410 lines
42 KiB
XML
410 lines
42 KiB
XML
<document ID-DOI="10.11646/zootaxa.5249.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687418" ID-ZooBank="81426096-9E0D-4E3D-9B1A-660B4D1BBB65" approvalRequired="9" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="5" approvalRequired_for_treatments="3" checkinTime="1677656213532" checkinUser="plazi" docAuthor="Páll-Gergely, Barna" docDate="2023" docId="8B1E6842FF90732CFF34FAA0FCA225AA" docLanguage="en" docName="zootaxa.5249.2.4.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alycaeus rolfbrandti Maassen 2006" docType="treatment" docVersion="2" lastPageNumber="259" masterDocId="7727103AFF95732AFFA3FFB9FFB2234B" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna" masterLastPageNumber="276" masterPageNumber="253" pageNumber="258" updateTime="1677656825642" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Páll-Gergely, Barna</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5249</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>253</mods:start>
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<mods:end>276</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.2.4</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5249.2.4</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7687418</mods:identifier>
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<mods:identifier type="ZooBank">81426096-9E0D-4E3D-9B1A-660B4D1BBB65</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:8B1E6842FF90732CFF34FAA0FCA225AA" httpUri="http://treatment.plazi.org/id/8B1E6842FF90732CFF34FAA0FCA225AA" lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258">
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<subSubSection box="[151,590,1305,1332]" pageId="5" pageNumber="258" type="nomenclature">
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<paragraph blockId="5.[151,590,1305,1367]" box="[151,590,1305,1332]" pageId="5" pageNumber="258">
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<heading bold="true" box="[151,590,1305,1332]" fontSize="11" level="1" pageId="5" pageNumber="258" reason="1">
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<taxonomicName ID-CoL="7PYMR" authority="Maassen, 2006" authorityName="Maassen" authorityYear="2006" box="[151,590,1305,1332]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
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<emphasis bold="true" box="[151,590,1305,1332]" pageId="5" pageNumber="258">
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<emphasis bold="true" box="[151,400,1305,1332]" italics="true" pageId="5" pageNumber="258">Alycaeus rolfbrandti</emphasis>
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<bibRefCitation author="Maassen, W. J. M." box="[407,590,1305,1331]" pageId="5" pageNumber="258" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen, 2006</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,354,1341,1367]" pageId="5" pageNumber="258" type="description">
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<paragraph blockId="5.[151,590,1305,1367]" box="[151,354,1341,1367]" pageId="5" pageNumber="258">
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(
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<figureCitation box="[159,271,1341,1367]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="5" pageNumber="258">Figs 1I–S</figureCitation>
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,
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<figureCitation box="[283,346,1341,1367]" captionStart="FIGURE 2" captionStartId="3.[152,255,918,942]" captionTargetBox="[151,1436,181,895]" captionTargetId="figure-15@3.[151,1436,181,895]" captionTargetPageId="3" captionText="FIGURE 2. Protoconch (A–C) and R2 ribs (D–F) of Alycaeus Baird, 1850 species.A and D: Alycaeus goliath n. sp. (holotype); B and E: Alycaeus eydouxi Venmans, 1956 (Cochinchina, coll. V.W. MaAndrew, NHMUK); C and F: Alycaeus rolfbrandti Maassen, 2006 (6L07p). All photos: B. Páll-Gergely. Scales represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7687422" httpUri="https://zenodo.org/record/7687422/files/figure.png" pageId="5" pageNumber="258">2C, F</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="5" pageNumber="258" type="reference_group">
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<paragraph blockId="5.[151,1144,1410,1464]" box="[151,722,1410,1434]" pageId="5" pageNumber="258">
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<treatmentCitationGroup box="[151,722,1410,1434]" pageId="5" pageNumber="258">
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<treatmentCitation author="Maassen, W. J. M." box="[151,622,1410,1434]" page="137" pageId="5" pageNumber="258" year="2006">
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<taxonomicName ID-CoL="7PYMR" authority="Maassen, 2006: 136 - 137" authorityName="Maassen" authorityPageNumber="136 - 137" authorityYear="2006" box="[151,622,1410,1434]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">
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<emphasis box="[151,359,1410,1434]" italics="true" pageId="5" pageNumber="258">Alycaeus rolfbrandti</emphasis>
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<bibRefCitation author="Maassen, W. J. M." box="[366,622,1410,1434]" pageId="5" pageNumber="258" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen, 2006: 136–137</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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, figs 6–9.
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph blockId="5.[151,1144,1410,1464]" box="[151,1144,1440,1464]" pageId="5" pageNumber="258">
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<treatmentCitationGroup box="[151,1144,1440,1464]" pageId="5" pageNumber="258">
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<emphasis box="[151,382,1440,1464]" italics="true" pageId="5" pageNumber="258">
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||
<taxonomicName ID-CoL="7PYMR" authorityName="Maassen" authorityYear="2006" box="[151,357,1440,1464]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="258" phylum="Mollusca" rank="species" species="rolfbrandti">Alycaeus rolfbrandti</taxonomicName>
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—
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</emphasis>
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Páll-Gergely
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<emphasis box="[517,568,1440,1464]" italics="true" pageId="5" pageNumber="258">et al.</emphasis>
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, 2017: 10, fig. 3B;
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<treatmentCitation author="Pall-Gergely, B. & Sajan, S. & Tripathy, B. & Meng, K. & Asami, T. & Ablett, J. D." box="[766,1056,1440,1464]" httpUri="http://treatment.plazi.org/id/A8258F43D119560F80E548AAF61749EE" page="23" pageId="5" pageNumber="258" year="2020">
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<bibRefCitation author="Pall-Gergely, B. & Sajan, S. & Tripathy, B. & Meng, K. & Asami, T. & Ablett, J. D." box="[766,1056,1440,1464]" pageId="5" pageNumber="258" pagination="1 - 220" refId="ref15744" refString="Pall-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. & Ablett, J. D. (2020) Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys, 981, 1 - 220. https: // doi. org / 10.3897 / zookeys. 981.53583" type="journal article" year="2020">
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Páll-Gergely
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<emphasis box="[901,953,1440,1464]" italics="true" pageId="5" pageNumber="258">et al.</emphasis>
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, 2020: 23
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</bibRefCitation>
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</treatmentCitation>
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, fig. 2B.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258" type="materials_examined">
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<paragraph blockId="5.[151,1437,1510,2041]" pageId="5" pageNumber="258">
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<emphasis bold="true" box="[151,326,1510,1537]" pageId="5" pageNumber="258">
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<typeStatus box="[151,210,1511,1537]" pageId="5" pageNumber="258">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus box="[333,389,1511,1537]" pageId="5" pageNumber="258">Type</typeStatus>
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specimens were not examined by us, but other shells from the original series were available (see under
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<typeStatus box="[274,490,1546,1572]" isOtherTypeStatus="true" pageId="5" pageNumber="258">additional material</typeStatus>
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||
).
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</paragraph>
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<paragraph blockId="5.[151,1437,1510,2041]" lastBlockId="6.[151,1437,150,1761]" lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258">
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<emphasis bold="true" box="[199,562,1582,1608]" pageId="5" pageNumber="258">Additional material examined.</emphasis>
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||
<collectingCountry box="[568,645,1583,1609]" name="Laos" pageId="5" pageNumber="258">LAOS</collectingCountry>
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: South-Central Laos,
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||
<collectingRegion box="[885,1138,1582,1608]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan Province</collectingRegion>
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, ca.
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||
<quantity box="[1186,1277,1582,1609]" metricMagnitude="4" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="258" unit="km" value="13.5">13.5 km</quantity>
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||
N of Thakhek (Muang
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<collectingRegion box="[248,397,1618,1644]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan</collectingRegion>
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), ca.
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<quantity box="[456,514,1618,1645]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="258" unit="km" value="5.0">5 km</quantity>
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SE of Ban Nakok, under limestone rocks in dry secondary forest under W facing cliff, limestone, clay, alt.
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<quantity box="[425,496,1655,1681]" metricMagnitude="2" metricUnit="m" metricValue="1.38" pageId="5" pageNumber="258" unit="m" value="138.0">138 m</quantity>
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,
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<geoCoordinate box="[505,650,1656,1683]" degrees="17" direction="north" minutes="30.984" orientation="latitude" pageId="5" pageNumber="258" precision="1" value="17.5164">17°30.984′N</geoCoordinate>
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,
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<geoCoordinate box="[660,812,1656,1683]" degrees="104" direction="east" minutes="47.682" orientation="longitude" pageId="5" pageNumber="258" precision="1" value="104.7947">104°47.682′E</geoCoordinate>
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(locality code: 9L07), leg.Abdou,A. & Muratov, I.
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<collectionCode box="[1371,1395,1655,1681]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="258" type="Museum">V</collectionCode>
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.,
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<date pageId="5" pageNumber="258" value="2007-11-30">30 November 2007</date>
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,
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||
<collectionCode box="[345,486,1691,1717]" pageId="5" pageNumber="258">MNHN-IM-</collectionCode>
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||
2012-27322/3 complete + 1 broken shell (
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<figureCitation box="[951,1039,1690,1717]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="5" pageNumber="258">Fig. 1Q</figureCitation>
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); South-Central Laos,
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<collectingRegion country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan Province</collectingRegion>
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||
, ca.
|
||
<quantity box="[306,378,1726,1753]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="258" unit="km" value="15.0">15 km</quantity>
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NE of Thakhek (Muang
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<collectingRegion box="[665,814,1726,1752]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan</collectingRegion>
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), ca.
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<quantity box="[874,967,1726,1753]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="258" unit="km" value="1.25">1.25 km</quantity>
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SE of Ban Nase, near large flooded cave under W facing cliff, limestone, clay, black soil on limestone, on and under rocks in dry secondary forest, alt.
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<quantity box="[1360,1432,1763,1789]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="258" unit="m" value="127.0">127 m</quantity>
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,
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<geoCoordinate box="[151,297,1800,1827]" degrees="17" direction="north" minutes="30.547" orientation="latitude" pageId="5" pageNumber="258" precision="1" value="17.509117">17°30.547′N</geoCoordinate>
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,
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<geoCoordinate box="[307,460,1800,1827]" degrees="104" direction="east" minutes="53.444" orientation="longitude" pageId="5" pageNumber="258" precision="1" value="104.89073">104°53.444′E</geoCoordinate>
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(locality code: 7L07), leg. Abdou, A. & Muratov, I.
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<collectionCode box="[1031,1055,1799,1825]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="258" type="Museum">V</collectionCode>
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||
.,
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||
<date box="[1070,1288,1798,1825]" pageId="5" pageNumber="258" value="2007-11-28">28 November 2007</date>
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||
,
|
||
<collectionCode box="[1298,1436,1799,1825]" pageId="5" pageNumber="258">MNHN-IM-</collectionCode>
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||
2012-27325/8 shells + some shell fragments (
|
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<figureCitation box="[661,749,1834,1861]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="5" pageNumber="258">Fig. 1O</figureCitation>
|
||
); South-Central Laos,
|
||
<collectingRegion box="[1002,1256,1834,1860]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan Province</collectingRegion>
|
||
, ca.
|
||
<quantity box="[1305,1362,1834,1861]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="258" unit="km" value="9.0">9 km</quantity>
|
||
NE of Thakhek (Muang
|
||
<collectingRegion box="[348,497,1870,1896]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan</collectingRegion>
|
||
), NW facing cliff, limestone, clay, black soil in limestone pockets, on and under rocks in dry secondary forest on and under, alt.
|
||
<quantity box="[606,678,1907,1933]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="258" unit="m" value="190.0">190 m</quantity>
|
||
,
|
||
<geoCoordinate box="[688,834,1908,1935]" degrees="17" direction="north" minutes="26.757" orientation="latitude" pageId="5" pageNumber="258" precision="1" value="17.44595">17°26.757′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[843,997,1908,1935]" degrees="104" direction="east" minutes="52.937" orientation="longitude" pageId="5" pageNumber="258" precision="1" value="104.882286">104°52.937′E</geoCoordinate>
|
||
(locality code: 6L07), leg. Abdou, A. & Muratov, I.
|
||
<collectionCode box="[273,297,1943,1969]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="258" type="Museum">V</collectionCode>
|
||
.,
|
||
<date box="[311,528,1942,1969]" pageId="5" pageNumber="258" value="2007-11-27">27 November 2007</date>
|
||
,
|
||
<collectionCode box="[538,680,1943,1969]" pageId="5" pageNumber="258">MNHN-IM-</collectionCode>
|
||
2012-27321/19 complete shells + some shell fragments (
|
||
<figureCitation box="[1302,1386,1942,1969]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="5" pageNumber="258">Figs 1P</figureCitation>
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||
,
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="3.[152,255,918,942]" captionTargetBox="[151,1436,181,895]" captionTargetId="figure-15@3.[151,1436,181,895]" captionTargetPageId="3" captionText="FIGURE 2. Protoconch (A–C) and R2 ribs (D–F) of Alycaeus Baird, 1850 species.A and D: Alycaeus goliath n. sp. (holotype); B and E: Alycaeus eydouxi Venmans, 1956 (Cochinchina, coll. V.W. MaAndrew, NHMUK); C and F: Alycaeus rolfbrandti Maassen, 2006 (6L07p). All photos: B. Páll-Gergely. Scales represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7687422" httpUri="https://zenodo.org/record/7687422/files/figure.png" pageId="5" pageNumber="258">2C, F</figureCitation>
|
||
); South-Central Laos,
|
||
<collectingRegion box="[424,678,1978,2004]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan Province</collectingRegion>
|
||
, ca.
|
||
<quantity box="[729,821,1978,2005]" metricMagnitude="4" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="258" unit="km" value="15.5">15.5 km</quantity>
|
||
ENE of Thakhek (Muang
|
||
<collectingRegion box="[1121,1270,1978,2004]" country="Laos" name="Khammouan" pageId="5" pageNumber="258">Khammouan</collectingRegion>
|
||
), on and under limestone rocks in dry secondary forest under NW facing cliff, limestone, clay, black soil in limestone pockets, alt.
|
||
<quantity box="[151,227,150,177]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="259" unit="m" value="155.0">155 m</quantity>
|
||
,
|
||
<geoCoordinate box="[241,387,152,179]" degrees="17" direction="north" minutes="26.817" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.44695">17°26.817′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[400,553,152,179]" degrees="104" direction="east" minutes="57.200" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.95333">104°57.200′E</geoCoordinate>
|
||
(locality code: 12L07), leg. Abdou, A. & Muratov, I.
|
||
<collectionCode box="[1169,1193,151,177]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="259" type="Museum">V</collectionCode>
|
||
.,
|
||
<date box="[1211,1432,151,177]" pageId="6" pageNumber="259" value="2007-12-01">01 December 2007</date>
|
||
,
|
||
<collectionCode box="[151,293,187,213]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27320/1 shell (
|
||
<figureCitation box="[523,606,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1S</figureCitation>
|
||
); South-Central Laos,
|
||
<collectingRegion box="[862,1116,187,213]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
, ca.
|
||
<quantity box="[1168,1239,186,213]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="259" unit="km" value="15.0">15 km</quantity>
|
||
ENE of Thakhek (Muang
|
||
<collectingRegion box="[247,396,223,249]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan</collectingRegion>
|
||
), in the upper entrance of the large cave on NW facing steep slope, limestone, clay, black soil on limestone, on and under limestone rocks in dry secondary forest, alt.
|
||
<quantity box="[946,1018,258,285]" metricMagnitude="2" metricUnit="m" metricValue="1.5699999999999998" pageId="6" pageNumber="259" unit="m" value="157.0">157 m</quantity>
|
||
,
|
||
<geoCoordinate box="[1028,1173,260,287]" degrees="17" direction="north" minutes="26.650" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.444166">17°26.650′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1184,1337,260,287]" degrees="104" direction="east" minutes="56.944" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.949066">104°56.944′E</geoCoordinate>
|
||
(locality code: 11L07), leg. Abdou, A. & Muratov, I.
|
||
<collectionCode box="[635,659,295,321]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="259" type="Museum">V</collectionCode>
|
||
.,
|
||
<date box="[675,891,295,321]" pageId="6" pageNumber="259" value="2007-12-01">01 December 2007</date>
|
||
,
|
||
<collectionCode box="[902,1043,294,321]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27324/5 damaged shells (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1R</figureCitation>
|
||
); Kalkberge ca.
|
||
<quantity box="[369,440,331,357]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="259" unit="km" value="20.0">20 km</quantity>
|
||
östl. Takek, leg. Brandt,
|
||
<date box="[720,940,331,357]" pageId="6" pageNumber="259" value="1963-09-08">08 September 1963</date>
|
||
,
|
||
<collectionCode box="[951,1007,331,357]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="259">SMF</collectionCode>
|
||
262540 (1 shell; labelled the
|
||
<typeStatus box="[1338,1436,331,357]" pageId="6" pageNumber="259">holotype</typeStatus>
|
||
of “
|
||
<taxonomicName authority="Brandt" authorityName="Brandt" box="[193,493,367,393]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="carinatus">
|
||
<emphasis box="[193,406,367,393]" italics="true" pageId="6" pageNumber="259">Alycaeus carinatus</emphasis>
|
||
Brandt
|
||
</taxonomicName>
|
||
”, but not mentioned by
|
||
<bibRefCitation author="Maassen, W. J. M." box="[755,918,367,393]" pageId="6" pageNumber="259" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen 2006</bibRefCitation>
|
||
,
|
||
<figureCitation box="[928,1036,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1I–L</figureCitation>
|
||
); Same data,
|
||
<collectionCode box="[1184,1240,367,393]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="259">SMF</collectionCode>
|
||
262541 (5 shells, labelled as
|
||
<typeStatus box="[274,381,403,429]" pageId="6" pageNumber="259">paratypes</typeStatus>
|
||
of “
|
||
<taxonomicName authority="Brandt" authorityName="Brandt" box="[428,726,403,429]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="carinatus">
|
||
<emphasis box="[428,640,403,429]" italics="true" pageId="6" pageNumber="259">Alycaeus carinatus</emphasis>
|
||
Brandt
|
||
</taxonomicName>
|
||
”, but not mentioned in
|
||
<bibRefCitation author="Maassen, W. J. M." box="[979,1142,403,429]" pageId="6" pageNumber="259" pagination="133 - 139" refId="ref15083" refString="Maassen, W. J. M. (2006) Remarks on Alycaeus species from South-East Asia, with the descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae). Basteria, 70 (4 / 6), 133 - 139." type="journal article" year="2006">Maassen 2006</bibRefCitation>
|
||
; consequently, these shells are part of the original series of
|
||
<taxonomicName authorityName="Maassen" authorityYear="2006" box="[514,668,439,465]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis box="[514,668,439,465]" italics="true" pageId="6" pageNumber="259">A. rolfbrandti</emphasis>
|
||
</taxonomicName>
|
||
, but they are not
|
||
<typeStatus box="[867,974,439,465]" pageId="6" pageNumber="259">paratypes</typeStatus>
|
||
since Maassen clearly had not examined them); South-Central Laos,
|
||
<collectingRegion box="[463,717,475,501]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
, ca.
|
||
<quantity box="[768,860,474,501]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="259" unit="km" value="10.5">10.5 km</quantity>
|
||
E of Thakhek (Muang
|
||
<collectingRegion box="[1122,1271,475,501]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan</collectingRegion>
|
||
), on and under rocks, cave deposits, in secondary forest under entrance and in large cave on NE facing steep slope, limestone, clay, black soil on limestone, alt.
|
||
<quantity box="[467,540,547,573]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="259" unit="m" value="160.0">160 m</quantity>
|
||
,
|
||
<geoCoordinate box="[552,697,548,575]" degrees="17" direction="north" minutes="24.340" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.405666">17°24.340′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[709,862,548,575]" degrees="104" direction="east" minutes="54.894" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.9149">104°54.894′E</geoCoordinate>
|
||
(locality code: 25L07), leg. Abdou, A. & Muratov, I.
|
||
<collectionCode box="[165,189,583,609]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="259" type="Museum">V</collectionCode>
|
||
.,
|
||
<date box="[208,431,583,609]" pageId="6" pageNumber="259" value="2007-12-09">09 December 2007</date>
|
||
,
|
||
<collectionCode box="[446,587,583,609]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27323/2 complete + 2 broken shells (
|
||
<figureCitation box="[1086,1183,583,609]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1M</figureCitation>
|
||
); South-Central Laos,
|
||
<collectingRegion box="[151,406,619,645]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
,
|
||
<quantity box="[418,476,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="259" unit="km" value="2.0">2 km</quantity>
|
||
NWW of Ban Na village, Pha Suang Cave – S entrance,
|
||
<geoCoordinate box="[1126,1270,620,647]" degrees="17" direction="north" minutes="33.054" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.5509">17°33.054′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1283,1436,620,647]" degrees="104" direction="east" minutes="52.414" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.873566">104°52.414′E</geoCoordinate>
|
||
(locality code:
|
||
<collectionCode box="[322,358,655,681]" pageId="6" pageNumber="259">JG</collectionCode>
|
||
3), leg. Grego, J.,
|
||
<date box="[562,767,655,681]" pageId="6" pageNumber="259" value="2017-02-09">09 February 2017</date>
|
||
, JG/9,
|
||
<collectionCode box="[848,905,655,681]" pageId="6" pageNumber="259">PGB</collectionCode>
|
||
/2; South-Central Laos,
|
||
<collectingRegion box="[1176,1432,655,681]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
, NE foot of Pha Soung Mountain, caverns among slope boulders,
|
||
<geoCoordinate box="[887,1033,692,719]" degrees="17" direction="north" minutes="33.108" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.5518">17°33.108′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1044,1198,692,719]" degrees="104" direction="east" minutes="52.301" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.87168">104°52.301′E</geoCoordinate>
|
||
(locality code:
|
||
<collectionCode box="[1375,1409,691,717]" pageId="6" pageNumber="259">JG</collectionCode>
|
||
4), leg. Grego, J.,
|
||
<date box="[321,526,727,753]" pageId="6" pageNumber="259" value="2017-02-08">08 February 2017</date>
|
||
, JG/4; South-Central Laos,
|
||
<collectingRegion box="[844,1099,727,753]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
,
|
||
<quantity box="[1113,1171,727,753]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="259" unit="km" value="6.0">6 km</quantity>
|
||
SSE of Ban Na, Tham Dan Makhia, lake cave entrance,
|
||
<geoCoordinate box="[521,666,764,791]" degrees="17" direction="north" minutes="30.616" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.510267">17°30.616′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[677,831,764,791]" degrees="104" direction="east" minutes="53.475" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="104.89125">104°53.475′E</geoCoordinate>
|
||
(locality code:
|
||
<collectionCode box="[1003,1038,763,789]" pageId="6" pageNumber="259">JG</collectionCode>
|
||
7), leg. Grego, J.,
|
||
<date box="[1232,1432,763,789]" pageId="6" pageNumber="259" value="2017-02-13">13 February 2017</date>
|
||
, JG/4;
|
||
<collectingRegion box="[220,487,799,825]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouane Province</collectingRegion>
|
||
, Nhommalath, bridge on road no. 12,
|
||
<quantity box="[920,1013,799,825]" metricMagnitude="4" metricUnit="m" metricValue="4.34" pageId="6" pageNumber="259" unit="km" value="43.4">43.4 km</quantity>
|
||
towards Nongchan, southern edge of Ban Khamhé, right side of the road,
|
||
<quantity box="[579,653,835,861]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="259" unit="m" value="210.0">210 m</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate box="[731,877,836,863]" degrees="17" direction="north" minutes="34.743" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.57905">17°34.743′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[891,1044,836,863]" degrees="105" direction="east" minutes="27.515" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="105.45858">105°27.515′E</geoCoordinate>
|
||
(locality code: 2019/108), leg. A. Hunyadi,
|
||
<date box="[263,487,871,897]" pageId="6" pageNumber="259" value="2019-09-30">30 September 2019</date>
|
||
, HA/10;
|
||
<collectingRegion box="[593,861,871,897]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouane Province</collectingRegion>
|
||
,
|
||
<quantity box="[874,946,871,897]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="259" unit="km" value="40.0">40 km</quantity>
|
||
from the centre of Thakhek, Mahaxay,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="259" unit="km" value="2.3">2.3 km</quantity>
|
||
southeast from Ban Na Coc, limestone wall,
|
||
<quantity box="[698,769,906,933]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="259" unit="m" value="155.0">155 m</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate box="[842,988,908,935]" degrees="17" direction="north" minutes="25.957" orientation="latitude" pageId="6" pageNumber="259" precision="1" value="17.432617">17°25.957′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[999,1153,908,935]" degrees="105" direction="east" minutes="09.669" orientation="longitude" pageId="6" pageNumber="259" precision="1" value="105.16115">105°09.669′E</geoCoordinate>
|
||
(locality code: 2019/95), leg. A. Hunyadi,
|
||
<date box="[350,575,943,969]" pageId="6" pageNumber="259" value="2019-09-27">27 September 2019</date>
|
||
, HA/4 (
|
||
<figureCitation box="[668,760,943,969]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1N</figureCitation>
|
||
); Bam Na Ka Yak (Nhoum) (exact locality unknown), leg. Saurin between 1950 and 1970,
|
||
<collectionCode box="[506,649,979,1005]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27296/1 shell; Nhommanth (exact locality unknown), leg. Saurin,
|
||
<collectionCode box="[151,293,1014,1041]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27295/31 shells (shells not keeled, identical to
|
||
<collectionCode box="[881,1024,1014,1041]" pageId="6" pageNumber="259">MNHN-IM-</collectionCode>
|
||
2012-27325).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="259" type="diagnosis">
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,322,1050,1076]" pageId="6" pageNumber="259">Diagnosis.</emphasis>
|
||
A very large alycaeid species with dominant, usually keeled last whorl (shell wider than high), finely reticulate protoconch, finely wrinkled/scaly R1, a very long R2 having ribs forming many connections with each other by calcareous projections, and a long R3 with low swelling.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1761]" box="[199,470,1158,1185]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,345,1158,1184]" pageId="6" pageNumber="259">Operculum:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="259" type="description">
|
||
<paragraph blockId="6.[151,1437,150,1761]" box="[199,1136,1194,1221]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,484,1194,1221]" pageId="6" pageNumber="259">Measurements (in mm).</emphasis>
|
||
D = 8.9–11.3, H = 6.4–8.9 (multiple populations, n = 27).
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,480,1230,1256]" pageId="6" pageNumber="259">Variation in specimens.</emphasis>
|
||
The examined shells show variability in terms of shell size and the sharpness of the keel.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="259" type="diagnosis">
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,466,1302,1328]" pageId="6" pageNumber="259">Differential diagnosis.</emphasis>
|
||
Differs from
|
||
<taxonomicName authorityName="Venmans" authorityYear="1956" box="[632,755,1303,1329]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="eydouxi">
|
||
<emphasis box="[632,755,1303,1329]" italics="true" pageId="6" pageNumber="259">A. eydouxi</emphasis>
|
||
</taxonomicName>
|
||
by the web-like structure of the R2 ribs, and the roughly sculptured protoconch. Moreover, most populations of
|
||
<taxonomicName authorityName="Maassen" authorityYear="2006" box="[783,938,1339,1365]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="rolfbrandti">
|
||
<emphasis box="[783,938,1339,1365]" italics="true" pageId="6" pageNumber="259">A. rolfbrandti</emphasis>
|
||
</taxonomicName>
|
||
have strongly keeled body whorl. See also under
|
||
<taxonomicName authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[222,332,1375,1401]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="259" phylum="Mollusca" rank="species" species="goliath" status="n. sp.">
|
||
<emphasis box="[222,332,1375,1401]" italics="true" pageId="6" pageNumber="259">A. goliath</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[339,402,1375,1401]" pageId="6" pageNumber="259">
|
||
<taxonomicNameLabel box="[339,402,1375,1401]" pageId="6" pageNumber="259" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="259" type="distribution">
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,353,1410,1436]" pageId="6" pageNumber="259">Distribution.</emphasis>
|
||
This species is known form the limestone region on the left bank of the Mekong River, east of Nakhon Phanom (
|
||
<collectingCountry box="[354,412,1447,1473]" name="Laos" pageId="6" pageNumber="259">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion box="[423,679,1447,1473]" country="Laos" name="Khammouan" pageId="6" pageNumber="259">Khammouan Province</collectingRegion>
|
||
). The furthest distance between the known populations is ca.
|
||
<quantity box="[1365,1436,1447,1473]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="259" unit="km" value="70.0">70 km</quantity>
|
||
(
|
||
<figureCitation box="[159,229,1483,1509]" captionStart="FIGURE 4" captionStartId="5.[152,255,1163,1187]" captionTargetBox="[151,1435,417,1139]" captionTargetId="figure-130@5.[151,1435,417,1139]" captionTargetPageId="5" captionText="FIGURE 4. Distribution of Alycaeus Baird, 1850 species in central Laos. Yellow circles: Alycaeus rolfbrandti Maassen, 2006; Red triangle: A. goliath n. sp. The map was made using Google Earth Pro." figureDoi="http://doi.org/10.5281/zenodo.7687426" httpUri="https://zenodo.org/record/7687426/files/figure.png" pageId="6" pageNumber="259">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="259" type="discussion">
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
<emphasis bold="true" box="[199,315,1519,1545]" pageId="6" pageNumber="259">Remarks.</emphasis>
|
||
This species does not require redescription due to the accurate original description. Only one addition has to be made: the R2 ribs either form projections to join each other, or they are regularly undulated, which results in a wire fence-like surface.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1761]" pageId="6" pageNumber="259">
|
||
The majority of the populations examined have typically keeled last whorl, although specimens of some examined populations have a bluntly keel, i.e., the body whorl is almost rounded (JG4, JG7 = 7L07), while it is rounded in one population (2019/95) (see
|
||
<figureCitation box="[636,762,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[152,255,1857,1881]" captionTargetBox="[151,1436,181,1834]" captionTargetId="figure-31@2.[151,1436,181,1834]" captionTargetPageId="2" captionText="FIGURE 1. Shells of Alycaeus Baird, 1850 species.A–D: Alycaeus goliath n. sp. (holotype); E–H: Alycaeus eydouxi Venmans, 1956 (SMF 109290); I–S: Alycaeus rolfbrandti Maassen, 2006. I–L: SMF 262540 = part of the original sample, M: 25L07, N: 2019/95 (syntopic with A. goliath n. sp.); O: 7L07, P: 6L07, Q: 9L07, R: 11L07, S: 12L07. All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687420" httpUri="https://zenodo.org/record/7687420/files/figure.png" pageId="6" pageNumber="259">Fig. 1M–S</figureCitation>
|
||
). I found no other differences between these morphotypes, therefore they are not separated at any taxonomic levels.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |