300 lines
35 KiB
XML
300 lines
35 KiB
XML
<document ID-CLB-Dataset="9622" ID-DOI="http://dx.doi.org/10.3897/zookeys.987.53648" ID-GBIF-Dataset="e14abbdb-e0a3-4c35-ad98-1cfd0cee46b5" ID-PMC="PMC7666096" ID-Pensoft-Pub="1313-2970-987-135" ID-Pensoft-UUID="A304608B226C5889BE8AF26E618669A3" ID-PubMed="33223888" ID-ZooBank="861F9C146E6D4222A41FFC56F978C2E8" ModsDocID="1313-2970-987-135" checkinTime="1604753978193" checkinUser="pensoft" docAuthor="Inoue, Shota, Nomura, Shuhei & Yin, Zi-Wei" docDate="2020" docId="897F5BB0F37F5E3393A813DA9D806DA6" docLanguage="en" docName="ZooKeys 987: 135-156" docOrigin="ZooKeys 987" docSource="http://dx.doi.org/10.3897/zookeys.987.53648" docTitle="Pseudophanias yaimensis Inoue, Nomura & Yin 2020, sp. nov." docType="treatment" docUuid="81ADE876-E798-467A-8625-03721012EF1B" docUuidSource="ZooBank" docVersion="6" id="A304608B226C5889BE8AF26E618669A3" lastPageNumber="135" masterDocId="A304608B226C5889BE8AF26E618669A3" masterDocTitle="Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="156" masterPageNumber="135" pageNumber="135" updateTime="1732761297298" updateUser="ExternalLinkService">
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<mods:mods id="C258B59946ED439002B4C2ACAE797F46" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B049BA22A981E66F8C483AC4BBC3C90D">
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<mods:title id="D5DE2DDDEFE42CD3CAAF583422DDF138">Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="14C7345C0B6A49493EDE546428BF6BD4" type="personal">
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<mods:roleTerm id="1D20D659F0D13A8945EA44C1F41C7023">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="44473E6283650E8837C33506FBE0C04F">Inoue, Shota</mods:namePart>
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<mods:affiliation id="8B2B26F09237FCC06AEDD6AE0470DD7B">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan & The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:nameIdentifier id="3BF48750E187DD85C188FD4FC775D09F" type="orcid">https://orcid.org/0000-0002-7005-6531</mods:nameIdentifier>
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<mods:nameIdentifier id="832A616A6495FEF530E7087A62200844" type="email">pselaphineman@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="694168E8492E4B97103A3737BC9FAB48" type="personal">
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<mods:roleTerm id="11A93823A830149A20C5BB2AFD974D4C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="43933E15D113E909F94EE69D21712449">Nomura, Shuhei</mods:namePart>
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<mods:affiliation id="6AB37C506EDE997DB9B149D38B49969E">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="1445869990B4599ED4C69A340DF9649F" type="personal">
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<mods:roleTerm id="18AEAF25A187D144A890F556DA4006FB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F3616E5FBB10E984D9A35766DCA58AC6">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="526D0951D5BD849F2C84767D4095B32F">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai 200234, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="422774BAC6C0FD51CB2EE7CF8B4CD943">text</mods:typeOfResource>
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<mods:title id="96B35E161D890E2367F2796AE41E3B75">ZooKeys</mods:title>
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<mods:part id="E730BBFE46159278AB9009C4D09FF5EC">
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<mods:date id="FC3B986F7422E13D9F2D97052A87642C">2020</mods:date>
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<mods:detail id="0B75821F51F77AB02E9FECFF03E0EB95" type="volume">
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<mods:number id="1978E573D4C76AD869053C0573ED6E2F">987</mods:number>
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<mods:start id="7EDDC1F4F42A700C282A0CE954BED2C9">135</mods:start>
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<mods:end id="80B2B698F3B7AACF240EE002D4446B0D">156</mods:end>
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<mods:location id="93D4B1E0D6C5A6B52094B9E4BB0026DC">
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<mods:url id="D026316B4C8AD0D7E3549AE829697788">http://dx.doi.org/10.3897/zookeys.987.53648</mods:url>
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<mods:classification id="C9F6540727EA28D9B8E9F010AE32F541">journal article</mods:classification>
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<mods:identifier id="9CBB7F57BA929B343EE8A1467127486E" type="DOI">http://dx.doi.org/10.3897/zookeys.987.53648</mods:identifier>
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<mods:identifier id="564A1D85B071B8D21E25399405132330" type="Pensoft-Pub">1313-2970-987-135</mods:identifier>
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<treatment id="897F5BB0F37F5E3393A813DA9D806DA6" ID-GBIF-Taxon="168821844" LSID="urn:lsid:zoobank.org:act:81ADE876-E798-467A-8625-03721012EF1B" httpUri="http://treatment.plazi.org/id/897F5BB0F37F5E3393A813DA9D806DA6" lastPageNumber="135" pageId="0" pageNumber="135" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Pselaphinae">
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<subSubSection id="1F5D726EA456C9E6952A548CA317A320" pageId="0" pageNumber="135" type="nomenclature">
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<paragraph id="C2470AE80A4A36B4892184136D9E89D1" pageId="0" pageNumber="135">
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<taxonomicName id="AC4236E977151348968D206F1F36E0AD" ID-CoL="8Y8T7" LSID="897F5BB0-F37F-5E33-93A8-13DA9D806DA6" authority="Inoue, Nomura & Yin" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis" status="sp. nov.">Pseudophanias yaimensis Inoue, Nomura & Yin</taxonomicName>
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<taxonomicNameLabel id="1932D220C1D6B7C778B6DF4414C407F8" pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
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<figureCitation id="DEF2A2F7320FF33CF4B4383F00F1AA7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Pseudophanias yaimensis A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure1" httpUri="https://binary.pensoft.net/fig/472456" pageId="0" pageNumber="135">Figs 1</figureCitation>
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<figureCitation id="F570BFAA273AF091A34B01EF234215F2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological details of Pseudophanias yaimensis A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135">, 2</figureCitation>
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<figureCitation id="9AE89B98E30F935C9DBE1D15B0E86195" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">, 3</figureCitation>
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<figureCitation id="A4ECFDFE812E3B8EC36E8D26A0F7F998" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Pseudophanias yaimensis A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure4" httpUri="https://binary.pensoft.net/fig/472459" pageId="0" pageNumber="135">, 4 [Japanese name: Yaima-tsumugata-arizukamushi]</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="56E0D94503DFF3725E335C08F8A80116" pageId="0" pageNumber="135" type="type material">
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<paragraph id="37E27378936C1A0389F027C948046EB8" pageId="0" pageNumber="135">Type material.</paragraph>
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<paragraph id="C6FA9EA0DB7B01C411C9F44F4EFE7C40" pageId="0" pageNumber="135">
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<emphasis id="2DB7082FB51B1E7F8AD66727FA4B0778" bold="true" italics="true" pageId="0" pageNumber="135">Holotype</emphasis>
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(NSMT): ♂, "Japan: [
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<normalizedToken id="170AE99A070749254335CC014A811A60" originalValue="Ryûkyû">Ryukyu</normalizedToken>
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], Ishigaki- / jima,
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<normalizedToken id="E2E8C628E55099F0369E46F6508B22F2" originalValue="Takeda-rindô">Takeda-rindo</normalizedToken>
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23 X 2007, Teruaki Ban leg. // HOLOTYPE (red) /♂,
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<taxonomicName id="5AB60DDE1F2EFF765700984F7EF3D900" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
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<emphasis id="FD60A573DBEABE3CF17F3BFA1F129E49" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020"
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<emphasis id="DE0A63EE9DAF14FE4BBC437D5B6B5866" bold="true" italics="true" pageId="0" pageNumber="135">Paratypes</emphasis>
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: Japan: 1 ♀, [
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<normalizedToken id="10B4BDC6D6FEDAD26A769762CA634378" originalValue="Ryûkyû">Ryukyu</normalizedToken>
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], Okinawa ken, Ishigaki-jima Is., Mt. Omoto-dake, 16 VIII 1991, K. Ogata leg. (NSMT); 1 ♂, Ishigaki-jima Is., Mt. Omoto-dake, (FIT), 14-20 V 2002, S. Hori leg. (NSMT); 1 ♂, 1 ♀, China: Taiwan, Taichung Co. (台中县), Guguan (谷关), 1238 m,
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<geoCoordinate id="C109E4A4ED7E93B92D668BF0505D199B" degrees="24.180691" direction="north" orientation="latitude" precision="1" value="24.18069">24.180691N</geoCoordinate>
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,
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<geoCoordinate id="26C678239C024785B00A9F34E2A6F9AF" degrees="120.944213" direction="east" orientation="longitude" precision="1" value="120.944214">120.944213E</geoCoordinate>
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, 29 III 2015, local collector, nest of
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<taxonomicName id="6E7667D50F4DDDC68115CE157E008534" baseAuthorityName="Nawa" baseAuthorityYear="1911" class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes parvonasutus" order="Blattodea" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="parvonasutus">
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<emphasis id="7557F5C44CA095C7EEC3E4BFE72BC479" italics="true" pageId="0" pageNumber="135">Nasutitermes parvonasutus</emphasis>
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</taxonomicName>
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(SNUC). Each paratype pinned with the following label: "PARATYPE (yellow) / ♂ (or ♀),
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<taxonomicName id="998532815F58CEEECE1BDF3C70BA407D" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
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<emphasis id="0C7DEA75E04AC9D66CF62E796FCBC3AA" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020".
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</paragraph>
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</subSubSection>
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<subSubSection id="258258785CF103308134DC4A3E87201E" pageId="0" pageNumber="135" type="diagnosis">
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<paragraph id="EF467C8A8FD33DD3E5FD381A6E87A1F1" pageId="0" pageNumber="135">Diagnosis.</paragraph>
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<paragraph id="4B225E2B4EE9E57DD0F318A411543A42" pageId="0" pageNumber="135">
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<taxonomicName id="AEE36DF97B49AF435834562C5B9AA9F8" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
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<emphasis id="5D510F37562D15207D0AC44B8C2E39B4" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
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</taxonomicName>
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is most similar to the Sumatran
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<taxonomicName id="80C7AE3A193382E0ACA8B5915C99485E" lsidName="P. robustus" pageId="0" pageNumber="135" rank="species" species="robustus">
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<emphasis id="8C9E82A2A1C81BF1D6283DB980A68E2C" italics="true" pageId="0" pageNumber="135">P. robustus</emphasis>
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</taxonomicName>
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Raffray, 1904, but can be distinguished by the distinctly smaller body size (3.00-3.20 mm in
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<taxonomicName id="51A1732490E9531F2B798E18C4433E7A" lsidName="P. robustus" pageId="0" pageNumber="135" rank="species" species="robustus">
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<emphasis id="72FEA643C129DCBEF5680F95E3038B82" italics="true" pageId="0" pageNumber="135">P. robustus</emphasis>
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</taxonomicName>
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), the angulate antennomere 11 at anterolateral margins in the male, the finely punctate head and pronotum, and the shorter discal carinae on tergite IV.
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</paragraph>
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</subSubSection>
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<subSubSection id="3FB121343DDFD761FFD294A05BBF8507" pageId="0" pageNumber="135" type="description">
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<paragraph id="FFC49FDFED24073F7CFEF73F0F47197B" pageId="0" pageNumber="135">Description.</paragraph>
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<paragraph id="6A5069C9294BC68C98066F1588BAB23E" pageId="0" pageNumber="135">
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<emphasis id="658DF1FA70D45D787FAE745204D69F31" bold="true" pageId="0" pageNumber="135">Male</emphasis>
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(Figs
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<figureCitation id="69D3B6E14327C8AE8A116A673966AE7A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Pseudophanias yaimensis A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure1" httpUri="https://binary.pensoft.net/fig/472456" pageId="0" pageNumber="135">1A</figureCitation>
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,
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<figureCitation id="B00004597A4E238973375717EE0FC79B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological details of Pseudophanias yaimensis A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135">2A</figureCitation>
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). Body length 2.16-2.32 mm. Dorsal surface polished and weakly shining, with dense setae.
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<emphasis id="C05BBA9D46E76EEFC0E364609E14D67F" bold="true" italics="true" pageId="0" pageNumber="135">Head</emphasis>
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(Fig.
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<figureCitation id="F2B7635758CD7C16666DC0B3B01D25CE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3A</figureCitation>
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) about as long as wide, HL 0.46 mm, HW 0.42-0.44 mm, same size as pronotum, nearly hexagonal, with dense setae; frontal rostrum with short longitudinal sulcus including large fovea; antennal tubercles prominent, with dense punctures which gradually disappear towards vertex; vertex polished, finely punctate, with pair of foveae; frontal, vertexal foveae glabrous; eyes prominent; postocular margins three times longer than length of eyes. Maxillary palpi (Fig.
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<figureCitation id="413BF1E534A56D34DD096DF02B8D581B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3B</figureCitation>
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) symmetrical; palpomere 1 minute; palpomere 2 elongate, narrowed in basal half; palpomere 3 small, widest at apices; palpomere 4 fusiform.
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<emphasis id="0361827B2E1775B82357A2466D949570" bold="true" italics="true" pageId="0" pageNumber="135">Antennae</emphasis>
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(Fig.
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<figureCitation id="DD430895DB1F1645F7C549FFA62F05E9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological details of Pseudophanias yaimensis A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135">2B</figureCitation>
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) elongate, 1.42 mm in length; antennal club formed by apical antennomere alone; antennomere 1 thick, elongate, 1.5 times longer than 2; 2 slightly longer than wide; 3-11 successively widened towards apices; 3-7 each slightly elongate; 8-10 quadrate; 11 enlarged, roundly broadened towards apices in inner margin, straightened towards apices in outer margin; each apical half of outer margin distinctly carinate, with angulate spine at anterolateral part.
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<emphasis id="950980C2936527437E752311C100E44D" bold="true" italics="true" pageId="0" pageNumber="135">Pronotum</emphasis>
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(Fig.
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<figureCitation id="985DE7C54CDC0D58E0CB7164D7993BD7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3C</figureCitation>
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) slightly longer than wide, PL 0.48 mm, PW 0.46 mm; widest at anterior one-third, weakly constricted from widest point towards base, polished, with coarse punctures along posterior margin, with a median and pair of lateral antebasal foveae, with distinct conical spine just anterior of median fovea.
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<emphasis id="5A482CE4AFCF58C789894CE98D970267" bold="true" italics="true" pageId="0" pageNumber="135">Metaventrite</emphasis>
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(Figs
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<figureCitation id="4056424ED23E9AEA3ED41658FCE3BE3A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological details of Pseudophanias yaimensis A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135">2D</figureCitation>
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,
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<figureCitation id="5708FEFEFF7B1139E4384E605C04DCC9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3D</figureCitation>
|
|
) finely punctate, moderately convex, but area just above metaventral apex impressed; that impression nearly trapezoidal, half as long as metaventral length, occupying 1/4 metaventral width; anterior margin of that impression straight, distinct.
|
|
<emphasis id="29DB4A21B7E45EAD2CB156A2CA870C5B" bold="true" italics="true" pageId="0" pageNumber="135">Elytra</emphasis>
|
|
(Fig.
|
|
<figureCitation id="86091415B2FA1ADA785049C337F9B3AB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3E</figureCitation>
|
|
) nearly trapezoidal, widest near posterior 1/4 much wider than long, EL 0.60-0.62 mm, EW 0.85-0.94 mm; dorsal surface polished, with thin, long setae, finely punctate; each elytron with two basal foveae; discal stria shallow, extending from basal fovea placed middle to posterior half.
|
|
<emphasis id="9FF1CBB9E2B1CB104BFEB29B238CE13C" bold="true" pageId="0" pageNumber="135">
|
|
<emphasis id="B1B42E157E9F522799BE0E6026F68015" italics="true" pageId="0" pageNumber="135">Legs</emphasis>
|
|
.
|
|
</emphasis>
|
|
All legs elongate and slender; femora each broadest near middle; tibiae each slightly broadened to apex, with dense yellow setae at apex; protibiae and mesotibiae nearly straight; metatibiae longest; tarsi each elongate, with tarsomeres 2 about as long as tarsomeres 3; mesotarsi (Fig.
|
|
<figureCitation id="58BBA0D83E04AF921DDD2FDB8FCE9C02" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3G</figureCitation>
|
|
) modified, broadest; tarsomeres 3 each with projection of 2/3 length of entire tarsomere; tarsal claws (Fig.
|
|
<figureCitation id="1CF2F12EA2A9477E503D872CAADAD67F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3H</figureCitation>
|
|
) each asymmetrical; posterior claws thin, short.
|
|
<emphasis id="FC0290D0A1D921968DFDFF1D160C2B89" bold="true" italics="true" pageId="0" pageNumber="135">Abdomen</emphasis>
|
|
(Fig.
|
|
<figureCitation id="41836D40919C685C73F8A6556C66B04E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Pseudophanias yaimensis, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G mesotarsus H mesotarsal claws. Scale bars: 0.2 mm (A, C-G); 0.1 mm (B); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135">3F</figureCitation>
|
|
) wider than long, widest at tergite IV, AL 0.62-0.76 mm, AW 0.85-0.90 mm; tergite IV longest, twice as long as tergite V, with pair of short longitudinal carinae about one-fifth as long as tergal length, with setose depression at base. Tergite and sternite VIII as in Fig.
|
|
<figureCitation id="380F820D704ED1FD0C08A4796433C058" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Pseudophanias yaimensis A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure4" httpUri="https://binary.pensoft.net/fig/472459" pageId="0" pageNumber="135">4D, E</figureCitation>
|
|
.
|
|
<emphasis id="716B677B7C6938AB8419B164B79D6402" bold="true" italics="true" pageId="0" pageNumber="135">Aedeagus</emphasis>
|
|
(Fig.
|
|
<figureCitation id="A82CE54A4795689840594A273878E6D7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Pseudophanias yaimensis A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure4" httpUri="https://binary.pensoft.net/fig/472459" pageId="0" pageNumber="135">4A-C</figureCitation>
|
|
) 0.39-0.40 mm in length, well-sclerotized, asymmetrical; parameres elongate, reaching near apex of median lobe; each paramere with three setae at apex; median lobe strongly widened towards apex in dorso-ventral view, constricted at median part in lateral view; ventral side of apical part asymmetrical, weakly sclerotized, nearly formed three pronged fork; medioapical part bended laterally; dorsal side of apical part asymmetrical, bifurcate from median part; endophallus indistinct.
|
|
</paragraph>
|
|
<caption id="0FBB4269C09F67B85FF0459863DFB36C" doi="10.3897/zookeys.987.53648.figure1" httpUri="https://binary.pensoft.net/fig/472456" pageId="0" pageNumber="135" start="Figure 1" startId="F1">
|
|
<paragraph id="07C086381D4E4A170B703A951A2EA7D4" pageId="0" pageNumber="135">
|
|
<emphasis id="E9B53986B9EE6968B7804CA5B98DF633" bold="true" pageId="0" pageNumber="135">Figure 1.</emphasis>
|
|
Dorsal habitus of
|
|
<taxonomicName id="F99DE02391DE2B47EC075C5F7B79DDD4" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
|
|
<emphasis id="0B0E37D3BBA05CAA7A75EEEC7AB1AA61" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="33669DA87E96D4F0E47144165E974ACA" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
male
|
|
<emphasis id="ED54F1F9CE6C22E4F1A8AE9DCD5213BA" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
female. Scale bar: 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="D0D0B56543608F98EA1FF5E1C557CD28" doi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135" start="Figure 2" startId="F2">
|
|
<paragraph id="8FC5386CB3D4A25B3F59D1D15F517A95" pageId="0" pageNumber="135">
|
|
<emphasis id="25F42716099E4AA72B62A998304D2E0D" bold="true" pageId="0" pageNumber="135">Figure 2.</emphasis>
|
|
Morphological details of
|
|
<taxonomicName id="D262C1BF2F2DE5E47460BD2DA6F912C9" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
|
|
<emphasis id="DE82CB741A090F4533F060EF5694EAAE" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="838CA8ADADD037DF10BB87C017BB4BDF" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
habitus in lateral view
|
|
<emphasis id="2AE148F46D340D0E1C85BC85DBD9D6DE" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
male antenna
|
|
<emphasis id="6F42B1A5F47FC0D27BE9085E4D678ABF" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
female antenna
|
|
<emphasis id="1CC5C3DBA40FEC005F71DB3F928D08A3" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
male metaventrite. Scale bars: 1.0 mm (
|
|
<emphasis id="617941ED5D573E6C639F57394FF009C5" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="6C32A1082D000BB0B6F929356F873A8E" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
); 0.2 mm (
|
|
<emphasis id="71AFD2F64331A389747F9002E79E5EC3" bold="true" pageId="0" pageNumber="135">B, C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="D4CE6C81BC10490A0260BF314F184A63" doi="10.3897/zookeys.987.53648.figure3" httpUri="https://binary.pensoft.net/fig/472458" pageId="0" pageNumber="135" start="Figure 3" startId="F3">
|
|
<paragraph id="6F70157091D220646564E2262E96318C" pageId="0" pageNumber="135">
|
|
<emphasis id="F440CDD67D6D1C739A093D6D7FDAAE18" bold="true" pageId="0" pageNumber="135">Figure 3.</emphasis>
|
|
SEM images of
|
|
<taxonomicName id="DD52793EF5B19C84BD695F5101FF817D" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
|
|
<emphasis id="BFD72B0B5224A146F6EDD134B930997E" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
|
|
</taxonomicName>
|
|
, male
|
|
<emphasis id="11029C6DD334FD6E9F98C0E80454D82F" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
head
|
|
<emphasis id="CCB173A208D89A319D9CFEBB63DF471B" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
maxillary palp
|
|
<emphasis id="0B4B57D3DFCE24B22B6711143C9D8A12" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
pronotum
|
|
<emphasis id="56C8B6DD59627A7CF3FCA17B1DDAF648" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
metaventrite
|
|
<emphasis id="61F5E31860870A7EB22805FFC1F56CA4" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
elytra
|
|
<emphasis id="FBEDB6E4C0C4E666F164EE0E63A66A36" bold="true" pageId="0" pageNumber="135">F</emphasis>
|
|
abdomen
|
|
<emphasis id="7295699BB1D971226A7D4E9DB6B13130" bold="true" pageId="0" pageNumber="135">G</emphasis>
|
|
mesotarsus
|
|
<emphasis id="A4E90ECD156B6CB2307B8196CE1E8C8E" bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
mesotarsal claws. Scale bars: 0.2 mm (
|
|
<emphasis id="874CC0CA07768BB2378258B8B139F6FF" bold="true" pageId="0" pageNumber="135">A, C-G</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="451EC7CE0D65152D7CAAF8587FEEBBC8" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
); 0.05 mm (
|
|
<emphasis id="7C3D277002F79BB03B02AAC3CE3AFEA5" bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="F4D7BA52907A8F9FA0C28321F40A0873" doi="10.3897/zookeys.987.53648.figure4" httpUri="https://binary.pensoft.net/fig/472459" pageId="0" pageNumber="135" start="Figure 4" startId="F4">
|
|
<paragraph id="484CDC0DDF6842AE273D81224DC875EC" pageId="0" pageNumber="135">
|
|
<emphasis id="36AA5338744D4E7A241562BD8A3E6EF5" bold="true" pageId="0" pageNumber="135">Figure 4.</emphasis>
|
|
Male genitalia of
|
|
<taxonomicName id="CC4BF1E32DFCFE03FAEFFC60CA5543D1" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias yaimensis" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yaimensis">
|
|
<emphasis id="BDFC76FC6E38A3700FDC25FAFC476F08" italics="true" pageId="0" pageNumber="135">Pseudophanias yaimensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="F325C4D77004B5A6E9FD190D07DE4D06" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
ventral view
|
|
<emphasis id="BCD48FE31C887E6358AF368F463CC437" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
lateral view
|
|
<emphasis id="B2D64A7BECAA41A7615049FF7010CA68" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
dorsal view
|
|
<emphasis id="389C2DB0C1536076B20141F7DF77A6A3" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
tergite VIII
|
|
<emphasis id="B87BDC5CF68FE6E7B7E07817F3009AA7" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
sternite VIII. Scale bars: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="4389A1FDDED37D048CCA49215D3F112D" pageId="0" pageNumber="135">
|
|
<emphasis id="EF095004D594D376A4B3A5A411CC8988" bold="true" pageId="0" pageNumber="135">Female</emphasis>
|
|
(Fig.
|
|
<figureCitation id="EECB4EE3292EFEA9958CF3BBE436FB5F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Pseudophanias yaimensis A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure1" httpUri="https://binary.pensoft.net/fig/472456" pageId="0" pageNumber="135">1B</figureCitation>
|
|
). BL 2.20-2.32 mm; HL 0.40-0.52 mm; HW 0.42-0.45 mm; PL 0.46-0.49 mm; PW 0.44-0.46 mm; EL 0.58-0.63 mm; EW 0.78-0.89 mm; AL 0.70-0.76 mm; AW 0.86-0.90 mm. Antennae (Fig.
|
|
<figureCitation id="3D0F10766F39253FFA01767334B8438C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological details of Pseudophanias yaimensis A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure2" httpUri="https://binary.pensoft.net/fig/472457" pageId="0" pageNumber="135">2C</figureCitation>
|
|
) with antennomeres 11 unmodified, each ovoid, without carina and angulate spine; eyes smaller than male; mesotarsi without projection.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59D6719351BCE1027C7893EAF5B28952" pageId="0" pageNumber="135" type="etymology">
|
|
<paragraph id="4CA2C0D986E0B3AE8D17BF41EC377181" pageId="0" pageNumber="135">Etymology.</paragraph>
|
|
<paragraph id="9D1A11077020FB3122C47FDC4D3B6331" pageId="0" pageNumber="135">Ishigaki Island, where the type locality of this species was discovered, is a part of the Yaeyama Islands. This specific epithet refers to Yaima which is a local dialect of the Yaeyama Islands.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="681A3578D100E34F2C590BABC0231833" pageId="0" pageNumber="135" type="distribution">
|
|
<paragraph id="100C478528DBECC004912E575EC5315E" pageId="0" pageNumber="135">Distribution.</paragraph>
|
|
<paragraph id="48ACDBFB7E50C4395051C3A3822B8D49" pageId="0" pageNumber="135">
|
|
Japan (
|
|
<normalizedToken id="FA0E296346F71D0D370067BEEE1818C1" originalValue="Ryûkyû">Ryukyu</normalizedToken>
|
|
: Ishigaki-jima Is.), China (Taiwan).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="67871FD5D64F4DEA2C67DF540A91AE3B" pageId="0" pageNumber="135" type="biology_ecology">
|
|
<paragraph id="F3D2EBEA23E5FC7274D6D96D5877923B" pageId="0" pageNumber="135">Biology.</paragraph>
|
|
<paragraph id="697B7F4A3933C44A3D48D99A5BCFFE9C" pageId="0" pageNumber="135">
|
|
Two paratypes from Taiwan were collected with the termite
|
|
<taxonomicName id="32FAD5847582DCE8604F865C66EC41D7" baseAuthorityName="Nawa" baseAuthorityYear="1911" class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes parvonasutus" order="Blattodea" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="parvonasutus">
|
|
<emphasis id="0483F86F25403F2947472916F5806677" italics="true" pageId="0" pageNumber="135">Nasutitermes parvonasutus</emphasis>
|
|
</taxonomicName>
|
|
Nawa, 1911. In Japan, one paratype was collected using a Flight Interception Trap (FIT).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="80273989438FE05D33ED16D17E9DE701" pageId="0" pageNumber="135" type="remarks">
|
|
<paragraph id="10E567566742785D92A688212678B098" pageId="0" pageNumber="135">Remarks.</paragraph>
|
|
<paragraph id="894A8443A9A2A0BF2BA102A4E089C308" pageId="0" pageNumber="135">
|
|
This species is distributed in
|
|
<normalizedToken id="D147345D60353C57B0FC37BE727F5815" originalValue="Yaeyama-shotô">Yaeyama-shoto</normalizedToken>
|
|
Islands, Japan and Taiwan, China. The two localities are close to each other and shared the same fauna for some insect groups. The two populations show slight difference in the morphology of the aedeagus. The lateral projections of the median lobe of the population of Taiwan are relatively longer and narrower than those of the population of Yaeyama. But the general appearance and especially the male sexual characters are otherwise almost identical. Therefore, we treat such a difference as interspecific variation.
|
|
</paragraph>
|
|
<paragraph id="0DDE686E8B9164872390EC80229FAEEC" pageId="0" pageNumber="135">
|
|
The Taiwanese specimens were collected from a nest of the
|
|
<taxonomicName id="7FD6C332480A6F772E277B72D2A07B51" baseAuthorityName="Nawa" baseAuthorityYear="1911" class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes parvonasutus" order="Blattodea" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="parvonasutus">
|
|
<emphasis id="099CD4422F8988230C3E6D6F70FB4404" italics="true" pageId="0" pageNumber="135">Nasutitermes parvonasutus</emphasis>
|
|
</taxonomicName>
|
|
termite. However, the Japanese specimens were collected from leaf litter samples or by FIT. Some pselaphine species are known to live under the bark and rotten wood with termites. Thus, more information is needed to recognize the possible termitophyly of the new species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |