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<document id="4566CEAE58F64F0FA548502B08FEF6A1" ID-CLB-Dataset="54218" ID-DOI="10.11646/zootaxa.5249.2.4" ID-GBIF-Dataset="34ef8754-e9dc-490a-affd-27027f406df2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687418" ID-ZooBank="81426096-9E0D-4E3D-9B1A-660B4D1BBB65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677656213532" checkinUser="plazi" docAuthor="Páll-Gergely, Barna" docDate="2023" docId="8B1E6842FF9F7327FF34FB36FED422C2" docLanguage="en" docName="zootaxa.5249.2.4.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pincerna costulosus" docType="treatment" docVersion="7" lastPageNumber="266" masterDocId="7727103AFF95732AFFA3FFB9FFB2234B" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna" masterLastPageNumber="276" masterPageNumber="253" pageNumber="263" updateTime="1699475959789" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C75A12C67A6343F22C4FE48D2166F9B4">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna</mods:title>
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<treatment id="8B1E6842FF9F7327FF34FB36FED422C2" ID-DOI="http://doi.org/10.5281/zenodo.7687460" ID-GBIF-Taxon="209384108" ID-Zenodo-Dep="7687460" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2" httpUri="http://treatment.plazi.org/id/8B1E6842FF9F7327FF34FB36FED422C2" lastPageId="13" lastPageNumber="266" pageId="10" pageNumber="263">
<subSubSection id="4BAD8ADFFF9F7320FF34FB36FCB427E3" box="[151,774,1166,1193]" pageId="10" pageNumber="263" type="nomenclature">
<paragraph id="0308D954FF9F7320FF34FB36FCB427E3" blockId="10.[151,774,1166,1229]" box="[151,774,1166,1193]" pageId="10" pageNumber="263">
<heading id="58406E38FF9F7320FF34FB36FCB427E3" bold="true" box="[151,774,1166,1193]" fontSize="11" level="1" pageId="10" pageNumber="263" reason="1">
<taxonomicName id="C4B7A2D7FF9F7320FF34FB36FCB427E3" ID-CoL="7PYC5" authority="(Bavay &amp; Dautzenberg, 1912)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[151,774,1166,1193]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF9F7320FF34FB36FCB427E3" bold="true" box="[151,774,1166,1193]" pageId="10" pageNumber="263">
<emphasis id="31C30546FF9F7320FF34FB36FE3927E2" bold="true" box="[151,395,1166,1193]" italics="true" pageId="10" pageNumber="263">Pincerna costulosus</emphasis>
(
<bibRefCitation id="6726A4A5FF9F7320FE39FB37FD4F27E3" author="Bavay, A. &amp; Dautzenberg, P." box="[410,765,1166,1193]" pageId="10" pageNumber="263" pagination="1 - 54" refId="ref13737" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de Coquilles nouvelles de I'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54. [https: // www. biodiversitylibrary. org / page / 16358455]" type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9F7320FF34FB0AFEB92786" box="[151,267,1203,1229]" pageId="10" pageNumber="263" type="description">
<paragraph id="0308D954FF9F7320FF34FB0AFEB92786" blockId="10.[151,774,1166,1229]" box="[151,267,1203,1229]" pageId="10" pageNumber="263">
(
<figureCitation id="9B8CC5D1FF9F7320FF3CFB0AFF552786" box="[159,231,1203,1229]" captionStart="FIGURE 8" captionStartId="11.[152,255,1975,1999]" captionTargetBox="[151,1436,885,1952]" captionTargetId="figure-310@11.[151,1436,885,1952]" captionTargetPageId="11" captionText="FIGURE 8. Shells of Pincerna costulosus (Bavay &amp; Dautzenberg, 1912).AD: MNHN-IM-2000-31786 (syntype); EH:Phong Tho, MNHN-IM-2012-27043 (probable syntype); IL: Northern Laos, MNHN-IM-2012-27218." figureDoi="http://doi.org/10.5281/zenodo.7687434" httpUri="https://zenodo.org/record/7687434/files/figure.png" pageId="10" pageNumber="263">Figs 8</figureCitation>
,
<figureCitation id="9B8CC5D1FF9F7320FF57FB0AFEB12786" box="[244,259,1203,1229]" captionStart="FIGURE 9" captionStartId="12.[152,255,1611,1635]" captionTargetBox="[151,1436,181,1587]" captionTargetId="figure-31@12.[151,1436,181,1587]" captionTargetPageId="12" captionText="FIGURE 9. Shells of Pincerna costulosus(Bavay &amp; Dautzenberg, 1912).AD: Laos, Phongsaly Province, JG29; EH:Holotype of Pincerna vallis Z.-Y. Chen &amp; Wu, 2020; IM: Pincerna cf. costulosus, Guangxi, China, 2009/104. Photos: B. Páll-Gergely (AD, IH), taken from the original description (EH)." figureDoi="http://doi.org/10.5281/zenodo.7687436" httpUri="https://zenodo.org/record/7687436/files/figure.png" pageId="10" pageNumber="263">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9F7320FF34FB4EFDCC2620" pageId="10" pageNumber="263" type="reference_group">
<paragraph id="0308D954FF9F7320FF34FB4EFCC42644" blockId="10.[151,1302,1271,1387]" box="[151,886,1271,1296]" pageId="10" pageNumber="263">
<treatmentCitationGroup id="23A7FE7AFF9F7320FF34FB4EFCC42644" box="[151,886,1271,1296]" pageId="10" pageNumber="263">
<treatmentCitation id="8216FF45FF9F7320FF34FB4EFD6B2644" author="Bavay, A. &amp; Dautzenberg, P." box="[151,729,1271,1296]" page="50" pageId="10" pageNumber="263" year="1912">
<taxonomicName id="C4B7A2D7FF9F7320FF34FB4EFD6B2644" ID-CoL="7PYC5" authority="Bavay &amp; Dautzenberg, 1912: 49 - 50" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="49 - 50" authorityYear="1912" box="[151,729,1271,1296]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF9F7320FF34FB4EFED12644" box="[151,355,1271,1295]" italics="true" pageId="10" pageNumber="263">Alycaeus costulosus</emphasis>
<bibRefCitation id="6726A4A5FF9F7320FECAFB41FD6B2644" author="Bavay, A. &amp; Dautzenberg, P." box="[361,729,1271,1296]" pageId="10" pageNumber="263" pagination="1 - 54" refId="ref13737" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de Coquilles nouvelles de I'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54. [https: // www. biodiversitylibrary. org / page / 16358455]" type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912: 4950</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 4, figs 14.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0308D954FF9F7320FF34FAAEFAA42665" blockId="10.[151,1302,1271,1387]" box="[151,1302,1302,1326]" pageId="10" pageNumber="263">
<treatmentCitationGroup id="23A7FE7AFF9F7320FF34FAAEFAA42665" box="[151,1302,1302,1326]" pageId="10" pageNumber="263">
<emphasis id="31C30546FF9F7320FF34FAAEFEC32665" box="[151,369,1302,1326]" italics="true" pageId="10" pageNumber="263">
<taxonomicName id="C4B7A2D7FF9F7320FF34FAAEFEE42665" ID-CoL="7SQ3L" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[151,342,1302,1326]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="costulosa">Pincerna costulosa</taxonomicName>
</emphasis>
Páll-Gergely
<emphasis id="31C30546FF9F7320FE5BFAAEFD992665" box="[504,555,1302,1326]" italics="true" pageId="10" pageNumber="263">et al.</emphasis>
, 2017: 10, fig. 3F; Páll-Gergely 2017: 214;
<treatmentCitation id="8216FF45FF9F7320FC4AFAAFFAA02665" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J. D." box="[1001,1298,1302,1326]" httpUri="http://treatment.plazi.org/id/7C36A7C8790D57B8B6981E693A7457CA" page="172" pageId="10" pageNumber="263" year="2020">
<bibRefCitation id="6726A4A5FF9F7320FC4AFAAFFAA02665" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J. D." box="[1001,1298,1302,1326]" pageId="10" pageNumber="263" pagination="1 - 220" refId="ref15744" refString="Pall-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. &amp; Ablett, J. D. (2020) Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys, 981, 1 - 220. https: // doi. org / 10.3897 / zookeys. 981.53583" type="journal article" year="2020">
Páll-Gergely
<emphasis id="31C30546FF9F7320FBD3FAAEFB162665" box="[1136,1188,1302,1326]" italics="true" pageId="10" pageNumber="263">et al.</emphasis>
2020: 172
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0308D954FF9F7320FF34FA8CFC7C2606" blockId="10.[151,1302,1271,1387]" box="[151,974,1333,1357]" pageId="10" pageNumber="263">
<taxonomicName id="C4B7A2D7FF9F7320FF34FA8CFE832606" ID-CoL="7SQ46" authorityName="Z. -Y. Chen &amp; M. Wu" authorityYear="2020" box="[151,305,1333,1357]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="31C30546FF9F7320FF34FA8CFE832606" box="[151,305,1333,1357]" italics="true" pageId="10" pageNumber="263">Pincerna vallis</emphasis>
</taxonomicName>
Z.-Y. Chen &amp; M. Wu, 2020: 42, figs 1, 4A, B, 5A.
<emphasis id="31C30546FF9F7320FC98FA8CFC7C2606" bold="true" box="[827,974,1333,1357]" pageId="10" pageNumber="263">new synonym</emphasis>
</paragraph>
<paragraph id="0308D954FF9F7320FF34FAEDFDCC2620" blockId="10.[151,1302,1271,1387]" box="[151,638,1363,1387]" pageId="10" pageNumber="263">
<treatmentCitationGroup id="23A7FE7AFF9F7320FF34FAEDFDCC2620" box="[151,638,1363,1387]" pageId="10" pageNumber="263">
<taxonomicName id="C4B7A2D7FF9F7320FF34FAEDFE832620" ID-CoL="7SQ46" authorityName="Z. -Y. Chen &amp; M. Wu" authorityYear="2020" box="[151,305,1363,1387]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="31C30546FF9F7320FF34FAEDFE832620" box="[151,305,1363,1387]" italics="true" pageId="10" pageNumber="263">Pincerna vallis</emphasis>
</taxonomicName>
<treatmentCitation id="8216FF45FF9F7320FEE8FAEAFDC82620" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J. D." box="[331,634,1363,1387]" httpUri="http://treatment.plazi.org/id/D8B3E62CE37851A2975A425355781F39" page="185" pageId="10" pageNumber="263" year="2020">
<bibRefCitation id="6726A4A5FF9F7320FEE8FAEAFDC82620" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J. D." box="[331,634,1363,1387]" pageId="10" pageNumber="263" pagination="1 - 220" refId="ref15744" refString="Pall-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. &amp; Ablett, J. D. (2020) Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys, 981, 1 - 220. https: // doi. org / 10.3897 / zookeys. 981.53583" type="journal article" year="2020">
Páll-Gergely
<emphasis id="31C30546FF9F7320FE71FAEDFDB72620" box="[466,517,1363,1387]" italics="true" pageId="10" pageNumber="263">et al.</emphasis>
, 2020: 185
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9F7321FF34FA23FAD4239E" lastPageId="11" lastPageNumber="264" pageId="10" pageNumber="263" type="materials_examined">
<paragraph id="0308D954FF9F7320FF34FA23FB682693" blockId="10.[151,1437,1433,2036]" pageId="10" pageNumber="263">
<emphasis id="31C30546FF9F7320FF34FA23FEFA26F8" bold="true" box="[151,328,1433,1460]" pageId="10" pageNumber="263">Type material.</emphasis>
<materialsCitation id="B3DFD309FF9F7320FEF1FA23FB642693" ID-GBIF-Occurrence="4039909359" collectionCode="MNHN" collectorName="Messager &amp; Staadt" country="Vietnam" location="Tonkin" municipality="Phong Tho" pageId="10" pageNumber="263" specimenCode="MNHN-IM-2000-31786, IM-2012-27043" specimenCount="2" typeStatus="syntype">
<collectingCountry id="7BA099C4FF9F7320FEF1FA23FE6A26FF" box="[338,472,1434,1460]" name="Vietnam" pageId="10" pageNumber="263">VIETNAM</collectingCountry>
:
<location id="06688F8FFF9F7320FE46FA20FD8526F8" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FE46FA20FD8526F8" box="[485,567,1433,1459]" country="Vietnam" municipality="Phong Tho" name="Tonkin" pageId="10" pageNumber="263">Tonkin</location>
,
<collectingMunicipality id="E36C432EFF9F7320FDE6FA20FD7426F8" box="[581,710,1433,1459]" pageId="10" pageNumber="263">Phong Tho</collectingMunicipality>
, leg.
<collectorName id="AE42BC82FF9F7320FCA4FA23FCC626FF" box="[775,884,1434,1460]" pageId="10" pageNumber="263">Messager</collectorName>
,
<specimenCode id="5311712FFF9F7320FC20FA20FB2626F8" box="[899,1172,1433,1459]" collectionCode="MNHN-IM-" pageId="10" pageNumber="263">MNHN-IM-2000-31786</specimenCode>
(
<typeStatus id="DC0C67F6FF9F7320FB04FA23FAB026FF" box="[1191,1282,1434,1460]" pageId="10" pageNumber="263" type="syntype">syntype</typeStatus>
,
<figureCitation id="9B8CC5D1FF9F7320FAB3FA20FA2226FF" box="[1296,1424,1433,1460]" captionStart="FIGURE 8" captionStartId="11.[152,255,1975,1999]" captionTargetBox="[151,1436,885,1952]" captionTargetId="figure-310@11.[151,1436,885,1952]" captionTargetPageId="11" captionText="FIGURE 8. Shells of Pincerna costulosus (Bavay &amp; Dautzenberg, 1912).AD: MNHN-IM-2000-31786 (syntype); EH:Phong Tho, MNHN-IM-2012-27043 (probable syntype); IL: Northern Laos, MNHN-IM-2012-27218." figureDoi="http://doi.org/10.5281/zenodo.7687434" httpUri="https://zenodo.org/record/7687434/files/figure.png" pageId="10" pageNumber="263">Fig. 8AD</figureCitation>
); Tonkin, Phong Tho, coll.
<collectorName id="AE42BC82FF9F7320FE16FA04FE4C269C" box="[437,510,1469,1495]" pageId="10" pageNumber="263">Staadt</collectorName>
, 1969,
<collectionCode id="65A64191FF9F7320FDECFA07FD172693" box="[591,677,1470,1496]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FD0FFA07FCED2693" box="[684,863,1470,1496]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27043</specimenCode>
/2 (probably syntype,
<figureCitation id="9B8CC5D1FF9F7320FBF7FA04FB7F2693" box="[1108,1229,1469,1496]" captionStart="FIGURE 8" captionStartId="11.[152,255,1975,1999]" captionTargetBox="[151,1436,885,1952]" captionTargetId="figure-310@11.[151,1436,885,1952]" captionTargetPageId="11" captionText="FIGURE 8. Shells of Pincerna costulosus (Bavay &amp; Dautzenberg, 1912).AD: MNHN-IM-2000-31786 (syntype); EH:Phong Tho, MNHN-IM-2012-27043 (probable syntype); IL: Northern Laos, MNHN-IM-2012-27218." figureDoi="http://doi.org/10.5281/zenodo.7687434" httpUri="https://zenodo.org/record/7687434/files/figure.png" pageId="10" pageNumber="263">Fig. 8EH</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0308D954FF9F7321FF64FA58FAD4239E" blockId="10.[151,1437,1433,2036]" lastBlockId="11.[151,1437,151,861]" lastPageId="11" lastPageNumber="264" pageId="10" pageNumber="263">
<materialsCitation id="B3DFD309FF9F7320FF64FA58FAAC256B" ID-GBIF-Occurrence="4039909329" collectionCode="V" collectorName="Abdou, A. &amp; Muratov, I." country="Laos" elevation="575" latitude="21.433983" location="Nam Leng" longLatPrecision="1" longitude="102.03609" pageId="10" pageNumber="263" specimenCount="1" stateProvince="Phongsali">
<emphasis id="31C30546FF9F7320FF64FA58FD8B26B0" bold="true" box="[199,569,1505,1531]" pageId="10" pageNumber="263">Additional material examined.</emphasis>
<collectingCountry id="7BA099C4FF9F7320FDE1FA5BFD3D26B7" box="[578,655,1506,1532]" name="Laos" pageId="10" pageNumber="263">LAOS</collectingCountry>
:
<collectingRegion id="C17317B6FF9F7320FD3FFA58FC3226B0" box="[668,896,1505,1531]" country="Laos" name="Phongsali" pageId="10" pageNumber="263">Phongsaly Province</collectingRegion>
, old forest on the northern bank of
<location id="06688F8FFF9F7320FABFFA5BFA2A26B7" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FABFFA5BFA2A26B7" box="[1308,1432,1506,1532]" country="Laos" latitude="21.433983" longLatPrecision="1" longitude="102.03609" name="Nam Leng" pageId="10" pageNumber="263" stateProvince="Phongsali">Nam Leng</location>
, 575 m,
<geoCoordinate id="6683BF93FF9F7320FF4DF9BEFECD2569" box="[238,383,1543,1570]" degrees="21" direction="north" minutes="26.039" orientation="latitude" pageId="10" pageNumber="263" precision="1" value="21.433983">21°26.039N</geoCoordinate>
,
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(locality code: 21L05), clay, shale, leg.
<collectorName id="AE42BC82FF9F7320FC4CF9BCFBD6256B" box="[1007,1124,1541,1568]" pageId="10" pageNumber="263">Abdou, A.</collectorName>
&amp;
<collectorName id="AE42BC82FF9F7320FB29F9BFFAB4256B" box="[1162,1286,1542,1568]" pageId="10" pageNumber="263">Muratov, I.</collectorName>
<collectionCode id="65A64191FF9F7320FAA5F9BFFAAC256B" box="[1286,1310,1542,1568]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="263" type="Museum">V</collectionCode>
</materialsCitation>
.,
<materialsCitation id="B3DFD309FF9F7320FA8CF9BCFE4A250F" ID-GBIF-Occurrence="4039909360" collectingDate="2005-03-15" collectionCode="MNHN" country="Laos" pageId="10" pageNumber="263" specimenCode="IM-2012-27217" specimenCount="1">
<date id="7709FF94FF9F7320FA8CF9BCFF60250F" pageId="10" pageNumber="263" value="2005-03-15">
<collectingDate id="674D067CFF9F7320FA8CF9BCFF60250F" pageId="10" pageNumber="263" value="2005-03-15">15 March 2005</collectingDate>
</date>
,
<collectionCode id="65A64191FF9F7320FF7CF993FE87250F" box="[223,309,1578,1604]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FE9EF993FE4A250F" box="[317,504,1578,1604]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27217</specimenCode>
</materialsCitation>
/1;
<materialsCitation id="B3DFD309FF9F7320FDB4F990FAC42523" ID-GBIF-Occurrence="4039909315" collectionCode="V" collectorName="Abdou, A. &amp; Muratov, I." country="Laos" elevation="560" latitude="21.427933" location="Nam Leng" longLatPrecision="1" longitude="102.01207" pageId="10" pageNumber="263" specimenCount="1" stateProvince="Phongsali">
<collectingRegion id="C17317B6FF9F7320FDB4F990FD482508" box="[535,762,1577,1603]" country="Laos" name="Phongsali" pageId="10" pageNumber="263">Phongsaly Province</collectingRegion>
, old forest just north of
<location id="06688F8FFF9F7320FBB1F993FB39250F" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FBB1F993FB39250F" box="[1042,1163,1578,1604]" country="Laos" latitude="21.427933" longLatPrecision="1" longitude="102.01207" name="Nam Leng" pageId="10" pageNumber="263" stateProvince="Phongsali">Nam Leng</location>
(river), ca.
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, 560 m,
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,
<geoCoordinate id="6683BF93FF9F7320FDA2F9F6FD282521" box="[513,666,1615,1642]" degrees="102" direction="east" minutes="00.724" orientation="longitude" pageId="10" pageNumber="263" precision="1" value="102.01207">102°00.724E</geoCoordinate>
(locality code: 31L04), clay, shale, leg.
<collectorName id="AE42BC82FF9F7320FBFCF9F4FB642523" box="[1119,1238,1613,1640]" pageId="10" pageNumber="263">Abdou, A.</collectorName>
,
<collectorName id="AE42BC82FF9F7320FB40F9F7FAEC2523" box="[1251,1374,1614,1640]" pageId="10" pageNumber="263">Muratov, I.</collectorName>
<collectionCode id="65A64191FF9F7320FAFDF9F7FAC42523" box="[1374,1398,1614,1640]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="263" type="Museum">V</collectionCode>
</materialsCitation>
., &amp;
<materialsCitation id="B3DFD309FF9F7320FF34F9C8FC2A25C7" ID-GBIF-Occurrence="4039909350" box="[151,920,1649,1676]" collectingDate="2004-12-07" collectionCode="MNHN" country="Laos" location="Phothihoumphanh" pageId="10" pageNumber="263" specimenCode="IM-2012-27220" specimenCount="1">
<location id="06688F8FFF9F7320FF34F9C8FED525C0" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FF34F9C8FED525C0" box="[151,359,1649,1675]" country="Laos" name="Phothihoumphanh" pageId="10" pageNumber="263">Phothihoumphanh</location>
, N.,
<date id="7709FF94FF9F7320FE3DF9CBFDCA25C7" box="[414,632,1649,1676]" pageId="10" pageNumber="263" value="2004-12-07">
<collectingDate id="674D067CFF9F7320FE3DF9CBFDCA25C7" box="[414,632,1649,1676]" pageId="10" pageNumber="263" value="2004-12-07">07 December 2004</collectingDate>
</date>
,
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<specimenCode id="5311712FFF9F7320FD42F9CBFC2A25C7" box="[737,920,1650,1676]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27220</specimenCode>
</materialsCitation>
/2;
<materialsCitation id="B3DFD309FF9F7320FC19F9C8FEF4259F" ID-GBIF-Occurrence="4039909327" collectionCode="V" collectorName="Abdou, A. &amp; Muratov, I." country="Laos" elevation="801" latitude="21.367567" location="SE of Boun Tai" longLatPrecision="1" longitude="102.000565" pageId="10" pageNumber="263" specimenCount="1" stateProvince="Phongsali">
<collectingRegion id="C17317B6FF9F7320FC19F9C8FB2E25C0" box="[954,1180,1649,1675]" country="Laos" name="Phongsali" pageId="10" pageNumber="263">Phongsaly Province</collectingRegion>
, old forest and banana plantation
<location id="06688F8FFF9F7320FEACF92FFE7025E4" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FEACF92FFE7025E4" box="[271,450,1685,1712]" country="Laos" latitude="21.367567" longLatPrecision="1" longitude="102.000565" name="SE of Boun Tai" pageId="10" pageNumber="263" stateProvince="Phongsali">SE of Boun Tai</location>
, 801 m,
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,
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(locality code: 23L04), clay, shale, leg.
<collectorName id="AE42BC82FF9F7320FA84F92CFA2E25FB" box="[1319,1436,1685,1712]" pageId="10" pageNumber="263">Abdou, A.</collectorName>
&amp;
<collectorName id="AE42BC82FF9F7320FF10F903FE9C259F" box="[179,302,1722,1748]" pageId="10" pageNumber="263">Muratov, I.</collectorName>
<collectionCode id="65A64191FF9F7320FE8DF903FEF4259F" box="[302,326,1722,1748]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="263" type="Museum">V</collectionCode>
</materialsCitation>
.,
<materialsCitation id="B3DFD309FF9F7320FEF6F903FCE1259F" ID-GBIF-Occurrence="4039909343" box="[341,851,1721,1748]" collectingDate="2004-12-02" collectionCode="MNHN" country="Laos" pageId="10" pageNumber="263" specimenCode="IM-2012-27219" specimenCount="1">
<date id="7709FF94FF9F7320FEF6F903FD9D259F" box="[341,559,1721,1748]" pageId="10" pageNumber="263" value="2004-12-02">
<collectingDate id="674D067CFF9F7320FEF6F903FD9D259F" box="[341,559,1721,1748]" pageId="10" pageNumber="263" value="2004-12-02">02 December 2004</collectingDate>
</date>
.,
<collectionCode id="65A64191FF9F7320FD9EF903FD21259F" box="[573,659,1722,1748]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FD3AF903FCE1259F" box="[665,851,1722,1748]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27219</specimenCode>
</materialsCitation>
/1;
<materialsCitation id="B3DFD309FF9F7320FCD3F900FE7E2457" ID-GBIF-Occurrence="4039909328" collectionCode="V" collectorName="Abdou, A. &amp; Muratov, I." country="Laos" elevation="614" latitude="21.432083" location="Nam Bun" longLatPrecision="1" longitude="101.98115" municipality="Nam Leng" pageId="10" pageNumber="263" specimenCount="1" stateProvince="Phongsali">
<collectingRegion id="C17317B6FF9F7320FCD3F900FBE32598" box="[880,1105,1721,1747]" country="Laos" name="Phongsali" pageId="10" pageNumber="263">Phongsaly Province</collectingRegion>
, NNE of
<collectingMunicipality id="E36C432EFF9F7320FB19F903FA83259F" box="[1210,1329,1722,1748]" pageId="10" pageNumber="263">Nam Leng</collectingMunicipality>
and
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confluence, 614 m, old forest near ravine, clay, shale,
<geoCoordinate id="6683BF93FF9F7320FC8FF966FC0F25B1" box="[812,957,1759,1786]" degrees="21" direction="north" minutes="25.925" orientation="latitude" pageId="10" pageNumber="263" precision="1" value="21.432083">21°25.925N</geoCoordinate>
,
<geoCoordinate id="6683BF93FF9F7320FC6AF966FBD125B1" box="[969,1123,1759,1786]" degrees="101" direction="east" minutes="58.869" orientation="longitude" pageId="10" pageNumber="263" precision="1" value="101.98115">101°58.869E</geoCoordinate>
(locality code: 27L05), leg.
<collectorName id="AE42BC82FF9F7320FF34F8B8FEBC2457" box="[151,270,1793,1820]" pageId="10" pageNumber="263">Abdou, A.</collectorName>
&amp;
<collectorName id="AE42BC82FF9F7320FE95F8BBFE062457" box="[310,436,1794,1820]" pageId="10" pageNumber="263">Muratov, I.</collectorName>
<collectionCode id="65A64191FF9F7320FE17F8BBFE7E2457" box="[436,460,1794,1820]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="263" type="Museum">V</collectionCode>
</materialsCitation>
.,
<materialsCitation id="B3DFD309FF9F7320FE7DF8BBFC002457" ID-GBIF-Occurrence="4039909335" box="[478,946,1793,1820]" collectingDate="2005-03-18" collectionCode="MNHN" country="Laos" pageId="10" pageNumber="263" specimenCode="IM-2012-27218" specimenCount="1">
<date id="7709FF94FF9F7320FE7DF8BBFD232457" box="[478,657,1793,1820]" pageId="10" pageNumber="263" value="2005-03-18">
<collectingDate id="674D067CFF9F7320FE7DF8BBFD232457" box="[478,657,1793,1820]" pageId="10" pageNumber="263" value="2005-03-18">18 March 2005</collectingDate>
</date>
,
<collectionCode id="65A64191FF9F7320FD3DF8BBFD462457" box="[670,756,1794,1820]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FD5EF8BBFC002457" box="[765,946,1794,1820]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27218</specimenCode>
</materialsCitation>
/3 (
<figureCitation id="9B8CC5D1FF9F7320FC7BF8B8FBFB2457" box="[984,1097,1793,1820]" captionStart="FIGURE 8" captionStartId="11.[152,255,1975,1999]" captionTargetBox="[151,1436,885,1952]" captionTargetId="figure-310@11.[151,1436,885,1952]" captionTargetPageId="11" captionText="FIGURE 8. Shells of Pincerna costulosus (Bavay &amp; Dautzenberg, 1912).AD: MNHN-IM-2000-31786 (syntype); EH:Phong Tho, MNHN-IM-2012-27043 (probable syntype); IL: Northern Laos, MNHN-IM-2012-27218." figureDoi="http://doi.org/10.5281/zenodo.7687434" httpUri="https://zenodo.org/record/7687434/files/figure.png" pageId="10" pageNumber="263">Fig. 8IL</figureCitation>
);
<materialsCitation id="B3DFD309FF9F7320FBC3F8B8FD48242F" ID-GBIF-Occurrence="4039909302" collectionCode="V" collectorName="Abdou, A. &amp; Muratov, I." country="Laos" elevation="1343" latitude="21.66105" location="Boun" longLatPrecision="1" longitude="102.08739" pageId="10" pageNumber="263" specimenCount="1" stateProvince="Phongsali">
<collectingRegion id="C17317B6FF9F7320FBC3F8B8FAF62450" box="[1120,1348,1793,1819]" country="Laos" name="Phongsali" pageId="10" pageNumber="263">Phongsaly Province</collectingRegion>
,
<locationDeviation id="12BAC076FF9F7320FAF1F8BBFEBF2474" location="Phongsaly" pageId="10" pageNumber="263">SW of Phongsaly</locationDeviation>
, near the road to
<location id="06688F8FFF9F7320FE6CF89FFDBE240B" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FE6CF89FFDBE240B" box="[463,524,1830,1856]" country="Laos" latitude="21.66105" longLatPrecision="1" longitude="102.08739" name="Boun" pageId="10" pageNumber="263" stateProvince="Phongsali">Boun</location>
Neua, 1343 m,
<geoCoordinate id="6683BF93FF9F7320FD1CF89EFCE32409" box="[703,849,1831,1858]" degrees="21" direction="north" minutes="39.663" orientation="latitude" pageId="10" pageNumber="263" precision="1" value="21.66105">21°39.663N</geoCoordinate>
,
<geoCoordinate id="6683BF93FF9F7320FCFFF89EFC472409" box="[860,1013,1831,1858]" degrees="102" direction="east" minutes="05.243" orientation="longitude" pageId="10" pageNumber="263" precision="1" value="102.08739">102°05.243E</geoCoordinate>
(locality code: 12L05), dry old forest in ravine, clay, shale, leg.
<collectorName id="AE42BC82FF9F7320FE66F8F0FD8E242F" box="[453,572,1865,1892]" pageId="10" pageNumber="263">Abdou, A.</collectorName>
&amp;
<collectorName id="AE42BC82FF9F7320FDC7F8F3FD50242F" box="[612,738,1866,1892]" pageId="10" pageNumber="263">Muratov, I.</collectorName>
<collectionCode id="65A64191FF9F7320FD41F8F3FD48242F" box="[738,762,1866,1892]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="263" type="Museum">V</collectionCode>
</materialsCitation>
.,
<materialsCitation id="B3DFD309FF9F7320FCAFF8F3FB572428" ID-GBIF-Occurrence="4039909361" box="[780,1253,1865,1892]" collectingDate="2005-03-10" collectionCode="MNHN" country="Laos" pageId="10" pageNumber="263" specimenCode="IM-2012-27216" specimenCount="1">
<date id="7709FF94FF9F7320FCAFF8F3FC0D242F" box="[780,959,1865,1892]" pageId="10" pageNumber="263" value="2005-03-10">
<collectingDate id="674D067CFF9F7320FCAFF8F3FC0D242F" box="[780,959,1865,1892]" pageId="10" pageNumber="263" value="2005-03-10">10 March 2005</collectingDate>
</date>
,
<collectionCode id="65A64191FF9F7320FC6FF8F3FB90242F" box="[972,1058,1866,1892]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FB88F8F0FB572428" box="[1067,1253,1865,1891]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27216</specimenCode>
</materialsCitation>
/1;
<materialsCitation id="B3DFD309FF9F7320FAA6F8F0FE94249B" ID-GBIF-Occurrence="4039909310" collectingDate="2017-02-23" collectionCode="JG" collectorName="Grego, J." country="Laos" elevation="1062" latitude="19.38775" location="6 km SW of Bank Knong" longLatPrecision="1" longitude="102.97535" pageId="10" pageNumber="263" specimenCode="JG29" specimenCount="1" stateProvince="Xiangkhoang Province">
<collectingRegion id="C17317B6FF9F7320FAA6F8F0FF4F24CC" inferredBySuffix="true" pageId="10" pageNumber="263">Xiangkhoang Province</collectingRegion>
, caverns among boulders at large sinkhole below small settlements (5 houses about
<location id="06688F8FFF9F7320FB6AF8D4FF5524E7" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FB6AF8D4FF5524E7" country="Laos" latitude="19.38775" longLatPrecision="1" longitude="102.97535" name="6 km SW of Bank Knong" pageId="10" pageNumber="263" stateProvince="Xiangkhoang Province">6 km SW of Bank Knong</location>
), 1062 m a.s.l.,
<geoCoordinate id="6683BF93FF9F7320FE02F82AFD8024E5" box="[417,562,1939,1966]" degrees="19" direction="north" minutes="23.265" orientation="latitude" pageId="10" pageNumber="263" precision="1" value="19.38775">19°23.265N</geoCoordinate>
,
<geoCoordinate id="6683BF93FF9F7320FD9CF82AFD6B24E5" box="[575,729,1939,1966]" degrees="102" direction="east" minutes="58.521" orientation="longitude" pageId="10" pageNumber="263" precision="1" value="102.97535">102°58.521E</geoCoordinate>
(locality code:
<specimenCode id="5311712FFF9F7320FC2DF82BFC7F24E7" box="[910,973,1938,1964]" collectionCode="JG" pageId="10" pageNumber="263">JG29</specimenCode>
), leg.
<collectorName id="AE42BC82FF9F7320FBB7F82BFBCC24E7" box="[1044,1150,1938,1964]" pageId="10" pageNumber="263">Grego, J.</collectorName>
,
<date id="7709FF94FF9F7320FB2FF82BFAE824E7" box="[1164,1370,1937,1964]" pageId="10" pageNumber="263" value="2017-02-23">
<collectingDate id="674D067CFF9F7320FB2FF82BFAE824E7" box="[1164,1370,1937,1964]" pageId="10" pageNumber="263" value="2017-02-23">23 February 2017</collectingDate>
</date>
, JG/1 (
<figureCitation id="9B8CC5D1FF9F7320FF3CF80CFEAF249B" box="[159,285,1973,2000]" captionStart="FIGURE 9" captionStartId="12.[152,255,1611,1635]" captionTargetBox="[151,1436,181,1587]" captionTargetId="figure-31@12.[151,1436,181,1587]" captionTargetPageId="12" captionText="FIGURE 9. Shells of Pincerna costulosus(Bavay &amp; Dautzenberg, 1912).AD: Laos, Phongsaly Province, JG29; EH:Holotype of Pincerna vallis Z.-Y. Chen &amp; Wu, 2020; IM: Pincerna cf. costulosus, Guangxi, China, 2009/104. Photos: B. Páll-Gergely (AD, IH), taken from the original description (EH)." figureDoi="http://doi.org/10.5281/zenodo.7687436" httpUri="https://zenodo.org/record/7687436/files/figure.png" pageId="10" pageNumber="263">Fig. 9AD</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="B3DFD309FF9F7320FE93F80FFB3F249B" ID-GBIF-Occurrence="4039909311" box="[304,1165,1973,2000]" collectionCode="MNHN" collectorName="Letellier" country="Vietnam" location="Lao Kay" municipality="Tonkin" pageId="10" pageNumber="263" specimenCode="IM-2012-27214" specimenCount="1">
<collectingCountry id="7BA099C4FF9F7320FE93F80FFE04249B" box="[304,438,1974,2000]" name="Vietnam" pageId="10" pageNumber="263">VIETNAM</collectingCountry>
:
<collectingMunicipality id="E36C432EFF9F7320FE62F80CFDA12484" box="[449,531,1973,1999]" pageId="10" pageNumber="263">Tonkin</collectingMunicipality>
,
<location id="06688F8FFF9F7320FDBCF80FFD30249B" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FDBCF80FFD30249B" box="[543,642,1974,2000]" country="Vietnam" municipality="Tonkin" name="Lao Kay" pageId="10" pageNumber="263">Lao Kay</location>
, coll.
<collectorName id="AE42BC82FF9F7320FD65F80CFC952484" box="[710,807,1973,1999]" pageId="10" pageNumber="263">Letellier</collectorName>
, 1949,
<collectionCode id="65A64191FF9F7320FCDAF80FFC7D249B" box="[889,975,1974,2000]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FC75F80FFB3F249B" box="[982,1165,1974,2000]" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012-27214</specimenCode>
</materialsCitation>
/
<materialsCitation id="B3DFD309FF9F7320FB35F80FFEC024BF" ID-GBIF-Occurrence="4039909353" collectionCode="MNHN" country="Vietnam" pageId="10" pageNumber="263" specimenCode="IM-2012- 27035" specimenCount="1">
1 (ex
<collectionCode id="65A64191FF9F7320FB71F80FFA9A249B" box="[1234,1320,1974,2000]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="263">MNHN</collectionCode>
<specimenCode id="5311712FFF9F7320FA8CF80FFF5224BF" collectionCode="IM-" pageId="10" pageNumber="263">IM-2012- 27035</specimenCode>
,
<taxonomicName id="C4B7A2D7FF9F7320FF49F860FED424B8" authorityName="Bavay &amp; Dautzenberg" authorityYear="1900" box="[234,358,2009,2035]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF9F7320FF49F860FED424B8" box="[234,358,2009,2035]" italics="true" pageId="10" pageNumber="263">vanbuensis</emphasis>
</taxonomicName>
)
</materialsCitation>
;
<materialsCitation id="B3DFD309FF9F7320FEDDF863FA8324BF" ID-GBIF-Occurrence="4039909334" box="[382,1329,2009,2036]" collectionCode="RBINS" collectorName="Messager" country="Vietnam" location="Lao Kay" pageId="10" pageNumber="263" specimenCode="RBINS/2" specimenCount="1">
<location id="06688F8FFF9F7320FEDDF863FE5224BF" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9F7320FEDDF863FE5224BF" box="[382,480,2010,2036]" country="Vietnam" name="Lao Kay" pageId="10" pageNumber="263">Lao Kay</location>
, leg.
<collectorName id="AE42BC82FF9F7320FDBAF863FD3624BF" box="[537,644,2010,2036]" pageId="10" pageNumber="263">Messager</collectorName>
no. 26,
<specimenCode id="5311712FFF9F7320FD7EF863FCF724BF" box="[733,837,2010,2036]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="10" pageNumber="263" type="Museum">RBINS/2</specimenCode>
(syntype mixed sample with
<taxonomicName id="C4B7A2D7FF9F7320FB28F863FA9424B8" authorityName="Bavay &amp; Dautzenberg" authorityYear="1900" box="[1163,1318,2009,2036]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="263" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF9F7320FB28F863FA9424B8" box="[1163,1318,2009,2036]" italics="true" pageId="10" pageNumber="263">A. vanbuensis</emphasis>
</taxonomicName>
)
</materialsCitation>
.
<materialsCitation id="B3DFD309FF9F7321FA9EF863FAD0239E" ID-GBIF-Occurrence="4039909362" collectingDate="2009-10-21" collectorName="Hunyadi, A." country="China" county="Hechi Shi" elevation="460" lastPageId="11" lastPageNumber="264" latitude="24.37845" location="Paoli Xiang" longLatPrecision="1" longitude="107.06375" municipality="Fengshan Xian" pageId="10" pageNumber="263" specimenCount="1">
<collectingCountry id="7BA099C4FF9F7320FA9EF863FA2B24BF" box="[1341,1433,2010,2036]" name="China" pageId="10" pageNumber="263">CHINA</collectingCountry>
,
<collectingCounty id="EA69A1D8FF9E7321FF34FF2EFEBB23FA" box="[151,265,151,177]" pageId="11" pageNumber="264">Hechi Shi</collectingCounty>
,
<collectingMunicipality id="E36C432EFF9E7321FEB6FF2EFE7223FA" box="[277,448,151,177]" pageId="11" pageNumber="264">Fengshan Xian</collectingMunicipality>
,
<location id="06688F8FFF9E7321FE6FFF2EFDE123FA" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9E7321FE6FFF2EFDE123FA" box="[460,595,151,177]" country="China" county="Hechi Shi" latitude="24.37845" longLatPrecision="1" longitude="107.06375" municipality="Fengshan Xian" name="Paoli Xiang" pageId="11" pageNumber="264">Paoli Xiang</location>
, southeastern edge of
<location id="06688F8FFF9E7321FCEEFF2EFC3B23FA" LSID="urn:lsid:plazi:treatment:8B1E6842FF9F7327FF34FB36FED422C2:06688F8FFF9E7321FCEEFF2EFC3B23FA" box="[845,905,151,177]" country="China" county="Hechi Shi" latitude="24.37845" longLatPrecision="1" longitude="107.06375" municipality="Fengshan Xian" name="Paoli" pageId="11" pageNumber="264">Paoli</location>
, cave, 460 m a.s.l.,
<geoCoordinate id="6683BF93FF9E7321FBC5FF21FB4523F8" box="[1126,1271,152,179]" degrees="24" direction="north" minutes="22.707" orientation="latitude" pageId="11" pageNumber="264" precision="1" value="24.37845">24°22.707N</geoCoordinate>
,
<geoCoordinate id="6683BF93FF9E7321FAA0FF21FA2E23F8" box="[1283,1436,152,179]" degrees="107" direction="east" minutes="03.825" orientation="longitude" pageId="11" pageNumber="264" precision="1" value="107.06375">107°03.825E</geoCoordinate>
(locality code: 2009/104), leg.
<collectorName id="AE42BC82FF9E7321FE53FF02FDCB239E" box="[496,633,187,213]" pageId="11" pageNumber="264">Hunyadi, A.</collectorName>
<date id="7709FF94FF9E7321FD23FF02FC8D239E" box="[640,831,187,213]" pageId="11" pageNumber="264" value="2009-10-21">
<collectingDate id="674D067CFF9E7321FD23FF02FC8D239E" box="[640,831,187,213]" pageId="11" pageNumber="264" value="2009-10-21">21 October 2009</collectingDate>
</date>
, HA/1 (identification questionable, see remarks)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9E7321FF64FF67FB052256" pageId="11" pageNumber="264" type="diagnosis">
<paragraph id="0308D954FF9E7321FF64FF67FB052256" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FF67FEF023B3" bold="true" box="[199,322,222,248]" pageId="11" pageNumber="264">Diagnosis.</emphasis>
A small to medium-sized alycaeid species with a shell that is slightly higher than wide, R1 strongly, regularly ribbed, R2 extremely short, R3 or normal length, smooth, with low, central swelling.
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9E7321FF64FE9FFE762186" pageId="11" pageNumber="264" type="description">
<paragraph id="0308D954FF9E7321FF64FE9FFAED2152" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FE9FFEEB220B" bold="true" box="[199,345,294,320]" pageId="11" pageNumber="264">Description.</emphasis>
Shell higher than wide, yellowish or off-white when fresh; protoconch finely granulate, rather matte, consisting of 1.5 or lightly less whorls; R1 glossy, of 2.5 whorls, with regular, rather strong ribs and extremely fine spiral striation; R2 extremely short, consisting of ca. 6 lighter stripes (=breathing tunnels) only; breathing tunnels not or only very slightly elevated from the shell surface; constriction between R2 and R3 rather shallow; R3 or normal length, with low swelling; R3 nearly smooth with some widely-spaced thread-like ribs; aperture rounded; outer peristome strongly expanded and slightly reflected; inner peristome thickened, protruding; boundary between inner and outer peristomes clearly visible; umbilicus entirely or nearly entirely closed by reflected peristome.
</paragraph>
<paragraph id="0308D954FF9E7321FF64FD9AFBE62176" blockId="11.[151,1437,151,861]" box="[199,1108,546,573]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FD9AFE562177" bold="true" box="[199,484,546,573]" pageId="11" pageNumber="264">Measurements (in mm).</emphasis>
D = 3.44.8, H = 3.94.9 (multiple populations, n = 7).
</paragraph>
<paragraph id="0308D954FF9E7321FF64FDFFFA2F21CE" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FDFFFEEA212B" bold="true" box="[199,344,582,608]" pageId="11" pageNumber="264">Operculum:</emphasis>
Thefragmentaryoperculumofasinglespecimen(MNHN-IM-2012-27214)wasexamined.Operculum semitransparent, thin, horny, concave, both outer and inner surface multispiral, no nipple found on the inner side.
</paragraph>
<paragraph id="0308D954FF9E7321FF64FD37FE762186" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FD37FE6D21E3" bold="true" box="[199,479,654,680]" pageId="11" pageNumber="264">Variation in specimens.</emphasis>
The typical specimens from northernmost
<collectingCountry id="7BA099C4FF9E7321FC60FD36FB9021E2" box="[963,1058,655,681]" name="Vietnam" pageId="11" pageNumber="264">Vietnam</collectingCountry>
are slightly larger than the newly collected in northern
<collectingCountry id="7BA099C4FF9E7321FE25FD0AFE722186" box="[390,448,691,717]" name="Laos" pageId="11" pageNumber="264">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9E7321FF64FD6FFB0C205E" pageId="11" pageNumber="264" type="diagnosis">
<paragraph id="0308D954FF9E7321FF64FD6FFB0C205E" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FD6FFE7C21BB" bold="true" box="[199,462,726,752]" pageId="11" pageNumber="264">Differential diagnosis.</emphasis>
This species is easily recognisable based on the combination of the shell shape (higher than wide), regularly ribbed R1, and very short R2. See also under
<taxonomicName id="C4B7A2D7FF9E7321FC25FD42FBCA205E" authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[902,1144,763,789]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="264" phylum="Mollusca" rank="species" species="acroptychia" status="sp. nov.">
<emphasis id="31C30546FF9E7321FC25FD42FBCA205E" box="[902,1144,763,789]" italics="true" pageId="11" pageNumber="264">Pincerna acroptychia</emphasis>
</taxonomicName>
<emphasis id="31C30546FF9E7321FBDCFD42FB0C205E" bold="true" box="[1151,1214,763,789]" pageId="11" pageNumber="264">
<taxonomicNameLabel id="2AF0B83DFF9E7321FBDCFD42FB0C205E" box="[1151,1214,763,789]" pageId="11" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF9E7321FF64FCA7FC632016" pageId="11" pageNumber="264" type="distribution">
<paragraph id="0308D954FF9E7321FF64FCA7FC632016" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF64FCA7FED32073" bold="true" box="[199,353,798,824]" pageId="11" pageNumber="264">Distribution.</emphasis>
This species is known from northern
<collectingCountry id="7BA099C4FF9E7321FC82FCA6FCE52072" box="[801,855,799,825]" name="Laos" pageId="11" pageNumber="264">Laos</collectingCountry>
(
<collectingRegion id="C17317B6FF9E7321FCC9FCA6FC522072" box="[874,992,799,825]" country="Laos" name="Phongsali" pageId="11" pageNumber="264">Phongsaly</collectingRegion>
and Xiangkhoang provinces), and the northernmost part of
<collectingCountry id="7BA099C4FF9E7321FE27FCFAFE512016" box="[388,483,835,861]" name="Vietnam" pageId="11" pageNumber="264">Vietnam</collectingCountry>
(
<collectingRegion id="C17317B6FF9E7321FE52FCFAFDE82016" box="[497,602,835,861]" country="Vietnam" name="Lai Chau" pageId="11" pageNumber="264">Lai Chau</collectingRegion>
and Lao Kay provinces) (
<figureCitation id="9B8CC5D1FF9E7321FCDDFCFAFC712016" box="[894,963,835,861]" captionStart="FIGURE 6" captionStartId="9.[152,255,1367,1391]" captionTargetBox="[151,1436,181,1343]" captionTargetId="figure-15@9.[151,1436,181,1343]" captionTargetPageId="9" captionText="FIGURE 6. Distribution of Pincerna Preston, 1907 species. Circles: Pincerna costulosus (Bavay &amp; Dautzenberg, 1912) (yellow circle indicates the type locality of Pincerna vallis Z.-Y. Chen &amp; M. Wu, 2020); Yellow triangle: Pincerna acroptychia n. sp.; Green rhomb: Pincerna yaanensis Z.-Y. Chen, 2022. The map was made using Google Earth Pro." figureDoi="http://doi.org/10.5281/zenodo.7687430" httpUri="https://zenodo.org/record/7687430/files/figure.png" pageId="11" pageNumber="264">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="57C889DCFF9E7321FF3BF80EFBD524BF" ID-DOI="http://doi.org/10.5281/zenodo.7687434" ID-Zenodo-Dep="7687434" httpUri="https://zenodo.org/record/7687434/files/figure.png" pageId="11" pageNumber="264" startId="11.[152,255,1975,1999]" targetBox="[151,1436,885,1952]" targetPageId="11">
<paragraph id="0308D954FF9E7321FF3BF80EFBD524BF" blockId="11.[151,1436,1975,2036]" pageId="11" pageNumber="264">
<emphasis id="31C30546FF9E7321FF3BF80EFEA42484" bold="true" box="[152,278,1975,1999]" pageId="11" pageNumber="264">FIGURE 8.</emphasis>
Shells of
<taxonomicName id="C4B7A2D7FF9E7321FED4F801FCC5249B" authority="(Bavay &amp; Dautzenberg, 1912)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[375,887,1976,2000]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="264" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF9E7321FED4F801FDF1249B" box="[375,579,1976,2000]" italics="true" pageId="11" pageNumber="264">Pincerna costulosus</emphasis>
(
<bibRefCitation id="6726A4A5FF9E7321FDEDF801FCC2249B" author="Bavay, A. &amp; Dautzenberg, P." box="[590,880,1976,2000]" pageId="11" pageNumber="264" pagination="1 - 54" refId="ref13737" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de Coquilles nouvelles de I'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54. [https: // www. biodiversitylibrary. org / page / 16358455]" type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
)
</taxonomicName>
.AD: MNHN-IM-2000-31786 (syntype); EH: Phong Tho, MNHN-IM-2012-27043 (probable syntype); IL: Northern Laos, MNHN-IM-2012-27218.
</paragraph>
</caption>
<caption id="57C889DCFF997326FF3BF9F2FD7525E0" ID-DOI="http://doi.org/10.5281/zenodo.7687436" ID-Zenodo-Dep="7687436" httpUri="https://zenodo.org/record/7687436/files/figure.png" pageId="12" pageNumber="265" startId="12.[152,255,1611,1635]" targetBox="[151,1436,181,1587]" targetPageId="12">
<paragraph id="0308D954FF997326FF3BF9F2FD7525E0" blockId="12.[151,1436,1611,1707]" pageId="12" pageNumber="265">
<emphasis id="31C30546FF997326FF3BF9F2FEA42528" bold="true" box="[152,278,1611,1635]" pageId="12" pageNumber="265">FIGURE 9.</emphasis>
Shells of
<taxonomicName id="C4B7A2D7FF997326FED5F9F5FCC62528" authority="(Bavay &amp; Dautzenberg, 1912)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[374,884,1611,1635]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FED5F9F5FDF32528" box="[374,577,1611,1635]" italics="true" pageId="12" pageNumber="265">Pincerna costulosus</emphasis>
(
<bibRefCitation id="6726A4A5FF997326FDE8F9F2FCDE2528" author="Bavay, A. &amp; Dautzenberg, P." box="[587,876,1611,1635]" pageId="12" pageNumber="265" pagination="1 - 54" refId="ref13737" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de Coquilles nouvelles de I'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54. [https: // www. biodiversitylibrary. org / page / 16358455]" type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
)
</taxonomicName>
.AD: Laos, Phongsaly Province, JG29; EH: Holotype of
<taxonomicName id="C4B7A2D7FF997326FF10F9C9FEFC25CC" authorityName="Z. -Y. Chen &amp; M. Wu" authorityYear="2020" box="[179,334,1647,1671]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="31C30546FF997326FF10F9C9FEFC25CC" box="[179,334,1647,1671]" italics="true" pageId="12" pageNumber="265">Pincerna vallis</emphasis>
</taxonomicName>
Z.-Y.
<bibRefCitation id="6726A4A5FF997326FE33F9D6FDFC25CC" author="Chen, Z. - Y. &amp; Wu, M." box="[400,590,1647,1671]" pageId="12" pageNumber="265" pagination="37 - 46" refId="ref14272" refString="Chen, Z. - Y. &amp; Wu, M. (2020) Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae). ZooKeys, 935, 37 - 46. https: // doi. org / 10.3897 / zookeys. 935.51414" type="journal article" year="2020">Chen &amp; Wu, 2020</bibRefCitation>
; IM:
<taxonomicName id="C4B7A2D7FF997326FD31F9C9FC3325CC" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[658,897,1647,1671]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" isUncertain="true" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FD31F9C9FD5D25CC" box="[658,751,1648,1671]" italics="true" pageId="12" pageNumber="265">Pincerna</emphasis>
cf.
<emphasis id="31C30546FF997326FCB4F9D6FC3325CC" box="[791,897,1647,1671]" italics="true" pageId="12" pageNumber="265">costulosus</emphasis>
</taxonomicName>
, Guangxi, China, 2009/104. Photos: B. Páll-Gergely (AD, IH), taken from the original description (EH).
</paragraph>
</caption>
<subSubSection id="4BAD8ADFFF997327FF64F960FED422C2" lastPageId="13" lastPageNumber="266" pageId="12" pageNumber="265" type="discussion">
<paragraph id="0308D954FF997326FF64F960FBD1242B" blockId="12.[151,1437,1753,2032]" pageId="12" pageNumber="265">
<emphasis id="31C30546FF997326FF64F960FE8925B8" bold="true" box="[199,315,1753,1779]" pageId="12" pageNumber="265">Remarks.</emphasis>
Some Indian
<taxonomicName id="C4B7A2D7FF997326FE75F960FD2125B8" box="[470,659,1753,1779]" pageId="12" pageNumber="265">
<emphasis id="31C30546FF997326FE75F960FD8925B8" box="[470,571,1753,1779]" italics="true" pageId="12" pageNumber="265">Cycloryx</emphasis>
species
</taxonomicName>
(e.g.
<taxonomicName id="C4B7A2D7FF997326FD71F963FB4E25B8" authority="(Godwin-Austen, 1914)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[722,1276,1753,1780]" class="Gastropoda" family="Alycaeidae" genus="Cycloryx" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="subSpecies" species="graphicus" subSpecies="dihingensis">
<emphasis id="31C30546FF997326FD71F963FC5A25B8" box="[722,1000,1753,1780]" italics="true" pageId="12" pageNumber="265">C. graphicus dihingensis</emphasis>
(Godwin-Austen, 1914)
</taxonomicName>
,
<taxonomicName id="C4B7A2D7FF997326FAA4F963FE19245C" authority="(Godwin-Austen, 1914)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Cycloryx" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="31C30546FF997326FAA4F963FA2F25B8" box="[1287,1437,1753,1780]" italics="true" pageId="12" pageNumber="265">C. thompsoni</emphasis>
(Godwin-Austen, 1914)
</taxonomicName>
,
<taxonomicName id="C4B7A2D7FF997326FE1EF947FC21245C" authority="(Godwin-Austen, 1914))" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[445,915,1789,1816]" class="Gastropoda" family="Alycaeidae" genus="Cycloryx" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="burrailensis">
<emphasis id="31C30546FF997326FE1EF947FDD9245C" box="[445,619,1789,1816]" italics="true" pageId="12" pageNumber="265">C. burrailensis</emphasis>
(Godwin-Austen, 1914))
</taxonomicName>
are so similar to
<taxonomicName id="C4B7A2D7FF997326FBD0F947FABB245C" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[1139,1289,1789,1815]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FBD0F947FABB245C" box="[1139,1289,1789,1815]" italics="true" pageId="12" pageNumber="265">P. costulosus</emphasis>
</taxonomicName>
, that it was challenging to find any notable differences by comparison under the microscope. This may indicate that Northern Lao, northern Vietnamese and Chinese
<taxonomicName id="C4B7A2D7FF997326FDF3F8FFFCBD2414" box="[592,783,1861,1887]" pageId="12" pageNumber="265">
<emphasis id="31C30546FF997326FDF3F8FFFD052414" box="[592,695,1862,1887]" italics="true" pageId="12" pageNumber="265">Pincerna</emphasis>
species
</taxonomicName>
belong to the genus
<taxonomicName id="C4B7A2D7FF997326FC54F8FCFBEE2414" box="[1015,1116,1861,1887]" class="Gastropoda" family="Alycaeidae" genus="Cycloryx" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="genus">
<emphasis id="31C30546FF997326FC54F8FCFBEE2414" box="[1015,1116,1861,1887]" italics="true" pageId="12" pageNumber="265">Cycloryx</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0308D954FF997327FF64F8D3FC3A23B2" blockId="12.[151,1437,1753,2032]" lastBlockId="13.[151,1437,151,393]" lastPageId="13" lastPageNumber="266" pageId="12" pageNumber="265">
<taxonomicName id="C4B7A2D7FF997326FF64F8D3FEC724C8" authorityName="Z. -Y. Chen &amp; M. Wu" authorityYear="2020" box="[199,373,1897,1923]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="31C30546FF997326FF64F8D3FEC724C8" box="[199,373,1897,1923]" italics="true" pageId="12" pageNumber="265">Pincerna vallis</emphasis>
</taxonomicName>
(
<figureCitation id="9B8CC5D1FF997326FE2BF8D0FDB624C8" box="[392,516,1897,1923]" captionStart="FIGURE 9" captionStartId="12.[152,255,1611,1635]" captionTargetBox="[151,1436,181,1587]" captionTargetId="figure-31@12.[151,1436,181,1587]" captionTargetPageId="12" captionText="FIGURE 9. Shells of Pincerna costulosus(Bavay &amp; Dautzenberg, 1912).AD: Laos, Phongsaly Province, JG29; EH:Holotype of Pincerna vallis Z.-Y. Chen &amp; Wu, 2020; IM: Pincerna cf. costulosus, Guangxi, China, 2009/104. Photos: B. Páll-Gergely (AD, IH), taken from the original description (EH)." figureDoi="http://doi.org/10.5281/zenodo.7687436" httpUri="https://zenodo.org/record/7687436/files/figure.png" pageId="12" pageNumber="265">Fig. 9EH</figureCitation>
), described from the Chaibuxi National Forest Park,
<collectingRegion id="C17317B6FF997326FBD2F8D0FA9424C8" box="[1137,1318,1897,1923]" country="China" name="Hubei" pageId="12" pageNumber="265">Hubei Province</collectingRegion>
,
<collectingCountry id="7BA099C4FF997326FA96F8D0FAC924C8" box="[1333,1403,1897,1923]" name="China" pageId="12" pageNumber="265">China</collectingCountry>
, is treated here as a junior synonym of
<taxonomicName id="C4B7A2D7FF997326FD92F837FD7124EC" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[561,707,1933,1959]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FD92F837FD7124EC" box="[561,707,1933,1959]" italics="true" pageId="12" pageNumber="265">P. costulosus</emphasis>
</taxonomicName>
. In its original description,
<bibRefCitation id="6726A4A5FF997326FC5DF834FB5E24EC" author="Chen, Z. - Y. &amp; Wu, M." box="[1022,1260,1933,1960]" pageId="12" pageNumber="265" pagination="37 - 46" refId="ref14272" refString="Chen, Z. - Y. &amp; Wu, M. (2020) Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae). ZooKeys, 935, 37 - 46. https: // doi. org / 10.3897 / zookeys. 935.51414" type="journal article" year="2020">Chen and Wu (2020)</bibRefCitation>
mentioned that
<taxonomicName id="C4B7A2D7FF997326FF34F80BFF432480" authorityName="Z. -Y. Chen &amp; M. Wu" authorityYear="2020" box="[151,241,1969,1995]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="31C30546FF997326FF34F80BFF432480" box="[151,241,1969,1995]" italics="true" pageId="12" pageNumber="265">P. vallis</emphasis>
</taxonomicName>
had a ribbed R3, which is smooth in
<taxonomicName id="C4B7A2D7FF997326FD02F80BFC812480" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[673,819,1969,1995]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FD02F80BFC812480" box="[673,819,1969,1995]" italics="true" pageId="12" pageNumber="265">P. costulosus</emphasis>
</taxonomicName>
. However, examination of several
<taxonomicName id="C4B7A2D7FF997326FB61F80BFAE62480" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[1218,1364,1969,1995]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FB61F80BFAE62480" box="[1218,1364,1969,1995]" italics="true" pageId="12" pageNumber="265">P. costulosus</emphasis>
</taxonomicName>
shells (including
<typeStatus id="DC0C67F6FF997326FEB2F86CFEC724A4" box="[273,373,2005,2031]" pageId="12" pageNumber="265" type="syntype">syntypes</typeStatus>
) revealed that
<taxonomicName id="C4B7A2D7FF997326FDB8F86FFD1E24A4" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[539,684,2005,2031]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="265" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF997326FDB8F86FFD1E24A4" box="[539,684,2005,2031]" italics="true" pageId="12" pageNumber="265">P. costulosus</emphasis>
</taxonomicName>
shows some variability in this respect. The range of other northern Vietnamese terrestrial operculate species also reach the Chinese
<collectingRegion id="C17317B6FF987327FCC6FF2EFBA423FA" box="[869,1046,151,177]" country="China" name="Hubei" pageId="13" pageNumber="266">Hubei Province</collectingRegion>
, such as
<taxonomicName id="C4B7A2D7FF987327FBDAFF2EFED8239E" authority="Mollendorff, 1885" authorityName="Mollendorff" authorityYear="1885" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="amoenus">
<emphasis id="31C30546FF987327FBDAFF2EFA2F23FB" box="[1145,1437,151,177]" italics="true" pageId="13" pageNumber="266">Pseudopomatias amoenus</emphasis>
<bibRefCitation id="6726A4A5FF987327FF34FF02FED8239E" author="Mollendorff, O. F. von" box="[151,362,186,213]" pageId="13" pageNumber="266" pagination="161 - 170" refId="ref15193" refString="Mollendorff, O. F. von (1885) Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 17 (11 - 12), 161 - 170." type="journal article" year="1885">Möllendorff, 1885</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C4B7A2D7FF987327FED4FF02FD05239E" authority="Pall-Gergely, 2015" authorityName="Pall-Gergely" authorityYear="2015" box="[375,695,186,213]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="nitens">
<emphasis id="31C30546FF987327FED4FF02FE65239F" box="[375,471,187,212]" italics="true" pageId="13" pageNumber="266">P. nitens</emphasis>
<bibRefCitation id="6726A4A5FF987327FE43FF02FD05239E" author="Pall-Gergely, B. &amp; Feher, Z. &amp; Hunyadi, A. &amp; Asami, T." box="[480,695,186,213]" pageId="13" pageNumber="266" pagination="1 - 49" refId="ref15492" refString="Pall-Gergely, B., Feher, Z., Hunyadi, A. &amp; Asami, T. (2015) Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa, 3937 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 3937.1.1" type="journal article" year="2015">Páll-Gergely, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C4B7A2D7FF987327FD67FF02FB30239E" authority="(Mollendorff, 1886)" baseAuthorityName="Mollendorff" baseAuthorityYear="1886" box="[708,1154,187,213]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="cristatus">
<emphasis id="31C30546FF987327FD67FF02FC24239F" box="[708,918,187,213]" italics="true" pageId="13" pageNumber="266">Dicharax cristatus</emphasis>
(
<bibRefCitation id="6726A4A5FF987327FC05FF02FBCB239E" author="Mollendorff, O. F. von" box="[934,1145,187,213]" pageId="13" pageNumber="266" pagination="156 - 210" refId="ref15225" refString="Mollendorff, O. F. von (1886) Materialen zur Fauna for China. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 13, 156 - 210." type="journal article" year="1886">Möllendorff, 1886</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="C4B7A2D7FF987327FB18FF02FE5823B2" authority="(Bavay &amp; Dautzenberg, 1900)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="31C30546FF987327FB18FF02FA2F239E" box="[1211,1437,187,213]" italics="true" pageId="13" pageNumber="266">Metalycaeus heudei</emphasis>
(Bavay &amp; Dautzenberg, 1900)
</taxonomicName>
(see
<bibRefCitation id="6726A4A5FF987327FD86FF66FC8623B2" author="Pall-Gergely, B. &amp; Feher, Z. &amp; Hunyadi, A. &amp; Asami, T." box="[549,820,222,249]" pageId="13" pageNumber="266" pagination="1 - 49" refId="ref15492" refString="Pall-Gergely, B., Feher, Z., Hunyadi, A. &amp; Asami, T. (2015) Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa, 3937 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 3937.1.1" type="journal article" year="2015">
Páll-Gergely
<emphasis id="31C30546FF987327FD19FF66FD4123B2" box="[698,755,223,249]" italics="true" pageId="13" pageNumber="266">et al.</emphasis>
2015
</bibRefCitation>
, 2017).
</paragraph>
<paragraph id="0308D954FF987327FF64FEBAFED422C2" blockId="13.[151,1437,151,393]" pageId="13" pageNumber="266">
The single shell collected in the Chinese
<collectingRegion id="C17317B6FF987327FD38FEBAFCD52256" box="[667,871,259,285]" country="China" name="Guangxi" pageId="13" pageNumber="266">Guangxi Province</collectingRegion>
(
<figureCitation id="9B8CC5D1FF987327FCD4FEBAFC5E2256" box="[887,1004,259,285]" captionStart="FIGURE 9" captionStartId="12.[152,255,1611,1635]" captionTargetBox="[151,1436,181,1587]" captionTargetId="figure-31@12.[151,1436,181,1587]" captionTargetPageId="12" captionText="FIGURE 9. Shells of Pincerna costulosus(Bavay &amp; Dautzenberg, 1912).AD: Laos, Phongsaly Province, JG29; EH:Holotype of Pincerna vallis Z.-Y. Chen &amp; Wu, 2020; IM: Pincerna cf. costulosus, Guangxi, China, 2009/104. Photos: B. Páll-Gergely (AD, IH), taken from the original description (EH)." figureDoi="http://doi.org/10.5281/zenodo.7687436" httpUri="https://zenodo.org/record/7687436/files/figure.png" pageId="13" pageNumber="266">Fig. 9IM</figureCitation>
) is larger than any other
<taxonomicName id="C4B7A2D7FF987327FAA8FEBAFA2F2256" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[1291,1437,259,285]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF987327FAA8FEBAFA2F2256" box="[1291,1437,259,285]" italics="true" pageId="13" pageNumber="266">P. costulosus</emphasis>
</taxonomicName>
shells (D:
<quantity id="C44F74B1FF987327FEA9FE9EFED3220A" box="[266,353,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="13" pageNumber="266" unit="mm" value="4.8">4.8 mm</quantity>
, H:
<quantity id="C44F74B1FF987327FE2CFE9EFE5B220A" box="[399,489,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="266" unit="mm" value="4.9">4.9 mm</quantity>
), and has stronger ribs than any other samples of the same species. It is provisionally identified as
<taxonomicName id="C4B7A2D7FF987327FE87FEF2FE01222E" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[292,435,331,357]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="31C30546FF987327FE87FEF2FE01222E" box="[292,435,331,357]" italics="true" pageId="13" pageNumber="266">P. costulosus</emphasis>
</taxonomicName>
, although more material from that area is necessary to understand the diversity of
<taxonomicName id="C4B7A2D7FF987327FA95FEF2FA2F222F" authorityName="Preston" authorityYear="1907" box="[1334,1437,331,356]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="266" phylum="Mollusca" rank="genus">
<emphasis id="31C30546FF987327FA95FEF2FA2F222F" box="[1334,1437,331,356]" italics="true" pageId="13" pageNumber="266">Pincerna</emphasis>
</taxonomicName>
in southern
<collectingCountry id="7BA099C4FF987327FEB8FED6FED022C2" box="[283,354,367,393]" name="China" pageId="13" pageNumber="266">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>