treatments-xml/data/8C/02/95/8C029544A90BFFCEFCC8FA4AFEF2CE19.xml
2024-06-21 12:43:25 +02:00

312 lines
39 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B5F5FD8DDC6AF40ADFAC25B00E7D2EEE" ID-DOI="10.1163/156854107779969646" ID-Zenodo-Dep="8111793" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1688432892703" checkinUser="carolina" docAuthor="Guerrero, Pablo &amp; Santiago, Reyes Peña-" docDate="2007" docId="8C029544A90BFFCEFCC8FA4AFEF2CE19" docLanguage="en" docName="Nematology.9.1.93-121.pdf" docOrigin="Nematology 9 (1)" docTitle="Enchodelus macrodorus Thorne 1939" docType="treatment" docVersion="6" lastPageNumber="116" masterDocId="703BED3CA918FFD9FFD1FF98FFBACA36" masterDocTitle="Redescription of Enchodelus species studied by Thorne in 1939 (Dorylaimida: Nordiidae)" masterLastPageNumber="121" masterPageNumber="93" pageNumber="112" updateTime="1688492984916" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="23293FD281A2C1B51CE26F23D73BF90D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B21774B8059A017DE7373FFFE801976A">
<mods:title id="49FF600E45B7785A0E2FBD626F0498C9">Redescription of Enchodelus species studied by Thorne in 1939 (Dorylaimida: Nordiidae)</mods:title>
</mods:titleInfo>
<mods:name id="5C723B94A0D5EC95FA791FB72CD191A4" type="personal">
<mods:role id="B657266C6243AA7A47766495105C9C40">
<mods:roleTerm id="61FD7333529D7E59D7B71AAA007FCC40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F85C0F8B58622ADB2C4D3B7D4F3FCADF">Guerrero, Pablo</mods:namePart>
<mods:affiliation id="8D192400C13380DF280DEF1EB6859338">Departamento de Biología Animal, Biología Vegetal y Ecología, Universidad de Jaén, Campus Las Lagunillas s / n, Edificio B 3, 23071 - Jaén, Spain</mods:affiliation>
</mods:name>
<mods:name id="95FC37712423E46DB110C60E493A1037" type="personal">
<mods:role id="CBCE9582CC89387FD4216DFC76EB3019">
<mods:roleTerm id="07DD73B14430E1A326D9261C1B63DB99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8EA7551BE2E5CDAB26A1403996E02130">Santiago, Reyes Peña-</mods:namePart>
<mods:affiliation id="0D57790C40AD35053BD2CBC516D2CF0A">Departamento de Biología Animal, Biología Vegetal y Ecología, Universidad de Jaén, Campus Las Lagunillas s / n, Edificio B 3, 23071 - Jaén, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="CD1BB4D507371865D9A620878A49A890">text</mods:typeOfResource>
<mods:relatedItem id="448CF59FE64853D6ECD35EA938835EA6" type="host">
<mods:titleInfo id="3C90CB265E2D7E626BD7224A933F7407">
<mods:title id="3394C88EAA85B3E6656198DAA9B7415A">Nematology</mods:title>
</mods:titleInfo>
<mods:part id="86EC3CC856E8D4C65A70DED194865EE9">
<mods:date id="2FBAC037EE31BD860AF10C9D4DBFD40F">2007</mods:date>
<mods:detail id="8DBC5986C47B76901D13A1D0ABFB8ED0" type="pubDate">
<mods:number id="CFCB87E5131862A0308A0DE4A240D02F">2007-12-31</mods:number>
</mods:detail>
<mods:detail id="06F555177D0D00A10076EE2679E76C2F" type="volume">
<mods:number id="D680AE0C99E0EBF42FB8389576ECF695">9</mods:number>
</mods:detail>
<mods:detail id="003EA4F753B7646A3CD9F3DA3A5A37D7" type="issue">
<mods:number id="C1F5EC08BD24CD75E3D24C8C41AE8FDB">1</mods:number>
</mods:detail>
<mods:extent id="15D6CC4C82144240E55BD06C156C3E58" unit="page">
<mods:start id="6D3718593EBD92D1C49558661444E312">93</mods:start>
<mods:end id="6A0F1EC1D687BC4F8D21B25A4B76EA45">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="70E86C6F5C4135161D146C14785A03DB">journal article</mods:classification>
<mods:identifier id="AA5243763C303337BF4938F522403666" type="DOI">10.1163/156854107779969646</mods:identifier>
<mods:identifier id="2FF206EE1FB46A811BFA4C29B70998E2" type="Zenodo-Dep">8111793</mods:identifier>
</mods:mods>
<treatment id="8C029544A90BFFCEFCC8FA4AFEF2CE19" ID-DOI="http://doi.org/10.5281/zenodo.8111782" ID-Zenodo-Dep="8111782" LSID="urn:lsid:plazi:treatment:8C029544A90BFFCEFCC8FA4AFEF2CE19" httpUri="http://treatment.plazi.org/id/8C029544A90BFFCEFCC8FA4AFEF2CE19" lastPageId="23" lastPageNumber="116" pageId="19" pageNumber="112">
<subSubSection id="4CB177D9A90BFFCAFCC8FA4AFB19CC9C" pageId="19" pageNumber="112" type="nomenclature">
<paragraph id="04142452A90BFFCAFCC8FA4AFB19CC9C" blockId="19.[768,1305,1490,1738]" pageId="19" pageNumber="112">
<heading id="5F5C933EA90BFFCAFCC8FA4AFBE5CC3A" bold="true" centered="true" fontSize="11" level="2" pageId="19" pageNumber="112" reason="0">
<taxonomicName id="C3AB5FD1A90BFFCAFCC8FA4AFBE5CC3A" authority="(de Man, 1880) Thorne, 1939" authorityName="Thorne" authorityYear="1939" baseAuthorityName="de Man" baseAuthorityYear="1880" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="19" pageNumber="112" phylum="Nematoda" rank="species" species="macrodorus">
<emphasis id="36DFF840A90BFFCAFCC8FA4AFBE5CC3A" bold="true" pageId="19" pageNumber="112">
<emphasis id="36DFF840A90BFFCAFCC8FA4AFB84CFDA" bold="true" box="[793,1086,1490,1516]" italics="true" pageId="19" pageNumber="112">Enchodelus macrodorus</emphasis>
(
<bibRefCitation id="603A59A3A90BFFCAFB9FFA4AFB42CFDA" author="DE MAN, J. G." box="[1102,1272,1490,1516]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="19" pageNumber="112" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
)
<bibRefCitation id="603A59A3A90BFFCAFC6BFA6AFBE5CC3A" author="THORNE, G." box="[954,1119,1522,1548]" journalOrPublisher="Capita Zoologica" pageId="19" pageNumber="112" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne, 1939</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
=
<taxonomicName id="C3AB5FD1A90BFFCAFCCFF988FB5CCC1C" authority="de Man, 1880" authorityName="de Man" authorityYear="1880" box="[798,1254,1552,1578]" class="Adenophorea" family="Dorylaimidae" genus="Dorylaimus" kingdom="Animalia" order="Dorylaimida" pageId="19" pageNumber="112" phylum="Nematoda" rank="species" species="macrodorus">
<emphasis id="36DFF840A90BFFCAFCCFF988FB83CC1F" box="[798,1081,1552,1577]" italics="true" pageId="19" pageNumber="112">Dorylaimus macrodorus</emphasis>
<bibRefCitation id="603A59A3A90BFFCAFB90F988FB5CCC1C" author="DE MAN, J. G." box="[1089,1254,1552,1578]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="19" pageNumber="112" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="C3AB5FD1A90BFFCAFCCFF9A8FB71CC5C" authority="(de Man, 1880) Ditlevsen, 1928" authorityName="Ditlevsen" authorityYear="1928" baseAuthorityName="de Man" baseAuthorityYear="1880" class="Adenophorea" family="Dorylaimidae" genus="Dorylaimus" kingdom="Animalia" order="Dorylaimida" pageId="19" pageNumber="112" phylum="Nematoda" rank="species" species="macrodorus" subGenus="Doryllium">
<emphasis id="36DFF840A90BFFCAFCCFF9A8FCEFCC5C" italics="true" pageId="19" pageNumber="112">Dorylaimus (Doryllium) macrodorus (</emphasis>
<bibRefCitation id="603A59A3A90BFFCAFC84F9C8FC47CC5C" author="DE MAN, J. G." box="[853,1021,1616,1642]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="19" pageNumber="112" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
<emphasis id="36DFF840A90BFFCAFC2CF9C8FBB2CC5C" box="[1021,1032,1616,1642]" italics="true" pageId="19" pageNumber="112">)</emphasis>
<bibRefCitation id="603A59A3A90BFFCAFBDEF9C8FB71CC5C" author="DITLEVSEN, H. J." box="[1039,1227,1616,1642]" editor="Jensen, A. S. &amp; Lundbeck, W. &amp; Mortensen, Th." journalOrPublisher="Zoology of the Faroes" pageId="19" pageNumber="112" pagination="1 - 28" part="13" refId="ref12962" refString="DITLEVSEN, H. J. (1928). Land- and freshwater nematodes. In: Jensen, A. S., Lundbeck, W. &amp; Mortensen, Th. (Eds). Zoology of the Faroes 13, pp. 1 - 28." title="Land- and freshwater nematodes" type="journal article" year="1928">Ditlevsen, 1928</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="C3AB5FD1A90BFFCAFCCFF9E8FB19CC9C" authority="(de Man, 1880) Thorne &amp; Swanger, 1936" authorityName="Thorne &amp; Swanger" authorityYear="1936" baseAuthorityName="de Man" baseAuthorityYear="1880" class="Adenophorea" family="Dorylaimidae" genus="Dorylaimellus" kingdom="Animalia" order="Dorylaimida" pageId="19" pageNumber="112" phylum="Nematoda" rank="species" species="macrodorus">
<emphasis id="36DFF840A90BFFCAFCCFF9E8FBD3CCBC" box="[798,1129,1648,1674]" italics="true" pageId="19" pageNumber="112">Dorylaimellus macrodorus (</emphasis>
<bibRefCitation id="603A59A3A90BFFCAFBB8F9E8FAB4CCBC" author="DE MAN, J. G." box="[1129,1294,1648,1674]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="19" pageNumber="112" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
<emphasis id="36DFF840A90BFFCAFADFF9E8FAA3CCBC" box="[1294,1305,1648,1674]" italics="true" pageId="19" pageNumber="112">)</emphasis>
<bibRefCitation id="603A59A3A90BFFCAFCA4F908FB19CC9C" author="THORNE, G. &amp; SWANGER, H. H." box="[885,1187,1680,1706]" journalOrPublisher="Capita Zoologica" pageId="19" pageNumber="112" pagination="1 - 220" part="6" refId="ref13314" refString="THORNE, G. &amp; SWANGER, H. H. (1936). A monograph of the nematode genera Dorylaimus Dujardin, Aporcelaimus n. g., Dorylaimoides n. g., and Pungentus n. g. Capita Zoologica 6, 1 - 220." title="A monograph of the nematode genera Dorylaimus Dujardin, Aporcelaimus n. g., Dorylaimoides n. g., and Pungentus n" type="journal article" year="1936">Thorne &amp; Swanger, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4CB177D9A90BFFCAFC6DF937FBE6CCFF" box="[956,1116,1711,1738]" pageId="19" pageNumber="112" type="description">
<paragraph id="04142452A90BFFCAFC6DF937FBE6CCFF" blockId="19.[768,1305,1490,1738]" box="[956,1116,1711,1738]" pageId="19" pageNumber="112">
<emphasis id="36DFF840A90BFFCAFC6DF937FC7DCCFF" box="[956,967,1711,1737]" italics="true" pageId="19" pageNumber="112">(</emphasis>
<figureCitation id="9C9038D7A90BFFCAFC16F928FBA4CCFC" box="[967,1054,1712,1738]" captionStart="Fig" captionStartId="20.[56,91,1652,1673]" captionTargetBox="[192,1205,144,1628]" captionTargetId="figure-4@20.[192,1206,144,1629]" captionTargetPageId="20" captionText="Fig. 10. Enchodelus macrodorus (de Man, 1880) Thorne, 1939 (female). A: Entire body; B: Anterior region; C: Reconstruction of superficial appearance of cephalic region; D: Vulval region; E: Pharyngeal region; F: Anterior genital branch; G: Tail. (Scale bars: A = 500 µm; B-D, G = 50 µm; E, F = 100 µm.)" figureDoi="http://doi.org/10.5281/zenodo.8111813" httpUri="https://zenodo.org/record/8111813/files/figure.png" pageId="19" pageNumber="112">Figs 10</figureCitation>
,
<figureCitation id="9C9038D7A90BFFCAFBE5F928FBEBCCFC" box="[1076,1105,1712,1738]" captionStart="Fig" captionStartId="22.[56,91,1619,1640]" captionTargetId="figure-6@21.[116,1281,144,1724]" captionTargetPageId="21" captionText="Fig. 11. Enchodelus macrodorus (de Man, 1880) Thorne, 1939 (female). A: Variability of anterior region; B: Expanded part of pharynx, showing dorsal nucleus (large arrow) and one of subventral nuclei (small arrow); C: Anterior genital branch showing sphincter (arrows); D: Base of pharynx; E: En face view of vulva; F: Egg; G: Variability of vulval region; H: Variability of tail. (Scale bars: A, D, E-H = 25 µm; B, C = 50 µm.)" figureDoi="http://doi.org/10.5281/zenodo.8111815" httpUri="https://zenodo.org/record/8111815/files/figure.png" pageId="19" pageNumber="112">11</figureCitation>
<emphasis id="36DFF840A90BFFCAFB80F937FBE6CCFF" box="[1105,1116,1711,1737]" italics="true" pageId="19" pageNumber="112">)</emphasis>
</paragraph>
</subSubSection>
<caption id="50D474DAA90CFFCDFFE9F9ECFDABCCF2" ID-DOI="http://doi.org/10.5281/zenodo.8111813" ID-Zenodo-Dep="8111813" httpUri="https://zenodo.org/record/8111813/files/figure.png" pageId="20" pageNumber="113" startId="20.[56,91,1652,1673]" targetBox="[192,1205,144,1628]" targetPageId="20" targetType="figure">
<paragraph id="04142452A90CFFCDFFE9F9ECFDABCCF2" blockId="20.[56,1343,1652,1733]" pageId="20" pageNumber="113">
<emphasis id="36DFF840A90CFFCDFFE9F9ECFF3CCCBF" bold="true" box="[56,134,1652,1673]" pageId="20" pageNumber="113">Fig. 10.</emphasis>
<taxonomicName id="C3AB5FD1A90CFFCDFF5FF9EDFF44CCBC" authorityName="Thorne" authorityYear="1939" box="[142,254,1653,1674]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="20" pageNumber="113" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
<emphasis id="36DFF840A90CFFCDFEAEF9EDFFFDCCF2" italics="true" pageId="20" pageNumber="113">
(
<bibRefCitation id="603A59A3A90CFFCDFE56F9EDFDA8CCBC" author="DE MAN, J. G." box="[391,530,1653,1674]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="20" pageNumber="113" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
)
<bibRefCitation id="603A59A3A90CFFCDFDF0F9EDFD1ECCBC" author="THORNE, G." box="[545,676,1653,1674]" journalOrPublisher="Capita Zoologica" pageId="20" pageNumber="113" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne, 1939</bibRefCitation>
(female). A: Entire body; B: Anterior region; C: Reconstruction of superficial appearance of cephalic region; D: Vulval region; E: Pharyngeal region; F: Anterior genital branch; G: Tail. (Scale bars: A
</emphasis>
=
<emphasis id="36DFF840A90CFFCDFFB4F937FEBECCF2" box="[101,260,1711,1732]" italics="true" pageId="20" pageNumber="113">500 µm; B-D, G</emphasis>
=
<emphasis id="36DFF840A90CFFCDFEF2F937FE20CCF2" box="[291,410,1711,1732]" italics="true" pageId="20" pageNumber="113">50 µm; E, F</emphasis>
= 100 µm.)
</paragraph>
</caption>
<subSubSection id="4CB177D9A90EFFCFFF82FF57FD3BCB90" pageId="22" pageNumber="115" type="materials_examined">
<paragraph id="04142452A90EFFCFFF82FF57FD3BCB90" blockId="22.[56,668,207,422]" pageId="22" pageNumber="115">
Nine
<specimenCount id="12ADEFDBA90EFFCFFF5FFF57FF59CAD0" box="[142,227,207,230]" pageId="22" pageNumber="115" type="female">females</specimenCount>
,
<materialsCitation id="B4C32E0FA90EFFCFFF3FFF57FF1ECB70" collectingDate="1932-06-22" country="United States of America" location="Utah" pageId="22" pageNumber="115" specimenCount="1" specimenCount-female="1" stateProvince="Utah">
<specimenCount id="12ADEFDBA90EFFCFFF3FFF57FEAFCAD0" box="[238,277,207,230]" pageId="22" pageNumber="115" type="female">five</specimenCount>
of them mounted on a slide labelled
<emphasis id="36DFF840A90EFFCFFFE9FF77FE85CB30" box="[56,319,239,262]" italics="true" pageId="22" pageNumber="115">
<taxonomicName id="C3AB5FD1A90EFFCFFFE9FF77FF0ECB30" authorityName="Thorne" authorityYear="1939" box="[56,180,239,262]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="22" pageNumber="115" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
</emphasis>
1h, and collected from Aspen Creek,
<collectingRegion id="C66FEAB0A90EFFCFFF5FFE97FF7ECB10" box="[142,196,271,294]" country="United States of America" name="Utah" pageId="22" pageNumber="115">Utah</collectingRegion>
,
<collectingCountry id="7CBC64C2A90EFFCFFF09FE97FEAACB10" box="[216,272,271,294]" name="United States of America" pageId="22" pageNumber="115">USA</collectingCountry>
, on
<date id="70150292A90EFFCFFE9CFE97FE4ACB10" box="[333,496,271,294]" pageId="22" pageNumber="115" value="1932-06-22">
<collectingDate id="6051FB7AA90EFFCFFE9CFE97FE4ACB10" box="[333,496,271,294]" pageId="22" pageNumber="115" value="1932-06-22">22 June 1932</collectingDate>
</date>
, in very good condition.
</materialsCitation>
<materialsCitation id="B4C32E0FA90EFFCFFF62FEB7FD3BCB90" collectedFrom="in sage brush soil" country="United States of America" location="Utah" pageId="22" pageNumber="115" specimenCount="1" specimenCount-female="1" stateProvince="Utah">
The other
<specimenCount id="12ADEFDBA90EFFCFFEE2FEB7FE78CB70" box="[307,450,303,326]" pageId="22" pageNumber="115" type="female">four females</specimenCount>
mounted on slide labelled
<emphasis id="36DFF840A90EFFCFFF47FED7FE20CB50" box="[150,410,335,358]" italics="true" pageId="22" pageNumber="115">
<taxonomicName id="C3AB5FD1A90EFFCFFF47FED7FEA8CB50" authorityName="Thorne" authorityYear="1939" box="[150,274,335,358]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="22" pageNumber="115" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
</emphasis>
1d, and collected from Strawberry Reservoir,
<collectingRegion id="C66FEAB0A90EFFCFFEFCFEF7FED9CBB0" box="[301,355,367,390]" country="United States of America" name="Utah" pageId="22" pageNumber="115">Utah</collectingRegion>
,
<collectingCountry id="7CBC64C2A90EFFCFFEA1FEF7FE12CBB0" box="[368,424,367,390]" name="United States of America" pageId="22" pageNumber="115">USA</collectingCountry>
,
<collectedFrom id="B7B767AFA90EFFCFFE64FEF7FDC3CBB0" box="[437,633,367,390]" pageId="22" pageNumber="115">in sage brush soil</collectedFrom>
, in September, 1924, three of them also in good condition.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CB177D9A90EFFCFFFE8FE4DFCE1CF58" pageId="22" pageNumber="115" type="description">
<paragraph id="04142452A90EFFCFFFE8FE4DFF40CBDA" blockId="22.[57,250,469,492]" box="[57,250,469,492]" pageId="22" pageNumber="115">
<heading id="5F5C933EA90EFFCFFFE8FE4DFF40CBDA" allCaps="true" box="[57,250,469,492]" fontSize="9" level="4" pageId="22" pageNumber="115" reason="3">
<smallCapsWord id="02F2B28EA90EFFCFFFE8FE4DFF40CBDA" baselines="486,487" box="[57,250,469,492]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Measurements" pageId="22" pageNumber="115">MEASUREMENTS</smallCapsWord>
</heading>
</paragraph>
<paragraph id="04142452A90EFFCFFF82FD94FF6FC815" blockId="22.[83,213,524,547]" box="[83,213,524,547]" pageId="22" pageNumber="115">
See
<tableCitation id="492911E9A90EFFCFFF51FD94FF6FC815" box="[128,213,524,547]" captionStart="Table 4" captionStartId="18.[56,112,154,175]" captionTargetPageId="18" captionText="Table 4. Morphometric data of Enchodelus hopedorus (Thorne, 1929) Thorne, 1939 and E. macrodorus (de Man, 1880) Thorne, 1939. Measurements are in µm (except L in mm), and in the form: mean ± standard deviation (range)." httpUri="http://table.plazi.org/id/50D474DAA90AFFCBFFE9FF02FC6FCAFB" pageId="22" pageNumber="115" tableUuid="50D474DAA90AFFCBFFE9FF02FC6FCAFB">Table 4.</tableCitation>
</paragraph>
<paragraph id="04142452A90EFFCFFFE8FDCAFF6EC85F" blockId="22.[57,212,594,617]" box="[57,212,594,617]" pageId="22" pageNumber="115">
<heading id="5F5C933EA90EFFCFFFE8FDCAFF6EC85F" allCaps="true" box="[57,212,594,617]" fontSize="9" level="4" pageId="22" pageNumber="115" reason="3">
<smallCapsWord id="02F2B28EA90EFFCFFFE8FDCAFF6EC85F" baselines="611,612" box="[57,212,594,617]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="22" pageNumber="115">DESCRIPTION</smallCapsWord>
</heading>
</paragraph>
<paragraph id="04142452A90EFFCFFFE9FD11FF3CC896" blockId="22.[56,134,649,672]" box="[56,134,649,672]" pageId="22" pageNumber="115">
<emphasis id="36DFF840A90EFFCFFFE9FD11FF3CC896" box="[56,134,649,672]" italics="true" pageId="22" pageNumber="115">Female</emphasis>
</paragraph>
<paragraph id="04142452A90EFFCFFF82FD2CFA96CF3B" blockId="22.[56,668,692,1546]" lastBlockId="22.[731,1343,153,1293]" pageId="22" pageNumber="115">
Moderately stout nematodes of medium size, 1.5- 1.7 mm long. Habitus after fixation slightly ventrad curved. Body cylindrical, tapering towards both ends but more so towards anterior. Cuticle 3.0-4.5
<emphasis id="36DFF840A90EFFCFFDC2FC8CFD9FC91D" box="[531,549,788,811]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m thick at anterior region, 3.5-4.5
<emphasis id="36DFF840A90EFFCFFEE5FCACFEFCC97D" box="[308,326,820,843]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m at mid-body and 8.5-11.0
<emphasis id="36DFF840A90EFFCFFDA5FCACFD3CC97D" box="[628,646,820,843]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m at tail; its outer layer practically smooth or bearing very fine transverse striations, and much thinner than inner, specially at level of tail. Lateral chord very narrow, less than one-tenth of mid-body diam., lacking any particular differentiation. Lateral pores obscure. Lip region rounded, offset by a marked depression, 2.5-3.5 times as broad as high and
<emphasis id="36DFF840A90EFFCFFF18FB8CFF58CE1D" box="[201,226,1044,1067]" italics="true" pageId="22" pageNumber="115">ca</emphasis>
one-third of body diam. at neck base. Lips amalgamated, perioral area hardly differentiated; labial and cephalic papillae visibly protruding from lip region contour. Amphid fovea cup-shaped, opening at level of cephalic depression and occupying one-half to two-thirds of corresponding body diam. Cheilostom a truncate cone, with thickened walls. Odontostyle long and slender, with distinct walls and narrow lumen; 11- 18 times as long as wide or 1.9-2.4 times as long as lip region diam.; aperture small,
<emphasis id="36DFF840A90EFFCFFE75FAACFE07CF7D" box="[420,445,1332,1355]" italics="true" pageId="22" pageNumber="115">ca</emphasis>
one-twelfth of total length. Odontophore bearing distinct flanges occupying
<emphasis id="36DFF840A90EFFCFFFE9FAEBFFEBCFBC" box="[56,81,1395,1418]" italics="true" pageId="22" pageNumber="115">ca</emphasis>
half of total length; scarcely longer (1.0-1.2 times) than odontostyle. Guiding ring double, at 25-28
<emphasis id="36DFF840A90EFFCFFD85FA0BFDDCCF9C" box="[596,614,1427,1450]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m or 1.3-1.5 times lip region diam. from anterior end. Pharynx consisting of slender but well muscled anterior portion expanding gradually into basal expansion at 59-62% of total neck length, reaching maximum diam. at 64- 68%; pharyngeal expansion occupying 35-38% of total neck length, and one-half to two-thirds of corresponding body diam. Pharyngeal gland nuclei located as follows: DN = 65-68; S
<subScript id="982F2617A90EFFCFFC5CFEBAFC2CCB04" attach="both" box="[909,918,290,306]" fontSize="7" pageId="22" pageNumber="115">1</subScript>
N
<subScript id="982F2617A90EFFCFFC7AFEBAFC0ECB04" attach="left" box="[939,948,290,306]" fontSize="7" pageId="22" pageNumber="115">1</subScript>
= 24-36; S
<subScript id="982F2617A90EFFCFFB90FEBAFBF0CB04" attach="both" box="[1089,1098,290,306]" fontSize="7" pageId="22" pageNumber="115">1</subScript>
N
<subScript id="982F2617A90EFFCFFB8EFEBAFBD2CB04" attach="left" box="[1119,1128,290,306]" fontSize="7" pageId="22" pageNumber="115">2</subScript>
= 27-37; S
<subScript id="982F2617A90EFFCFFB24FEBAFB44CB04" attach="left" box="[1269,1278,290,306]" fontSize="7" pageId="22" pageNumber="115">2</subScript>
N
<subScript id="982F2617A90EFFCFFAC2FEBAFAA6CB04" attach="left" box="[1299,1308,290,306]" fontSize="7" pageId="22" pageNumber="115">1</subScript>
= 54-59; S
<subScript id="982F2617A90EFFCFFCECFEDAFCFCCB64" attach="both" box="[829,838,322,338]" fontSize="7" pageId="22" pageNumber="115">2</subScript>
N
<subScript id="982F2617A90EFFCFFC8AFEDAFCDECB64" attach="left" box="[859,868,322,338]" fontSize="7" pageId="22" pageNumber="115">2</subScript>
= 57-61. Base of pharyngeal expansion surrounded by weak membrane-like structure. Cardia very small, rounded conoid, and wider than long (12- 19 × 9-16
<emphasis id="36DFF840A90EFFCFFC85FE00FCDCCB99" box="[852,870,408,431]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m). Genital system didelphic-amphidelphic, with both branches equally and well developed. Ovaries relatively large, 83-188
<emphasis id="36DFF840A90EFFCFFC06FE40FC53CBD9" box="[983,1001,472,495]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m long, but usually not reaching sphincter level; oocytes first in two or more rows, then one. Oviduct 97-159
<emphasis id="36DFF840A90EFFCFFC07FD80FC52C819" box="[982,1000,536,559]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m long, consisting of slender portion with prismatic cells and well developed
<emphasis id="36DFF840A90EFFCFFADEFDA0FC94C859" italics="true" pageId="22" pageNumber="115">pars dilatata</emphasis>
with distinct lumen. Sphincter distinct, located between oviduct and uterus. Uterus short, tripartite: wider proximal portion having distinct lumen followed by a narrower intermediate region with narrow lumen joining a small spheroid
<emphasis id="36DFF840A90EFFCFFC40FD40FBD7C8D9" box="[913,1133,728,751]" italics="true" pageId="22" pageNumber="115">pars dilatata distalis</emphasis>
; length 61-143
<emphasis id="36DFF840A90EFFCFFAC6FD4FFA93C8D8" box="[1303,1321,727,750]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m or 1.1-2.0 times body diam. at its level. Sperm not observed in genital tract. Two eggs observed inside both genital branches of one female, measuring 57 × 92 and 55 × 87
<emphasis id="36DFF840A90EFFCFFC70FCCFFC09C958" box="[929,947,855,878]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m.
<taxonomicName id="C3AB5FD1A90EFFCFFC0AFCCFFB9FC958" authorityName="Megerle" authorityYear="1811" box="[987,1061,855,878]" genus="Vagina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="115" phylum="Mollusca" rank="genus">Vagina</taxonomicName>
extending inwards to
<emphasis id="36DFF840A90EFFCFFAF4FCCFFA84C958" box="[1317,1342,855,878]" italics="true" pageId="22" pageNumber="115">ca</emphasis>
half (47-57%) of body diam.;
<emphasis id="36DFF840A90EFFCFFBFCFCEFFB6DC9B8" box="[1069,1239,887,910]" italics="true" pageId="22" pageNumber="115">pars proximalis</emphasis>
17-21 × 21-27
<emphasis id="36DFF840A90EFFCFFCF7FC0FFC82C998" box="[806,824,919,942]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m, with almost straight or somewhat sigmoid walls, and enveloped by moderately developed circular musculature;
<emphasis id="36DFF840A90EFFCFFCBEFC4FFBAFC9D8" box="[879,1045,983,1006]" italics="true" pageId="22" pageNumber="115">pars refringens</emphasis>
with (in lateral view) two close trapezoidal sclerotisations with a combined width of 19-20
<emphasis id="36DFF840A90EFFCFFCF3FB8EFC8ECE1B" box="[802,820,1046,1069]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m;
<emphasis id="36DFF840A90EFFCFFC89FB8FFC67CE18" box="[856,989,1047,1070]" italics="true" pageId="22" pageNumber="115">pars distalis</emphasis>
short, 4.0-6.0
<emphasis id="36DFF840A90EFFCFFBAFFB8EFB2ACE1B" box="[1150,1168,1046,1069]" italics="true" pageId="22" pageNumber="115">µ</emphasis>
m long. Vulva a transverse slit. Prerectum long, 3.9-5.8 anal body diam. Rectum almost equal to anal body diam. Tail short rounded to hemispheroid. Two pairs of caudal pores, both subterminal, one subdorsal, another subventral; most specimens bearing some small irregularities between inner and outer layers of cuticle, mainly at ventral side of the tail, sometimes appearing as vesicles in dorsal view.
</paragraph>
<paragraph id="04142452A90EFFCFFD0AFAB4FCA8CF75" blockId="22.[731,786,1324,1347]" box="[731,786,1324,1347]" pageId="22" pageNumber="115">
<emphasis id="36DFF840A90EFFCFFD0AFAB4FCA8CF75" box="[731,786,1324,1347]" italics="true" pageId="22" pageNumber="115">Male</emphasis>
</paragraph>
<paragraph id="04142452A90EFFCFFD27FACFFCE1CF58" blockId="22.[758,859,1367,1390]" box="[758,859,1367,1390]" pageId="22" pageNumber="115">Not seen.</paragraph>
</subSubSection>
<subSubSection id="4CB177D9A90EFFCEFD0DFA05FDC3CBE6" lastPageId="23" lastPageNumber="116" pageId="22" pageNumber="115" type="diagnosis">
<paragraph id="04142452A90EFFCFFD0DFA05FCE6CF82" blockId="22.[732,860,1437,1460]" box="[732,860,1437,1460]" pageId="22" pageNumber="115">
<heading id="5F5C933EA90EFFCFFD0DFA05FCE6CF82" allCaps="true" box="[732,860,1437,1460]" fontSize="9" level="4" pageId="22" pageNumber="115" reason="3">
<smallCapsWord id="02F2B28EA90EFFCFFD0DFA05FCE6CF82" baselines="1454,1455" box="[732,860,1437,1460]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="22" pageNumber="115">DIAGNOSIS</smallCapsWord>
</heading>
</paragraph>
<paragraph id="04142452A90EFFCEFD27FA4BFDC3CBE6" blockId="22.[731,1342,1491,1546]" lastBlockId="23.[56,667,153,464]" lastPageId="23" lastPageNumber="116" pageId="22" pageNumber="115">
<emphasis id="36DFF840A90EFFCFFD27FA4BFC46CFDC" box="[758,1020,1491,1514]" italics="true" pageId="22" pageNumber="115">
<taxonomicName id="C3AB5FD1A90EFFCFFD27FA4BFCC8CFDC" authorityName="Thorne" authorityYear="1939" box="[758,882,1491,1514]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="22" pageNumber="115" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
</emphasis>
is distinguished by its body 1.5-1.7 mm long, lip region offset by marked depression and 17.5-20
<emphasis id="36DFF840A90FFFCEFF16FF01FF63CA86" box="[199,217,153,176]" italics="true" pageId="23" pageNumber="116">µ</emphasis>
m diam., odontostyle 38-43
<emphasis id="36DFF840A90FFFCEFDC9FF01FD90CA86" box="[536,554,153,176]" italics="true" pageId="23" pageNumber="116">µ</emphasis>
m long or 1.9-2.4 times the lip region diam., odontophore provided with distinct basal flanges and slightly longer (1.0-1.2 times) than odontostyle, neck length 303-345
<emphasis id="36DFF840A90FFFCEFDE5FF61FDFCCB26" box="[564,582,249,272]" italics="true" pageId="23" pageNumber="116">µ</emphasis>
m long, pharyngeal expansion 111-130
<emphasis id="36DFF840A90FFFCEFE40FE81FE19CB06" box="[401,419,281,304]" italics="true" pageId="23" pageNumber="116">µ</emphasis>
m long and occupying 35-38% of total neck length, female genital system amphidelphic, uterus tripartite,
<emphasis id="36DFF840A90FFFCEFE44FEC1FD21CB46" box="[405,667,345,368]" italics="true" pageId="23" pageNumber="116">pars refringens vaginae</emphasis>
with two sclerotisations, vulva in form of a transverse slit and uniquely anterior (V = 41-44), and tail short rounded to hemispherical (18-24
<emphasis id="36DFF840A90FFFCEFEECFE21FEF5CBE6" box="[317,335,441,464]" italics="true" pageId="23" pageNumber="116">µ</emphasis>
m, c = 68-91,
<superScript id="F3DE891AA90FFFCEFE3BFE21FE41CBF3" attach="left" box="[490,507,436,464]" fontSize="7" pageId="23" pageNumber="116"></superScript>
= 0.5-0.6).
</paragraph>
</subSubSection>
<caption id="50D474DAA90EFFCFFFE9F9CBFE3ACCF6" ID-DOI="http://doi.org/10.5281/zenodo.8111815" ID-Zenodo-Dep="8111815" httpUri="https://zenodo.org/record/8111815/files/figure.png" pageId="22" pageNumber="115" startId="22.[56,91,1619,1640]" targetBox="[117,1281,145,1723]" targetPageId="21" targetType="figure">
<paragraph id="04142452A90EFFCFFFE9F9CBFE3ACCF6" blockId="22.[56,1343,1619,1728]" pageId="22" pageNumber="115">
<emphasis id="36DFF840A90EFFCFFFE9F9CBFF38CC5E" bold="true" box="[56,130,1619,1640]" pageId="22" pageNumber="115">Fig. 11.</emphasis>
<taxonomicName id="C3AB5FD1A90EFFCFFF56F9CBFF4DCC5E" authorityName="Thorne" authorityYear="1939" box="[135,247,1619,1640]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="22" pageNumber="115" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
<emphasis id="36DFF840A90EFFCFFEA3F9CBFEC1CC95" italics="true" pageId="22" pageNumber="115">
(
<bibRefCitation id="603A59A3A90EFFCFFEA8F9CBFE44CC5E" author="DE MAN, J. G." box="[377,510,1619,1640]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="22" pageNumber="115" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man, 1880</bibRefCitation>
)
<bibRefCitation id="603A59A3A90EFFCFFDDBF9CBFD30CC5E" author="THORNE, G." box="[522,650,1619,1640]" journalOrPublisher="Capita Zoologica" pageId="22" pageNumber="115" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne, 1939</bibRefCitation>
(female). A: Variability of anterior region; B: Expanded part of pharynx, showing dorsal nucleus (large arrow) and one of subventral nuclei (small arrow); C: Anterior genital branch showing sphincter (arrows); D: Base of pharynx; E:
</emphasis>
En face
<emphasis id="36DFF840A90EFFCFFE01F916FFC7CCF6" italics="true" pageId="22" pageNumber="115">view of vulva; F: Egg; G: Variability of vulval region; H: Variability of tail. (Scale bars: A, D, E-H</emphasis>
=
<emphasis id="36DFF840A90EFFCFFF4DF933FEAECCF6" box="[156,276,1707,1728]" italics="true" pageId="22" pageNumber="115">25 µm; B, C</emphasis>
= 50 µm.)
</paragraph>
</caption>
<subSubSection id="4CB177D9A90FFFCEFFE8FD98FEF2CE19" pageId="23" pageNumber="116" type="discussion">
<paragraph id="04142452A90FFFCEFFE8FD98FF11C821" blockId="23.[57,171,512,535]" box="[57,171,512,535]" pageId="23" pageNumber="116">
<heading id="5F5C933EA90FFFCEFFE8FD98FF11C821" allCaps="true" box="[57,171,512,535]" fontSize="9" level="4" pageId="23" pageNumber="116" reason="3">
<smallCapsWord id="02F2B28EA90FFFCEFFE8FD98FF11C821" baselines="529,530" box="[57,171,512,535]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Remarks" pageId="23" pageNumber="116">REMARKS</smallCapsWord>
</heading>
</paragraph>
<paragraph id="04142452A90FFFCEFF82FDA0FDC0C959" blockId="23.[56,668,568,1071]" pageId="23" pageNumber="116">
<bibRefCitation id="603A59A3A90FFFCEFF82FDA0FF48C879" author="THORNE, G." box="[83,242,568,591]" journalOrPublisher="Capita Zoologica" pageId="23" pageNumber="116" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne (1939)</bibRefCitation>
wrote that this species was “common at elevations of 8,000 to 12,000 feet in Wasatch and Uinta Mountains, Utah, U.S.A.”, and, although he did not mention the two localities from where the specimens studied herein were collected, it seems more than likely that this material belongs to that studied by him since his morphometric and morphological details fit perfectly with those given above.
<bibRefCitation id="603A59A3A90FFFCEFEDAFC80FE12C919" author="THORNE, G." box="[267,424,792,815]" journalOrPublisher="Capita Zoologica" pageId="23" pageNumber="116" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne (1939)</bibRefCitation>
described the male of the species but unfortunately nomale was found in the two available slides from the USDA Nematode Collection.
</paragraph>
<paragraph id="04142452A90FFFCEFF82FCE0FEB0C9D9" blockId="23.[56,668,568,1071]" pageId="23" pageNumber="116">
The present description and that of
<bibRefCitation id="603A59A3A90FFFCEFE0FFCE0FDC6C9B9" author="THORNE, G." box="[478,636,888,911]" journalOrPublisher="Capita Zoologica" pageId="23" pageNumber="116" pagination="1 - 261" part="8" refId="ref13246" refString="THORNE, G. (1939). A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica 8, 1 - 261." title="A monograph of the nematodes of the superfamily Dorylaimoidea" type="journal article" year="1939">Thorne (1939)</bibRefCitation>
of American specimens of
<emphasis id="36DFF840A90FFFCEFEE9FC00FE6EC999" box="[312,468,920,943]" italics="true" pageId="23" pageNumber="116">E. macrodorus</emphasis>
apparently fit very well that of
<bibRefCitation id="603A59A3A90FFFCEFF69FC20FEE3C9F9" author="DE MAN, J. G." box="[184,345,952,975]" journalOrPublisher="Tijdschrift Nederlandsche Dierkundige Vereeniging" pageId="23" pageNumber="116" pagination="1 - 104" part="5" refId="ref13148" refString="DE MAN, J. G. (1880). Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche Dierkundige Vereeniging 5, 1 - 104." title="Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden" type="journal article" year="1880">de Man (1880)</bibRefCitation>
, although, of course, the latter is not very detailed.
</paragraph>
<paragraph id="04142452A90FFFCEFF82FC60FEF2CE19" blockId="23.[56,668,568,1071]" pageId="23" pageNumber="116">
<emphasis id="36DFF840A90FFFCEFF82FC60FEEDCE39" box="[83,343,1016,1039]" italics="true" pageId="23" pageNumber="116">
<taxonomicName id="C3AB5FD1A90FFFCEFF82FC60FF75CE39" authorityName="Thorne" authorityYear="1939" box="[83,207,1016,1039]" class="Adenophorea" family="Dorylaimidae" genus="Enchodelus" kingdom="Animalia" order="Dorylaimida" pageId="23" pageNumber="116" phylum="Nematoda" rank="genus">Enchodelus</taxonomicName>
macrodorus
</emphasis>
seems to be very widely distributed in Holartic areas.
</paragraph>
</subSubSection>
</treatment>
</document>