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<document id="0B32777E86EAEE5228F6A69C866605FF" ID-DOI="10.37520/aemnp.2024.022" ID-ISSN="1804-6487" ID-Zenodo-Dep="14655471" ID-ZooBank="F8D54B80-AB41-4D03-A03B-F084FFD34890" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736956398870" checkinUser="felipe" docAuthor="Montoya, Augusto León" docDate="2024" docId="8A0687C2FFB1FFFCCDFBA64CFE56FA16" docLanguage="en" docName="ActaEntMusNatPra.64.2.307-325.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 64 (2)" docSource="https://doi.org/10.37520/aemnp.2024.022" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Lycopale radioheadi Montoya, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="318" masterDocId="763FFFBAFFB9FFF7CE35A378FF9BFFE3" masterDocTitle="Description of two new species of Lycopale (Diptera: Syrphidae) from the northwestern Tropical Andes hotspot, with the redescription of Lycopale magnifica" masterLastPageNumber="325" masterPageNumber="307" pageNumber="315" updateTime="1736957796906" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="C611145210EE78F33D66F5950837A2CC">Description of two new species of Lycopale (Diptera: Syrphidae) from the northwestern Tropical Andes hotspot, with the redescription of Lycopale magnifica</mods:title>
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<mods:namePart id="6965C442318670C46E8E93115C76E89C">Montoya, Augusto León</mods:namePart>
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<mods:title id="3013D7E95E5404FBCBDA01C537A9192D">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E6A69935A4ACB29839D0354C1C2B427A">2024</mods:date>
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<mods:title id="08288168B56963802F1908F67ED1D368">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="8E09BBB14750C7BFC6DB2461167C3C7A">2024-10-24</mods:number>
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<mods:classification id="C0E4247B1CEB82317602BB302F89C452">journal article</mods:classification>
<mods:identifier id="D4517E0B8FC9B43E34EEE025CFBD48DE" type="DOI">10.37520/aemnp.2024.022</mods:identifier>
<mods:identifier id="A57D3D38FC94825CABCEEF8B2DC663B6" type="ISSN">1804-6487</mods:identifier>
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<treatment id="8A0687C2FFB1FFFCCDFBA64CFE56FA16" LSID="urn:lsid:plazi:treatment:8A0687C2FFB1FFFCCDFBA64CFE56FA16" httpUri="http://treatment.plazi.org/id/8A0687C2FFB1FFFCCDFBA64CFE56FA16" lastPageId="11" lastPageNumber="318" pageId="8" pageNumber="315">
<subSubSection id="4AB5655FFFB1FFFFCDFBA64CFA91FAAF" box="[974,1290,1332,1356]" pageId="8" pageNumber="315" type="nomenclature">
<paragraph id="021036D4FFB1FFFFCDFBA64CFA91FAAF" blockId="8.[974,1290,1332,1356]" box="[974,1290,1332,1356]" pageId="8" pageNumber="315">
<heading id="595881B8FFB1FFFFCDFBA64CFA91FAAF" box="[974,1290,1332,1356]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="315" reason="3">
<emphasis id="30DBEAC6FFB1FFFFCDFBA64CFA91FAAF" bold="true" box="[974,1290,1332,1356]" pageId="8" pageNumber="315">
<taxonomicName id="C5AF4D57FFB1FFFFCDFBA64CFB35FAAF" authorityName="Montoya" authorityYear="2024" box="[974,1198,1332,1356]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="315" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB1FFFFCDFBA64CFB35FAAF" bold="true" box="[974,1198,1332,1356]" italics="true" pageId="8" pageNumber="315">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB1FFFFCA81A64DFA91FAAF" box="[1204,1290,1333,1356]" pageId="8" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB1FFFFCA25A62BFB5CFA85" box="[1040,1223,1363,1382]" pageId="8" pageNumber="315" type="description">
<paragraph id="021036D4FFB1FFFFCA25A62BFB5CFA85" blockId="8.[826,1438,1363,2083]" box="[1040,1223,1363,1382]" pageId="8" pageNumber="315">
(
<figureCitation id="9A942A51FFB1FFFFCA23A62BFBF8FA85" box="[1046,1123,1363,1382]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[149,180,1672,1691]" captionStartId-1="10.[151,182,1207,1226]" captionTargetBox-0="[150,1436,190,1657]" captionTargetBox-1="[151,1436,188,1190]" captionTargetId-0="figure-16@9.[150,1436,189,1658]" captionTargetId-1="figure-13@10.[151,1436,187,1190]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." captionText-1="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14655491" figureDoi-1="http://doi.org/10.5281/zenodo.14655493" httpUri-0="https://zenodo.org/record/14655491/files/figure.png" httpUri-1="https://zenodo.org/record/14655493/files/figure.png" pageId="8" pageNumber="315">Figs 7–8</figureCitation>
,
<figureCitation id="9A942A51FFB1FFFFCA59A62BFB18FA85" box="[1132,1155,1363,1382]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="8" pageNumber="315">9h</figureCitation>
,
<figureCitation id="9A942A51FFB1FFFFCAB9A62BFB5AFA85" box="[1164,1217,1363,1382]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[151,182,1241,1260]" captionStartId-1="17.[151,182,1002,1021]" captionTargetBox-0="[151,1436,189,1220]" captionTargetBox-1="[152,1417,197,958]" captionTargetId-0="figure-402@16.[151,1436,189,1233]" captionTargetId-1="figure-598@17.[151,1436,189,977]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 10. Distributional map of Lycopale species." captionText-1="Fig. 11. A – detail of distributional map of Lycopale in Colombia and Ecuador. B–C – study area in: B – Paramo Santa Inés Belmira; C – Paramo Sonsón. Sampling sites where the type material of the new species was collected (see MONTOYA et al. 2021)." figureDoi-0="http://doi.org/10.5281/zenodo.14655499" figureDoi-1="http://doi.org/10.5281/zenodo.14655501" httpUri-0="https://zenodo.org/record/14655499/files/figure.png" httpUri-1="https://zenodo.org/record/14655501/files/figure.png" pageId="8" pageNumber="315">10–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB1FFFECD09A60EFD2BF8B8" lastPageId="9" lastPageNumber="316" pageId="8" pageNumber="315" type="materials_examined">
<paragraph id="021036D4FFB1FFFECD09A60EFDA5F8D0" blockId="8.[826,1438,1363,2083]" lastBlockId="9.[151,761,1771,2075]" lastPageId="9" lastPageNumber="316" pageId="8" pageNumber="315">
<emphasis id="30DBEAC6FFB1FFFFCD09A60EFC53FA6A" bold="true" box="[828,968,1398,1417]" pageId="8" pageNumber="315">Type material.</emphasis>
<materialsCitation id="B2C73C89FFB1FFFFCDE4A60EFBCAF9ED" collectingDate="2018-09-01" collectingDateMax="2018-09-12" collectingDateMin="2018-09-01" collectionCode="CEUA" collectorName="A. M. Echeverry &amp; A. L. Montoya" elevation="3034" location="Original" pageId="8" pageNumber="315" specimenCode="CEUA 103529" specimenCount="2" stateProvince="Antioquia" typeStatus="holotype">
<typeStatus id="DD148876FFB1FFFFCDE4A60EFBB5FA6A" box="[977,1070,1398,1417]" pageId="8" pageNumber="315" type="holotype">HOLOTYPE</typeStatus>
: Adult J, pinned.
<location id="0770600FFFB1FFFFCAD5A60EFAB1FA6A" LSID="urn:lsid:plazi:treatment:8A0687C2FFB1FFFCCDFBA64CFE56FA16:0770600FFFB1FFFFCAD5A60EFAB1FA6A" box="[1248,1322,1398,1417]" name="Original" pageId="8" pageNumber="315" stateProvince="Antioquia">Original</location>
label: “COLOMBIA,
<collectingRegion id="C06BF836FFB1FFFFCDAEA6E8FC68FA40" box="[923,1011,1424,1443]" country="Colombia" name="Antioquia" pageId="8" pageNumber="315">Antioquia</collectingRegion>
, Sonsón, Vereda Nori Cerro Nori ” / “ 5,812861, -75,268444,
<quantity id="C5579B31FFB1FFFFCD9CA6D3FBB9FA5D" box="[937,1058,1451,1470]" metricMagnitude="3" metricUnit="m" metricValue="3.0335" metricValueMax="3.045" metricValueMin="3.022" pageId="8" pageNumber="315" unit="m" value="3033.5" valueMax="3045.0" valueMin="3022.0">
<elevation id="8982D1E7FFB1FFFFCD9CA6D3FBB9FA5D" box="[937,1058,1451,1470]" metricMagnitude="3" metricUnit="m" metricValue="3.0335" metricValueMax="3.045" metricValueMin="3.022" pageId="8" pageNumber="315" unit="m" value="3033.5" valueMax="3045.0" valueMin="3022.0">3022–3045 m</elevation>
</quantity>
[eters], Paramo, Red entomológica [Net] ” / “
<date id="76111014FFB1FFFFCD71A6BEFC2EFA3A" box="[836,949,1478,1497]" pageId="8" pageNumber="315" value="2018-09-01" valueMax="2018-09-12" valueMin="2018-09-01">
<collectingDate id="6655E9FCFFB1FFFFCD71A6BEFC2EFA3A" box="[836,949,1478,1497]" pageId="8" pageNumber="315" value="2018-09-01" valueMax="2018-09-12" valueMin="2018-09-01">1–12.ix.2018</collectingDate>
</date>
,
<collectorName id="AF5A5302FFB1FFFFCD8EA6BEFBDFFA3A" box="[955,1092,1478,1497]" pageId="8" pageNumber="315">A.M.Echeverry</collectorName>
;
<collectorName id="AF5A5302FFB1FFFFCA7FA6BEFB5FFA3A" box="[1098,1220,1478,1497]" pageId="8" pageNumber="315">A.L.Montoya</collectorName>
”.“
<typeStatus id="DD148876FFB1FFFFCAECA6BEFAD1FA3A" box="[1241,1354,1478,1497]" pageId="8" pageNumber="315" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="C5AF4D57FFB1FFFFCB6DA6BEFC36FA10" authorityName="Montoya" authorityYear="2024" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="315" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB1FFFFCB6DA6BEFC36FA10" italics="true" pageId="8" pageNumber="315">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB1FFFFCD84A698FC68FA10" box="[945,1011,1504,1523]" pageId="8" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
/ Montoya 2023” [red, handwritten except the first line] (
<specimenCode id="52099EAFFFB1FFFFCD5BA683FC6BF9ED" box="[878,1008,1531,1550]" collectionCode="CEUA" pageId="8" pageNumber="315">CEUA 103529</specimenCode>
, dissected)
</materialsCitation>
.
<typeStatus id="DD148876FFB1FFFFCA6CA683FB29F9ED" box="[1113,1202,1531,1550]" pageId="8" pageNumber="315" type="paratype">PARATYPES</typeStatus>
:
<emphasis id="30DBEAC6FFB1FFFFCA88A683FAA6F9ED" bold="true" box="[1213,1341,1531,1550]" pageId="8" pageNumber="315">
<collectingCountry id="7AB87644FFB1FFFFCA88A683FAA2F9ED" box="[1213,1337,1531,1550]" name="Colombia" pageId="8" pageNumber="315">COLOMBIA</collectingCountry>
:
</emphasis>
<collectingRegion id="C06BF836FFB1FFFFCB77A683FA01F9ED" box="[1346,1434,1531,1550]" country="Colombia" name="Antioquia" pageId="8" pageNumber="315">Antioquia</collectingRegion>
, Belmira, Paramo Santa Inés, Vereda Río Arriba, Sector Laguna de Sabanas,
<geoCoordinate id="679B5013FFB1FFFFCD43A548FC53F9A0" box="[886,968,1584,1603]" degrees="6.640931" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="6.640931">6.640931</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDFAA548FBA8F9A0" box="[975,1075,1584,1603]" degrees="75.665335" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.66534">-75.665335</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA0FA548FB2AF9A0" box="[1082,1201,1584,1603]" metricMagnitude="3" metricUnit="m" metricValue="2.9" metricValueMax="2.95" metricValueMin="2.85" pageId="8" pageNumber="315" unit="m" value="2900.0" valueMax="2950.0" valueMin="2850.0">2850–2950 m</quantity>
, Bosque, Red entomológica [Net],
<date id="76111014FFB1FFFFCD46A533FC6AF9BD" box="[883,1009,1611,1630]" pageId="8" pageNumber="315" value="2017-09-16" valueMax="2017-09-27" valueMin="2017-09-16">16–27.ix.2017</date>
, A. L. Montoya (1 J,
<collectionCode id="64BEAE11FFB1FFFFCA9EA533FB7FF9BD" box="[1195,1252,1611,1630]" pageId="8" pageNumber="315">CEUA</collectionCode>
103258); … Paramo, flying around
<taxonomicName id="C5AF4D57FFB1FFFFCD80A51EFBF1F99A" box="[949,1130,1638,1657]" class="Magnoliopsida" family="Asteraceae" genus="Taraxacum" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="315" phylum="Tracheophyta" rank="species" species="officinale">
<emphasis id="30DBEAC6FFB1FFFFCD80A51EFBF1F99A" box="[949,1130,1638,1657]" italics="true" pageId="8" pageNumber="315">Taraxacum officinale</emphasis>
</taxonomicName>
(L.) Weber ex F.H.Wigg., Prim. Fl. Holsat.,
<date id="76111014FFB1FFFFCDB4A5F8FB96F970" box="[897,1037,1664,1683]" pageId="8" pageNumber="315" value="2017-03-25" valueMax="2017-04-05" valueMin="2017-03-25">25.iii.–5.iv.2017</date>
,A. L. Montoya; C. Rodríguez; J. D.Carmona (1 J,
<collectionCode id="64BEAE11FFB1FFFFCD63A5E3FC14F94D" box="[854,911,1691,1710]" pageId="8" pageNumber="315">CEUA</collectionCode>
93560); Cabaña Cabildo Verde, El Morro-Alto de La Gallina,
<geoCoordinate id="679B5013FFB1FFFFCD09A5CEFC15F92A" box="[828,910,1718,1737]" degrees="6.632639" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="6.632639">6.632639</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDA2A5CEFC60F92A" box="[919,1019,1718,1737]" degrees="75.645267" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.64526">-75.645267</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA30A5CEFBE5F92A" box="[1029,1150,1718,1737]" metricMagnitude="3" metricUnit="m" metricValue="3.185" metricValueMax="3.2" metricValueMin="3.17" pageId="8" pageNumber="315" unit="m" value="3185.0" valueMax="3200.0" valueMin="3170.0">3170–3200 m</quantity>
, Paramo, Trampa de Emergencia [Emergency trap], Decomposing wood log,
<date id="76111014FFB1FFFFCA83A5A8FAA9F900" box="[1206,1330,1744,1763]" pageId="8" pageNumber="315" value="2016-04-12" valueMax="2016-04-14" valueMin="2016-04-12">12–14.iv.2016</date>
, A. L. Montoya; J. Torres; J. D. Carmona (
<specimenCount id="14A9FD5DFFB1FFFFCA77A593FBC5F91D" box="[1090,1118,1765,1790]" count="1" pageId="8" pageNumber="315" type="female">1 ♀</specimenCount>
,
<collectionCode id="64BEAE11FFB1FFFFCA5CA593FB39F91D" box="[1129,1186,1771,1790]" pageId="8" pageNumber="315">CEUA</collectionCode>
95143); Bello, San Félix, Las Baldías,
<geoCoordinate id="679B5013FFB1FFFFCDB3A47EFC56F8FA" box="[902,973,1798,1817]" degrees="6.33931" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="6.33931">6.33931</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDE0A47EFBB4F8FA" box="[981,1071,1798,1817]" degrees="75.60801" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.60801">-75.60801</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA03A47EFB35F8FA" box="[1078,1198,1798,1817]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.15" metricValueMin="2.95" pageId="8" pageNumber="315" unit="m" value="3050.0" valueMax="3150.0" valueMin="2950.0">2950–3150 m</quantity>
, Paramo, Red entomológica [Net],
<date id="76111014FFB1FFFFCD46A458FC50F8D0" box="[883,971,1824,1843]" pageId="8" pageNumber="315" value="2013-02-17">17.ii.2013</date>
, A. L. Montoya (1 J,
<collectionCode id="64BEAE11FFB1FFFFCABDA458FB5AF8D0" box="[1160,1217,1824,1843]" pageId="8" pageNumber="315">CEUA</collectionCode>
73227); Bello, San Félix, Las Baldías,
<geoCoordinate id="679B5013FFB1FFFFCD91A443FC71F8AD" box="[932,1002,1851,1870]" degrees="6.33931" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="6.33931">6.33931</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDC4A443FBDCF8AD" box="[1009,1095,1851,1870]" degrees="75.60801" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.60801">-75.60801</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA7BA443FB59F8AD" box="[1102,1218,1851,1870]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.15" metricValueMin="2.95" pageId="8" pageNumber="315" unit="m" value="3050.0" valueMax="3150.0" valueMin="2950.0">2950–3150 m</quantity>
, Paramo,Red entomológica [Net],
<date id="76111014FFB1FFFFCDBEA42EFC46F88A" box="[907,989,1878,1897]" pageId="8" pageNumber="315" value="2015-05-23">23.v.2015</date>
(1J,
<collectionCode id="64BEAE11FFB1FFFFCA3BA42EFBDDF88A" box="[1038,1094,1878,1897]" pageId="8" pageNumber="315">CEUA</collectionCode>
87185); Sonsón,Vereda Nori Cerro Nori,
<geoCoordinate id="679B5013FFB1FFFFCD09A408FC15F860" box="[828,910,1904,1923]" degrees="5.809667" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="5.809667">5.809667</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDACA408FC65F860" box="[921,1022,1904,1923]" degrees="75.269028" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.26903">-75.269028</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA3CA408FB1FF860" box="[1033,1156,1904,1923]" metricMagnitude="3" metricUnit="m" metricValue="2.8465" metricValueMax="2.853" metricValueMin="2.84" pageId="8" pageNumber="315" unit="m" value="2846.5" valueMax="2853.0" valueMin="2840.0">2840–2853 m</quantity>
, Bosque, Trampa malaise suelo [Malaise trap ground],
<date id="76111014FFB1FFFFCDC9A4F3FB10F87D" box="[1020,1163,1931,1950]" pageId="8" pageNumber="315" value="2018-03-31" valueMax="2018-07-07" valueMin="2018-03-31">31.iii–7.vii.2018</date>
,A.L. Montoya; J. P.Carmona (
<specimenCount id="14A9FD5DFFB1FFFFCBA6A4F3FCD1F85B" count="1" pageId="8" pageNumber="315" type="female">1 ♀</specimenCount>
,
<collectionCode id="64BEAE11FFB1FFFFCD60A4DEFC15F85A" box="[853,910,1958,1977]" pageId="8" pageNumber="315">CEUA</collectionCode>
103427); Sonsón, Vereda San Francisco, Cerro Las Palomas A,
<geoCoordinate id="679B5013FFB1FFFFCD09A4B8FC15F830" box="[828,910,1984,2003]" degrees="5.725443" direction="north" orientation="latitude" pageId="8" pageNumber="315" precision="1" value="5.725443">5.725443</geoCoordinate>
,
<geoCoordinate id="679B5013FFB1FFFFCDA3A4B8FC62F830" box="[918,1017,1984,2003]" degrees="75.263638" direction="west" orientation="longitude" pageId="8" pageNumber="315" precision="1" value="-75.26364">-75.263638</geoCoordinate>
,
<quantity id="C5579B31FFB1FFFFCA37A4B8FBE2F830" box="[1026,1145,1984,2003]" metricMagnitude="3" metricUnit="m" metricValue="2.9535" metricValueMax="2.98" metricValueMin="2.927" pageId="8" pageNumber="315" unit="m" value="2953.5" valueMax="2980.0" valueMin="2927.0">2927–2980 m</quantity>
, Potrero, Red entomológica [Net],
<date id="76111014FFB1FFFFCD09A4A3FC33F80D" box="[828,936,2011,2030]" pageId="8" pageNumber="315" value="2019-02-02" valueMax="2019-02-19" valueMin="2019-02-02">2–19.ii.2019</date>
, J. Sauceda; I. Ceballos; M.I. Salinas; D.Uchima &amp;A.Mejía (1 J,
<collectionCode id="64BEAE11FFB1FFFFCD5FA48EFC38F7EA" box="[874,931,2038,2057]" pageId="8" pageNumber="315">CEUA</collectionCode>
114094, ZFMK-DNA-FD119585551, GenBank PP979582.
<emphasis id="30DBEAC6FFB1FFFFCD09AB68FC35F7C0" bold="true" box="[828,942,2064,2083]" pageId="8" pageNumber="315">
<collectingCountry id="7AB87644FFB1FFFFCD09AB68FC31F7C0" box="[828,938,2064,2083]" name="Ecuador" pageId="8" pageNumber="315">ECUADOR</collectingCountry>
:
</emphasis>
<collectingRegion id="C06BF836FFB1FFFFCD84AB68FBABF7C0" box="[945,1072,2064,2083]" country="Ecuador" name="Napo" pageId="8" pageNumber="315">Napo Province</collectingRegion>
,Reserva Ecológica Antisana, Jumandy Inca trail, from Virgen de Guacamayos to Sector Sarayacu, from
<geoCoordinate id="679B5013FFB0FFFECCAEA593FD6EF91D" box="[667,757,1771,1790]" degrees="0.625461" direction="south" orientation="latitude" pageId="9" pageNumber="316" precision="1" value="-0.625461">-0.625461</geoCoordinate>
,
<geoCoordinate id="679B5013FFB0FFFECEA2A47EFF62F8FA" box="[151,249,1798,1817]" degrees="77.841658" direction="west" orientation="longitude" pageId="9" pageNumber="316" precision="1" value="-77.84166">-77.841658</geoCoordinate>
to
<geoCoordinate id="679B5013FFB0FFFECF23A47EFEF4F8FA" box="[278,367,1798,1817]" degrees="0.692165" direction="south" orientation="latitude" pageId="9" pageNumber="316" precision="1" value="-0.692165">-0.692165</geoCoordinate>
,
<geoCoordinate id="679B5013FFB0FFFECF4DA47EFE47F8FA" box="[376,476,1798,1817]" degrees="77.820256" direction="west" orientation="longitude" pageId="9" pageNumber="316" precision="1" value="-77.82026">-77.820256</geoCoordinate>
,
<quantity id="C5579B31FFB0FFFECFD0A47EFDC3F8FA" box="[485,600,1798,1817]" metricMagnitude="3" metricUnit="m" metricValue="1.7825" metricValueMax="2.26" metricValueMin="1.305" pageId="9" pageNumber="316" unit="m" value="1782.5" valueMax="2260.0" valueMin="1305.0">1305–2260m</quantity>
,
<date id="76111014FFB0FFFECC54A47EFD2FF8FA" box="[609,692,1798,1817]" pageId="9" pageNumber="316" value="2019-09-02">2.ix.2019</date>
, leg. X. Mengual (
<specimenCount id="14A9FD5DFFB0FFFECEC4A458FE94F8D0" box="[241,271,1818,1843]" count="1" pageId="9" pageNumber="316" type="female">1 ♀</specimenCount>
, ZFMK-DIP-00068601, INABIO).
</paragraph>
<caption id="56D0665CFFB0FFFECEA0A5F0FA88F955" ID-DOI="http://doi.org/10.5281/zenodo.14655491" ID-Zenodo-Dep="14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316" startId="9.[149,180,1672,1691]" targetBox="[150,1436,190,1657]" targetPageId="9" targetType="figure">
<paragraph id="021036D4FFB0FFFECEA0A5F0FA88F955" blockId="9.[149,1438,1672,1718]" pageId="9" pageNumber="316">
Fig. 7.
<taxonomicName id="C5AF4D57FFB0FFFECEE6A5F0FE18F978" authorityName="Montoya" authorityYear="2024" box="[211,387,1672,1691]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB0FFFECEE6A5F0FE18F978" box="[211,387,1672,1691]" italics="true" pageId="9" pageNumber="316">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB0FFFECFBDA5F0FE50F978" box="[392,459,1672,1691]" pageId="9" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="021036D4FFB0FFFECEA2A43CFD2BF8B8" blockId="9.[151,761,1771,2075]" box="[151,688,1860,1883]" pageId="9" pageNumber="316">
<emphasis id="30DBEAC6FFB0FFFECEA2A43CFF6BF8B8" bold="true" box="[151,240,1860,1883]" pageId="9" pageNumber="316">Length.</emphasis>
Body
<quantity id="C5579B31FFB0FFFECF03A43CFE4EF8B8" box="[310,469,1860,1883]" metricMagnitude="-2" metricUnit="m" metricValue="1.0399999999999998" metricValueMax="1.06" metricValueMin="1.02" pageId="9" pageNumber="316" unit="mm" value="10.399999999999999" valueMax="10.6" valueMin="10.2">10.2–10.6 mm</quantity>
; wing
<quantity id="C5579B31FFB0FFFECC2EA43CFD37F8B8" box="[539,684,1860,1883]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="10.8" metricValueMin="7.4" pageId="9" pageNumber="316" unit="mm" value="9.1" valueMax="10.8" valueMin="7.4">7.4–10.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB0FFFECEA2A41CFB90F801" pageId="9" pageNumber="316" type="diagnosis">
<paragraph id="021036D4FFB0FFFECEA2A41CFB90F801" blockId="9.[151,761,1771,2075]" lastBlockId="9.[828,1437,1771,2082]" pageId="9" pageNumber="316">
<emphasis id="30DBEAC6FFB0FFFECEA2A41CFE94F898" bold="true" box="[151,271,1892,1915]" pageId="9" pageNumber="316">Diagnosis.</emphasis>
<taxonomicName id="C5AF4D57FFB0FFFECF2FA41CFD9AF898" authorityName="Montoya" authorityYear="2024" box="[282,513,1892,1915]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB0FFFECF2FA41CFD9AF898" box="[282,513,1892,1915]" italics="true" pageId="9" pageNumber="316">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB0FFFECC39A41CFDFFF898" box="[524,612,1892,1915]" pageId="9" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="C5AF4D57FFB0FFFECEA2A4FCFEAEF878" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[151,309,1924,1947]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB0FFFECEA2A4FCFEAEF878" box="[151,309,1924,1947]" italics="true" pageId="9" pageNumber="316">Lycopale lizeri</emphasis>
</taxonomicName>
but can be separated from it by facial vitta and gena yellow (
<figureCitation id="9A942A51FFB0FFFECF62A4DCFE5FF858" box="[343,452,1956,1979]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7a–b</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECFE5A4DCFE70F858" box="[464,491,1956,1979]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
); antenna orange, except for dorsally brown postpedicel (
<figureCitation id="9A942A51FFB0FFFECFD6A4BCFDD6F838" box="[483,589,1988,2011]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7a–b</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECC62A4BCFDE9F838" box="[599,626,1988,2011]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
); postpronotum black pruinose (
<figureCitation id="9A942A51FFB0FFFECF46A49CFE7BF818" box="[371,480,2020,2043]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECFD9A49CFD9CF818" box="[492,519,2020,2043]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECC26A49CFDAAF818" box="[531,561,2020,2043]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="9" pageNumber="316">9h</figureCitation>
); scutum with two submedial and one medial grayish white vittae reaching 3/4 of scutum length (
<figureCitation id="9A942A51FFB0FFFECA05A593FB04F8E1" box="[1072,1183,1771,1794]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECA99A593FB53F8E1" box="[1196,1224,1771,1794]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECAE0A593FB6FF8E1" box="[1237,1268,1771,1794]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="9" pageNumber="316">9h</figureCitation>
); pre-scutellum extensively black, without vitta (
<figureCitation id="9A942A51FFB0FFFECA94A473FB68F8C1" box="[1185,1267,1803,1826]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="9" pageNumber="316">Fig. 9h</figureCitation>
); wing hyaline, except for light orange brown anteriorly (
<figureCitation id="9A942A51FFB0FFFECACFA453FAFCF8A1" box="[1274,1383,1835,1858]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECB46A453FA15F8A1" box="[1395,1422,1835,1858]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
); metafemur bright orange, except for light reddish brown on apical 1/2 (
<figureCitation id="9A942A51FFB0FFFECDE7A413FBA7F861" box="[978,1084,1899,1922]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7a–c</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECA72A413FBF9F861" box="[1095,1122,1899,1922]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
); second tergum orange, with basal maculae occupying the full lateral width and extending medially into a triangular basomedial black vitta (
<figureCitation id="9A942A51FFB0FFFECD76A4B3FC2BF801" box="[835,944,1995,2018]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECD8FA4B3FC4DF801" box="[954,982,1995,2018]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECDD4A4B3FC64F801" box="[993,1023,1995,2018]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="9" pageNumber="316">9h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB0FFFDCD09A493FBACF89E" lastPageId="10" lastPageNumber="317" pageId="9" pageNumber="316" type="description">
<paragraph id="021036D4FFB0FFFDCD09A493FE26FA1E" blockId="9.[828,1437,1771,2082]" lastBlockId="10.[151,761,1318,2077]" lastPageId="10" lastPageNumber="317" pageId="9" pageNumber="316">
<emphasis id="30DBEAC6FFB0FFFECD09A493FB8BF7E1" bold="true" box="[828,1040,2026,2050]" pageId="9" pageNumber="316">
Description.
<emphasis id="30DBEAC6FFB0FFFECDE4A492FB8BF7E1" box="[977,1040,2026,2050]" italics="true" pageId="9" pageNumber="316">Male.</emphasis>
</emphasis>
Head (
<figureCitation id="9A942A51FFB0FFFECA53A493FB4EF7E1" box="[1126,1237,2027,2050]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="9" pageNumber="316">Figs 7a–b</figureCitation>
,
<figureCitation id="9A942A51FFB0FFFECAD1A493FA9BF7E1" box="[1252,1280,2027,2050]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="9" pageNumber="316">8a</figureCitation>
). Face orange yellow with broad orange medial vitta, yellow pilose; gena orange, yellow pilose and pruinose; frontal triangle brown, black and yellow pilose; vertical triangle black, black pilose; eyes bare, narrowly dichoptic, separated from eye margin by at least ocellus width; occiput white pruinose and pilose, except for a few black piles dorsally. Antenna: orange, except for postpedicel brown on dorsal 1/3; arista light yellow and pubescent.
</paragraph>
<caption id="56D0665CFFB3FFFDCEA2A7CFFD7EFB06" ID-DOI="http://doi.org/10.5281/zenodo.14655493" ID-Zenodo-Dep="14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317" startId="10.[151,182,1207,1226]" targetBox="[151,1436,188,1190]" targetPageId="10" targetType="figure">
<paragraph id="021036D4FFB3FFFDCEA2A7CFFD7EFB06" blockId="10.[151,1436,1207,1253]" pageId="10" pageNumber="317">
Fig. 8.
<taxonomicName id="C5AF4D57FFB3FFFDCEE1A7CFFE1FFB29" authorityName="Montoya" authorityYear="2024" box="[212,388,1207,1226]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB3FFFDCEE1A7CFFE1FFB29" box="[212,388,1207,1226]" italics="true" pageId="10" pageNumber="317">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB3FFFDCFBCA7CFFE57FB29" box="[393,460,1207,1226]" pageId="10" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="021036D4FFB3FFFDCE80A57EFD7BF91E" blockId="10.[151,761,1318,2077]" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCE80A57EFE99F9FE" box="[181,258,1542,1565]" italics="true" pageId="10" pageNumber="317">Thorax</emphasis>
(
<figureCitation id="9A942A51FFB3FFFDCF27A57EFEE4F9FE" box="[274,383,1542,1565]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="10" pageNumber="317">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCFBEA57EFE3DF9FE" box="[395,422,1542,1565]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">8a</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCF87A57EFE4BF9FE" box="[434,464,1542,1565]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="10" pageNumber="317">9h</figureCitation>
). Dull black; postpronotum yellowish pilose; scutum with two submedial and one medial grayish white vitta from base, reaching 3/4 scutum length, yellow and black pile intermixed; pleuron dark grey pruinose, yellow pilose, except for dorsally black pilose posterior anepisternum; scutellum orange, black pilose; calypter white, border brown, brown pilose marginally; plumula orange; halter pedicel white; capitulum white.
</paragraph>
<paragraph id="021036D4FFB3FFFDCE80A47EFDB8F8BE" blockId="10.[151,761,1318,2077]" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCE80A47EFF77F8FE" box="[181,236,1798,1821]" italics="true" pageId="10" pageNumber="317">Wing</emphasis>
(
<figureCitation id="9A942A51FFB3FFFDCEC9A47EFEF2F8FE" box="[252,361,1798,1821]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="10" pageNumber="317">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCF40A47EFE0AF8FE" box="[373,401,1798,1821]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">8a</figureCitation>
). Hyaline except for light orange brownish anteriorly; extensively microtrichose; tegula black pilose, basicosta orange pilose.
</paragraph>
<paragraph id="021036D4FFB3FFFDCE80A41EFBADFADE" blockId="10.[151,761,1318,2077]" lastBlockId="10.[828,1437,1318,2077]" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCE80A41EFF7CF89E" box="[181,231,1894,1917]" italics="true" pageId="10" pageNumber="317">Legs</emphasis>
(
<figureCitation id="9A942A51FFB3FFFDCEC0A41EFEC4F89E" box="[245,351,1894,1917]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="10" pageNumber="317">Figs 7a–c</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCF5CA41EFE1EF89E" box="[361,389,1894,1917]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">8a</figureCitation>
). Orange, orange pilose, coxae and trochanters dark grey pruinose and golden pilose; pro- and mesofemur golden pilose, except for black pilose on posterodorsal margin; metafemur bright orange, orange pilose, except for black pile ventrally, in addition to some black pile on apicodorsal 1/4; hind tibia with some black pile on basoventral 1/3.
</paragraph>
<paragraph id="021036D4FFB3FFFDCD6FA63EFC49F95E" blockId="10.[828,1437,1318,2077]" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCD6FA63EFC25FABE" box="[858,958,1350,1373]" italics="true" pageId="10" pageNumber="317">Abdomen</emphasis>
(
<figureCitation id="9A942A51FFB3FFFDCDFEA63EFBAEFABE" box="[971,1077,1350,1373]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="10" pageNumber="317">Figs 7b–c</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCA0BA63EFBC2FABE" box="[1086,1113,1350,1373]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">8a</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCA57A63EFB1BFABE" box="[1122,1152,1350,1373]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="10" pageNumber="317">9h</figureCitation>
). Orange, orange and black pilose; first tergum black, grey pruinose, long orange pilose; second tergum orange, with basal maculae occupying full lateral width and extending medially into triangular basomedial black vitta, orange pilose, except for black pile on apicolateral margin and medially; third to fourth terga with orange, with triangular diamond-shape basomedial black vitta, orange pilose, except for black pile on apicolateral margin and medially; fifth tergum orange, orange pilose; sterna orange, orange pilose; male genitalia as
<figureCitation id="9A942A51FFB3FFFDCD09A5FEFC3DF97E" box="[828,934,1670,1693]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">Figs 8c–e</figureCitation>
, orange to brown, grayish pruinose and orange golden pilose.
</paragraph>
<paragraph id="021036D4FFB3FFFDCD6FA5BDFBACF89E" blockId="10.[828,1437,1318,2077]" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCD6FA5BDFC36F93E" bold="true" box="[858,941,1733,1757]" italics="true" pageId="10" pageNumber="317">Female</emphasis>
(
<figureCitation id="9A942A51FFB3FFFDCD8AA5BEFBB6F93E" box="[959,1069,1734,1757]" captionStart="Fig" captionStartId="9.[149,180,1672,1691]" captionTargetBox="[150,1436,190,1657]" captionTargetId="figure-16@9.[150,1436,189,1658]" captionTargetPageId="9" captionText="Fig. 7. Lycopale radioheadi sp. nov. A–C – male, holotype (CEUA 103529): A – head, frontal view; B – lateral view; C – abdomen posterodorsal view. D–F – female, paratype (CEUA 103427): D – head, frontal view; E – lateral view; F – abdomen posterodorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14655491" httpUri="https://zenodo.org/record/14655491/files/figure.png" pageId="10" pageNumber="317">Figs 7d–f</figureCitation>
,
<figureCitation id="9A942A51FFB3FFFDCA0EA5BEFBC3F93E" box="[1083,1112,1734,1757]" captionStart="Fig" captionStartId="10.[151,182,1207,1226]" captionTargetBox="[151,1436,188,1190]" captionTargetId="figure-13@10.[151,1436,187,1190]" captionTargetPageId="10" captionText="Fig. 8. Lycopale radioheadi sp. nov. A – male, dorsal view; B – female, dorsal view. C–E – male genitalia: C – whole genitalia, lateral view; D – epandrium, dorsal view; E – hypandrium, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14655493" httpUri="https://zenodo.org/record/14655493/files/figure.png" pageId="10" pageNumber="317">8b</figureCitation>
). Similar to male, except for normal sexual dimorphism and the following: antennae darker than in male; abdominal maculae triangular, approximately equilateral, comparatively shorter than in male; fifth tergum with triangular macula. Body length
<quantity id="C5579B31FFB3FFFDCB0AA43EFCF2F89E" metricMagnitude="-3" metricUnit="m" metricValue="9.899999999999999" metricValueMax="10.1" metricValueMin="9.7" pageId="10" pageNumber="317" unit="mm" value="9.899999999999999" valueMax="10.1" valueMin="9.7">9.7–10.1 mm</quantity>
; wing
<quantity id="C5579B31FFB3FFFDCD9AA41EFBA8F89E" box="[943,1075,1894,1917]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="9.9" metricValueMin="7.3" pageId="10" pageNumber="317" unit="mm" value="8.6" valueMax="9.9" valueMin="7.3">7.3–9.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB3FFFCCD09A4FEFE46FCB6" lastPageId="11" lastPageNumber="318" pageId="10" pageNumber="317" type="diagnosis">
<paragraph id="021036D4FFB3FFFCCD09A4FEFE46FCB6" blockId="10.[828,1437,1318,2077]" lastBlockId="11.[151,760,190,1525]" lastPageId="11" lastPageNumber="318" pageId="10" pageNumber="317">
<emphasis id="30DBEAC6FFB3FFFDCD09A4FEFBA9F87E" bold="true" box="[828,1074,1926,1949]" pageId="10" pageNumber="317">Differential diagnosis.</emphasis>
<taxonomicName id="C5AF4D57FFB3FFFDCA03A4FEFA8AF87E" authorityName="Montoya" authorityYear="2024" box="[1078,1297,1926,1949]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB3FFFDCA03A4FEFA8AF87E" box="[1078,1297,1926,1949]" italics="true" pageId="10" pageNumber="317">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB3FFFDCB23A4FEFAFCF87E" box="[1302,1383,1926,1949]" pageId="10" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
keys out to
<taxonomicName id="C5AF4D57FFB3FFFDCDB0A4DEFBBDF85E" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[901,1062,1958,1981]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB3FFFDCDB0A4DEFBBDF85E" box="[901,1062,1958,1981]" italics="true" pageId="10" pageNumber="317">Lycopale lizeri</emphasis>
</taxonomicName>
in the keys of
<bibRefCitation id="663E4B25FFB3FFFDCAE4A4DEFA02F85E" author="THOMPSON F. C." box="[1233,1433,1958,1981]" pageId="10" pageNumber="317" pagination="1 - 16" refId="ref14948" refString="THOMPSON F. C. 2012: Fabulous flower flies for famous fly fanatics (Diptera: Syrphidae). A tribute to the dipterists of the Canadian National Collection. Canadian Entomologist 144: 1 - 16." type="journal article" year="2012">THOMPSON (2012)</bibRefCitation>
.
<taxonomicName id="C5AF4D57FFB3FFFDCD09A4BEFB86F83E" authorityName="Montoya" authorityYear="2024" box="[828,1053,1990,2013]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB3FFFDCD09A4BEFB86F83E" box="[828,1053,1990,2013]" italics="true" pageId="10" pageNumber="317">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB3FFFDCA12A4BEFBE6F83E" box="[1063,1149,1990,2013]" pageId="10" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from
<taxonomicName id="C5AF4D57FFB3FFFDCBB2A4BEFCE8F81E" baseAuthorityName="Brethes" baseAuthorityYear="1914" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB3FFFDCBB2A4BEFCE8F81E" italics="true" pageId="10" pageNumber="317">L. lizeri</emphasis>
</taxonomicName>
by facial vitta and gena yellow (
<emphasis id="30DBEAC6FFB3FFFDCAE8A49EFABAF81E" box="[1245,1313,2022,2045]" italics="true" pageId="10" pageNumber="317">versus</emphasis>
facial vitta and gena black in
<taxonomicName id="C5AF4D57FFB3FFFDCA3CAB7EFBC4F7FE" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[1033,1119,2054,2077]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB3FFFDCA3CAB7EFBC4F7FE" box="[1033,1119,2054,2077]" italics="true" pageId="10" pageNumber="317">L. lizeri</emphasis>
</taxonomicName>
); antenna orange, except for dorsally brown postpedicel (
<emphasis id="30DBEAC6FFB2FFFCCFFBA3C6FD89FF36" box="[462,530,190,213]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
antenna ferruginous, yellowish in
<taxonomicName id="C5AF4D57FFB2FFFCCF15A3A6FEE8FF16" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[288,371,222,245]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCF15A3A6FEE8FF16" box="[288,371,222,245]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
); postpronotum black pruinose,
<figureCitation id="9A942A51FFB2FFFCCCF8A3A6FF29FEF6" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9h</figureCitation>
(
<emphasis id="30DBEAC6FFB2FFFCCEF4A386FE9EFEF6" box="[193,261,254,277]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
postpronotum yellow pruinose in
<taxonomicName id="C5AF4D57FFB2FFFCCC5AA386FD5AFEF6" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[623,705,254,277]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCC5AA386FD5AFEF6" box="[623,705,254,277]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
,
<figureCitation id="9A942A51FFB2FFFCCCF8A386FF2FFED6" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9e</figureCitation>
); scutum with two submedial and one medial grayish white vittae from base, reaching 3/4 scutum length,
<figureCitation id="9A942A51FFB2FFFCCCFBA246FF29FE96" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9h</figureCitation>
(
<emphasis id="30DBEAC6FFB2FFFCCEF7A226FE9DFE96" box="[194,262,350,373]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
scutum with three submedial golden pruinose vittae throughout the entire scutum length, confluent with a transverse vitta before scutellum base in
<taxonomicName id="C5AF4D57FFB2FFFCCC5FA2E6FD24FE56" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[618,703,414,437]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCC5FA2E6FD24FE56" box="[618,703,414,437]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
,
<figureCitation id="9A942A51FFB2FFFCCCF8A2E6FF2FFE36" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9e</figureCitation>
); pre-scutellum extensively black, without vitta,
<figureCitation id="9A942A51FFB2FFFCCCFBA2C6FF29FE16" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9h</figureCitation>
(
<emphasis id="30DBEAC6FFB2FFFCCEF3A2A6FE90FE16" box="[198,267,478,501]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
pre-scutellum orange, rectangular and confluent with longitudinal vittae in
<taxonomicName id="C5AF4D57FFB2FFFCCFC9A286FDCBFDF6" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[508,592,510,533]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCFC9A286FDCBFDF6" box="[508,592,510,533]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
,
<figureCitation id="9A942A51FFB2FFFCCC6AA286FD35FDF6" box="[607,686,510,533]" captionStart="Fig" captionStartId="14.[151,182,1695,1714]" captionTargetBox="[152,1435,189,1661]" captionTargetId="figure-13@14.[151,1436,188,1673]" captionTargetPageId="14" captionText="Fig. 9. Lycopale thoracic and abdominal patterns, dorsal view:A – Lycopale magnifica (Bigot, 1880); B – L. rectilinea (Hull, 1942); C – L. chrysotaenia (Fluke, 1937); D – L. woodi Thompson, 2012; E – L. lizeri (Brèthes, 1914); F – L. lipoflava (Fluke, 1937); G – L. mendozai sp. nov.; H – L. radioheadi sp. nov. The pile has been omitted." figureDoi="http://doi.org/10.5281/zenodo.14655495" httpUri="https://zenodo.org/record/14655495/files/figure.png" pageId="11" pageNumber="318">Fig. 9e</figureCitation>
); wing hyaline, except for light orange brown anteriorly (
<emphasis id="30DBEAC6FFB2FFFCCC81A166FD63FDD6" box="[692,760,542,565]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
wing hyaline in
<taxonomicName id="C5AF4D57FFB2FFFCCF08A146FE15FDB6" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[317,398,574,597]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCF08A146FE15FDB6" box="[317,398,574,597]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
); metafemur bright orange, except for light reddish brown on apical 1/2 (
<emphasis id="30DBEAC6FFB2FFFCCC0DA126FDE7FD96" box="[568,636,606,629]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
metafemur brownish basally and orange on apical 1/2 or more in
<taxonomicName id="C5AF4D57FFB2FFFCCCD7A106FF55FD56" baseAuthorityName="Brethes" baseAuthorityYear="1914" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCCD7A106FF55FD56" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
); second tergum orange, with basal maculae occupying the full lateral width and extending medially into a triangular basomedial black vitta (
<emphasis id="30DBEAC6FFB2FFFCCC38A1A6FDCAFD16" box="[525,593,734,757]" italics="true" pageId="11" pageNumber="318">versus</emphasis>
second tergum black grey, covered with a deeply black pubescence except for a transverse median line which widens on the edges and in the midline in
<taxonomicName id="C5AF4D57FFB2FFFCCF4EA046FE55FCB6" baseAuthorityName="Brethes" baseAuthorityYear="1914" box="[379,462,830,853]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="lizeri">
<emphasis id="30DBEAC6FFB2FFFCCF4EA046FE55FCB6" box="[379,462,830,853]" italics="true" pageId="11" pageNumber="318">L. lizeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB2FFFCCEA2A026FD1DFB96" pageId="11" pageNumber="318" type="etymology">
<paragraph id="021036D4FFB2FFFCCEA2A026FD1DFB96" blockId="11.[151,760,190,1525]" pageId="11" pageNumber="318">
<emphasis id="30DBEAC6FFB2FFFCCEA2A026FE83FC96" bold="true" box="[151,280,862,885]" pageId="11" pageNumber="318">Etymology.</emphasis>
The specific epithet
<taxonomicName id="C5AF4D57FFB2FFFCCC36A026FDE3FC96" authorityName="Montoya" authorityYear="2024" box="[515,632,862,885]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="radioheadi">
<emphasis id="30DBEAC6FFB2FFFCCC36A026FDE3FC96" box="[515,632,862,885]" italics="true" pageId="11" pageNumber="318">radioheadi</emphasis>
</taxonomicName>
is given in honor of the English rock band Radiohead (formed in 1985), for their extraordinary music, considered one of the most influential bands in contemporary alternative rock. Radiohead is an environmental activist band that raises awareness of climate change and socio-political issues. Their melodies were a source of inspiration and an excellent company while conducting this study. The species name is a masculine noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="4AB5655FFFB2FFFCCEA2A706FE56FA16" pageId="11" pageNumber="318" type="distribution">
<paragraph id="021036D4FFB2FFFCCEA2A706FE56FA16" blockId="11.[151,760,190,1525]" pageId="11" pageNumber="318">
<emphasis id="30DBEAC6FFB2FFFCCEA2A706FD8AFB76" bold="true" box="[151,529,1150,1173]" pageId="11" pageNumber="318">Distribution and natural history.</emphasis>
<taxonomicName id="C5AF4D57FFB2FFFCCC2CA706FD63FB76" authorityName="Montoya" authorityYear="2024" box="[537,760,1150,1173]" class="Insecta" family="Syrphidae" genus="Lycopale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="radioheadi" status="sp. nov.">
<emphasis id="30DBEAC6FFB2FFFCCC2CA706FD63FB76" box="[537,760,1150,1173]" italics="true" pageId="11" pageNumber="318">Lycopale radioheadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BE857BDFFB2FFFCCEA2A7E6FF74FB56" box="[151,239,1182,1205]" pageId="11" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the
<typeStatus id="DD148876FFB2FFFCCC25A7E6FDA4FB56" box="[528,575,1182,1205]" pageId="11" pageNumber="318">type</typeStatus>
localities in the
<collectingRegion id="C06BF836FFB2FFFCCEA2A7C6FE88FB36" box="[151,275,1214,1237]" country="Colombia" name="Magdalena" pageId="11" pageNumber="318">Magdalena</collectingRegion>
Valley and Eastern Mountain range Real Montane Forests ecoregions. The species is restricted to pristine forests and Paramo ecosystems (sky islands) in the Paramo complexes of Santa Ines Belmira and Sonsón at elevations between 2840 and
<quantity id="C5579B31FFB2FFFCCFC4A646FDD8FAB6" box="[497,579,1342,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.02" pageId="11" pageNumber="318" unit="m" value="3020.0">3020 m</quantity>
in isolated areas in northwestern Central Mountain range in
<collectingCountry id="7AB87644FFB2FFFCCC57A626FD50FA96" box="[610,715,1374,1397]" name="Colombia" pageId="11" pageNumber="318">Colombia</collectingCountry>
and eastern slope of Eastern Mountain range in
<collectingCountry id="7AB87644FFB2FFFCCC57A606FD20FA76" box="[610,699,1406,1429]" name="Ecuador" pageId="11" pageNumber="318">Ecuador</collectingCountry>
(
<figureCitation id="9A942A51FFB2FFFCCCFDA606FF47FA56" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[151,182,1241,1260]" captionStartId-1="17.[151,182,1002,1021]" captionTargetBox-0="[151,1436,189,1220]" captionTargetBox-1="[152,1417,197,958]" captionTargetId-0="figure-402@16.[151,1436,189,1233]" captionTargetId-1="figure-598@17.[151,1436,189,977]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 10. Distributional map of Lycopale species." captionText-1="Fig. 11. A – detail of distributional map of Lycopale in Colombia and Ecuador. B–C – study area in: B – Paramo Santa Inés Belmira; C – Paramo Sonsón. Sampling sites where the type material of the new species was collected (see MONTOYA et al. 2021)." figureDoi-0="http://doi.org/10.5281/zenodo.14655499" figureDoi-1="http://doi.org/10.5281/zenodo.14655501" httpUri-0="https://zenodo.org/record/14655499/files/figure.png" httpUri-1="https://zenodo.org/record/14655501/files/figure.png" pageId="11" pageNumber="318">Figs 10–11</figureCitation>
). The habitats where the specimens were collected are covered mainly by shrubby plants and inhabited by endemic species (
<figureCitation id="9A942A51FFB2FFFCCF66A6A6FE5BFA16" box="[339,448,1502,1525]" captionStart="Fig" captionStartId="2.[151,182,2000,2019]" captionTargetBox="[151,1429,658,1975]" captionTargetId="figure-178@2.[151,1437,655,1987]" captionTargetPageId="2" captionText="Fig. 1A–G. The type locality of Lycopale radioheadi sp. nov. in Paramo (above) and pristine Andean forests (middle) in Antioquia, Colombia. C – Sky Islands. H – type material of Lycopale mendozai sp. nov. was collected inside a forest flying around the Native Bromeliad, Guzmania coriostachya (Griseb.) Mez (Bromeliaceae) (below). I – habitat of Lycopale magnifica (Bigot, 1880) in the Western Mountain range in Colombia." figureDoi="http://doi.org/10.5281/zenodo.14655473" httpUri="https://zenodo.org/record/14655473/files/figure.png" pageId="11" pageNumber="318">Figs 1a–g</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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