaine from the Wealden of Belgium. Memoire de l'Institut Royal des Sciences Naturelles de Belgique 166: 1 - 33." type="journal article" year="1971">Alvin, 1971</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="52886574FF9BFFF0FE4DFBA91C4BFBE3" box="[391,516,1077,1103]" pageId="7" pageNumber="460" type="description">
<paragraph id="1A2D36FFFF9BFFF0FE4DFBA91C4BFBE3" blockId="7.[207,701,1012,1103]" box="[391,516,1077,1103]" pageId="7" pageNumber="460">
(
<figureCitation id="82A92A7AFF9BFFF0FE5AFBA91FB2FBE3" box="[400,509,1077,1103]" captionStart="FIG" captionStartId="6.[132,143,1827,1844]" captionTargetBox="[143,1444,216,1800]" captionTargetId="figure-274@6.[692,1243,773,1334]" captionTargetPageId="6" captionText="FIG. 3. — Ferns and conifers from the Cenomanian deposits of La Gripperie-Saint-Symphorien:A-D, Matonia sp.,fragment of elongated and narrow pinnule bearing sunken, indusiate and circular sori; specimen (A, B) and its counterpart (C, D); photograph (A, C) and interpretative sketches (B, D); IGR-23246; E, sketch of a pinnule of Matonia sp. in transverse section, showing the persistence of indusia on both, the specimen and its counterpart; F, G, Weichselia reticulata (Stokes &amp; Webb, 1824) Fontaine emend. Alvin, 1971, pinna fragment bearing alternate, entire, oblong and leathery pinnules; photograph (F) and interpretative sketch (G); IGR-23247; H-J, Geinitzia reichenbachii (Geinitz, 1842) Hollick &amp; Jeffrey, 1909, photograph (H) and interpretative sketches (I) of a leafy shoot bearing helically arranged, elongated and falcate leaves; sketches of several transverse sections in a leave showing a rhomboidal to triangular shape, drew from broken surfaces of leaves (J); IGR-23248; K, L, Pagiophyllum sp., leafy axis bearing helically arranged, highly adpressed and imbricated scale-like leaves; photograph (K) and interpretative sketches (L); IGR-23250.Abbreviations:I.In., imprint of the indusium;I.St., insertion of the stalk;Mi., midvein;P.In., persisting indusium;So., sorus; St., stalk. Scale bars: A-I, K, L, 1 cm; J, 1 mm." pageId="7" pageNumber="460">Fig. 3F, G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52886574FF9BFFF0FF4EFBF21EBBFB0D" pageId="7" pageNumber="460" type="reference_group">
<paragraph id="1A2D36FFFF9BFFF0FF4EFBF21EBBFB0D" blockId="7.[132,774,1134,1185]" pageId="7" pageNumber="460">
<treatmentCitationGroup id="3A8211D1FF9BFFF0FF4EFBF21EBBFB0D" pageId="7" pageNumber="460">
<taxonomicName id="DD924D7CFF9BFFF0FF4EFBF21C18FB2A" authority="Stokes &amp; Webb, 1824: 423" authorityName="Stokes &amp; Webb" authorityPageNumber="423" authorityYear="1824" box="[132,599,1134,1158]" class="Polypodiopsida" family="Weichseliaceae" genus="Weichselia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="7" pageNumber="460" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="28E6EAEDFF9BFFF0FF4EFBF21F0CFB2A" box="[132,323,1134,1158]" italics="true" pageId="7" pageNumber="460">Weichselia reticulata</emphasis>
Stokes &amp; Webb, 1824: 423
</taxonomicName>
.
<treatmentCitation id="9B3310EEFF9BFFF0FDB9FBF21D49FB2A" author="ALVIN K. L." box="[627,774,1134,1158]" page="27" pageId="7" pageNumber="460" year="1971">
<bibRefCitation id="7E034B0EFF9BFFF0FDB9FBF21D49FB2A" author="ALVIN K. L." box="[627,774,1134,1158]" pageId="7" pageNumber="460" pagination="1 - 33" refId="ref6305" refString="ALVIN K. L. 1971. - Weichselia reticulata (Stokes et Webb) Fontaine from the Wealden of Belgium. Memoire de l'Institut Royal des Sciences Naturelles de Belgique 166: 1 - 33." type="journal article" year="1971">Alvin 1971: 27</bibRefCitation>
</treatmentCitation>
[cum syn.].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="52886574FF9BFFF0FF4EFB241C85FB7C" box="[132,714,1208,1232]" pageId="7" pageNumber="460" type="materials_examined">
<paragraph id="1A2D36FFFF9BFFF0FF4EFB241C85FB7C" blockId="7.[132,714,1208,1232]" box="[132,714,1208,1232]" pageId="7" pageNumber="460">
<smallCapsWord id="1CCBA023FF9BFFF0FF4EFB241EA8FB62" baselines="1225,1226" box="[132,231,1208,1231]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="7" pageNumber="460">MATERIAL</smallCapsWord>
. —
<materialsCitation id="AAFA3CA2FF9BFFF0FED9FB241C05FB7C" ID-GBIF-Occurrence="4985328312" box="[275,586,1208,1232]" pageId="7" pageNumber="460" specimenCode="IGR-23245_B" specimenCount="1">
<specimenCount id="0C94FD76FF9BFFF0FED9FB241FE4FB7C" box="[275,427,1208,1232]" count="2" pageId="7" pageNumber="460" type="generic">Two specimens</specimenCount>
:
<specimenCode id="4A349E84FF9BFFF0FE7EFB241C0AFB7C" box="[436,581,1208,1232]" pageId="7" pageNumber="460">IGR-23245_B</specimenCode>
;
</materialsCitation>
<materialsCitation id="AAFA3CA2FF9BFFF0FD9AFB241C85FB7C" ID-GBIF-Occurrence="4985328309" box="[592,714,1208,1232]" pageId="7" pageNumber="460" specimenCode="IGR-23247" specimenCount="1">
<specimenCode id="4A349E84FF9BFFF0FD9AFB241C89FB7C" box="[592,710,1208,1232]" pageId="7" pageNumber="460">IGR-23247</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52886574FF9BFFF0FF4EFB691C43F921" pageId="7" pageNumber="460" type="description">
<paragraph id="1A2D36FFFF9BFFF0FF4EFB691F5AFAA1" blockId="7.[131,777,1269,1677]" box="[132,277,1269,1295]" pageId="7" pageNumber="460">
<smallCapsWord id="1CCBA023FF9BFFF0FF4EFB691F5AFAA1" baselines="1288,1288" box="[132,277,1269,1295]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="460">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="1A2D36FFFF9BFFF0FF4EFA881C43F921" blockId="7.[131,777,1269,1677]" pageId="7" pageNumber="460">
The material consists of fragmented secondary pinnae (
<figureCitation id="82A92A7AFF9BFFF0FF44FAA81F4AFAE2" box="[142,261,1332,1358]" captionStart="FIG" captionStartId="6.[132,143,1827,1844]" captionTargetBox="[143,1444,216,1800]" captionTargetId="figure-274@6.[692,1243,773,1334]" captionTargetPageId="6" captionText="FIG. 3. — Ferns and conifers from the Cenomanian deposits of La Gripperie-Saint-Symphorien:A-D, Matonia sp.,fragment of elongated and narrow pinnule bearing sunken, indusiate and circular sori; specimen (A, B) and its counterpart (C, D); photograph (A, C) and interpretative sketches (B, D); IGR-23246; E, sketch of a pinnule of Matonia sp. in transverse section, showing the persistence of indusia on both, the specimen and its counterpart; F, G, Weichselia reticulata (Stokes &amp; Webb, 1824) Fontaine emend. Alvin, 1971, pinna fragment bearing alternate, entire, oblong and leathery pinnules; photograph (F) and interpretative sketch (G); IGR-23247; H-J, Geinitzia reichenbachii (Geinitz, 1842) Hollick &amp; Jeffrey, 1909, photograph (H) and interpretative sketches (I) of a leafy shoot bearing helically arranged, elongated and falcate leaves; sketches of several transverse sections in a leave showing a rhomboidal to triangular shape, drew from broken surfaces of leaves (J); IGR-23248; K, L, Pagiophyllum sp., leafy axis bearing helically arranged, highly adpressed and imbricated scale-like leaves; photograph (K) and interpretative sketches (L); IGR-23250.Abbreviations:I.In., imprint of the indusium;I.St., insertion of the stalk;Mi., midvein;P.In., persisting indusium;So., sorus; St., stalk. Scale bars: A-I, K, L, 1 cm; J, 1 mm." pageId="7" pageNumber="460">Fig. 3F, G</figureCitation>
). The largest pinna fragment (IGR-23247) is
<quantity id="DD6A9B1AFF9BFFF0FF4CFAC81E9BFAC2" box="[134,212,1364,1390]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="460" unit="mm" value="20.0">20 mm</quantity>
long and
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wide. The pinnae bear up to 230µm long and up to 165 µm wide alternate, entire, oblong and leathery pinnules. They are attached by a broad base to the rachis and display a rounded apex and an entire margin. The pinnules are slightly inclined forward, forming an angle of 60-70° with the rachis. Adjacent pinnules are not in contact between them, being inserted at
<quantity id="DD6A9B1AFF9BFFF0FE32F98F1CC2F982" box="[504,653,1555,1582]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="7" pageNumber="460" unit="mm" value="175.0" valueMax="190.0" valueMin="160.0">160-190 mm</quantity>
from each other. The main vein of pinnules is straight and ends before the apex. The secondary veins display anastomosing lateral venation forming a polygonal mesh.
</paragraph>
</subSubSection>
<subSubSection id="52886574FF9BFFF0FF4EF92F1FD0F840" pageId="7" pageNumber="460" type="discussion">
<paragraph id="1A2D36FFFF9BFFF0FF4EF92F1EA7F967" blockId="7.[130,776,1715,2028]" box="[132,232,1715,1741]" pageId="7" pageNumber="460">
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</paragraph>
<paragraph id="1A2D36FFFF9BFFF0FF4EF94E1FD0F840" blockId="7.[130,776,1715,2028]" pageId="7" pageNumber="460">
The gross morphology of pinnules as well as the reticulate venation allow to ascribe these specimens to
<taxonomicName id="DD924D7CFF9BFFF0FD5DF96E1EA6F880" authorityName="Stokes &amp; Webb" authorityYear="1824" class="Polypodiopsida" family="Weichseliaceae" genus="Weichselia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="7" pageNumber="460" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="28E6EAEDFF9BFFF0FD5DF96E1EA6F880" italics="true" pageId="7" pageNumber="460">Weichselia reticulata</emphasis>
</taxonomicName>
which is a common fern in the Cretaceous floras from Europe (
<bibRefCitation id="7E034B0EFF9BFFF0FE84F8AE1C32F8E0" author="BLANCO-MORENO C. &amp; GOMEZ B. &amp; BUSCALIONI A." box="[334,637,1842,1868]" pageId="7" pageNumber="460" pagination="571 - 578" refId="ref6466" refString="BLANCO-MORENO C., GOMEZ B. &amp; BUSCALIONI A. 2018. - Palaeobiogeographic and metric analysis of the Mesozoic fern Weichselia. Geobios 51 (6): 571 - 578. https: // doi. org / 10.1016 / j. geobios. 2018.05.001" type="journal article" year="2018">
Blanco-Moreno
<emphasis id="28E6EAEDFF9BFFF0FDCFF8AF1C73F8E0" box="[517,572,1842,1868]" italics="true" pageId="7" pageNumber="460">et al.</emphasis>
2018
</bibRefCitation>
). In Northern
<collectingCountry id="6285766FFF9BFFF0FF7BF8CF1EB2F8C1" box="[177,253,1875,1901]" name="France" pageId="7" pageNumber="460">France</collectingCountry>
,
<taxonomicName id="DD924D7CFF9BFFF0FEC1F8CF1FDDF8C0" authorityName="Stokes &amp; Webb" authorityYear="1824" box="[267,402,1874,1900]" class="Polypodiopsida" family="Weichseliaceae" genus="Weichselia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="7" pageNumber="460" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="28E6EAEDFF9BFFF0FEC1F8CF1FDDF8C0" box="[267,402,1874,1900]" italics="true" pageId="7" pageNumber="460">W. reticulata</emphasis>
</taxonomicName>
was previously reported from the Barremian-Aptian of the Paris Basin (
<bibRefCitation id="7E034B0EFF9BFFF0FDC6F8EE1CF2F820" author="CARPENTIER A." box="[524,701,1906,1932]" pageId="7" pageNumber="460" refId="ref6665" refString="CARPENTIER A. 1927. - La flore wealdienne de Feron-Glageon (Nord). Memoires de la Societe Geologique du Nord 1: 1 e 151." type="journal volume" year="1927">Carpentier 1927</bibRefCitation>
,
<bibRefCitation id="7E034B0EFF9BFFF0FD02F8EE1D4FF820" author="CARPENTIER A." box="[712,768,1906,1932]" pageId="7" pageNumber="460" refId="ref6693" refString="CARPENTIER A. 1929. - Recherches sur les vegetaux fossiles des argiles eocretaciques du Pays de Bray. Bulletin de la Societe Geologique de France 4: 89 e 96." type="journal volume" year="1929">1929</bibRefCitation>
;
<bibRefCitation id="7E034B0EFF9BFFF0FF4EF80E1F3FF800" author="DECONINCK J. - F. &amp; BOUE D. &amp; AMEDRO F. &amp; BAUDIN F. &amp; BRUNEAU L. &amp; HURET E. &amp; LANDREIN P. &amp; MOREAU J. - D. &amp; SANTONI A. L." box="[132,368,1938,1964]" pageId="7" pageNumber="460" refId="ref6916" refString="DECONINCK J. - F., BOUE D., AMEDRO F., BAUDIN F., BRUNEAU L., HURET E., LANDREIN P., MOREAU J. - D. &amp; SANTONI A. L. 2021. - First record of early Aptian Oceanic Anoxic Event 1 a from the Paris Basin (France) - Climate signals on a terrigenous shelf. Cretaceous Research 125: 104846. https: // doi. org / 10.1016 / j. cretres. 2021.104846" type="journal volume" year="2021">
Deconinck
<emphasis id="28E6EAEDFF9BFFF0FECBF80F1F7CF800" box="[257,307,1938,1964]" italics="true" pageId="7" pageNumber="460">et al.</emphasis>
2021
</bibRefCitation>
). In Charente-Maritime, this fern was reported from the Cenomanian clay of the Puy-Puy Lagerstätte (
<bibRefCitation id="7E034B0EFF9BFFF0FF04F84E1FC0F840" author="LE DIOURON T." box="[206,399,2001,2028]" pageId="7" pageNumber="460" refId="ref7733" refString="LE DIOURON T. 2005. - Les vegetaux fossiles des argiles de l'Albien terminal et du Cenomanien basal de la carriere de Puy-Puy (Tonnay- Charente, Charente-Maritime). Implications paleoenvironnementales. Diplome d'Etude Approfondies Universite de Rennes I, Rennes, 75 p. (in French)." type="book" year="2005">Le Diouron 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>