251 lines
30 KiB
XML
251 lines
30 KiB
XML
<document id="FF3005B8247CA424A1BFF2916EACACF2" ID-CLB-Dataset="54218" ID-DOI="10.11646/zootaxa.5249.2.4" ID-GBIF-Dataset="34ef8754-e9dc-490a-affd-27027f406df2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687418" ID-ZooBank="81426096-9E0D-4E3D-9B1A-660B4D1BBB65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677656213532" checkinUser="plazi" docAuthor="Páll-Gergely, Barna" docDate="2023" docId="8B1E6842FF91732FFF34FCFBFADD22C2" docLanguage="en" docName="zootaxa.5249.2.4.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alycaeus goliath Pall-Gergely 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="258" masterDocId="7727103AFF95732AFFA3FFB9FFB2234B" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna" masterLastPageNumber="276" masterPageNumber="253" pageNumber="257" updateTime="1699475959789" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E2C8B8DA0CF3AD22031E2B941743C173">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna</mods:title>
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<mods:name id="BCBE055284222E2D070E6C889D2C0969" type="personal">
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<mods:namePart id="CE96481B6724B7D945B8D4FCA77095F2">Páll-Gergely, Barna</mods:namePart>
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<mods:typeOfResource id="9FFE950873B517ACE7EDFC5903992067">text</mods:typeOfResource>
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<mods:title id="1792C2DBCF0BB0780E68066059591478">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B35C29530FAC6340D6D8D2306DE96CE4">
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<mods:date id="C9D7BF67B784DF9014FC098E4ED51E1D">2023</mods:date>
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<mods:number id="F5C3707440E49D2E8658BD09976FAB1C">2023-03-01</mods:number>
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<mods:number id="4155223964CED2CF32B9963FB646004A">5249</mods:number>
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<mods:url id="805D8C0C9995A2F7A54D766CDEDA21A7">http://dx.doi.org/10.11646/zootaxa.5249.2.4</mods:url>
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<mods:classification id="90B17064E26BE60C8C9C6A4122C03562">journal article</mods:classification>
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<mods:identifier id="76031A617A735A212907270E05A89700" type="CLB-Dataset">54218</mods:identifier>
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<mods:identifier id="B6E94E9D2DFBD0A32A43E2B26E803655" type="DOI">10.11646/zootaxa.5249.2.4</mods:identifier>
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<mods:identifier id="E8A8AAD3D7CE77467679F87C2B9DA78C" type="GBIF-Dataset">34ef8754-e9dc-490a-affd-27027f406df2</mods:identifier>
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<treatment id="8B1E6842FF91732FFF34FCFBFADD22C2" ID-DOI="http://doi.org/10.5281/zenodo.7688578" ID-GBIF-Taxon="209384101" ID-Zenodo-Dep="7688578" LSID="urn:lsid:plazi:treatment:8B1E6842FF91732FFF34FCFBFADD22C2" httpUri="http://treatment.plazi.org/id/8B1E6842FF91732FFF34FCFBFADD22C2" lastPageId="5" lastPageNumber="258" pageId="4" pageNumber="257">
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<subSubSection id="4BAD8ADFFF91732EFF34FCFBFDE22016" box="[151,592,834,861]" pageId="4" pageNumber="257" type="nomenclature">
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<paragraph id="0308D954FF91732EFF34FCFBFDE22016" blockId="4.[151,592,834,897]" box="[151,592,834,861]" pageId="4" pageNumber="257">
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<heading id="58406E38FF91732EFF34FCFBFDE22016" bold="true" box="[151,592,834,861]" fontSize="11" level="1" pageId="4" pageNumber="257" reason="1">
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<emphasis id="31C30546FF91732EFF34FCFBFDE22016" bold="true" box="[151,592,834,861]" pageId="4" pageNumber="257">
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||
<taxonomicName id="C4B7A2D7FF91732EFF34FCFBFDB42016" ID-CoL="d48f6e3a-3222-49bf-a257-a8b3e173d091" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2023" box="[151,518,834,861]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="257" phylum="Mollusca" rank="species" species="goliath" status="sp. nov.">
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<emphasis id="31C30546FF91732EFF34FCFBFED32016" bold="true" box="[151,353,834,861]" italics="true" pageId="4" pageNumber="257">Alycaeus goliath</emphasis>
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Páll-Gergely
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</taxonomicName>
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<taxonomicNameLabel id="2AF0B83DFF91732EFDAEFCFAFDE22016" box="[525,592,835,861]" pageId="4" pageNumber="257" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BAD8ADFFF91732EFF34FCDEFECA20CA" box="[151,376,871,897]" pageId="4" pageNumber="257" type="description">
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<paragraph id="0308D954FF91732EFF34FCDEFECA20CA" blockId="4.[151,592,834,897]" box="[151,376,871,897]" pageId="4" pageNumber="257">
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||
(
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<figureCitation id="9B8CC5D1FF91732EFF3CFCDEFEAC20CA" box="[159,286,871,897]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="4" pageNumber="257">Figs 1A–D</figureCitation>
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,
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<figureCitation id="9B8CC5D1FF91732EFE89FCDEFEDD20CA" box="[298,367,871,897]" captionStart="FIGURE 2" captionStartId="3.[152,255,918,942]" captionTargetBox="[151,1436,181,895]" captionTargetId="figure-15@3.[151,1436,181,895]" captionTargetPageId="3" captionText="FIGURE 2. Protoconch (A–C) and R2 ribs (D–F) of Alycaeus Baird, 1850 species.A and D: Alycaeus goliath n. sp. (holotype); B and E: Alycaeus eydouxi Venmans, 1956 (Cochinchina, coll. V.W. MaAndrew, NHMUK); C and F: Alycaeus rolfbrandti Maassen, 2006 (6L07p). All photos: B. Páll-Gergely. Scales represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7687422" httpUri="https://zenodo.org/record/7687422/files/figure.png" pageId="4" pageNumber="257">2A, D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4BAD8ADFFF91732EFF34FC16FA3627A2" pageId="4" pageNumber="257" type="materials_examined">
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<paragraph id="0308D954FF91732EFF34FC16FE13278E" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="257">
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<emphasis id="31C30546FF91732EFF34FC16FEFB2083" bold="true" box="[151,329,942,969]" pageId="4" pageNumber="257">Type material.</emphasis>
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<materialsCitation id="B3DFD309FF91732EFEF7FC16FDB02712" ID-GBIF-Occurrence="4039909339" collectingDate="2007-11-25" collectionCode="V" collectorName="Abdou, A. & Muratov, I." country="Laos" elevation="150" latitude="17.4326" location="Muang" longLatPrecision="1" longitude="105.16115" municipality="South-Central Laos" pageId="4" pageNumber="257" specimenCode="MNHN-IM- 2012-27319" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
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<emphasis id="31C30546FF91732EFEF7FC16FE782082" bold="true" box="[340,458,943,969]" pageId="4" pageNumber="257">
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||
<typeStatus id="DC0C67F6FF91732EFEF7FC16FE772082" box="[340,453,943,969]" pageId="4" pageNumber="257" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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(SH: 11.4 mm, SW: 13.9 mm),
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<collectingCountry id="7BA099C4FF91732EFCE8FC16FC2A2082" box="[843,920,943,969]" name="Laos" pageId="4" pageNumber="257">LAOS</collectingCountry>
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:
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<collectingMunicipality id="E36C432EFF91732EFC04FC16FB352082" box="[935,1159,943,969]" pageId="4" pageNumber="257">South-Central Laos</collectingMunicipality>
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,
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<collectingRegion id="C17317B6FF91732EFB35FC16FA2A2082" box="[1174,1432,943,969]" country="Laos" name="Khammouan" pageId="4" pageNumber="257">Khammouan Province</collectingRegion>
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, ca.
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<locationDeviation id="12BAC076FF91732EFF61FC6AFE6E20A6" box="[194,476,979,1005]" location="Thakhek" pageId="4" pageNumber="257">37 km ENE of Thakhek</locationDeviation>
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(
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<location id="06688F8FFF91732EFE4DFC6AFD8D20A6" LSID="urn:lsid:plazi:treatment:8B1E6842FF91732FFF34FCFBFADD22C2:06688F8FFF91732EFE4DFC6AFD8D20A6" box="[494,575,979,1005]" country="Laos" latitude="17.4326" longLatPrecision="1" longitude="105.16115" municipality="South-Central Laos" name="Muang" pageId="4" pageNumber="257" stateProvince="Khammouan">Muang</location>
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Khammouan), ca.
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<locationDeviation id="12BAC076FF91732EFC82FC6BFBE920A6" box="[801,1115,978,1005]" location="Mahaxay" pageId="4" pageNumber="257">4.5 km WNW of Mahaxay</locationDeviation>
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, on and under rocks in dry secondary forest, under a cliff facing east, limestone, clay, black soil in limestone pockets, cave deposits, alt. 150 m,
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<geoCoordinate id="6683BF93FF91732EFF34FBA5FE9A277C" box="[151,296,1052,1079]" degrees="17" direction="north" minutes="25.956" orientation="latitude" pageId="4" pageNumber="257" precision="1" value="17.4326">17°25.956′N</geoCoordinate>
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,
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||
<geoCoordinate id="6683BF93FF91732EFE90FBA5FE7E277C" box="[307,460,1052,1079]" degrees="105" direction="east" minutes="09.669" orientation="longitude" pageId="4" pageNumber="257" precision="1" value="105.16115">105°09.669′E</geoCoordinate>
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(locality code: 3L07), leg.
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<collectorName id="AE42BC82FF91732EFD54FBA2FCD8277E" box="[759,874,1051,1077]" pageId="4" pageNumber="257">Abdou, A.</collectorName>
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&
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<collectorName id="AE42BC82FF91732EFC2FFBA2FBB4277E" box="[908,1030,1051,1077]" pageId="4" pageNumber="257">Muratov, I.</collectorName>
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||
<collectionCode id="65A64191FF91732EFBA5FBA2FBAC277E" box="[1030,1054,1051,1077]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="257" type="Museum">V</collectionCode>
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||
.,
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<date id="7709FF94FF91732EFB8FFBA3FAB0277E" box="[1068,1282,1050,1077]" pageId="4" pageNumber="257" value="2007-11-25">
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<collectingDate id="674D067CFF91732EFB8FFBA3FAB0277E" box="[1068,1282,1050,1077]" pageId="4" pageNumber="257" value="2007-11-25">25 November 2007</collectingDate>
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</date>
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(
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<specimenCode id="5311712FFF91732EFAB3FBA2FE902712" collectionCode="MNHN-IM-" pageId="4" pageNumber="257">MNHN-IM-2012-27319</specimenCode>
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||
,
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<figureCitation id="9B8CC5D1FF91732EFE88FB86FE162712" box="[299,420,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="4" pageNumber="257">Figs 1A–E</figureCitation>
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,
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<figureCitation id="9B8CC5D1FF91732EFE0DFB86FE432712" box="[430,497,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[152,255,918,942]" captionTargetBox="[151,1436,181,895]" captionTargetId="figure-15@3.[151,1436,181,895]" captionTargetPageId="3" captionText="FIGURE 2. Protoconch (A–C) and R2 ribs (D–F) of Alycaeus Baird, 1850 species.A and D: Alycaeus goliath n. sp. (holotype); B and E: Alycaeus eydouxi Venmans, 1956 (Cochinchina, coll. V.W. MaAndrew, NHMUK); C and F: Alycaeus rolfbrandti Maassen, 2006 (6L07p). All photos: B. Páll-Gergely. Scales represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7687422" httpUri="https://zenodo.org/record/7687422/files/figure.png" pageId="4" pageNumber="257">2A, D</figureCitation>
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||
)
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</materialsCitation>
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.
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<materialsCitation id="B3DFD309FF91732EFDAEFB86FED92736" ID-GBIF-Occurrence="4039909342" collectingDate="2007-11-25" collectionCode="V" collectorName="Abdou, A. & Muratov, I." country="Laos" elevation="150" latitude="17.4326" location="Muang" longLatPrecision="1" longitude="105.16115" municipality="South-Central Laos" pageId="4" pageNumber="257" specimenCode="MNHN-IM-2012-27318" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
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||
<emphasis id="31C30546FF91732EFDAEFB86FD3C2712" bold="true" box="[525,654,1087,1113]" pageId="4" pageNumber="257">
|
||
<typeStatus id="DC0C67F6FF91732EFDAEFB86FD3B2712" box="[525,649,1087,1113]" pageId="4" pageNumber="257" type="paratype">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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Same data as for holotype,
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||
<specimenCode id="5311712FFF91732EFC1FFB86FB7C2712" box="[956,1230,1087,1113]" collectionCode="MNHN-IM-" pageId="4" pageNumber="257">MNHN-IM-2012-27318</specimenCode>
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||
(3 complete shells + 8 broken shells)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="B3DFD309FF91732EFEDAFBDAFE2F278E" ID-GBIF-Occurrence="4039909324" collectingDate="2019-09-27" collectorName="A. Hunyadi" country="Laos" county="Thakhek" elevation="155" latitude="17.432617" location="Ban Na Coc" longLatPrecision="1" longitude="105.16115" municipality="Mahaxay" pageId="4" pageNumber="257" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
|
||
<collectingRegion id="C17317B6FF91732EFEDAFBDAFDCB2736" box="[377,633,1123,1149]" country="Laos" name="Khammouan" pageId="4" pageNumber="257">Khammouan Province</collectingRegion>
|
||
, 40 km from the centre of
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<collectingCounty id="EA69A1D8FF91732EFC11FBDAFBA52736" box="[946,1047,1123,1149]" pageId="4" pageNumber="257">Thakhek</collectingCounty>
|
||
,
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||
<collectingMunicipality id="E36C432EFF91732EFB87FBDAFB3F2736" box="[1060,1165,1123,1149]" pageId="4" pageNumber="257">Mahaxay</collectingMunicipality>
|
||
, 2.3 km southeast from
|
||
<location id="06688F8FFF91732EFF34FB3EFE9027EA" LSID="urn:lsid:plazi:treatment:8B1E6842FF91732FFF34FCFBFADD22C2:06688F8FFF91732EFF34FB3EFE9027EA" box="[151,290,1159,1185]" country="Laos" county="Thakhek" latitude="17.432617" longLatPrecision="1" longitude="105.16115" municipality="Mahaxay" name="Ban Na Coc" pageId="4" pageNumber="257" stateProvince="Khammouan">Ban Na Coc</location>
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, limestone wall, 155 m a.s.l.,
|
||
<geoCoordinate id="6683BF93FF91732EFDCFFB31FD4C27E8" box="[620,766,1160,1187]" degrees="17" direction="north" minutes="25.957" orientation="latitude" pageId="4" pageNumber="257" precision="1" value="17.432617">17°25.957′N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="6683BF93FF91732EFCAAFB31FC1027E8" box="[777,930,1160,1187]" degrees="105" direction="east" minutes="09.669" orientation="longitude" pageId="4" pageNumber="257" precision="1" value="105.16115">105°09.669′E</geoCoordinate>
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||
(locality code: 2019/95), leg.
|
||
<collectorName id="AE42BC82FF91732EFB53FB3EFAC727EA" box="[1264,1397,1159,1185]" pageId="4" pageNumber="257">A. Hunyadi</collectorName>
|
||
,
|
||
<date id="7709FF94FF91732EFA23FB3EFEE3278E" pageId="4" pageNumber="257" value="2019-09-27">
|
||
<collectingDate id="674D067CFF91732EFA23FB3EFEE3278E" pageId="4" pageNumber="257" value="2019-09-27">27 September 2019</collectingDate>
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||
</date>
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||
, HA/1
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||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="0308D954FF91732EFF64FB77FA3627A2" blockId="4.[151,1437,942,2013]" box="[199,1412,1230,1257]" pageId="4" pageNumber="257">
|
||
<materialsCitation id="B3DFD309FF91732EFF64FB77FA3227A2" ID-GBIF-Occurrence="4039909322" box="[199,1408,1230,1257]" collectingDate="2007-11-25" collectionCode="V" collectorName="Abdou, A. & Muratov, I." country="Laos" elevation="150" latitude="17.4326" location="Muang" longLatPrecision="1" longitude="105.16115" municipality="South-Central Laos" pageId="4" pageNumber="257" specimenCode="MNHN-IM-2012-27318" specimenCount="1" stateProvince="Khammouan">
|
||
<emphasis id="31C30546FF91732EFF64FB77FD8427A3" bold="true" box="[199,566,1230,1256]" pageId="4" pageNumber="257">Additional material examined:</emphasis>
|
||
Same data as for holotype, some shell fragments,
|
||
<specimenCode id="5311712FFF91732EFBC9FB76FA3227A2" box="[1130,1408,1231,1257]" collectionCode="MNHN-IM-" pageId="4" pageNumber="257">MNHN-IM-2012-27318</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF91732EFF64FB4AFF42267A" pageId="4" pageNumber="257" type="etymology">
|
||
<paragraph id="0308D954FF91732EFF64FB4AFF42267A" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64FB4AFEFC2646" bold="true" box="[199,334,1267,1293]" pageId="4" pageNumber="257">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="C4B7A2D7FF91732EFE6BFB4AFDA52646" authorityName="Pall-Gergely" authorityYear="2023" box="[456,535,1267,1293]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="257" phylum="Mollusca" rank="species" species="goliath">
|
||
<emphasis id="31C30546FF91732EFE6BFB4AFDA52646" box="[456,535,1267,1293]" italics="true" pageId="4" pageNumber="257">goliath</emphasis>
|
||
</taxonomicName>
|
||
(to be used as a noun in apposition) refers to the remarkably large size of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF91732EFF64FA83FB0226D6" pageId="4" pageNumber="257" type="diagnosis">
|
||
<paragraph id="0308D954FF91732EFF64FA83FB0226D6" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64FA83FEF0261F" bold="true" box="[199,322,1338,1364]" pageId="4" pageNumber="257">Diagnosis.</emphasis>
|
||
An exceptionally large alycaeid species with dominant last whorl (shell width larger than shell height), smooth protoconch, finely reticulated R1, very long R2 having ribs that form many connections with each other by calcareous projections, long R3 with low swelling, and a protruding inner peristome.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF91732EFF64FA1FFAC424DE" pageId="4" pageNumber="257" type="description">
|
||
<paragraph id="0308D954FF91732EFF64FA1FFA3B2462" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64FA1FFEEB268B" bold="true" box="[199,345,1446,1472]" pageId="4" pageNumber="257">Description.</emphasis>
|
||
Shell exceptionally large, pinkish, conical with wide last whorl; shell wider than high; protoconch consists of ca. 2 whorls, it is nearly smooth, with only indication of extremely fine ribs on the last quarter whorl; R1 consists of ca. 3 whorls; whorls bulging; sculpture of R1 dominated by weak, dense, rather irregular radial ribs and much weaker spiral striation; R2 and R3 slightly longer than half whorl combined; R2 slightly longer than R3; R2 with sharp, dense ribs, which form many connections with each other by calcareous projections, resulting in a web-like surface continuous between tube and umbilicus; tube situated deep in suture, its surface is very rough, irregularly segmented, although internally probably continuous; constriction between R2 and R3 deep but short; blunt, low swelling on R3 situated just after constriction; aperture rounded with slight upper incision; outer peristome strongly expanded and slightly reflected, inner peristome strong, sharp, protruding, clearly separated from the outer peristome, especially in the palatal region; outer peristome attached to penultimate whorl, but inner peristome does not; peristome whitish, inner side of aperture orange in colour; umbilicus open, shows all whorls.
|
||
</paragraph>
|
||
<paragraph id="0308D954FF91732EFF64F88BFE662406" blockId="4.[151,1437,942,2013]" box="[199,468,1842,1869]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64F88BFEE52407" bold="true" box="[199,343,1842,1868]" pageId="4" pageNumber="257">Operculum.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="0308D954FF91732EFF64F8EEFC24243A" blockId="4.[151,1437,942,2013]" box="[199,918,1878,1905]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64F8EEFE56243B" bold="true" box="[199,484,1878,1905]" pageId="4" pageNumber="257">Measurements (in mm).</emphasis>
|
||
D = 13.8–15.2, H = 11.1–12.0 (n = 5).
|
||
</paragraph>
|
||
<paragraph id="0308D954FF91732EFF64F8C3FAC424DE" blockId="4.[151,1437,942,2013]" box="[199,1398,1914,1941]" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64F8C3FE6C24DF" bold="true" box="[199,478,1914,1940]" pageId="4" pageNumber="257">Variation in specimens.</emphasis>
|
||
There are no notable conchological differences between the examined specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF91732FFF64F827FBA1220A" lastPageId="5" lastPageNumber="258" pageId="4" pageNumber="257" type="diagnosis">
|
||
<paragraph id="0308D954FF91732FFF64F827FBA1220A" blockId="4.[151,1437,942,2013]" lastBlockId="5.[151,1437,151,393]" lastPageId="5" lastPageNumber="258" pageId="4" pageNumber="257">
|
||
<emphasis id="31C30546FF91732EFF64F827FE7C24F3" bold="true" box="[199,462,1950,1976]" pageId="4" pageNumber="257">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="C4B7A2D7FF91732EFE75F826FD2324F2" authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[470,657,1951,1977]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="257" phylum="Mollusca" rank="species" species="goliath" status="sp. nov.">
|
||
<emphasis id="31C30546FF91732EFE75F826FD2324F2" box="[470,657,1951,1977]" italics="true" pageId="4" pageNumber="257">Alycaeus goliath</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31C30546FF91732EFD3AF826FD6A24F2" bold="true" box="[665,728,1951,1977]" pageId="4" pageNumber="257">
|
||
<taxonomicNameLabel id="2AF0B83DFF91732EFD3AF826FD6A24F2" box="[665,728,1951,1977]" pageId="4" pageNumber="257" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="C4B7A2D7FF91732EFCCDF826FC5624F2" authorityName="Venmans" authorityYear="1956" box="[878,996,1951,1977]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="257" phylum="Mollusca" rank="species" species="eydouxi">
|
||
<emphasis id="31C30546FF91732EFCCDF826FC5624F2" box="[878,996,1951,1977]" italics="true" pageId="4" pageNumber="257">A. eydouxi</emphasis>
|
||
</taxonomicName>
|
||
by the larger size, the more protruding inner peristome, the comparatively less bulging R2, and most importantly, the R2 ribs which form connections with their neighbours, resulting in a web-like structure.
|
||
<taxonomicName id="C4B7A2D7FF90732FFD6AFF2EFBEE23FA" authority="Maassen, 2006" authorityName="Maassen" authorityYear="2006" box="[713,1116,151,177]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis id="31C30546FF90732FFD6AFF2EFC1D23FA" box="[713,943,151,177]" italics="true" pageId="5" pageNumber="258">Alycaeus rolfbrandti</emphasis>
|
||
<bibRefCitation id="6726A4A5FF90732FFC16FF2EFBEE23FA" author="Maassen, W. J. M." box="[949,1116,151,177]" pageId="5" pageNumber="258" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
differs from the new species by the smaller shell size and the irregularly ribbed, squamous protoconch. Although most known populations of
|
||
<taxonomicName id="C4B7A2D7FF90732FFA27FF02FEA323B2" authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis id="31C30546FF90732FFA27FF02FEA323B2" italics="true" pageId="5" pageNumber="258">A. rolfbrandti</emphasis>
|
||
</taxonomicName>
|
||
have keeled shells, three examined samples have a rounded edge of the body whorl. Therefore, although this is a useful shell character, this not sufficient to distinguish the two species. The
|
||
<taxonomicName id="C4B7A2D7FF90732FFB9AFEBAFAAC2256" authorityName="Maassen" authorityYear="2006" box="[1081,1310,259,285]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis id="31C30546FF90732FFB9AFEBAFAAC2256" box="[1081,1310,259,285]" italics="true" pageId="5" pageNumber="258">Alycaeus rolfbrandti</emphasis>
|
||
</taxonomicName>
|
||
population occurring syntopically with
|
||
<taxonomicName id="C4B7A2D7FF90732FFE71FE9EFDF2220A" authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[466,576,295,321]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="goliath" status="sp. nov.">
|
||
<emphasis id="31C30546FF90732FFE71FE9EFDF2220A" box="[466,576,295,321]" italics="true" pageId="5" pageNumber="258">A. goliath</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31C30546FF90732FFDEBFE9EFD34220A" bold="true" box="[584,646,295,321]" pageId="5" pageNumber="258">
|
||
<taxonomicNameLabel id="2AF0B83DFF90732FFDEBFE9EFD34220A" box="[584,646,295,321]" pageId="5" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has rounded body whorl (
|
||
<figureCitation id="9B8CC5D1FF90732FFC0FFE9EFBB7220A" box="[940,1029,295,321]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="5" pageNumber="258">Fig. 1N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF90732FFF64FEF3FA2E222E" box="[199,1436,330,357]" pageId="5" pageNumber="258" type="distribution">
|
||
<paragraph id="0308D954FF90732FFF64FEF3FA2E222E" blockId="5.[151,1437,151,393]" box="[199,1436,330,357]" pageId="5" pageNumber="258">
|
||
<emphasis id="31C30546FF90732FFF64FEF3FED3222F" bold="true" box="[199,353,330,356]" pageId="5" pageNumber="258">Distribution.</emphasis>
|
||
This species is known from the
|
||
<typeStatus id="DC0C67F6FF90732FFD14FEF2FD55222E" box="[695,743,331,357]" pageId="5" pageNumber="258">type</typeStatus>
|
||
locality only, where it lives together with
|
||
<taxonomicName id="C4B7A2D7FF90732FFB09FEF2FAF2222E" authorityName="Maassen" authorityYear="2006" box="[1194,1344,331,357]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis id="31C30546FF90732FFB09FEF2FAF2222E" box="[1194,1344,331,357]" italics="true" pageId="5" pageNumber="258">A. rolfbrandti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9B8CC5D1FF90732FFAEFFEF2FA3D222E" box="[1356,1423,331,357]" captionStart="FIGURE 4" captionStartId="5.[152,255,1163,1187]" captionTargetBox="[151,1435,417,1139]" captionTargetId="figure-130@5.[151,1435,417,1139]" captionTargetPageId="5" captionText="FIGURE 4. Distribution of Alycaeus Baird, 1850 species in central Laos. Yellow circles: Alycaeus rolfbrandti Maassen, 2006; Red triangle: A. goliath n. sp. The map was made using Google Earth Pro." figureDoi="http://doi.org/10.5281/zenodo.7687426" httpUri="https://zenodo.org/record/7687426/files/figure.png" pageId="5" pageNumber="258">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAD8ADFFF90732FFF64FED6FADD22C2" box="[199,1391,367,393]" pageId="5" pageNumber="258" type="discussion">
|
||
<paragraph id="0308D954FF90732FFF64FED6FADD22C2" blockId="5.[151,1437,151,393]" box="[199,1391,367,393]" pageId="5" pageNumber="258">
|
||
<emphasis id="31C30546FF90732FFF64FED6FE8922C2" bold="true" box="[199,315,367,393]" pageId="5" pageNumber="258">Remarks.</emphasis>
|
||
This is the largest species of the family, followed by
|
||
<taxonomicName id="C4B7A2D7FF90732FFC32FED6FAD922C2" authority="(Maassen, 2006)" baseAuthorityName="Maassen" baseAuthorityYear="2006" box="[913,1387,367,393]" class="Gastropoda" family="Alycaeidae" genus="Stomacosmethis" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="christae">
|
||
<emphasis id="31C30546FF90732FFC32FED6FB1422C2" box="[913,1190,367,393]" italics="true" pageId="5" pageNumber="258">Stomacosmethis christae</emphasis>
|
||
(
|
||
<bibRefCitation id="6726A4A5FF90732FFB16FED6FAD122C2" author="Maassen, W. J. M." box="[1205,1379,367,393]" pageId="5" pageNumber="258" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |